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<document id="62763E8D805630925F28276C05F3E08F" ID-DOI="10.21248/contrib.entomol.64.1.1-28" ID-ISSN="0005-805X" ID-Zenodo-Dep="4753531" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1620837658217" checkinUser="carolina" docAuthor="Assing, Volker" docDate="2014" docId="03FF87F6FFB1FFA2FF0E88298A86F898" docLanguage="en" docName="ContribEntomol.64.1.1-28.pdf" docOrigin="Beiträge Zur Entomologie = Contributions to Entomology 64 (1)" docSource="https://www.contributions-to-entomology.org/article/view/1848" docStyle="DocumentStyle:05A48F5F6902F879680B6E64765AA512.2:ContribEntomol.2013-2016.journal_article" docStyleId="05A48F5F6902F879680B6E64765AA512" docStyleName="ContribEntomol.2013-2016.journal_article" docStyleVersion="2" docTitle="Lathrobium imberbe Assing 2014, sp. n." docType="treatment" docVersion="3" lastPageNumber="21" masterDocId="FFC6FF8EFFA5FFB6FFED8A3A8E48FFB9" masterDocTitle="New species and records of Lathrobium from the Palaearctic region, primarily from Nepal (Coleoptera: Staphylinidae: Paederinae)" masterLastPageNumber="28" masterPageNumber="1" pageNumber="21" updateTime="1698916626789" updateUser="plazi">
<mods:mods id="CCB562C96B125D54397CAF17238879D6" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="CDC945A135FE48D64A79826ACE481812">
<mods:title id="A6E7A60B51715416FE959C282A336BA4">New species and records of Lathrobium from the Palaearctic region, primarily from Nepal (Coleoptera: Staphylinidae: Paederinae)</mods:title>
</mods:titleInfo>
<mods:name id="A10AE638B57BC808DCE9A1654953D635" type="personal">
<mods:role id="191195E8856E34A83F3D787A0999EEE3">
<mods:roleTerm id="9A11097D7C32523A803AB3C8F72F2BD2">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5691448E8E03FE3E2EEF81328F7C11D5">Assing, Volker</mods:namePart>
</mods:name>
<mods:typeOfResource id="1857D4ED6574CDE8D2124970CE48EF23">text</mods:typeOfResource>
<mods:relatedItem id="2DE4197E82CA5839E8A8AC9AE3C91245" type="host">
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<mods:title id="5806E5745D968E7B3202A0116170BCD7">Beiträge Zur Entomologie = Contributions to Entomology</mods:title>
</mods:titleInfo>
<mods:part id="1A524B960B55D6EA5EC6DFDF4F353754">
<mods:date id="A087F2E535F295076E3648E5A25683E1">2014</mods:date>
<mods:detail id="E746D882B2230AD8CD842F6CBF24720F" type="pubDate">
<mods:number id="7B53842D83606BDFA621990EAB6036BE">2014-07-31</mods:number>
</mods:detail>
<mods:detail id="6CC639926BAA2B72EBC7098307D1FF26" type="volume">
<mods:number id="DD307061C37B7C2ACA3128CA658782B2">64</mods:number>
</mods:detail>
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<mods:number id="6BEBCFB820E8C79D9349BE26D108A648">1</mods:number>
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<mods:start id="5B3FE8ABF8180324A79EFB5E9FA60039">1</mods:start>
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<mods:url id="D8A001EAC0CF6C6D87CF7D8396F3E129">https://www.contributions-to-entomology.org/article/view/1848</mods:url>
</mods:location>
<mods:classification id="BE85636FD3F5B5320E2BB1F7BC0054EE">journal article</mods:classification>
<mods:identifier id="00A58E98BFAEE5983A55D7231967F938" type="DOI">10.21248/contrib.entomol.64.1.1-28</mods:identifier>
<mods:identifier id="F96DAC1F5B0C977EF659D2E73849A951" type="ISSN">0005-805X</mods:identifier>
<mods:identifier id="0719F3AB0CF311A2E570CB7B8F160FB4" type="Zenodo-Dep">4753531</mods:identifier>
</mods:mods>
<treatment id="03FF87F6FFB1FFA2FF0E88298A86F898" LSID="urn:lsid:plazi:treatment:03FF87F6FFB1FFA2FF0E88298A86F898" httpUri="http://treatment.plazi.org/id/03FF87F6FFB1FFA2FF0E88298A86F898" lastPageNumber="21" pageId="20" pageNumber="21">
<subSubSection id="C34C656BFFB1FFA2FF0E88298FBAFD97" box="[227,498,531,558]" pageId="20" pageNumber="21" type="nomenclature">
<paragraph id="8BE936E0FFB1FFA2FF0E88298FBAFD97" blockId="20.[227,498,531,591]" box="[227,498,531,558]" pageId="20" pageNumber="21">
<heading id="D0A1818CFFB1FFA2FF0E88298FBAFD97" box="[227,498,531,558]" centered="true" fontSize="12" level="2" pageId="20" pageNumber="21" reason="2">
<taxonomicName id="4C564D63FFB1FFA2FF0E88298FFAFD97" ID-CoL="3SK25" authority="Assing, 2014" authorityName="Assing" authorityYear="2014" box="[227,434,531,558]" class="Insecta" family="Staphylinidae" genus="Lathrobium" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="imberbe" status="sp. nov.">
