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<document id="FBDD07ED8C951A3AB2D285FA5D66C6BE" ID-CLB-Dataset="26448" ID-DOI="10.11646/zootaxa.4619.2.9" ID-GBIF-Dataset="1e87ced1-1631-463d-92d4-508d30dbbc48" ID-ISSN="1175-5326" ID-Zenodo-Dep="3249675" ID-ZooBank="6268CC00-B103-4110-BDE5-AF539836DB86" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1560917068779" checkinUser="plazi" docAuthor="Tan, Ming Kai, Japir, Razy, Chung, Arthur Y. C. &amp; Robillard, Tony" docDate="2019" docId="03FF87FDFFEFFFD84DD2D2C7FCEAFD2F" docLanguage="en" docName="zootaxa.4619.2.9.pdf" docOrigin="Zootaxa 4619 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cardiodactylus borneoe Robillard &amp; Gorochov 2014" docType="treatment" docVersion="11" lastPageNumber="362" masterDocId="FFC6FF85FFE7FFD74D45D420FFECFFA8" masterDocTitle="Crickets of the subfamily Eneopterinae (Orthoptera: Grylloidea) from Sandakan, Sabah: one new species and calling songs of a sympatric species" masterLastPageNumber="363" masterPageNumber="347" pageNumber="355" updateTime="1698735472763" updateUser="ExternalLinkService">
<mods:mods id="6986E0E170FA111B776B96D605AC93E1" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="39E3E1FB2E6F9A10C086FE4909B07364">
<mods:title id="A6590FBD242F6438BCCE742F84537571">Crickets of the subfamily Eneopterinae (Orthoptera: Grylloidea) from Sandakan, Sabah: one new species and calling songs of a sympatric species</mods:title>
</mods:titleInfo>
<mods:name id="FFDB7310AC3DF15C08D2128101A9867A" type="personal">
<mods:role id="2B4BE801D7FB0CD0537DF0CFC371FE52">
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<mods:namePart id="9FC91E6996ACC04CD7E860A6293C37EC">Tan, Ming Kai</mods:namePart>
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<mods:name id="CB6BD5B7E6328F138474E4665435340B" type="personal">
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<mods:namePart id="85CEB8368333014DCA54E92A8A6AC772">Japir, Razy</mods:namePart>
</mods:name>
<mods:name id="19B094EEF491C0DE464E4C0CF939AD9A" type="personal">
<mods:role id="CD451D751FBA92030CF1C45B138A2F18">
<mods:roleTerm id="2E87301F786D7EF22F79E7400DE5DD9D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="07D403F143C012228A369379122AB420">Chung, Arthur Y. C.</mods:namePart>
</mods:name>
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<mods:role id="B4FD98FB8B5BBBD94F6BDA0F41D1AC7C">
<mods:roleTerm id="57D4E3EA0BAB3CF9BAD6AAE95DF06E5B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="47AC8711A6A8CC76E2D622EB981A3B11">Robillard, Tony</mods:namePart>
</mods:name>
<mods:typeOfResource id="22444483DF7BE54C14D3D536C292007E">text</mods:typeOfResource>
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<mods:title id="413C0516175A6BF06CEECBF21B7C7AB6">Zootaxa</mods:title>
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<mods:part id="A8F29128E0A4B5E7252261DC3F44BD01">
<mods:date id="937F1B7AAFC29DAE3E6E66CA36D52BF1">2019</mods:date>
<mods:detail id="2A141CC35C131FDD74D851BF51826136" type="pubDate">
<mods:number id="4F24B7EEAC11826473F15099B2953821">2019-06-19</mods:number>
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<mods:number id="3DF298A917B0E49FEC71614A99333BBD">4619</mods:number>
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<mods:number id="4C6761559896ED004857860B2F08B55D">2</mods:number>
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<mods:classification id="D31BB53C85F44A857EE6C311F72918B8">journal article</mods:classification>
<mods:identifier id="2FC421232334B9F7B8C684CB0B3FE70D" type="CLB-Dataset">26448</mods:identifier>
<mods:identifier id="FDEA20A461E967E9B9C4488D2002514B" type="DOI">10.11646/zootaxa.4619.2.9</mods:identifier>
<mods:identifier id="AECCF924B6AAE9D87473CC0D99F8D843" type="GBIF-Dataset">1e87ced1-1631-463d-92d4-508d30dbbc48</mods:identifier>
<mods:identifier id="180C9D224F3772759CFFB794945FE520" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="C9C76FD830E07F8F57461DB2333849D8" type="Zenodo-Dep">3249675</mods:identifier>
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<treatment id="03FF87FDFFEFFFD84DD2D2C7FCEAFD2F" ID-DOI="http://doi.org/10.5281/zenodo.5690110" ID-GBIF-Taxon="158297477" ID-Zenodo-Dep="5690110" LSID="urn:lsid:plazi:treatment:03FF87FDFFEFFFD84DD2D2C7FCEAFD2F" httpUri="http://treatment.plazi.org/id/03FF87FDFFEFFFD84DD2D2C7FCEAFD2F" lastPageId="15" lastPageNumber="362" pageId="8" pageNumber="355">
<subSubSection id="C34C6560FFEFFFDF4DD2D2C7FCCDF8A9" box="[151,801,1767,1794]" pageId="8" pageNumber="355" type="nomenclature">
<paragraph id="8BE936EBFFEFFFDF4DD2D2C7FCCDF8A9" blockId="8.[151,801,1767,1829]" box="[151,801,1767,1794]" pageId="8" pageNumber="355">
<heading id="D0A18187FFEFFFDF4DD2D2C7FCCDF8A9" bold="true" box="[151,801,1767,1794]" fontSize="11" level="1" pageId="8" pageNumber="355" reason="1">
