235 lines
31 KiB
XML
235 lines
31 KiB
XML
<document id="01978B37AD095BFA5ABC8EF3443C9E02" ID="10.11646/zootaxa.4258.2.5SLASH10907" ID-CLB-Dataset="33155" ID-DOI="10.11646/zootaxa.4258.2.5" ID-GBIF-Dataset="ac8dfde6-c270-402e-835d-cd37633fdfca" ID-ISSN="1175-5326" ID-Zenodo-Dep="569442" ID-ZooBank="312CBC35-CBB7-438C-AF7D-F0B373A460BC" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1493281446952" checkinUser="plazi" docAuthor="Sehnal, Richard" docDate="2017" docId="03FF87E2E117FFA718F7FC2BFC97F85E" docLanguage="en" docName="zootaxa.4258.2.5SLASH10907.pdf" docOrigin="Zootaxa 4258 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Entypophana biapicata Moser 1913" docType="treatment" docVersion="7" lastPageNumber="161" masterDocId="FFC6FF9AE114FFA31860FFFFFFE5FF8A" masterDocTitle="Review of the central African leaf chafer genus Entypophana Moser, 1913 (Coleoptera: Scarabaeidae: Melolonthinae: Schizonychini)" masterLastPageNumber="171" masterPageNumber="157" pageNumber="160" updateTime="1698439120373" updateUser="ExternalLinkService">
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<mods:title id="774CD53F5C34334DE613AD42CB56824E">Review of the central African leaf chafer genus Entypophana Moser, 1913 (Coleoptera: Scarabaeidae: Melolonthinae: Schizonychini)</mods:title>
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<mods:namePart id="7C10CF6D0E66B0777A9BFD52A832D237">Sehnal, Richard</mods:namePart>
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<mods:title id="576A315A305B8E4191E9E06404A1BC1A">Zootaxa</mods:title>
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<mods:part id="820BF4840C2FEAB0975D6D7FAC868B1B">
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<mods:date id="8B2E24D74123852CDC915C7DAE80B82F">2017</mods:date>
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<mods:number id="3FD2146BD7029949290272C09A831A0D">4258</mods:number>
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<mods:identifier id="E18E5B54EA1D55C99F4F5413BDDE585E" type="DOI">10.11646/zootaxa.4258.2.5</mods:identifier>
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<treatment id="03FF87E2E117FFA718F7FC2BFC97F85E" ID-DOI="http://doi.org/10.5281/zenodo.6034950" ID-GBIF-Taxon="128810298" ID-Zenodo-Dep="6034950" LSID="urn:lsid:plazi:treatment:03FF87E2E117FFA718F7FC2BFC97F85E" httpUri="http://treatment.plazi.org/id/03FF87E2E117FFA718F7FC2BFC97F85E" lastPageId="4" lastPageNumber="161" pageId="3" pageNumber="160">
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<subSubSection id="C34C657FE117FFA018F7FC2BFE07FB85" pageId="3" pageNumber="160" type="nomenclature">
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<paragraph id="8BE936F4E117FFA018F7FC2BFDA8FC64" blockId="3.[151,589,980,1039]" box="[151,589,980,1006]" pageId="3" pageNumber="160">
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<heading id="D0A18198E117FFA018F7FC2BFDA8FC64" bold="true" box="[151,589,980,1006]" fontSize="11" level="1" pageId="3" pageNumber="160" reason="1">
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<taxonomicName id="4C564D77E117FFA018F7FC2BFDA8FC64" authority="Moser, 1913" authorityName="Moser" authorityYear="1913" box="[151,589,980,1006]" class="Insecta" family="Melolonthidae" genus="Entypophana" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="160" phylum="Arthropoda" rank="species" species="biapicata">
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<emphasis id="B922EAE6E117FFA018F7FC2BFDA8FC64" bold="true" box="[151,589,980,1006]" pageId="3" pageNumber="160">
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<emphasis id="B922EAE6E117FFA018F7FC2BFE4AFC64" bold="true" box="[151,431,980,1006]" italics="true" pageId="3" pageNumber="160">Entypophana biapicata</emphasis>
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<bibRefCitation id="EFC74B05E117FFA019D6FC2BFDA8FC64" author="Moser" box="[438,589,980,1006]" pageId="3" pageNumber="160" refString="Moser, J. (1913) Neue afrikanische Schizonychinen. Annales de la Societe Entomologique de Belgique, 57, 283 - 296." type="journal article" year="1913">Moser, 1913</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="8BE936F4E117FFA018F7FC09FE07FB85" blockId="3.[151,589,980,1039]" box="[151,482,1014,1039]" pageId="3" pageNumber="160">
