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<document id="39EBCC907CB041519DCAA206DEB3BDCF" ID-DOI="10.5281/zenodo.276621" ID-GBIF-Dataset="a6eb9c5a-9b0b-47df-a207-6c5ee54acebd" ID-ISSN="1175-5326" ID-Zenodo-Dep="276621" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460225034079" checkinUser="plazi" docAuthor="Latha, Mathews, Cuccodoro, Giulio &amp; Sabu, Thomas K." docDate="2011" docId="03FF87EFFFD3B857FF723446E08AF9BB" docLanguage="en" docName="zt02745p029.pdf" docOrigin="Zootaxa 2745" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Ochicanthon besucheti Cuccodoro, sp. nov." docType="treatment" docVersion="8" lastPageNumber="3" masterDocId="FFC6FF97FFD1B855FFE53643E06BFFEF" masterDocTitle="Taxonomy of the dung beetle genus Ochicanthon Vaz-de-Mello (Coleoptera: Scarabaeidae: Scarabaeinae) of the Indian subcontinent, with notes on distribution patterns and flightlessness" masterLastPageNumber="29" masterPageNumber="1" pageNumber="3" updateTime="1698238191473" updateUser="plazi">
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<mods:title id="8C5BCF62C0C819EDEBF10823388C07C4">Taxonomy of the dung beetle genus Ochicanthon Vaz-de-Mello (Coleoptera: Scarabaeidae: Scarabaeinae) of the Indian subcontinent, with notes on distribution patterns and flightlessness</mods:title>
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<mods:namePart id="1395DC22C47F5C3AC8542AA4476CFAE8">Latha, Mathews</mods:namePart>
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<mods:namePart id="7B3E809BA65210D7953F72D4D5B44484">Cuccodoro, Giulio</mods:namePart>
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<mods:namePart id="4CCB7ECC2C694E406768F593AE9B757F">Sabu, Thomas K.</mods:namePart>
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<mods:title id="DC6E08083C16975A229DDA4B2B3F8618">Zootaxa</mods:title>
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<mods:date id="F4807D4468CD1DEC2F3369697E0D79EC">2011</mods:date>
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<mods:number id="1A362107A17A9AD60286AE1C8AEFE968">2745</mods:number>
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<treatment id="03FF87EFFFD3B857FF723446E08AF9BB" ID-DOI="http://doi.org/10.5281/zenodo.6182036" ID-GBIF-Taxon="119386107" ID-Zenodo-Dep="6182036" LSID="urn:lsid:plazi:treatment:03FF87EFFFD3B857FF723446E08AF9BB" httpUri="http://treatment.plazi.org/id/03FF87EFFFD3B857FF723446E08AF9BB" lastPageNumber="3" pageId="2" pageNumber="3">
<subSubSection id="C34C6572FFD3B857FF723446E179FDD0" pageId="2" pageNumber="3" type="nomenclature">
<paragraph id="8BE936F9FFD3B857FF723446E2C9FDF0" blockId="2.[151,674,517,575]" box="[151,674,517,543]" pageId="2" pageNumber="3">
<heading id="D0A18195FFD3B857FF723446E2C9FDF0" bold="true" box="[151,674,517,543]" fontSize="11" level="1" pageId="2" pageNumber="3" reason="1">
<emphasis id="B922EAEBFFD3B857FF723446E2C9FDF0" bold="true" box="[151,674,517,543]" pageId="2" pageNumber="3">
<taxonomicName id="4C564D7AFFD3B857FF723446E256FDF0" ID-CoL="48CV3" authority="Cuccodoro" authorityName="Cuccodoro" box="[151,573,517,543]" class="Insecta" family="Scarabaeidae" genus="Ochicanthon" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="besucheti" status="sp. nov.">
<emphasis id="B922EAEBFFD3B857FF723446E1C5FDF0" bold="true" box="[151,430,517,543]" italics="true" pageId="2" pageNumber="3">Ochicanthon besucheti</emphasis>