<taxonomicName id="4C564D68FFEFFFDF4DD2D2C7FCCDF8A9" authority="Robillard &amp; Gorochov, 2014" authorityName="Robillard &amp; Gorochov" authorityYear="2014" box="[151,801,1767,1794]" class="Insecta" family="Haglotettigoniidae" genus="Cardiodactylus" kingdom="Animalia" order="Orthoptera" pageId="8" pageNumber="355" phylum="Arthropoda" rank="species" species="borneoe">
<emphasis id="B922EAF9FFEFFFDF4DD2D2C7FCCDF8A9" bold="true" box="[151,801,1767,1794]" pageId="8" pageNumber="355">
<emphasis id="B922EAF9FFEFFFDF4DD2D2C7FE5BF8AA" bold="true" box="[151,439,1767,1794]" italics="true" pageId="8" pageNumber="355">Cardiodactylus borneoe</emphasis>
<bibRefCitation id="EFC74B1AFFEFFFDF4CFBD2C7FCCDF8A9" author="Robillard, T. &amp; Gorochov, A. V. &amp; Poulain, S. &amp; Suhardjono, Y. R." box="[446,801,1767,1793]" pageId="8" pageNumber="355" pagination="1 - 104" refId="ref5214" refString="Robillard, T., Gorochov, A. V., Poulain, S. &amp; Suhardjono, Y. R. (2014) Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa, 3854 (1), 1 - 104. https: // doi. org / 10.11646 / zootaxa. 3854.1.1" type="journal article" year="2014">Robillard &amp; Gorochov, 2014</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34C6560FFEFFFDF4DD2D32BFAAAF8C0" pageId="8" pageNumber="355" type="description">
<paragraph id="8BE936EBFFEFFFDF4DD2D32BFEC3F88D" blockId="8.[151,801,1767,1829]" box="[151,303,1803,1829]" pageId="8" pageNumber="355">
(
<figureCitation id="136D2A6EFFEFFFDF4DDAD32BFF11F88D" box="[159,253,1803,1829]" captionStart="FIGURE 11" captionStartId="13.[151,250,1968,1993]" captionTargetBox="[164,1424,166,1923]" captionTargetId="figure@13.[164,1424,166,1949]" captionTargetPageId="13" captionText="FIGURE 11. Cardiodactylus borneoe Robillard &amp; Gorochov, 2014 from Sandakan: female in its natural microhabitat when alive (A); male genitalia in dorsal (B), ventral (C) and lateral (D) views." httpUri="https://zenodo.org/record/3249695/files/figure.png" pageId="8" pageNumber="355">Figs. 11</figureCitation>
,
<figureCitation id="136D2A6EFFEFFFDF4C4CD32BFECAF88D" box="[265,294,1803,1829]" captionStart="FIGURE 12" captionStartId="14.[151,250,1952,1977]" captionTargetBox="[170,1428,221,1864]" captionTargetId="figure@14.[151,1436,219,1888]" captionTargetPageId="14" captionText="FIGURE 12. Calling song of Cardiodactylus borneoe Robillard &amp; Gorochov, 2014 from Sandakan: oscillogram of 3 echemes (A), detailed oscillogram of 1 echeme (B), spectrogram of 1 echeme (C), power spectrum (D)." httpUri="https://zenodo.org/record/3249697/files/figure.png" pageId="8" pageNumber="355">12</figureCitation>
)
</paragraph>
<paragraph id="8BE936EBFFEFFFDF4DD2D36FFAAAF8C0" blockId="8.[151,1350,1871,1896]" box="[151,1350,1871,1896]" pageId="8" pageNumber="355">
<bibRefCitation id="EFC74B1AFFEFFFDF4DD2D36FFE75F8CF" author="Robillard, T. &amp; Gorochov, A. V. &amp; Poulain, S. &amp; Suhardjono, Y. R." box="[151,409,1871,1896]" pageId="8" pageNumber="355" pagination="1 - 104" refId="ref5214" refString="Robillard, T., Gorochov, A. V., Poulain, S. &amp; Suhardjono, Y. R. (2014) Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa, 3854 (1), 1 - 104. https: // doi. org / 10.11646 / zootaxa. 3854.1.1" type="journal article" year="2014">
Robillard
<emphasis id="B922EAF9FFEFFFDF4DB8D370FEDDF8CF" box="[253,305,1871,1895]" italics="true" pageId="8" pageNumber="355">et al.</emphasis>
, 2014: 12
</bibRefCitation>
;
<bibRefCitation id="EFC74B1AFFEFFFDF4CE6D370FD82F8C0" author="Dong, J. &amp; Kergoat, G. J. &amp; Vicente, N. &amp; Rahmadi, C. &amp; Xu, S. &amp; Robillard, T." box="[419,622,1871,1896]" pageId="8" pageNumber="355" pagination="1 - 14" refId="ref4849" refString="Dong, J., Kergoat, G. J., Vicente, N., Rahmadi, C., Xu, S. &amp; Robillard, T. (2018) Biogeographic patterns and diversification dynamics of the genus Cardiodactylus Saussure (Orthoptera, Grylloidea, Eneopterinae) in Southeast Asia. Molecular Phylogenetics and Evolution, 129, 1 - 14. https: // doi. org / 10.1016 / j. ympev. 2018.06.001" type="journal article" year="2018">
Dong
<emphasis id="B922EAF9FFEFFFDF4CA7D370FDF9F8CF" box="[482,533,1871,1895]" italics="true" pageId="8" pageNumber="355">et al.</emphasis>
, 2018: 6
</bibRefCitation>
(molecular phylogeny);
<bibRefCitation id="EFC74B1AFFEFFFDF4E2CD36FFBD5F8CF" author="Cigliano, M. M. &amp; Braun, H. &amp; Eades, D. C. &amp; Otte, D." box="[873,1081,1871,1896]" pageId="8" pageNumber="355" refId="ref4692" refString="Cigliano, M. M., Braun, H., Eades, D. C. &amp; Otte, D. (2019) Orthoptera Species File Online. Version 5 (5.0). Available from: http: // orthoptera. speciesfile. org / HomePage / Orthoptera / HomePage. aspx (accessed 8 March 2019)" type="url" year="2019">
Cigliano
<emphasis id="B922EAF9FFEFFFDF4E82D370FC16F8CF" box="[967,1018,1871,1895]" italics="true" pageId="8" pageNumber="355">et al.</emphasis>
, 2019
</bibRefCitation>
(
<taxonomicName id="4C564D68FFEFFFDF4903D36FFB5AF8C0" box="[1094,1206,1871,1896]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="8" pageNumber="355" phylum="Arthropoda" rank="order">Orthoptera</taxonomicName>
Species File).