<emphasis id="B922EAF2FFB1FFA2FF0E88298FFAFD97" box="[227,434,531,558]" italics="true" pageId="20" pageNumber="21">Lathrobium imberbe</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2115789FFB1FFA2FE5488298FBAFD97" box="[441,498,531,558]" pageId="20" pageNumber="21" rank="species">sp. n.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34C656BFFB1FFA2FF0E88038F29FDF6" box="[227,353,569,591]" pageId="20" pageNumber="21" type="description">
<paragraph id="8BE936E0FFB1FFA2FF0E88038F29FDF6" blockId="20.[227,498,531,591]" box="[227,353,569,591]" pageId="20" pageNumber="21">
(
<figureCitation id="136D2A65FFB1FFA2FF0A88038F13FDF6" box="[231,347,569,591]" captionStart="Figs 115–129" captionStartId="21.[151,191,1967,1989]" captionTargetBox="[155,1394,219,1952]" captionTargetId="figure-117@21.[151,1398,219,1958]" captionTargetPageId="21" captionText="Figs 115–129: Lathrobium imberbe sp. n. (115–122) and L. aculeatum COIFFAIT (123–129): habitus (115, 123); forebody (116, 124); male sternite VII (117, 125); male sternite VIII (118, 126); aedeagus in lateral and in ventral view (119–120, 127–128); female sternite VIII (121, 129); female tergites IX–X (122). Scale bars: 115–116, 123–124: 1.0 mm; 117–122, 125–129: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4753558" httpUri="https://zenodo.org/record/4753558/files/figure.png" pageId="20" pageNumber="21">Figs 115–122</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C34C656BFFB1FFA2FF5088538D5BFCF8" pageId="20" pageNumber="21" type="materials_examined">
<paragraph id="8BE936E0FFB1FFA2FF5088538CC1FD58" blockId="20.[189,795,617,833]" pageId="20" pageNumber="21">
<emphasis id="B922EAF2FFB1FFA2FF5088538F1BFD38" bold="true" box="[189,339,617,641]" pageId="20" pageNumber="21">Type material</emphasis>
:
<materialsCitation id="3B3E3CBDFFB1FFA2FEB288538CC1FD58" collectingDate="1995-05-10" collectionCode="V, NME" collectorName="A. Weigel" country="Nepal" elevation="3400" location="Garpung Khola" municipality="Kagmara Lekh" pageId="20" pageNumber="21" specimenCount="2" stateProvince="Karnali" typeStatus="holotype">
<typeStatus id="54ED8842FFB1FFA2FEB288538F8AFD38" box="[351,450,617,641]" pageId="20" pageNumber="21" type="holotype">Holotype</typeStatus>
: “
<collectingCountry id="F3417670FFB1FFA2FE1C88538C0BFD38" box="[497,579,617,641]" name="Nepal" pageId="20" pageNumber="21">NEPAL</collectingCountry>
Prov.
<collectingRegion id="4992F802FFB1FFA2FD6988538C9DFD38" box="[644,725,617,641]" country="Nepal" name="Karnali" pageId="20" pageNumber="21">Karnali</collectingRegion>
, Distr. Dolpa,
<collectingMunicipality id="6B8DAC9AFFB1FFA2FEFF88B08FF0FD18" box="[274,440,649,674]" pageId="20" pageNumber="21">Kagmara Lekh</collectingMunicipality>
,
<location id="8E89603BFFB1FFA2FE2488B38C30FD18" LSID="urn:lsid:plazi:treatment:03FF87F6FFB1FFA2FF0E88298A86F898:8E89603BFFB1FFA2FE2488B38C30FD18" box="[457,632,649,673]" country="Nepal" municipality="Kagmara Lekh" name="Garpung Khola" pageId="20" pageNumber="21" stateProvince="Karnali">Garpung Khola</location>
,
<quantity id="4CAE9B05FFB1FFA2FD6788B38D52FD18" box="[650,794,649,673]" metricMagnitude="3" metricUnit="m" metricValue="3.4" metricValueMax="3.8" metricValueMin="3.0" pageId="20" pageNumber="21" unit="m" value="3400.0" valueMax="3800.0" valueMin="3000.0">
<elevation id="007BD1D3FFB1FFA2FD6788B38D52FD18" box="[650,794,649,673]" metricMagnitude="3" metricUnit="m" metricValue="3.4" metricValueMax="3.8" metricValueMin="3.0" pageId="20" pageNumber="21" unit="m" value="3400.0" valueMax="3800.0" valueMin="3000.0">3000–3800 m</elevation>
</quantity>
<date id="FFE81020FFB1FFA2FF5088938F6DFD78" box="[189,293,681,705]" pageId="20" pageNumber="21" value="1995-05-10">
<collectingDate id="EFACE9C8FFB1FFA2FF5088938F6DFD78" box="[189,293,681,705]" pageId="20" pageNumber="21" value="1995-05-10">10.V.1995</collectingDate>
</date>
leg.
<collectorName id="26A35336FFB1FFA2FE8A88938F9EFD78" box="[359,470,681,705]" pageId="20" pageNumber="21">A. Weigel</collectorName>
/
<typeStatus id="54ED8842FFB1FFA2FE1088938C24FD78" box="[509,620,681,705]" pageId="20" pageNumber="21" type="holotype">Holotypus</typeStatus>
<taxonomicName id="4C564D63FFB1FFA2FD7388938F59FD58" authority="Assing, 2014" authorityName="Assing" authorityYear="2014" class="Insecta" family="Staphylinidae" genus="Lathrobium" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="imberbe" status="sp. nov.">
<emphasis id="B922EAF2FFB1FFA2FD7388938F59FD58" italics="true" pageId="20" pageNumber="21">Lathrobium imberbe</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2115789FFB1FFA2FEFA88F08F07FD58" box="[279,335,714,737]" pageId="20" pageNumber="21" rank="species">sp. n.</taxonomicNameLabel>
, det.