Cuccodoro
</taxonomicName>
<taxonomicNameLabel id="A2115790FFD3B857FDA13446E2C9FDF0" box="[580,674,517,543]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BE936F9FFD3B857FF723464E179FDD0" blockId="2.[151,674,517,575]" box="[151,274,551,575]" pageId="2" pageNumber="3">
(
<figureCitation id="136D2A7CFFD3B857FF7A3464E163FDD0" box="[159,264,551,575]" captionStart="FIGURES 1 5" captionStartId="3.[151,264,1953,1976]" captionTargetBox="[215,1365,195,1896]" captionTargetId="figure@3.[215,1366,195,1926]" captionTargetPageId="3" captionText="FIGURES 1 5. Ochicanthon besucheti (holotype). Dorsal habitus (1); base of pronotum and elytra (2); mesosternum and metasternum (3); aedeagus, ventral view (4); aedeagus, lateral left view (5). Scale bars = 1 mm." httpUri="https://zenodo.org/record/276622/files/figure.png" pageId="2" pageNumber="3">Figs. 15</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C34C6572FFD3B857FF72342CE523FBB3" pageId="2" pageNumber="3" type="description">
<paragraph id="8BE936F9FFD3B857FF72342CE24FFBFB" blockId="2.[151,1436,623,1620]" pageId="2" pageNumber="3">
<emphasis id="B922EAEBFFD3B857FF72342CE142FD67" bold="true" box="[151,297,623,648]" pageId="2" pageNumber="3">Description.</emphasis>
Body (
<figureCitation id="136D2A7CFFD3B857FE9E342CE1D7FD68" box="[379,444,623,647]" captionStart="FIGURES 1 5" captionStartId="3.[151,264,1953,1976]" captionTargetBox="[215,1365,195,1896]" captionTargetId="figure@3.[215,1366,195,1926]" captionTargetPageId="3" captionText="FIGURES 1 5. Ochicanthon besucheti (holotype). Dorsal habitus (1); base of pronotum and elytra (2); mesosternum and metasternum (3); aedeagus, ventral view (4); aedeagus, lateral left view (5). Scale bars = 1 mm." httpUri="https://zenodo.org/record/276622/files/figure.png" pageId="2" pageNumber="3">Fig. 1</figureCitation>
) predominantly black. Punctation on dorsal surface of head setiferous and annular separated by intervals not exceeding their diameter. Pronotum in lateral view weakly convex, outline at base almost in line with that of elytral suture; laterobasal paramarginal ridge well marked, extended on slightly more than one third of pronotal length. Pronotal punctation (
<figureCitation id="136D2A7CFFD3B857FD7C349FE2B6FD1B" box="[665,733,732,756]" captionStart="FIGURES 1 5" captionStartId="3.[151,264,1953,1976]" captionTargetBox="[215,1365,195,1896]" captionTargetId="figure@3.[215,1366,195,1926]" captionTargetPageId="3" captionText="FIGURES 1 5. Ochicanthon besucheti (holotype). Dorsal habitus (1); base of pronotum and elytra (2); mesosternum and metasternum (3); aedeagus, ventral view (4); aedeagus, lateral left view (5). Scale bars = 1 mm." httpUri="https://zenodo.org/record/276622/files/figure.png" pageId="2" pageNumber="3">Fig. 2</figureCitation>
) consisting of annular setiferous punctures separated by interval not exceeding their diameter, becoming sparser medially and larger posteriorly; setae fine and very short, almost indistinct. Elytra in lateral view moderately convex; suture not raised; striae shallow, wide, consisting of chains of fairly confluent discoidal depressions (