</paragraph>
</subSubSection>
<subSubSection id="C34C6560FFEFFFD84DD2D3B5FD0DFE7B" lastPageId="15" lastPageNumber="362" pageId="8" pageNumber="355" type="materials_examined">
<paragraph id="8BE936EBFFEFFFD84DD2D3B5FEF5FF53" blockId="8.[151,1437,1941,2040]" lastBlockId="15.[151,1437,152,647]" lastPageId="15" lastPageNumber="362" pageId="8" pageNumber="355">
<emphasis id="B922EAF9FFEFFFDF4DD2D3B5FE68F818" bold="true" box="[151,388,1941,1968]" pageId="8" pageNumber="355">Material examined:</emphasis>
<materialsCitation id="3B3E3CB6FFEFFFDF4CC9D3B5FBF2F818" ID-GBIF-Occurrence="2268317743" box="[396,1054,1941,1968]" collectorName="Rainforest Discovery Centre" country="Malaysia" location="Sandakan" pageId="8" pageNumber="355" specimenCount="1" stateProvince="Sabah">
<collectingCountry id="F341767BFFEFFFDF4CC9D3B5FE15F818" box="[396,505,1941,1968]" name="Malaysia" pageId="8" pageNumber="355">
<emphasis id="B922EAF9FFEFFFDF4CC9D3B5FE15F818" bold="true" box="[396,505,1941,1968]" pageId="8" pageNumber="355">Malaysia</emphasis>
</collectingCountry>
,
<collectingRegion id="4992F809FFEFFFDF4F4DD3B6FDA3F818" box="[520,591,1942,1968]" country="Malaysia" name="Sabah" pageId="8" pageNumber="355">Sabah</collectingRegion>
,
<location id="8E896030FFEFFFDF4F19D3B6FD20F818" LSID="urn:lsid:plazi:treatment:03FF87FDFFEFFFD84DD2D2C7FCEAFD2F:8E896030FFEFFFDF4F19D3B6FD20F818" box="[604,716,1942,1968]" country="Malaysia" name="Sandakan" pageId="8" pageNumber="355" stateProvince="Sabah">Sandakan</location>
,
<collectorName id="26A3533DFFEFFFDF4F9DD3B6FBF2F818" box="[728,1054,1942,1968]" pageId="8" pageNumber="355">Rainforest Discovery Centre</collectorName>
</materialsCitation>
,
<materialsCitation id="3B3E3CB6FFEFFFDF496FD3B6FADEF87C" ID-GBIF-Occurrence="2268317735" collectingDate="2019-01-08" collectionCode="SDK, ZRC" collectorName="M. K. Tan &amp; S. T. Toh" country="Malaysia" elevation="5" latitude="5.8741" longLatPrecision="1" longitude="117.94366" pageId="8" pageNumber="355" specimenCount="1" specimenCount-female="1" stateProvince="Sabah">
<specimenCount id="9D50FD62FFEFFFDF496FD3B6FB60F818" box="[1066,1164,1942,1968]" pageId="8" pageNumber="355" type="female">1 female</specimenCount>
(
<collectionCode id="ED47AE2EFFEFFFDF49D9D3B5FB36F818" box="[1180,1242,1941,1968]" pageId="8" pageNumber="355">SDK</collectionCode>
.19.6),
<geoCoordinate id="EE62502CFFEFFFDF4863D3B5FA74F818" box="[1318,1432,1941,1968]" degrees="5.87410" direction="north" orientation="latitude" pageId="8" pageNumber="355" precision="1" value="5.8741">N5.87410</geoCoordinate>
,
<geoCoordinate id="EE62502CFFEFFFDF4DD2D399FECEF87B" box="[151,290,1977,2003]" degrees="117.94366" direction="east" orientation="longitude" pageId="8" pageNumber="355" precision="1" value="117.94366">E117.94366</geoCoordinate>
, 50.2±
<elevation id="007BD1D8FFEFFFDF4C2BD399FE5EF87C" box="[366,434,1977,2004]" metricMagnitude="0" metricUnit="m" metricValue="5.1" pageId="8" pageNumber="355" unit="m" value="5.1">
<quantity id="4CAE9B0EFFEFFFDF4C2BD399FE5EF87C" box="[366,434,1977,2004]" metricMagnitude="0" metricUnit="m" metricValue="5.1" pageId="8" pageNumber="355" unit="m" value="5.1">5.1 m</quantity>
</elevation>
.a.s.l.,
<collectingDate id="EFACE9C3FFEFFFDF4CB6D39AFD73F87C" box="[499,671,1978,2004]" pageId="8" pageNumber="355" value="2019-01-08">
<date id="FFE8102BFFEFFFDF4CB6D39AFD73F87C" box="[499,671,1978,2004]" pageId="8" pageNumber="355" value="2019-01-08">8 January 2019</date>
</collectingDate>
, 2102 hours, on foliage, coll.