<collectionCode id="ED47AE25FFB1FFA2FE6588F08FD2FD58" box="[392,410,714,737]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="20" pageNumber="21" type="Museum">V</collectionCode>
. Assing 2013” (
<collectionCode id="ED47AE25FFB1FFA2FDD088F38C36FD58" box="[573,638,713,737]" country="Germany" httpUri="http://grbio.org/cool/87iw-xebj" name="Sammlung des Naturkundemseum Erfurt" pageId="20" pageNumber="21">NME</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BE936E0FFB1FFA2FF5088D08D5BFCF8" blockId="20.[189,795,617,833]" pageId="20" pageNumber="21">
<materialsCitation id="3B3E3CBDFFB1FFA2FF5088D08D58FCF8" collectingDate="1995-05-12" collectionCode="NME" collectorName="J. Weipert" country="Nepal" elevation="3800" location="Bachtal" municipality="Kagmara Lekh" pageId="20" pageNumber="21" specimenCount="1" stateProvince="Karnali" typeStatus="paratype">
<typeStatus id="54ED8842FFB1FFA2FF5088D08F6FFCBB" box="[189,295,746,770]" pageId="20" pageNumber="21" type="paratype">Paratypes</typeStatus>
: 1 : same data as holotype (cAss); 1 : “
<collectingCountry id="F3417670FFB1FFA2FF2B89338F52FC98" box="[198,282,777,801]" name="Nepal" pageId="20" pageNumber="21">NEPAL</collectingCountry>
, Prov.
<collectingRegion id="4992F802FFB1FFA2FE9B89338F8FFC98" box="[374,455,777,801]" country="Nepal" name="Karnali" pageId="20" pageNumber="21">Karnali</collectingRegion>
, Garpung Khola E Hurikot,
<location id="8E89603BFFB1FFA2FF5089138F46FCF8" LSID="urn:lsid:plazi:treatment:03FF87F6FFB1FFA2FF0E88298A86F898:8E89603BFFB1FFA2FF5089138F46FCF8" box="[189,270,809,833]" country="Nepal" name="Bachtal" pageId="20" pageNumber="21" stateProvince="Karnali">Bachtal</location>
,
<quantity id="4CAE9B05FFB1FFA2FEF589138F20FCF8" box="[280,360,809,833]" metricMagnitude="3" metricUnit="m" metricValue="3.8" pageId="20" pageNumber="21" unit="m" value="3800.0">
<elevation id="007BD1D3FFB1FFA2FEF589138F20FCF8" box="[280,360,809,833]" metricMagnitude="3" metricUnit="m" metricValue="3.8" pageId="20" pageNumber="21" unit="m" value="3800.0">3800 m</elevation>
</quantity>
NN,
<date id="FFE81020FFB1FFA2FE4C89138C43FCF8" box="[417,523,809,833]" pageId="20" pageNumber="21" value="1995-05-12">
<collectingDate id="EFACE9C8FFB1FFA2FE4C89138C43FCF8" box="[417,523,809,833]" pageId="20" pageNumber="21" value="1995-05-12">12.V.1995</collectingDate>
</date>
, leg.:
<collectorName id="26A35336FFB1FFA2FDAB89108CF8FCF8" box="[582,688,809,833]" pageId="20" pageNumber="21">J. Weipert</collectorName>
” (
<collectionCode id="ED47AE25FFB1FFA2FD2A89138D40FCF8" box="[711,776,809,833]" country="Germany" httpUri="http://grbio.org/cool/87iw-xebj" name="Sammlung des Naturkundemseum Erfurt" pageId="20" pageNumber="21">NME</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34C656BFFB1FFA2FF5089538CFFFC78" pageId="20" pageNumber="21" type="etymology">
<paragraph id="8BE936E0FFB1FFA2FF5089538CFFFC78" blockId="20.[189,794,873,961]" pageId="20" pageNumber="21">
<emphasis id="B922EAF2FFB1FFA2FF5089538F7CFC38" bold="true" box="[189,308,873,897]" pageId="20" pageNumber="21">Etymology</emphasis>
: The specific epithet (Latin, adjective: without beard) alludes to the absence of a comb of palisade setae at the posterior margin of the male sternite VII.
</paragraph>
</subSubSection>
<subSubSection id="C34C656BFFB1FFA2FF5089D38D8EFDD8" pageId="20" pageNumber="21" type="description">
<paragraph id="8BE936E0FFB1FFA2FF5089D38CFEFBF8" blockId="20.[189,795,1001,2081]" pageId="20" pageNumber="21">
<emphasis id="B922EAF2FFB1FFA2FF5089D38F77FBB8" bold="true" box="[189,319,1001,1025]" pageId="20" pageNumber="21">Description</emphasis>
: Body length 5.0–
<quantity id="4CAE9B05FFB1FFA2FDFE89D38C2EFBB8" box="[531,614,1001,1025]" metricMagnitude="-3" metricUnit="m" metricValue="5.8" pageId="20" pageNumber="21" unit="mm" value="5.8">5.8 mm</quantity>
; length of forebody
<quantity id="4CAE9B05FFB1FFA2FEED8E338FC9FB98" box="[256,385,1033,1057]" metricMagnitude="-3" metricUnit="m" metricValue="2.75" metricValueMax="2.9" metricValueMin="2.6" pageId="20" pageNumber="21" unit="mm" value="2.75" valueMax="2.9" valueMin="2.6">2.6–2.9 mm</quantity>
. Habitus as in
<figureCitation id="136D2A65FFB1FFA2FDD58E338CC7FB98" box="[568,655,1033,1057]" captionStart="Figs 115–129" captionStartId="21.[151,191,1967,1989]" captionTargetBox="[155,1394,219,1952]" captionTargetId="figure-117@21.[151,1398,219,1958]" captionTargetPageId="21" captionText="Figs 115–129: Lathrobium imberbe sp. n. (115–122) and L. aculeatum COIFFAIT (123–129): habitus (115, 123); forebody (116, 124); male sternite VII (117, 125); male sternite VIII (118, 126); aedeagus in lateral and in ventral view (119–120, 127–128); female sternite VIII (121, 129); female tergites IX–X (122). Scale bars: 115–116, 123–124: 1.0 mm; 117–122, 125–129: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4753558" httpUri="https://zenodo.org/record/4753558/files/figure.png" pageId="20" pageNumber="21">Fig. 115</figureCitation>
. Coloration: whole body, including the appendages, reddish.