<figureCitation id="136D2A7CFFD3B857FD513504E292FCB0" box="[692,761,839,863]" captionStart="FIGURES 1 5" captionStartId="3.[151,264,1953,1976]" captionTargetBox="[215,1365,195,1896]" captionTargetId="figure@3.[215,1366,195,1926]" captionTargetPageId="3" captionText="FIGURES 1 5. Ochicanthon besucheti (holotype). Dorsal habitus (1); base of pronotum and elytra (2); mesosternum and metasternum (3); aedeagus, ventral view (4); aedeagus, lateral left view (5). Scale bars = 1 mm." httpUri="https://zenodo.org/record/276622/files/figure.png" pageId="2" pageNumber="3">Fig. 2</figureCitation>
); interstriae fairly flat, even, covered with annular setiferous punctures scattered irregularly and separated by interval nearly equal to their diameter; background microsculpture smooth; setae fine, straight and recumbent; lateral ridge posteriorly extended to level of apex of third stria. Mesosternum impunctate; meso-metasternal line arcuate; metasternum coarsely and uniformly punctate (
<figureCitation id="136D2A7CFFD3B857FB1035F7E551FC23" box="[1269,1338,948,972]" captionStart="FIGURES 1 5" captionStartId="3.[151,264,1953,1976]" captionTargetBox="[215,1365,195,1896]" captionTargetId="figure@3.[215,1366,195,1926]" captionTargetPageId="3" captionText="FIGURES 1 5. Ochicanthon besucheti (holotype). Dorsal habitus (1); base of pronotum and elytra (2); mesosternum and metasternum (3); aedeagus, ventral view (4); aedeagus, lateral left view (5). Scale bars = 1 mm." httpUri="https://zenodo.org/record/276622/files/figure.png" pageId="2" pageNumber="3">Fig. 3</figureCitation>
). Pygidium covered with shallow setiferous punctures becoming finer and sparser ventrally. Protrochanter with outer margin arcuate.
<collectingCountry id="F3417669FFD3B857FEA935BFE1EFFBFB" box="[332,388,1020,1044]" name="India" pageId="2" pageNumber="3">Hind</collectingCountry>
wings absent.
</paragraph>
<paragraph id="8BE936F9FFD3B857FF20325CE475FBD8" blockId="2.[151,1436,623,1620]" box="[197,1054,1055,1080]" pageId="2" pageNumber="3">
Male lacking obvious secondary sexual characters. Aedeagus as in
<figureCitation id="136D2A7CFFD3B857FC53325CE470FBD8" box="[950,1051,1055,1079]" captionStart="FIGURES 1 5" captionStartId="3.[151,264,1953,1976]" captionTargetBox="[215,1365,195,1896]" captionTargetId="figure@3.[215,1366,195,1926]" captionTargetPageId="3" captionText="FIGURES 1 5. Ochicanthon besucheti (holotype). Dorsal habitus (1); base of pronotum and elytra (2); mesosternum and metasternum (3); aedeagus, ventral view (4); aedeagus, lateral left view (5). Scale bars = 1 mm." httpUri="https://zenodo.org/record/276622/files/figure.png" pageId="2" pageNumber="3">Figs 45</figureCitation>
.
</paragraph>
<paragraph id="8BE936F9FFD3B857FF203207E523FBB3" blockId="2.[151,1436,623,1620]" box="[197,1352,1092,1116]" pageId="2" pageNumber="3">
<emphasis id="B922EAEBFFD3B857FF203207E11FFBB3" bold="true" box="[197,372,1092,1116]" pageId="2" pageNumber="3">Measurements</emphasis>
(mm; n = 4): TL = 3.34.5; BW = 2.22.7; PL = 1.01.2; PW = 1.92.3; EL = 1.92.2.
</paragraph>
</subSubSection>
<subSubSection id="C34C6572FFD3B857FF20322BE20DFAFF" pageId="2" pageNumber="3" type="materials_examined">
<paragraph id="8BE936F9FFD3B857FF20322BE459FB4B" blockId="2.[151,1436,623,1620]" pageId="2" pageNumber="3">
<emphasis id="B922EAEBFFD3B857FF20322BE11AFB6F" bold="true" box="[197,369,1127,1152]" pageId="2" pageNumber="3">
<typeStatus id="54ED885BFFD3B857FF20322BE095FB6F" box="[197,254,1128,1152]" pageId="2" pageNumber="3">Type</typeStatus>
material.