<collectorName id="26A3533DFFEFFFDF4EA0D39AFBBBF87C" box="[997,1111,1978,2004]" pageId="8" pageNumber="355">M. K. Tan</collectorName>
&amp;
<collectorName id="26A3533DFFEFFFDF493DD39AFB0CF87C" box="[1144,1248,1978,2004]" pageId="8" pageNumber="355">S. T. Toh</collectorName>
(
<collectionCode id="ED47AE2EFFEFFFDF49ABD39AFAC6F87C" box="[1262,1322,1978,2004]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="8" pageNumber="355">ZRC</collectionCode>
)
</materialsCitation>
<materialsCitation id="3B3E3CB6FFEFFFD84877D39AFD1EFF1B" ID-GBIF-Occurrence="2268317742" collectingDate="2019-01-09" collectionCode="SDK, FRC" collectorName="M. K. Tan &amp; R. Japir &amp; Momin Binti &amp; J. Y. Lee" country="Malaysia" elevation="5" lastPageId="15" lastPageNumber="362" latitude="5.87542" longLatPrecision="1" longitude="117.94231" pageId="8" pageNumber="355" specimenCount="1" specimenCount-female="1" stateProvince="Sabah">
;
<specimenCount id="9D50FD62FFEFFFDF4879D39AFA70F87C" box="[1340,1436,1978,2004]" pageId="8" pageNumber="355" type="female">1 female</specimenCount>
(
<collectionCode id="ED47AE2EFFEFFFDF4DDAD3FEFF30F850" box="[159,220,2014,2040]" pageId="8" pageNumber="355">SDK</collectionCode>
.19.21,
<figureCitation id="136D2A6EFFEFFFDF4C75D3FEFE7BF850" box="[304,407,2014,2040]" captionStart="FIGURE 11" captionStartId="13.[151,250,1968,1993]" captionTargetBox="[164,1424,166,1923]" captionTargetId="figure@13.[164,1424,166,1949]" captionTargetPageId="13" captionText="FIGURE 11. Cardiodactylus borneoe Robillard &amp; Gorochov, 2014 from Sandakan: female in its natural microhabitat when alive (A); male genitalia in dorsal (B), ventral (C) and lateral (D) views." httpUri="https://zenodo.org/record/3249695/files/figure.png" pageId="8" pageNumber="355">Fig. 11A</figureCitation>
)
<geoCoordinate id="EE62502CFFEFFFDF4CEFD3FDFDF0F850" box="[426,540,2013,2040]" degrees="5.87542" direction="north" orientation="latitude" pageId="8" pageNumber="355" precision="1" value="5.87542">N5.87542</geoCoordinate>
,
<geoCoordinate id="EE62502CFFEFFFDF4F6FD3FEFD59F850" box="[554,693,2014,2040]" degrees="117.94231" direction="east" orientation="longitude" pageId="8" pageNumber="355" precision="1" value="117.94231">E117.94231</geoCoordinate>
, 31.1±
<elevation id="007BD1D8FFEFFFDF4E46D3FEFCA2F850" box="[771,846,2014,2040]" metricMagnitude="0" metricUnit="m" metricValue="4.8" pageId="8" pageNumber="355" unit="m" value="4.8">
<quantity id="4CAE9B0EFFEFFFDF4E46D3FEFCA2F850" box="[771,846,2014,2040]" metricMagnitude="0" metricUnit="m" metricValue="4.8" pageId="8" pageNumber="355" unit="m" value="4.8">4.8 m</quantity>
</elevation>
.a.s.l.,
<collectingDate id="EFACE9C3FFEFFFDF4ED6D3FEFBABF850" box="[915,1095,2014,2040]" pageId="8" pageNumber="355" value="2019-01-09">
<date id="FFE8102BFFEFFFDF4ED6D3FEFBABF850" box="[915,1095,2014,2040]" pageId="8" pageNumber="355" value="2019-01-09">9 January 2019</date>
</collectingDate>
, 1930 hours, on foliage, coll.