</paragraph>
<paragraph id="8BE936E0FFB1FFA2FF508E738F30FA98" blockId="20.[189,795,1001,2081]" pageId="20" pageNumber="21">
Head (
<figureCitation id="136D2A65FFB1FFA2FEEB8E738F15FBD8" box="[262,349,1097,1121]" captionStart="Figs 115–129" captionStartId="21.[151,191,1967,1989]" captionTargetBox="[155,1394,219,1952]" captionTargetId="figure-117@21.[151,1398,219,1958]" captionTargetPageId="21" captionText="Figs 115–129: Lathrobium imberbe sp. n. (115–122) and L. aculeatum COIFFAIT (123–129): habitus (115, 123); forebody (116, 124); male sternite VII (117, 125); male sternite VIII (118, 126); aedeagus in lateral and in ventral view (119–120, 127–128); female sternite VIII (121, 129); female tergites IX–X (122). Scale bars: 115–116, 123–124: 1.0 mm; 117–122, 125–129: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4753558" httpUri="https://zenodo.org/record/4753558/files/figure.png" pageId="20" pageNumber="21">Fig. 116</figureCitation>
) 1.03–1.05 times as long as broad; punctation sparse and moderately coarse; interstices on average distinctly broader than diameter of punctures, with shallow microreticulation. Eyes small, composed of approximately 10 ommatidia, approximately one-fifth as long as postocular region in dorsal view. Antenna
<quantity id="4CAE9B05FFB1FFA2FF508F338F74FA98" box="[189,316,1289,1313]" metricMagnitude="-3" metricUnit="m" metricValue="1.45" metricValueMax="1.5" metricValueMin="1.4" pageId="20" pageNumber="21" unit="mm" value="1.45" valueMax="1.5" valueMin="1.4">1.4–1.5 mm</quantity>
long.
</paragraph>
<paragraph id="8BE936E0FFB1FFA2FF508F108FF7FA18" blockId="20.[189,795,1001,2081]" pageId="20" pageNumber="21">
Pronotum (
<figureCitation id="136D2A65FFB1FFA2FED68F138FDAFAF8" box="[315,402,1321,1345]" captionStart="Figs 115–129" captionStartId="21.[151,191,1967,1989]" captionTargetBox="[155,1394,219,1952]" captionTargetId="figure-117@21.[151,1398,219,1958]" captionTargetPageId="21" captionText="Figs 115–129: Lathrobium imberbe sp. n. (115–122) and L. aculeatum COIFFAIT (123–129): habitus (115, 123); forebody (116, 124); male sternite VII (117, 125); male sternite VIII (118, 126); aedeagus in lateral and in ventral view (119–120, 127–128); female sternite VIII (121, 129); female tergites IX–X (122). Scale bars: 115–116, 123–124: 1.0 mm; 117–122, 125–129: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4753558" httpUri="https://zenodo.org/record/4753558/files/figure.png" pageId="20" pageNumber="21">Fig. 116</figureCitation>
) slender, 1.25–1.30 times as long as broad and about as broad as head; punctation similar to that of head; midline broadly impunctate; interstices without microsculpture.
</paragraph>
<paragraph id="8BE936E0FFB1FFA2FF508F938C52F9F8" blockId="20.[189,795,1001,2081]" pageId="20" pageNumber="21">
Elytra (
<figureCitation id="136D2A65FFB1FFA2FEFD8F938F20FA78" box="[272,360,1449,1473]" captionStart="Figs 115–129" captionStartId="21.[151,191,1967,1989]" captionTargetBox="[155,1394,219,1952]" captionTargetId="figure-117@21.[151,1398,219,1958]" captionTargetPageId="21" captionText="Figs 115–129: Lathrobium imberbe sp. n. (115–122) and L. aculeatum COIFFAIT (123–129): habitus (115, 123); forebody (116, 124); male sternite VII (117, 125); male sternite VIII (118, 126); aedeagus in lateral and in ventral view (119–120, 127–128); female sternite VIII (121, 129); female tergites IX–X (122). Scale bars: 115–116, 123–124: 1.0 mm; 117–122, 125–129: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4753558" httpUri="https://zenodo.org/record/4753558/files/figure.png" pageId="20" pageNumber="21">Fig. 116</figureCitation>
) rather broad, 0.56–0.60 times as long as pronotum; punctation distinct and moderately dense; interstices without distinct microsculpture. Hind wings completely reduced. Protarsomeres I–IV with weakly pronounced sexual dimorphism.