</emphasis>
<typeStatus id="54ED885BFFD3B857FE9E322BE188FB6F" box="[379,483,1128,1152]" pageId="2" pageNumber="3" type="holotype">Holotype</typeStatus>
(male,
<collectionCode id="ED47AE3CFFD3B857FDDC322BE2FEFB6F" box="[569,661,1128,1152]" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="2" pageNumber="3">MHNG</collectionCode>
): “
<collectingCountry id="F3417669FFD3B857FD57322BE29BFB6F" box="[690,752,1128,1152]" name="India" pageId="2" pageNumber="3">India</collectingCountry>
, Meghalaya, Khasi Hills, Shillong,
<quantity id="4CAE9B1CFFD3B857FB66322BE572FB6F" box="[1155,1305,1128,1152]" metricMagnitude="3" metricUnit="m" metricValue="1.9" metricValueMax="1.95" metricValueMin="1.85" pageId="2" pageNumber="3" unit="m" value="1900.0" valueMax="1950.0" valueMin="1850.0">18501950m</quantity>
,
<date id="FFE81039FFD3B857FAC0322BE5FCFB6F" box="[1317,1431,1128,1152]" pageId="2" pageNumber="3" value="1978-10-25">25.x.1978</date>
, leg. Besuchet &amp; Löbl, #27” [sifting in primary forest, Shillong Peak, slope North].
</paragraph>
<paragraph id="8BE936F9FFD3B857FF2032F2E20DFAFF" blockId="2.[151,1436,623,1620]" pageId="2" pageNumber="3">
<typeStatus id="54ED885BFFD3B857FF2032F2E159FB26" box="[197,306,1201,1225]" pageId="2" pageNumber="3" type="paratype">Paratypes</typeStatus>
(3,
<collectionCode id="ED47AE3CFFD3B857FE8532F3E1D7FB27" box="[352,444,1200,1224]" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="2" pageNumber="3">MHNG</collectionCode>
): Same data as
<typeStatus id="54ED885BFFD3B857FD8E32F3E2BBFB27" box="[619,720,1200,1224]" pageId="2" pageNumber="3" type="holotype">holotype</typeStatus>
,
<specimenCount id="9D50FD70FFD3B857FD3932F3E342FB27" box="[732,809,1200,1224]" pageId="2" pageNumber="3" type="male">1 male</specimenCount>
and
<specimenCount id="9D50FD70FFD3B857FC8732F3E3A3FB27" box="[866,968,1200,1224]" pageId="2" pageNumber="3" type="female">1 female</specimenCount>
; “
<collectingCountry id="F3417669FFD3B857FC3B32F3E477FB27" box="[990,1052,1200,1224]" name="India" pageId="2" pageNumber="3">India</collectingCountry>
, Meghalaya, Khasi hills, Shillong Peak,
<quantity id="4CAE9B1CFFD3B857FF3B3297E145FB03" box="[222,302,1236,1260]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="2" pageNumber="3" unit="m" value="1800.0">1800m</quantity>
,
<date id="FFE81039FFD3B857FED93297E1C0FB03" box="[316,427,1236,1260]" pageId="2" pageNumber="3" value="2000-10-21">21.x.2000</date>
&lt;
<geoCoordinate id="EE62503EFFD3B857FE233297E235FB03" box="[454,606,1236,1260]" direction="north" orientation="latitude" pageId="2" pageNumber="3" precision="15" value="25.541666">25°3230 N</geoCoordinate>
;
<geoCoordinate id="EE62503EFFD3B857FD893297E369FB03" box="[620,770,1236,1260]" direction="east" orientation="longitude" pageId="2" pageNumber="3" precision="15" value="91.84861">91°5055 E</geoCoordinate>
&gt;, leg. Carlton, Cuccodoro, Erne &amp; Leschen, #9a” [sifting leaf litter in mixed Pine forest],
<specimenCount id="9D50FD70FFD3B857FE1832BBE209FAFF" box="[509,610,1272,1296]" pageId="2" pageNumber="3" type="female">1 female</specimenCount>
.