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<figureCitation id="136D2A71E117FFA018F7FC09FEF3FB85" box="[151,278,1014,1039]" captionStart="FIGURES 2" captionStartId="4.[151,264,909,932]" captionTargetBox="[157,1424,206,882]" captionTargetId="figure@4.[150,1436,193,889]" captionTargetPageId="4" captionText="FIGURES 2 A – D. Entypophana biapicata Moser, 1913 male, lectotype; length 17.2 mm. A, dorsal view; B, detail of head, anterior view; C, head and pronotum, dorsal view; D, labels. Not to scale." httpUri="https://zenodo.org/record/569444/files/figure.png" pageId="3" pageNumber="160">Figs 2A–D</figureCitation>
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,
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<figureCitation id="136D2A71E117FFA01940FC08FEA6FB84" box="[288,323,1015,1038]" captionStart="FIGURES 7" captionStartId="11.[151,264,1701,1724]" captionTargetBox="[185,1411,848,1668]" captionTargetId="figure@11.[177,1414,841,1674]" captionTargetPageId="11" captionText="FIGURES 7 A – F. Aedeagus in dorsal view. A, Entypophana apicata Moser, 1913, holotype; B, Entypophana biapicata Moser, 1913, lectotype; C, Entypophana hulstaerti Burgeon, 1946, lectotype; D, Entypophana lujai Moser, 1917, lectotype; E, Entypophana maynei Burgeon, 1946, lectotype; F, Entypophana njiapanda new species, holotype. Scale bar: 1.0 mm." httpUri="https://zenodo.org/record/569449/files/figure.png" pageId="3" pageNumber="160">7B</figureCitation>
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||
,
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<figureCitation id="136D2A71E117FFA0192FFC08FE97FB85" box="[335,370,1015,1039]" captionStart="FIGURES 8" captionStartId="12.[151,264,1326,1349]" captionTargetBox="[170,1427,419,1302]" captionTargetId="figure@12.[151,1437,411,1305]" captionTargetPageId="12" captionText="FIGURES 8 A – F. Aedeagus in lateral view. A, Entypophana apicata Moser, 1913, holotype; B, Entypophana biapicata Moser, 1913, lectotype; C, Entypophana hulstaerti Burgeon, 1946, lectotype; D, Entypophana lujai Moser, 1917, lectotype; E, Entypophana maynei Burgeon, 1946, lectotype; F, Entypophana njiapanda new species, holotype. Scale bar: 1.0 mm." httpUri="https://zenodo.org/record/569450/files/figure.png" pageId="3" pageNumber="160">8B</figureCitation>
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,
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<figureCitation id="136D2A71E117FFA0191EFC08FE44FB85" box="[382,417,1015,1039]" captionStart="FIGURES 9" captionStartId="13.[151,264,925,948]" captionTargetBox="[159,1429,206,892]" captionTargetId="figure@13.[151,1436,195,904]" captionTargetPageId="13" captionText="FIGURES 9 A – F. Inner mesotarsal claw. A, Entypophana apicata Moser, 1913, holotype; B, Entypophana biapicata Moser, 1913, lectotype; C, Entypophana hulstaerti Burgeon, 1946, lectotype; D, Entypophana lujai Moser, 1917, lectotype; E, Entypophana maynei Burgeon, 1946, lectotype; F, Entypophana njiapanda new species, holotype. Scale bar: 1.0 mm." httpUri="https://zenodo.org/record/569451/files/figure.png" pageId="3" pageNumber="160">9B</figureCitation>
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,
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<figureCitation id="136D2A71E117FFA019CDFC08FE3AFB85" box="[429,479,1015,1039]" captionStart="FIGURES 10" captionStartId="13.[151,264,1927,1950]" captionTargetBox="[151,1435,1059,1906]" captionTargetId="figure@13.[151,1435,1059,1906]" captionTargetPageId="13" captionText="FIGURES 10 A – F. Map with currently known geographical distribution. A, Entypophana apicata Moser, 1913; B, Entypophana biapicata Moser, 1913; C, Entypophana hulstaerti Burgeon, 1946; D, Entypophana lujai Moser, 1917; E, Entypophana maynei Burgeon, 1946; F, Entypophana njiapanda new species." httpUri="https://zenodo.org/record/569452/files/figure.png" pageId="3" pageNumber="160">10B</figureCitation>