</paragraph>
<paragraph id="8BE936E0FFB1FFA2FF508C738F05F8B8" blockId="20.[189,795,1001,2081]" pageId="20" pageNumber="21">Abdomen only slightly broader than elytra; punctation not very fine and moderately dense; interstices with shallow microsculpture; posterior margin of tergite VII without palisade fringe; posterior margin of tergite VIII weakly convex, without appreciable sexual dimorphism.</paragraph>
<paragraph id="8BE936E0FFB1FFA2FF508D308D33FEF8" blockId="20.[189,795,1001,2081]" lastBlockId="20.[831,1437,233,609]" pageId="20" pageNumber="21">
: protarsomeres I–IV moderately dilated; sternite VII (
<figureCitation id="136D2A65FFB1FFA2FF288D138F54F8F8" box="[197,284,1833,1857]" captionStart="Figs 115–129" captionStartId="21.[151,191,1967,1989]" captionTargetBox="[155,1394,219,1952]" captionTargetId="figure-117@21.[151,1398,219,1958]" captionTargetPageId="21" captionText="Figs 115–129: Lathrobium imberbe sp. n. (115–122) and L. aculeatum COIFFAIT (123–129): habitus (115, 123); forebody (116, 124); male sternite VII (117, 125); male sternite VIII (118, 126); aedeagus in lateral and in ventral view (119–120, 127–128); female sternite VIII (121, 129); female tergites IX–X (122). Scale bars: 115–116, 123–124: 1.0 mm; 117–122, 125–129: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4753558" httpUri="https://zenodo.org/record/4753558/files/figure.png" pageId="20" pageNumber="21">Fig. 117</figureCitation>
) distinctly transverse and with sparse unmodified pubescence, posterior margin broadly concave; sternite VIII (
<figureCitation id="136D2A65FFB1FFA2FEBE8D538FE2F838" box="[339,426,1897,1921]" captionStart="Figs 115–129" captionStartId="21.[151,191,1967,1989]" captionTargetBox="[155,1394,219,1952]" captionTargetId="figure-117@21.[151,1398,219,1958]" captionTargetPageId="21" captionText="Figs 115–129: Lathrobium imberbe sp. n. (115–122) and L. aculeatum COIFFAIT (123–129): habitus (115, 123); forebody (116, 124); male sternite VII (117, 125); male sternite VIII (118, 126); aedeagus in lateral and in ventral view (119–120, 127–128); female sternite VIII (121, 129); female tergites IX–X (122). Scale bars: 115–116, 123–124: 1.0 mm; 117–122, 125–129: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4753558" httpUri="https://zenodo.org/record/4753558/files/figure.png" pageId="20" pageNumber="21">Fig. 118</figureCitation>
) as long as broad, in the middle with extremely short, fine, and sparse pubescence, posterior excision moderately deep (approximately 0.2 times as deep as length of sternite) and slightly asymmetric; aedeagus (
<figureCitation id="136D2A65FFB1FFA2FEDB8DD38F8EF7B8" box="[310,454,2025,2049]" captionStart="Figs 115–129" captionStartId="21.[151,191,1967,1989]" captionTargetBox="[155,1394,219,1952]" captionTargetId="figure-117@21.[151,1398,219,1958]" captionTargetPageId="21" captionText="Figs 115–129: Lathrobium imberbe sp. n. (115–122) and L. aculeatum COIFFAIT (123–129): habitus (115, 123); forebody (116, 124); male sternite VII (117, 125); male sternite VIII (118, 126); aedeagus in lateral and in ventral view (119–120, 127–128); female sternite VIII (121, 129); female tergites IX–X (122). Scale bars: 115–116, 123–124: 1.0 mm; 117–122, 125–129: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4753558" httpUri="https://zenodo.org/record/4753558/files/figure.png" pageId="20" pageNumber="21">Figs 119–120</figureCitation>
)
<quantity id="4CAE9B05FFB1FFA2FE328DD38C76F7B8" box="[479,574,2025,2049]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" pageId="20" pageNumber="21" unit="mm" value="1.25">1.25 mm</quantity>
long and apically weakly asymmetric; ventral process shaped like a stout spine; dorsal plate short and lamellate, in lateral view very weakly curved; internal sac without sclerotized structures.