<collectorName id="26A3533DFFE8FFD84DD2D4B9FEFCFF1B" box="[151,272,153,179]" pageId="15" pageNumber="362">M. K. Tan</collectorName>
,
<collectorName id="26A3533DFFE8FFD84C59D4B9FE9AFF1B" box="[284,374,153,179]" pageId="15" pageNumber="362">R. Japir</collectorName>
,
<collectorName id="26A3533DFFE8FFD84CC7D4B9FDF8FF1B" box="[386,532,153,179]" pageId="15" pageNumber="362">Momin Binti</collectorName>
&amp;
<collectorName id="26A3533DFFE8FFD84F7CD4B9FD73FF1B" box="[569,671,153,179]" pageId="15" pageNumber="362">J. Y. Lee</collectorName>
(
<collectionCode id="ED47AE2EFFE8FFD84FEAD4B9FD06FF1B" box="[687,746,153,179]" country="USA" httpUri="http://grbio.org/cool/z9c1-g2b0" name="Fusarium Research Center" pageId="15" pageNumber="362">FRC</collectionCode>
)
</materialsCitation>
<materialsCitation id="3B3E3CB6FFE8FFD84FB7D4B9FEF9FF53" ID-GBIF-Occurrence="2268317741" collectingDate="2019-01-09" collectionCode="SDK, ZRC" collectorName="M. K. Tan &amp; R. Japir &amp; Momin Binti &amp; J. Y. Lee" country="Malaysia" elevation="5" latitude="5.87547" longLatPrecision="1" longitude="117.9411" pageId="15" pageNumber="362" specimenCount="1" specimenCount-male="1" stateProvince="Sabah">
;
<specimenCount id="9D50FD62FFE8FFD84FBBD4B9FCA6FF1B" box="[766,842,153,179]" pageId="15" pageNumber="362" type="male">1 male</specimenCount>
(
<collectionCode id="ED47AE2EFFE8FFD84E1FD4B9FC7AFF1B" box="[858,918,153,179]" pageId="15" pageNumber="362">SDK</collectionCode>
.19.36,
<figureCitation id="136D2A6EFFE8FFD84EADD4B9FBD8FF1B" box="[1000,1076,153,179]" captionStart="FIGURE 9" captionStartId="11.[151,250,2000,2025]" captionTargetBox="[151,1436,221,1944]" captionTargetId="figure@11.[151,1436,221,1944]" captionTargetPageId="11" captionText="FIGURE 9. Lebinthus sandakan sp. nov. juveniles in their natural microhabitats when alive." httpUri="https://zenodo.org/record/3249691/files/figure.png" pageId="15" pageNumber="362">Figs. 9</figureCitation>
B–9D),
<geoCoordinate id="EE62502CFFE8FFD849CAD4B8FAEEFF1B" box="[1167,1282,152,179]" degrees="5.87547" direction="north" orientation="latitude" pageId="15" pageNumber="362" precision="1" value="5.87547">N5.87547</geoCoordinate>
,
<geoCoordinate id="EE62502CFFE8FFD84848D4B9FA74FF1B" box="[1293,1432,153,179]" degrees="117.94110" direction="east" orientation="longitude" pageId="15" pageNumber="362" precision="1" value="117.9411">E117.94110</geoCoordinate>
, 34.1±
<elevation id="007BD1D8FFE8FFD84D92D49CFEF3FF7F" box="[215,287,188,215]" metricMagnitude="0" metricUnit="m" metricValue="5.4" pageId="15" pageNumber="362" unit="m" value="5.4">
<quantity id="4CAE9B0EFFE8FFD84D92D49CFEF3FF7F" box="[215,287,188,215]" metricMagnitude="0" metricUnit="m" metricValue="5.4" pageId="15" pageNumber="362" unit="m" value="5.4">5.4 m</quantity>
</elevation>
.a.s.l.,
<collectingDate id="EFACE9C3FFE8FFD84C24D49DFDE3FF7F" box="[353,527,189,215]" pageId="15" pageNumber="362" value="2019-01-09">
<date id="FFE8102BFFE8FFD84C24D49DFDE3FF7F" box="[353,527,189,215]" pageId="15" pageNumber="362" value="2019-01-09">9 January 2019</date>
</collectingDate>
, 2046 hours, under foliage calling, coll.
<collectorName id="26A3533DFFE8FFD84E96D49DFBA4FF7F" box="[979,1096,189,215]" pageId="15" pageNumber="362">M. K. Tan</collectorName>
,
<collectorName id="26A3533DFFE8FFD84916D49DFB40FF7F" box="[1107,1196,189,215]" pageId="15" pageNumber="362">R. Japir</collectorName>
,
<collectorName id="26A3533DFFE8FFD849F2D49DFAA5FF7F" box="[1207,1353,189,215]" pageId="15" pageNumber="362">Momin Binti</collectorName>
&amp;
<collectorName id="26A3533DFFE8FFD84828D49DFF2DFF53" pageId="15" pageNumber="362">J. Y. Lee</collectorName>
(
<collectionCode id="ED47AE2EFFE8FFD84D95D4C1FEE0FF53" box="[208,268,225,251]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="15" pageNumber="362">ZRC</collectionCode>
)
</materialsCitation>
.
</paragraph>
<caption id="DF296663FFEEFFDE4DD2D3FFFB3CF850" box="[151,1232,2015,2040]" httpUri="https://zenodo.org/record/3249687/files/figure.png" pageId="9" pageNumber="356" startId="9.[151,250,2015,2040]" targetBox="[253,1334,181,1995]" targetPageId="9">
<paragraph id="8BE936EBFFEEFFDE4DD2D3FFFB3CF850" blockId="9.[151,1232,2015,2040]" box="[151,1232,2015,2040]" pageId="9" pageNumber="356">
<emphasis id="B922EAF9FFEEFFDE4DD2D3FFFEFFF850" bold="true" box="[151,275,2015,2040]" pageId="9" pageNumber="356">FIGURE 7.</emphasis>
<taxonomicName id="4C564D68FFEEFFDE4C5CD3FFFE09F85F" authority="Tan &amp; Japir &amp; Chung &amp; Robillard, 2019" authorityName="Tan &amp; Japir &amp; Chung &amp; Robillard" authorityYear="2019" box="[281,485,2015,2039]" class="Insecta" family="Haglotettigoniidae" genus="Lebinthus" kingdom="Animalia" order="Orthoptera" pageId="9" pageNumber="356" phylum="Arthropoda" rank="species" species="sandakan" status="sp. nov.">
<emphasis id="B922EAF9FFEEFFDE4C5CD3FFFE09F85F" box="[281,485,2015,2039]" italics="true" pageId="9" pageNumber="356">Lebinthus sandakan</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2115782FFEEFFDE4CAED3C0FDD2F850" box="[491,574,2016,2040]" pageId="9" pageNumber="356" rank="species">
<emphasis id="B922EAF9FFEEFFDE4CAED3C0FDD2F850" bold="true" box="[491,574,2016,2040]" pageId="9" pageNumber="356">sp. nov.</emphasis>
</taxonomicNameLabel>
males in their natural microhabitats when alive. Holotype is (B).