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.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C34C657FE117FFA018F7FBC0FE3DFBF9" pageId="3" pageNumber="160" type="reference_group">
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<paragraph id="8BE936F4E117FFA018F7FBC0FE3DFBF9" blockId="3.[151,1436,1086,1139]" pageId="3" pageNumber="160">
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<treatmentCitationGroup id="AB4611DAE117FFA018F7FBC0FE3DFBF9" pageId="3" pageNumber="160">
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<taxonomicName id="4C564D77E117FFA018F7FBC0FDA9FBDE" authority="Moser, 1913: 295" authorityName="Moser" authorityPageNumber="295" authorityYear="1913" box="[151,588,1086,1108]" class="Insecta" family="Melolonthidae" genus="Entypophana" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="160" phylum="Arthropoda" rank="species" species="biapicata">
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||
<emphasis id="B922EAE6E117FFA018F7FBC0FE6DFBDE" box="[151,392,1087,1108]" italics="true" pageId="3" pageNumber="160">Entypophana biapicata</emphasis>
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<treatmentCitation id="0AF710E5E117FFA019F2FBC0FDA9FBDE" author="Moser" box="[402,588,1086,1108]" page="295" pageId="3" pageNumber="160" year="1913">
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<bibRefCitation id="EFC74B05E117FFA019F2FBC0FDFDFBDE" author="Moser" box="[402,536,1086,1108]" pageId="3" pageNumber="160" refString="Moser, J. (1913) Neue afrikanische Schizonychinen. Annales de la Societe Entomologique de Belgique, 57, 283 - 296." type="journal article" year="1913">Moser, 1913</bibRefCitation>
|
||
: 295
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||
</treatmentCitation>
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||
</taxonomicName>
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||
(description);
|
||
<treatmentCitation id="0AF710E5E117FFA01A89FBC0FC7BFBDE" author="Moser" box="[745,926,1086,1108]" page="256" pageId="3" pageNumber="160" year="1917">
|
||
<bibRefCitation id="EFC74B05E117FFA01A89FBC0FC8FFBDE" author="Moser" box="[745,874,1086,1108]" pageId="3" pageNumber="160" refString="Moser, J. (1917) Neue afrikanische Melolonthiden. Deutsche Entomologische Zeitschrift, 1917, 183 - 256." type="journal article" year="1917">Moser 1917</bibRefCitation>
|
||
: 256
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||
</treatmentCitation>
|
||
(comparison with
|
||
<taxonomicName id="4C564D77E117FFA01C09FBC0FB52FBDE" box="[1129,1207,1087,1108]" class="Insecta" family="Melolonthidae" genus="Entypophana" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="160" phylum="Arthropoda" rank="species" species="lujai">
|
||
<emphasis id="B922EAE6E117FFA01C09FBC0FB52FBDE" box="[1129,1207,1087,1108]" italics="true" pageId="3" pageNumber="160">E. lujai</emphasis>
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||
</taxonomicName>
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||
);
|
||
<treatmentCitation id="0AF710E5E117FFA01CB1FBC0FA79FBDE" author="Burgeon" box="[1233,1436,1086,1108]" page="262" pageId="3" pageNumber="160" year="1946">
|
||
<bibRefCitation id="EFC74B05E117FFA01CB1FBC0FA8DFBDE" author="Burgeon" box="[1233,1384,1086,1108]" pageId="3" pageNumber="160" refString="Burgeon, L. (1946) Melolonthini et Pachydemini du Congo belge. Revue de Zoologie et de Botanique Africains, 39, 230 - 273." type="journal article" year="1946">Burgeon 1946</bibRefCitation>
|
||
: 262
|
||
</treatmentCitation>
|
||
(comparison with
|
||
<taxonomicName id="4C564D77E117FFA0191FFBA1FE2CFBF8" box="[383,457,1117,1139]" class="Insecta" family="Melolonthidae" genus="Entypophana" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="160" phylum="Arthropoda" rank="species" species="lujai">
|
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<emphasis id="B922EAE6E117FFA0191FFBA1FE2CFBF8" box="[383,457,1117,1139]" italics="true" pageId="3" pageNumber="160">E. lujai</emphasis>
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</taxonomicName>
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||
).