</paragraph>
<paragraph id="8BE936E0FFB1FFA2FCD28B708D8EFDD8" blockId="20.[831,1437,233,609]" pageId="20" pageNumber="21">
: protarsomeres I–IV moderately dilated, slightly less so than in male; sternite VIII (
<figureCitation id="136D2A65FFB1FFA2FB778B538ABAFE38" box="[1178,1266,361,385]" captionStart="Figs 115–129" captionStartId="21.[151,191,1967,1989]" captionTargetBox="[155,1394,219,1952]" captionTargetId="figure-117@21.[151,1398,219,1958]" captionTargetPageId="21" captionText="Figs 115–129: Lathrobium imberbe sp. n. (115–122) and L. aculeatum COIFFAIT (123–129): habitus (115, 123); forebody (116, 124); male sternite VII (117, 125); male sternite VIII (118, 126); aedeagus in lateral and in ventral view (119–120, 127–128); female sternite VIII (121, 129); female tergites IX–X (122). Scale bars: 115–116, 123–124: 1.0 mm; 117–122, 125–129: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4753558" httpUri="https://zenodo.org/record/4753558/files/figure.png" pageId="20" pageNumber="21">Fig. 121</figureCitation>
) approximately 1.15 times as long as broad and with moderately sparse pubescence, posterior margin strongly convex; tergite IX (
<figureCitation id="136D2A65FFB1FFA2FCAA8BF38DD7FE58" box="[839,927,457,481]" captionStart="Figs 115–129" captionStartId="21.[151,191,1967,1989]" captionTargetBox="[155,1394,219,1952]" captionTargetId="figure-117@21.[151,1398,219,1958]" captionTargetPageId="21" captionText="Figs 115–129: Lathrobium imberbe sp. n. (115–122) and L. aculeatum COIFFAIT (123–129): habitus (115, 123); forebody (116, 124); male sternite VII (117, 125); male sternite VIII (118, 126); aedeagus in lateral and in ventral view (119–120, 127–128); female sternite VIII (121, 129); female tergites IX–X (122). Scale bars: 115–116, 123–124: 1.0 mm; 117–122, 125–129: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4753558" httpUri="https://zenodo.org/record/4753558/files/figure.png" pageId="20" pageNumber="21">Fig. 122</figureCitation>
) undivided in the middle, postero-lateral processes short and apically acute, slightly extending beyond apex of tergite X; tergite X (
<figureCitation id="136D2A65FFB1FFA2FB5088338B5DFD98" box="[1213,1301,521,545]" captionStart="Figs 115–129" captionStartId="21.[151,191,1967,1989]" captionTargetBox="[155,1394,219,1952]" captionTargetId="figure-117@21.[151,1398,219,1958]" captionTargetPageId="21" captionText="Figs 115–129: Lathrobium imberbe sp. n. (115–122) and L. aculeatum COIFFAIT (123–129): habitus (115, 123); forebody (116, 124); male sternite VII (117, 125); male sternite VIII (118, 126); aedeagus in lateral and in ventral view (119–120, 127–128); female sternite VIII (121, 129); female tergites IX–X (122). Scale bars: 115–116, 123–124: 1.0 mm; 117–122, 125–129: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4753558" httpUri="https://zenodo.org/record/4753558/files/figure.png" pageId="20" pageNumber="21">Fig. 122</figureCitation>
) flat, of oval shape, and distinctly longer than antero-median portion of tergite IX.
</paragraph>
</subSubSection>
<subSubSection id="C34C656BFFB1FFA2FCD288B38AD0F998" pageId="20" pageNumber="21" type="discussion">
<paragraph id="8BE936E0FFB1FFA2FCD288B38A53FA9B" blockId="20.[831,1437,649,1570]" pageId="20" pageNumber="21">
<emphasis id="B922EAF2FFB1FFA2FCD288B38A8DFD18" bold="true" box="[831,1221,649,673]" pageId="20" pageNumber="21">Comment and comparative notes</emphasis>
: According to the species group concept proposed in
<bibRefCitation id="EFC74B11FFB1FFA2FB1D88938BD0FD78" author="ASSING, V." box="[1264,1432,681,705]" pageId="20" pageNumber="21" pagination="142 - 209" refId="ref18797" refString="ASSING, V. 2012: A revision of the Lathrobium species of the Himalaya (Coleoptera: Staphylinidae: Paederinae). - Bonn Zoological Bulletin 61 (2): 142 - 209." type="journal article" year="2012">ASSING (2012)</bibRefCitation>
,
<taxonomicName id="4C564D63FFB1FFA2FCD288F08DE6FD58" authorityName="Assing" authorityYear="2014" box="[831,942,713,737]" class="Insecta" family="Staphylinidae" genus="Lathrobium" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="imberbe">
<emphasis id="B922EAF2FFB1FFA2FCD288F08DE6FD58" box="[831,942,713,737]" italics="true" pageId="20" pageNumber="21">L. imberbe</emphasis>
</taxonomicName>
would have to be attributed to the
<taxonomicName id="4C564D63FFB1FFA2FADB88F08D22FCB8" authorityName="COIFFAIT" authorityYear="1982" class="Insecta" family="Staphylinidae" genus="Lathrobium" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="aculeatum">
<emphasis id="B922EAF2FFB1FFA2FADB88F08D22FCB8" italics="true" pageId="20" pageNumber="21">L. aculeatum</emphasis>
</taxonomicName>
group, which previously included two species from Central
<collectingCountry id="F3417670FFB1FFA2FC3189338A55FC98" box="[988,1053,777,801]" name="Nepal" pageId="20" pageNumber="21">Nepal</collectingCountry>
,
<taxonomicName id="4C564D63FFB1FFA2FBC089308B20FC98" authority="COIFFAIT, 1982" authorityName="COIFFAIT" authorityYear="1982" box="[1069,1384,777,801]" class="Insecta" family="Staphylinidae" genus="Lathrobium" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="aculeatum">