</paragraph>
</caption>
<caption id="DF296663FFEDFFDD4DD2D3F0FBD6F841" box="[151,1082,2000,2025]" httpUri="https://zenodo.org/record/3249689/files/figure.png" pageId="10" pageNumber="357" startId="10.[151,250,2000,2025]" targetBox="[289,1298,166,1980]" targetPageId="10">
<paragraph id="8BE936EBFFEDFFDD4DD2D3F0FBD6F841" blockId="10.[151,1082,2000,2025]" box="[151,1082,2000,2025]" pageId="10" pageNumber="357">
<emphasis id="B922EAF9FFEDFFDD4DD2D3F0FEFFF841" bold="true" box="[151,275,2000,2025]" pageId="10" pageNumber="357">FIGURE 8.</emphasis>
<taxonomicName id="4C564D68FFEDFFDD4C5CD3F0FE09F840" authority="Tan &amp; Japir &amp; Chung &amp; Robillard, 2019" authorityName="Tan &amp; Japir &amp; Chung &amp; Robillard" authorityYear="2019" box="[281,485,2000,2024]" class="Insecta" family="Haglotettigoniidae" genus="Lebinthus" kingdom="Animalia" order="Orthoptera" pageId="10" pageNumber="357" phylum="Arthropoda" rank="species" species="sandakan" status="sp. nov.">
<emphasis id="B922EAF9FFEDFFDD4C5CD3F0FE09F840" box="[281,485,2000,2024]" italics="true" pageId="10" pageNumber="357">Lebinthus sandakan</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2115782FFEDFFDD4CAED3F1FDD2F841" box="[491,574,2001,2025]" pageId="10" pageNumber="357" rank="species">
<emphasis id="B922EAF9FFEDFFDD4CAED3F1FDD2F841" bold="true" box="[491,574,2001,2025]" pageId="10" pageNumber="357">sp. nov.</emphasis>
</taxonomicNameLabel>
females in their natural microhabitats when alive.
</paragraph>
</caption>
<caption id="DF296663FFECFFDC4DD2D3F0FBABF841" box="[151,1095,2000,2025]" httpUri="https://zenodo.org/record/3249691/files/figure.png" pageId="11" pageNumber="358" startId="11.[151,250,2000,2025]" targetBox="[151,1436,221,1944]" targetPageId="11">
<paragraph id="8BE936EBFFECFFDC4DD2D3F0FBABF841" blockId="11.[151,1095,2000,2025]" box="[151,1095,2000,2025]" pageId="11" pageNumber="358">
<emphasis id="B922EAF9FFECFFDC4DD2D3F0FEFFF841" bold="true" box="[151,275,2000,2025]" pageId="11" pageNumber="358">FIGURE 9.</emphasis>
<taxonomicName id="4C564D68FFECFFDC4C5CD3F1FE09F841" authority="Tan &amp; Japir &amp; Chung &amp; Robillard, 2019" authorityName="Tan &amp; Japir &amp; Chung &amp; Robillard" authorityYear="2019" box="[281,485,2001,2025]" class="Insecta" family="Haglotettigoniidae" genus="Lebinthus" kingdom="Animalia" order="Orthoptera" pageId="11" pageNumber="358" phylum="Arthropoda" rank="species" species="sandakan" status="sp. nov.">
<emphasis id="B922EAF9FFECFFDC4C5CD3F1FE09F841" box="[281,485,2001,2025]" italics="true" pageId="11" pageNumber="358">Lebinthus sandakan</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2115782FFECFFDC4CAED3F1FDD2F841" box="[491,574,2001,2025]" pageId="11" pageNumber="358" rank="species">
<emphasis id="B922EAF9FFECFFDC4CAED3F1FDD2F841" bold="true" box="[491,574,2001,2025]" pageId="11" pageNumber="358">sp. nov.</emphasis>
</taxonomicNameLabel>
juveniles in their natural microhabitats when alive.
</paragraph>
</caption>
<caption id="DF296663FFEBFFDB4DD2D355FEFFF87E" httpUri="https://zenodo.org/record/3249693/files/figure.png" pageId="12" pageNumber="359" startId="12.[151,250,1909,1934]" targetBox="[180,1417,222,1824]" targetPageId="12">
<paragraph id="8BE936EBFFEBFFDB4DD2D355FEFFF87E" blockId="12.[151,1437,1909,2006]" pageId="12" pageNumber="359">
<emphasis id="B922EAF9FFEBFFDB4DD2D355FECEF826" bold="true" box="[151,290,1909,1934]" pageId="12" pageNumber="359">FIGURE 10.</emphasis>
Calling song of
<taxonomicName id="4C564D68FFEBFFDB4C90D356FD4FF826" authority="Tan &amp; Japir &amp; Chung &amp; Robillard, 2019" authorityName="Tan &amp; Japir &amp; Chung &amp; Robillard" authorityYear="2019" box="[469,675,1910,1934]" class="Insecta" family="Haglotettigoniidae" genus="Lebinthus" kingdom="Animalia" order="Orthoptera" pageId="12" pageNumber="359" phylum="Arthropoda" rank="species" species="sandakan" status="sp. nov.">
<emphasis id="B922EAF9FFEBFFDB4C90D356FD4FF826" box="[469,675,1910,1934]" italics="true" pageId="12" pageNumber="359">Lebinthus sandakan</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2115782FFEBFFDB4FE9D356FCEDF826" box="[684,769,1910,1934]" pageId="12" pageNumber="359" rank="species">
<emphasis id="B922EAF9FFEBFFDB4FE9D356FCEDF826" bold="true" box="[684,769,1910,1934]" pageId="12" pageNumber="359">sp. nov.</emphasis>
</taxonomicNameLabel>
of paratype: oscillogram of 1 echeme (A), detailed oscillogram of trill part of echeme (B) and corresponding spectrogram (C); power spectra of beginning of syllable (D), of ending part of syllable (E).