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||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34C657FE117FFA018F7FB60FE0AFA05" pageId="3" pageNumber="160" type="materials_examined">
|
||
<paragraph id="8BE936F4E117FFA018F7FB60FACDFB32" blockId="3.[151,1437,1183,2001]" box="[151,1320,1183,1208]" pageId="3" pageNumber="160">
|
||
<materialsCitation id="3B3E3CA9E117FFA018F7FB60FACDFB32" ID-GBIF-Occurrence="1499294120" box="[151,1320,1183,1208]" country="Tanzania" county="Usambara" location="Usambara Mountains" municipality="Neu Bethel" pageId="3" pageNumber="160" specimenCount="1" stateProvince="Tanga region" typeStatus="holotype">
|
||
<emphasis id="B922EAE6E117FFA018F7FB60FED0FB32" bold="true" box="[151,309,1183,1208]" pageId="3" pageNumber="160">
|
||
<typeStatus id="54ED8856E117FFA018F7FB60FF37FB32" box="[151,210,1183,1208]" pageId="3" pageNumber="160">Type</typeStatus>
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locality.
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||
</emphasis>
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||
“
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<collectingCounty id="62884E78E117FFA01926FB60FE58FB32" box="[326,445,1183,1208]" pageId="3" pageNumber="160">Usambara</collectingCounty>
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||
,
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||
<collectingMunicipality id="6B8DAC8EE117FFA019A8FB5FFDACFB32" box="[456,585,1183,1208]" pageId="3" pageNumber="160">Neu Bethel</collectingMunicipality>
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||
) [=
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||
<collectingCountry id="F3417664E117FFA01A17FB60FD3AFB32" box="[631,735,1183,1208]" name="Tanzania" pageId="3" pageNumber="160">Tanzania</collectingCountry>
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||
,
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||
<collectingRegion id="4992F816E117FFA01A8AFB5FFC9AFB32" box="[746,895,1183,1208]" country="Tanzania" name="Tanga" pageId="3" pageNumber="160">Tanga region</collectingRegion>
|
||
,
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||
<location id="8E89602FE117FFA01BEAFB60FB9BFB32" LSID="urn:lsid:plazi:treatment:03FF87E2E117FFA718F7FC2BFC97F85E:8E89602FE117FFA01BEAFB60FB9BFB32" box="[906,1150,1183,1208]" country="Tanzania" county="Usambara" municipality="Neu Bethel" name="Usambara Mountains" pageId="3" pageNumber="160" stateProvince="Tanga region">Usambara Mountains</location>
|
||
, near
|
||
<location id="8E89602FE117FFA01CA0FB5FFAF7FB32" LSID="urn:lsid:plazi:treatment:03FF87E2E117FFA718F7FC2BFC97F85E:8E89602FE117FFA01CA0FB5FFAF7FB32" box="[1216,1298,1184,1208]" country="Tanzania" county="Usambara" municipality="Neu Bethel" name="Gonya" pageId="3" pageNumber="160" stateProvince="Tanga region">Gonya</location>
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||
]”.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BE936F4E117FFA018A7FB3BFA7EFA8A" blockId="3.[151,1437,1183,2001]" pageId="3" pageNumber="160">
|
||
<emphasis id="B922EAE6E117FFA018A7FB3BFE9EFB57" bold="true" box="[199,379,1220,1245]" pageId="3" pageNumber="160">Type material.</emphasis>
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||
Lectotype (by present designation), ♂, labelled: “Usambara / Neu Bethel [printed] / X [handwritten] 190 [p] 3 [handwritten, white label] //
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||
<taxonomicName id="4C564D77E117FFA01A84FB18FC9DFA8A" box="[740,888,1255,1280]" class="Insecta" family="Melolonthidae" genus="Entypophana" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="160" phylum="Arthropoda" rank="genus">Entypophana</taxonomicName>
|
||
/
|
||
<taxonomicName id="4C564D77E117FFA01BEDFB18FC17FA8A" box="[909,1010,1255,1280]" class="Insecta" family="Melolonthidae" genus="Entypophana" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="160" phylum="Arthropoda" rank="species" species="biapicata">biapicata</taxonomicName>
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||
/ Type ♂ Mos [printed, white label]”.