<emphasis id="B922EAF2FFB1FFA2FBC089308AFCFC98" box="[1069,1204,777,801]" italics="true" pageId="20" pageNumber="21">L. aculeatum</emphasis>
COIFFAIT, 1982
</taxonomicName>
and
<taxonomicName id="4C564D63FFB1FFA2FCD289108A20FCF8" authority="ASSING, 2012" authorityName="ASSING" authorityYear="2012" box="[831,1128,809,833]" class="Insecta" family="Staphylinidae" genus="Lathrobium" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="spiculatum">
<emphasis id="B922EAF2FFB1FFA2FCD289108D85FCF8" box="[831,973,809,833]" italics="true" pageId="20" pageNumber="21">L. spiculatum</emphasis>
<bibRefCitation id="EFC74B11FFB1FFA2FC3889138A20FCF8" author="ASSING, V." box="[981,1128,809,833]" pageId="20" pageNumber="21" pagination="142 - 209" refId="ref18797" refString="ASSING, V. 2012: A revision of the Lathrobium species of the Himalaya (Coleoptera: Staphylinidae: Paederinae). - Bonn Zoological Bulletin 61 (2): 142 - 209." type="journal article" year="2012">ASSING, 2012</bibRefCitation>
</taxonomicName>
. These two species probably represent adelphotaxa and are mainly distinguished from the species of the
<taxonomicName id="4C564D63FFB1FFA2FBEF89508ADBFC38" authorityName="COIFFAIT" authorityYear="1981" box="[1026,1171,874,897]" class="Insecta" family="Staphylinidae" genus="Lathrobium" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="pectinatum">
<emphasis id="B922EAF2FFB1FFA2FBEF89508ADBFC38" box="[1026,1171,874,897]" italics="true" pageId="20" pageNumber="21">L. pectinatum</emphasis>
</taxonomicName>
group by the absence of combs of palisade setae and a more slender aedeagus. Regarding the morphology of the aedeagus,
<taxonomicName id="4C564D63FFB1FFA2FAC389908BD4FC78" authorityName="Assing" authorityYear="2014" box="[1326,1436,937,961]" class="Insecta" family="Staphylinidae" genus="Lathrobium" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="imberbe">
<emphasis id="B922EAF2FFB1FFA2FAC389908BD4FC78" box="[1326,1436,937,961]" italics="true" pageId="20" pageNumber="21">L. imberbe</emphasis>
</taxonomicName>
is intermediate between the species of the
<taxonomicName id="4C564D63FFB1FFA2FAE189F08BD4FC58" authorityName="COIFFAIT" authorityYear="1981" box="[1292,1436,970,993]" class="Insecta" family="Staphylinidae" genus="Lathrobium" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="pectinatum">
<emphasis id="B922EAF2FFB1FFA2FAE189F08BD4FC58" box="[1292,1436,970,993]" italics="true" pageId="20" pageNumber="21">L. pectinatum</emphasis>
</taxonomicName>
group and those previously attributed to the
<taxonomicName id="4C564D63FFB1FFA2FAF989D08BD4FBB8" authorityName="COIFFAIT" authorityYear="1982" box="[1300,1436,1001,1025]" class="Insecta" family="Staphylinidae" genus="Lathrobium" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="aculeatum">
<emphasis id="B922EAF2FFB1FFA2FAF989D08BD4FBB8" box="[1300,1436,1001,1025]" italics="true" pageId="20" pageNumber="21">L. aculeatum</emphasis>
</taxonomicName>
group. With respect to other external and male sexual characters the species of both groups are very similar. These observations and supporting zoogeographic evidence make it seem likely that the absence of combs of palisade setae at the posterior margin of the male sternite VII is actually a secondary reduction that may have occurred convergently. Consequently,
<taxonomicName id="4C564D63FFB1FFA2FAC58EF08BDEFB58" authorityName="Assing" authorityYear="2014" box="[1320,1430,1225,1249]" class="Insecta" family="Staphylinidae" genus="Lathrobium" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="imberbe">
<emphasis id="B922EAF2FFB1FFA2FAC58EF08BDEFB58" box="[1320,1430,1225,1249]" italics="true" pageId="20" pageNumber="21">L. imberbe</emphasis>
</taxonomicName>
,
<taxonomicName id="4C564D63FFB1FFA2FCD28ED08D8FFAB8" authorityName="COIFFAIT" authorityYear="1982" box="[831,967,1257,1281]" class="Insecta" family="Staphylinidae" genus="Lathrobium" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="aculeatum">
<emphasis id="B922EAF2FFB1FFA2FCD28ED08D8FFAB8" box="[831,967,1257,1281]" italics="true" pageId="20" pageNumber="21">L. aculeatum</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C564D63FFB1FFA2FBF58ED08AEDFAB8" authorityName="ASSING" authorityYear="2012" box="[1048,1189,1257,1281]" class="Insecta" family="Staphylinidae" genus="Lathrobium" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="spiculatum">
<emphasis id="B922EAF2FFB1FFA2FBF58ED08AEDFAB8" box="[1048,1189,1257,1281]" italics="true" pageId="20" pageNumber="21">L. spiculatum</emphasis>
</taxonomicName>
are assigned to the
<taxonomicName id="4C564D63FFB1FFA2FCD28F308D98FA98" authorityName="COIFFAIT" authorityYear="1981" box="[831,976,1290,1313]" class="Insecta" family="Staphylinidae" genus="Lathrobium" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="pectinatum">
<emphasis id="B922EAF2FFB1FFA2FCD28F308D98FA98" box="[831,976,1290,1313]" italics="true" pageId="20" pageNumber="21">L. pectinatum</emphasis>
</taxonomicName>
group.