</paragraph>
</caption>
<caption id="DF296663FFEAFFDA4DD2D390FC9EF845" httpUri="https://zenodo.org/record/3249695/files/figure.png" pageId="13" pageNumber="360" startId="13.[151,250,1968,1993]" targetBox="[164,1424,166,1923]" targetPageId="13">
<paragraph id="8BE936EBFFEAFFDA4DD2D390FC9EF845" blockId="13.[151,1436,1968,2029]" pageId="13" pageNumber="360">
<emphasis id="B922EAF9FFEAFFDA4DD2D390FECDF861" bold="true" box="[151,289,1968,1993]" pageId="13" pageNumber="360">FIGURE 11.</emphasis>
<taxonomicName id="4C564D68FFEAFFDA4C6FD391FCBFF861" authority="Robillard &amp; Gorochov, 2014" authorityName="Robillard &amp; Gorochov" authorityYear="2014" box="[298,851,1968,1993]" class="Insecta" family="Haglotettigoniidae" genus="Cardiodactylus" kingdom="Animalia" order="Orthoptera" pageId="13" pageNumber="360" phylum="Arthropoda" rank="species" species="borneoe">
<emphasis id="B922EAF9FFEAFFDA4C6FD391FDCDF861" box="[298,545,1969,1993]" italics="true" pageId="13" pageNumber="360">Cardiodactylus borneoe</emphasis>
<bibRefCitation id="EFC74B1AFFEAFFDA4F6CD390FCBFF861" author="Robillard, T. &amp; Gorochov, A. V. &amp; Poulain, S. &amp; Suhardjono, Y. R." box="[553,851,1968,1993]" pageId="13" pageNumber="360" pagination="1 - 104" refId="ref5214" refString="Robillard, T., Gorochov, A. V., Poulain, S. &amp; Suhardjono, Y. R. (2014) Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa, 3854 (1), 1 - 104. https: // doi. org / 10.11646 / zootaxa. 3854.1.1" type="journal article" year="2014">Robillard &amp; Gorochov, 2014</bibRefCitation>
</taxonomicName>
from Sandakan: female in its natural microhabitat when alive (A); male genitalia in dorsal (B), ventral (C) and lateral (D) views.
</paragraph>
</caption>
<caption id="DF296663FFE9FFD94DD2D380FBBFF875" httpUri="https://zenodo.org/record/3249697/files/figure.png" pageId="14" pageNumber="361" startId="14.[151,250,1952,1977]" targetBox="[170,1428,221,1864]" targetPageId="14">
<paragraph id="8BE936EBFFE9FFD94DD2D380FBBFF875" blockId="14.[151,1436,1952,2013]" pageId="14" pageNumber="361">
<emphasis id="B922EAF9FFE9FFD94DD2D380FECCF811" bold="true" box="[151,288,1952,1977]" pageId="14" pageNumber="361">FIGURE 12.</emphasis>
Calling song of
<taxonomicName id="4C564D68FFE9FFD94C8CD381FC07F811" authority="Robillard &amp; Gorochov, 2014" authorityName="Robillard &amp; Gorochov" authorityYear="2014" box="[457,1003,1952,1977]" class="Insecta" family="Haglotettigoniidae" genus="Cardiodactylus" kingdom="Animalia" order="Orthoptera" pageId="14" pageNumber="361" phylum="Arthropoda" rank="species" species="borneoe">
<emphasis id="B922EAF9FFE9FFD94C8CD381FD53F811" box="[457,703,1953,1977]" italics="true" pageId="14" pageNumber="361">Cardiodactylus borneoe</emphasis>
<bibRefCitation id="EFC74B1AFFE9FFD94F80D380FC07F811" author="Robillard, T. &amp; Gorochov, A. V. &amp; Poulain, S. &amp; Suhardjono, Y. R." box="[709,1003,1952,1977]" pageId="14" pageNumber="361" pagination="1 - 104" refId="ref5214" refString="Robillard, T., Gorochov, A. V., Poulain, S. &amp; Suhardjono, Y. R. (2014) Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa, 3854 (1), 1 - 104. https: // doi. org / 10.11646 / zootaxa. 3854.1.1" type="journal article" year="2014">Robillard &amp; Gorochov, 2014</bibRefCitation>
</taxonomicName>
from Sandakan: oscillogram of 3 echemes (A), detailed oscillogram of 1 echeme (B), spectrogram of 1 echeme (C), power spectrum (D).
</paragraph>
</caption>
<paragraph id="8BE936EBFFE8FFD84D82D525FBDFFE23" blockId="15.[151,1437,152,647]" pageId="15" pageNumber="362">
Calling song (
<figureCitation id="136D2A6EFFE8FFD84C22D525FE50FEB7" box="[359,444,261,287]" captionStart="FIGURE 12" captionStartId="14.[151,250,1952,1977]" captionTargetBox="[170,1428,221,1864]" captionTargetId="figure@14.[151,1436,219,1888]" captionTargetPageId="14" captionText="FIGURE 12. Calling song of Cardiodactylus borneoe Robillard &amp; Gorochov, 2014 from Sandakan: oscillogram of 3 echemes (A), detailed oscillogram of 1 echeme (B), spectrogram of 1 echeme (C), power spectrum (D)." httpUri="https://zenodo.org/record/3249697/files/figure.png" pageId="15" pageNumber="362">Fig. 12</figureCitation>
): At 28.4°C and RH 74%, the calling song of
<taxonomicName id="4C564D68FFE8FFD84E83D525FBA8FEB7" authorityName="Robillard &amp; Gorochov" authorityYear="2014" box="[966,1092,261,287]" class="Insecta" family="Haglotettigoniidae" genus="Cardiodactylus" kingdom="Animalia" order="Orthoptera" pageId="15" pageNumber="362" phylum="Arthropoda" rank="species" species="borneoe">
<emphasis id="B922EAF9FFE8FFD84E83D525FBA8FEB7" box="[966,1092,261,287]" italics="true" pageId="15" pageNumber="362">C. borneoe</emphasis>
</taxonomicName>
consists of doublets (or rarely triplets) of long syllables including strong amplitude modulation forming pulse-like structures. Echemes (doublet) last 660 ± 75 ms with a period of 3.5 ± 1.2 s. Within doublets, syllables last 196 ± 43 ms, with a period of 372 ± 47 ms. Dominant frequency value is 17.6 ± 0.4 kHz with no clear harmonic structure.