</paragraph>
</subSubSection>
<subSubSection id="C34C6572FFD3B857FF203358E180FAB7" pageId="2" pageNumber="3" type="distribution">
<paragraph id="8BE936F9FFD3B857FF203358E180FAB7" blockId="2.[151,1436,623,1620]" pageId="2" pageNumber="3">
<emphasis id="B922EAEBFFD3B857FF203358E227FADB" bold="true" box="[197,588,1307,1332]" pageId="2" pageNumber="3">Distribution and natural history.</emphasis>
North East
<collectingCountry id="F3417669FFD3B857FD36335FE366FADB" box="[723,781,1308,1332]" name="India" pageId="2" pageNumber="3">India</collectingCountry>
(Upper montane forest in Shillong Peak in the Meghalaya subtropical forests ecoregion
<emphasis id="B922EAEBFFD3B857FE3F337CE188FAB7" bold="true" box="[474,483,1343,1368]" pageId="2" pageNumber="3">)</emphasis>
.
</paragraph>
</subSubSection>
<subSubSection id="C34C6572FFD3B857FF203327E4C4FA93" box="[197,1199,1380,1405]" pageId="2" pageNumber="3" type="etymology">
<paragraph id="8BE936F9FFD3B857FF203327E4C4FA93" blockId="2.[151,1436,623,1620]" box="[197,1199,1380,1405]" pageId="2" pageNumber="3">
<emphasis id="B922EAEBFFD3B857FF203327E12FFA93" bold="true" box="[197,324,1380,1404]" pageId="2" pageNumber="3">Etymology</emphasis>
. Named after Claude Besuchet, Geneva, who collected part of the
<typeStatus id="54ED885BFFD3B857FBD53326E40BFA92" box="[1072,1120,1381,1405]" pageId="2" pageNumber="3">type</typeStatus>
series.
</paragraph>
</subSubSection>
<subSubSection id="C34C6572FFD3B857FF2033C4E08AF9BB" pageId="2" pageNumber="3" type="diagnosis">
<paragraph id="8BE936F9FFD3B857FF2033C4E08AF9BB" blockId="2.[151,1436,623,1620]" pageId="2" pageNumber="3">
<emphasis id="B922EAEBFFD3B857FF2033C4E154FA4F" bold="true" box="[197,319,1415,1440]" pageId="2" pageNumber="3">Diagnosis.</emphasis>
Among the four wingless species of
<taxonomicName id="4C564D7AFFD3B857FD0633CBE31FFA4F" box="[739,884,1416,1440]" class="Insecta" family="Scarabaeidae" genus="Ochicanthon" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="B922EAEBFFD3B857FD0633CBE31FFA4F" box="[739,884,1416,1440]" italics="true" pageId="2" pageNumber="3">Ochicanthon</emphasis>
</taxonomicName>
(
<taxonomicName id="4C564D7AFFD3B857FC6133CBE47AFA4F" box="[900,1041,1416,1440]" class="Insecta" family="Scarabaeidae" genus="Ochicanthon" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="besucheti">
<emphasis id="B922EAEBFFD3B857FC6133CBE47AFA4F" box="[900,1041,1416,1440]" italics="true" pageId="2" pageNumber="3">O. besucheti</emphasis>
</taxonomicName>
,
<taxonomicName id="4C564D7AFFD3B857FBC533CBE4B3FA4F" box="[1056,1240,1416,1440]" class="Insecta" family="Scarabaeidae" genus="Ochicanthon" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="devagiriensis">
<emphasis id="B922EAEBFFD3B857FBC533CBE4B3FA4F" box="[1056,1240,1416,1440]" italics="true" pageId="2" pageNumber="3">O. devagiriensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C564D7AFFD3B857FB0333CBE522FA4F" box="[1254,1353,1416,1440]" class="Insecta" family="Scarabaeidae" genus="Ochicanthon" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="loebli">