</paragraph>
<paragraph id="8BE936E0FFB1FFA2FCD28F138AD0F998" blockId="20.[831,1437,649,1570]" pageId="20" pageNumber="21">
<taxonomicName id="4C564D63FFB1FFA2FCD28F138A6EFAF8" authorityName="Assing" authorityYear="2014" box="[831,1062,1321,1345]" class="Insecta" family="Staphylinidae" genus="Lathrobium" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="imberbe">
<emphasis id="B922EAF2FFB1FFA2FCD28F138A6EFAF8" box="[831,1062,1321,1345]" italics="true" pageId="20" pageNumber="21">Lathrobium imberbe</emphasis>
</taxonomicName>
is readily distinguished from
<taxonomicName id="4C564D63FFB1FFA2FCD28F708D8FFAD8" authorityName="COIFFAIT" authorityYear="1982" box="[831,967,1353,1377]" class="Insecta" family="Staphylinidae" genus="Lathrobium" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="aculeatum">
<emphasis id="B922EAF2FFB1FFA2FCD28F708D8FFAD8" box="[831,967,1353,1377]" italics="true" pageId="20" pageNumber="21">L. aculeatum</emphasis>
</taxonomicName>
and
<taxonomicName id="4C564D63FFB1FFA2FC128F708AC4FAD8" authorityName="ASSING" authorityYear="2012" box="[1023,1164,1353,1377]" class="Insecta" family="Staphylinidae" genus="Lathrobium" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="spiculatum">
<emphasis id="B922EAF2FFB1FFA2FC128F708AC4FAD8" box="[1023,1164,1353,1377]" italics="true" pageId="20" pageNumber="21">L. spiculatum</emphasis>
</taxonomicName>
, with which it shares the unmodified chaetotaxy of the male sternite VII, by the different morphology of the aedeagus. Most other species of the
<taxonomicName id="4C564D63FFB1FFA2FC668F908A53FA78" authorityName="COIFFAIT" authorityYear="1981" box="[907,1051,1450,1473]" class="Insecta" family="Staphylinidae" genus="Lathrobium" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="pectinatum">
<emphasis id="B922EAF2FFB1FFA2FC668F908A53FA78" box="[907,1051,1450,1473]" italics="true" pageId="20" pageNumber="21">L. pectinatum</emphasis>
</taxonomicName>
group have conspicuous combs of palisade setae on the male sternite VII. For illustrations of the previously described representatives of the
<taxonomicName id="4C564D63FFB1FFA2FAA58FD08DCCF998" authorityName="COIFFAIT" authorityYear="1981" class="Insecta" family="Staphylinidae" genus="Lathrobium" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="pectinatum">
<emphasis id="B922EAF2FFB1FFA2FAA58FD08DCCF998" italics="true" pageId="20" pageNumber="21">L. pectinatum</emphasis>
</taxonomicName>
group see
<bibRefCitation id="EFC74B11FFB1FFA2FC1A8C338ADDF998" author="ASSING, V." box="[1015,1173,1545,1569]" pageId="20" pageNumber="21" pagination="142 - 209" refId="ref18797" refString="ASSING, V. 2012: A revision of the Lathrobium species of the Himalaya (Coleoptera: Staphylinidae: Paederinae). - Bonn Zoological Bulletin 61 (2): 142 - 209." type="journal article" year="2012">ASSING (2012)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34C656BFFB1FFA2FCD28C738A86F898" pageId="20" pageNumber="21" type="distribution">
<paragraph id="8BE936E0FFB1FFA2FCD28C738A86F898" blockId="20.[831,1436,1609,1825]" pageId="20" pageNumber="21">
<emphasis id="B922EAF2FFB1FFA2FCD28C738AEAF9D8" bold="true" box="[831,1186,1609,1633]" pageId="20" pageNumber="21">Distribution and natural history</emphasis>
: The species was found in two geographically close localities in the Garpung Khola valley in Dolpa District,
<collectingRegion id="4992F802FFB1FFA2FB4D8CB38B11F918" box="[1184,1369,1673,1697]" country="Nepal" name="Karnali" pageId="20" pageNumber="21">Karnali Province</collectingRegion>
, West
<collectingCountry id="F3417670FFB1FFA2FCD28C938D36F978" box="[831,894,1705,1729]" name="Nepal" pageId="20" pageNumber="21">Nepal</collectingCountry>
(
<figureCitation id="136D2A65FFB1FFA2FC618C938D9CF978" box="[908,980,1705,1729]" captionStart="Map 2" captionStartId="10.[189,234,892,913]" captionTargetBox="[189,1436,232,875]" captionTargetId="figure-553@10.[189,1436,229,875]" captionTargetPageId="10" captionText="Map 2:Distributions of Lathrobium species inNepal:L. flexaculeatum (white square),L. rectaculeatum (blackstar),L. curvaculeatum (white star), L. hebes (black square), L. imberbe (black triangle), L. plicatum (black diamond), L. calcaratum (circles; white circles: new records; black circle: type locality), L. fodens (white triangles), L. aculeatum (white diamonds)." figureDoi="http://doi.org/10.5281/zenodo.4753548" httpUri="https://zenodo.org/record/4753548/files/figure.png" pageId="20" pageNumber="21">Map 2</figureCitation>
). One
<typeStatus id="54ED8842FFB1FFA2FBF68C908A30F97B" box="[1051,1144,1706,1730]" pageId="20" pageNumber="21" type="paratype">paratype</typeStatus>
was collected at an altitude of
<quantity id="4CAE9B05FFB1FFA2FCB18CF38DE5F958" box="[860,941,1737,1761]" metricMagnitude="3" metricUnit="m" metricValue="3.8" pageId="20" pageNumber="21" unit="m" value="3800.0">3800 m</quantity>
, the remaining type specimens at an elevation between 3000 and
<quantity id="4CAE9B05FFB1FFA2FBF98CD38A2DF8B8" box="[1044,1125,1769,1793]" metricMagnitude="3" metricUnit="m" metricValue="3.8" pageId="20" pageNumber="21" unit="m" value="3800.0">3800 m</quantity>
, together with an unnamed species (
<taxonomicName id="4C564D63FFB1FFA2FC778D338A03F898" box="[922,1099,1801,1825]" class="Insecta" family="Staphylinidae" genus="Lathrobium" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="undefined-1">
<emphasis id="B922EAF2FFB1FFA2FC778D338A5EF898" box="[922,1046,1801,1825]" italics="true" pageId="20" pageNumber="21">Lathrobium</emphasis>
sp. 1
</taxonomicName>
; see below).
</paragraph>
</subSubSection>
</treatment>
</document>
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