</paragraph>
<paragraph id="8BE936EBFFE8FFD84D82D5B5FD0DFE7B" blockId="15.[151,1437,152,647]" pageId="15" pageNumber="362">
Habitat and life history traits.
<taxonomicName id="4C564D68FFE8FFD84F5DD5B5FD79FE07" authorityName="Robillard &amp; Gorochov" authorityYear="2014" box="[536,661,405,431]" class="Insecta" family="Haglotettigoniidae" genus="Cardiodactylus" kingdom="Animalia" order="Orthoptera" pageId="15" pageNumber="362" phylum="Arthropoda" rank="species" species="borneoe">
<emphasis id="B922EAF9FFE8FFD84F5DD5B5FD79FE07" box="[536,661,405,431]" italics="true" pageId="15" pageNumber="362">C. borneoe</emphasis>
</taxonomicName>
inhabits the dipterocarp forest in Sandakan and is typically found on or under the foliage of tree sapling or shrub at night.
</paragraph>
</subSubSection>
<subSubSection id="C34C6560FFE8FFD84D82D5FDFCEAFD2F" pageId="15" pageNumber="362" type="discussion">
<paragraph id="8BE936EBFFE8FFD84D82D5FDFCEAFD2F" blockId="15.[151,1437,152,647]" pageId="15" pageNumber="362">
<emphasis id="B922EAF9FFE8FFD84D82D5FDFEDCFE5F" bold="true" box="[199,304,477,503]" pageId="15" pageNumber="362">Remark.</emphasis>
The calling song of
<taxonomicName id="4C564D68FFE8FFD84F54D5FDFD61FE5F" authorityName="Robillard &amp; Gorochov" authorityYear="2014" box="[529,653,477,503]" class="Insecta" family="Haglotettigoniidae" genus="Cardiodactylus" kingdom="Animalia" order="Orthoptera" pageId="15" pageNumber="362" phylum="Arthropoda" rank="species" species="borneoe">
<emphasis id="B922EAF9FFE8FFD84F54D5FDFD61FE5F" box="[529,653,477,503]" italics="true" pageId="15" pageNumber="362">C. borneoe</emphasis>
</taxonomicName>
differs from that of
<taxonomicName id="4C564D68FFE8FFD84E2FD5FDFA70FE5F" authority="Tan &amp; Robillard, 2014" authorityName="Tan &amp; Robillard" authorityYear="2014" box="[874,1436,477,503]" class="Insecta" family="Haglotettigoniidae" genus="Cardiodactylus" kingdom="Animalia" order="Orthoptera" pageId="15" pageNumber="362" phylum="Arthropoda" rank="species" species="admirabilis">
<emphasis id="B922EAF9FFE8FFD84E2FD5FDFB70FE5F" box="[874,1180,477,503]" italics="true" pageId="15" pageNumber="362">Cardiodactylus admirabilis</emphasis>
<bibRefCitation id="EFC74B1AFFE8FFD849E7D5FDFA70FE5F" author="Tan, M. K. &amp; Robillard, T." box="[1186,1436,477,503]" pageId="15" pageNumber="362" pagination="364 - 376" refId="ref5373" refString="Tan, M. K. &amp; Robillard, T. (2014) A new species of Cardiodactylus (Orthoptera: Grylloidea: Eneopterinae) from Singapore. Zootaxa, 3764 (3), 364 - 376. http: // dx. doi. org / 10.11646 / zootaxa. 3764.3.6" type="journal article" year="2014">Tan &amp; Robillard, 2014</bibRefCitation>
</taxonomicName>
from
<collectingCountry id="F341767BFFE8FFD84D91D621FEAAFDB3" box="[212,326,513,539]" name="Singapore" pageId="15" pageNumber="362">Singapore</collectingCountry>
by consisting of doublets with longer echeme duration (instead of triplets with shorter echeme duration of 205.6 ms) and higher dominant frequency (instead of 14.3 kHz). This is despite of the similarities in the morphology and genitalia of these two species. Another morphologically similar species is
<taxonomicName id="4C564D68FFE8FFD84930D669FEAAFD2F" authority="Robillard, 2011" authorityName="Robillard" authorityYear="2011" class="Insecta" family="Haglotettigoniidae" genus="Cardiodactylus" kingdom="Animalia" order="Orthoptera" pageId="15" pageNumber="362" phylum="Arthropoda" rank="species" species="thailandia">
<emphasis id="B922EAF9FFE8FFD84930D669FA70FDCB" box="[1141,1436,585,611]" italics="true" pageId="15" pageNumber="362">Cardiodactylus thailandia</emphasis>
Robillard, 2011
</taxonomicName>
from
<collectingCountry id="F341767BFFE8FFD84CCFD64DFE02FD2F" box="[394,494,621,647]" name="Thailand" pageId="15" pageNumber="362">Thailand</collectingCountry>
but its song is unknown.
</paragraph>
</subSubSection>
</treatment>
</document>
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