<emphasis id="B922EAEBFFD3B857FB0333CBE522FA4F" box="[1254,1353,1416,1440]" italics="true" pageId="2" pageNumber="3">O. loebli</emphasis>
</taxonomicName>
and
<emphasis id="B922EAEBFFD3B857FA6433CBE14EFA2B" italics="true" pageId="2" pageNumber="3">
<taxonomicName id="4C564D7AFFD3B857FA6433CBE175FA2B" class="Insecta" family="Scarabaeidae" genus="Ochicanthon" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="vazdemelloi">O. vazdemelloi</taxonomicName>
)
</emphasis>
in the Indian subcontinent,
<taxonomicName id="4C564D7AFFD3B857FD8133EEE29BFA2B" box="[612,752,1452,1476]" class="Insecta" family="Scarabaeidae" genus="Ochicanthon" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="besucheti">
<emphasis id="B922EAEBFFD3B857FD8133EEE29BFA2B" box="[612,752,1452,1476]" italics="true" pageId="2" pageNumber="3">O. besucheti</emphasis>
</taxonomicName>
and
<taxonomicName id="4C564D7AFFD3B857FCCC33EEE3E7FA2B" box="[809,908,1452,1476]" class="Insecta" family="Scarabaeidae" genus="Ochicanthon" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="loebli">
<emphasis id="B922EAEBFFD3B857FCCC33EEE3E7FA2B" box="[809,908,1452,1476]" italics="true" pageId="2" pageNumber="3">O. loebli</emphasis>
</taxonomicName>
possess elytral interstriae with annular setiferous punctures not arranged in two rows. It is the only wingless
<taxonomicName id="4C564D7AFFD3B857FCBA3393E39BFA07" box="[863,1008,1488,1512]" class="Insecta" family="Scarabaeidae" genus="Ochicanthon" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="B922EAEBFFD3B857FCBA3393E39BFA07" box="[863,1008,1488,1512]" italics="true" pageId="2" pageNumber="3">Ochicanthon</emphasis>
</taxonomicName>
recorded from northeast of the Indian subcontinent.
<taxonomicName id="4C564D7AFFD3B857FED633B7E25FF9E3" box="[307,564,1524,1548]" class="Insecta" family="Scarabaeidae" genus="Ochicanthon" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="besucheti">
<emphasis id="B922EAEBFFD3B857FED633B7E25FF9E3" box="[307,564,1524,1548]" italics="true" pageId="2" pageNumber="3">Ochicanthon besucheti</emphasis>
</taxonomicName>
strongly resembles
<taxonomicName id="4C564D7AFFD3B857FCF633B6E304F9E3" box="[787,879,1525,1548]" class="Insecta" family="Scarabaeidae" genus="Ochicanthon" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="ernei">
<emphasis id="B922EAEBFFD3B857FCF633B6E304F9E3" box="[787,879,1525,1548]" italics="true" pageId="2" pageNumber="3">O. ernei</emphasis>
</taxonomicName>
and
<taxonomicName id="4C564D7AFFD3B857FC4133B6E446F9E3" box="[932,1069,1524,1548]" class="Insecta" family="Scarabaeidae" genus="Ochicanthon" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="tristoides">
<emphasis id="B922EAEBFFD3B857FC4133B6E446F9E3" box="[932,1069,1524,1548]" italics="true" pageId="2" pageNumber="3">O. tristoides</emphasis>
</taxonomicName>
, from northeast
<collectingCountry id="F3417669FFD3B857FB0133B7E54BF9E3" box="[1252,1312,1524,1548]" name="India" pageId="2" pageNumber="3">India</collectingCountry>
, but is easily distinguishable by the absence of wings, aedeagal shape and coarsely punctate central area of the metasternal shield.
</paragraph>
</subSubSection>
</treatment>
</document>