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||
</paragraph>
|
||
<paragraph id="8BE936F4E117FFA018A7FAF3FDE5FAAE" blockId="3.[151,1437,1183,2001]" box="[199,512,1292,1317]" pageId="3" pageNumber="160">
|
||
<emphasis id="B922EAE6E117FFA018A7FAF3FE69FAAF" bold="true" box="[199,396,1292,1317]" pageId="3" pageNumber="160">Type depository.</emphasis>
|
||
ZMHUB.
|
||
</paragraph>
|
||
<paragraph id="8BE936F4E117FFA018A7FAD0FE0AFA05" blockId="3.[151,1437,1183,2001]" pageId="3" pageNumber="160">
|
||
<materialsCitation id="3B3E3CA9E117FFA018A7FAD0FC37FAC2" ID-GBIF-Occurrence="1499294126" box="[199,978,1327,1352]" collectorName="Addition" country="Tanzania" location="Amani" pageId="3" pageNumber="160" specimenCount="6">
|
||
<emphasis id="B922EAE6E117FFA018A7FAD0FDF9FAC2" bold="true" box="[199,540,1327,1352]" pageId="3" pageNumber="160">
|
||
<collectorName id="26A35322E117FFA018A7FAD0FED4FAC2" box="[199,305,1327,1352]" pageId="3" pageNumber="160">Addition</collectorName>
|
||
material examined
|
||
</emphasis>
|
||
(
|
||
<specimenCount id="9D50FD7DE117FFA01A4EFAD0FD25FAC2" box="[558,704,1327,1352]" pageId="3" pageNumber="160" type="generic">6 specimens</specimenCount>
|
||
).
|
||
<emphasis id="B922EAE6E117FFA01AB4FAD0FCAEFAC2" bold="true" box="[724,843,1327,1352]" pageId="3" pageNumber="160">
|
||
<collectingCountry id="F3417664E117FFA01AB4FAD0FCA3FAC2" box="[724,838,1327,1352]" name="Tanzania" pageId="3" pageNumber="160">Tanzania</collectingCountry>
|
||
:
|
||
</emphasis>
|
||
<location id="8E89602FE117FFA01B35FACFFC41FAC2" LSID="urn:lsid:plazi:treatment:03FF87E2E117FFA718F7FC2BFC97F85E:8E89602FE117FFA01B35FACFFC41FAC2" box="[853,932,1328,1352]" country="Tanzania" name="Amani" pageId="3" pageNumber="160">Amani</location>
|
||
, 25.
|
||
</materialsCitation>
|
||
-
|
||
<date id="FFE81034E117FFA01BB9FACFFBA6FAC2" box="[985,1091,1328,1352]" pageId="3" pageNumber="160" value="06-12-30">30.12.06</date>
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||
, Vosseler G, 2 ♂ in ZMHUB; Amani, 19.-
|
||
<date id="FFE81034E117FFA01976FAABFE64FAE6" box="[278,385,1364,1388]" pageId="3" pageNumber="160" value="06-11-22">22.11.06</date>
|
||
, Vosseler G, 1 ♀ in ZMHUB; Usambara, Kwai, P.Weise S, 2 ♀ in ZMHUB; Bumbuli,
|
||
<date id="FFE81034E117FFA01D25FAABFA78FAE1" box="[1349,1437,1364,1388]" pageId="3" pageNumber="160">22. XII.</date>
|
||
0 2, Meinkop, 1 ♀ in ZMHUB.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34C657FE117FFA718A7FA63FC62F8E2" lastPageId="4" lastPageNumber="161" pageId="3" pageNumber="160" type="description">
|
||
<paragraph id="8BE936F4E117FFA018A7FA63FDD0F9CE" blockId="3.[151,1437,1183,2001]" pageId="3" pageNumber="160">
|
||
<emphasis id="B922EAE6E117FFA018A7FA63FDEDFA3F" bold="true" box="[199,520,1436,1461]" pageId="3" pageNumber="160">
|
||
Redescription of
|
||
<typeStatus id="54ED8856E117FFA019FDFA63FDEDFA3F" box="[413,520,1436,1461]" pageId="3" pageNumber="160" type="lectotype">lectotype</typeStatus>
|
||
</emphasis>
|
||
(
|
||
<specimenCount id="9D50FD7DE117FFA01A7EFA63FDD0FA3E" box="[542,565,1436,1460]" pageId="3" pageNumber="160" type="male">♂</specimenCount>
|
||
). Body length
|
||
<quantity id="4CAE9B11E117FFA01A8BFA63FCB3FA3E" box="[747,854,1436,1460]" metricMagnitude="-2" metricUnit="m" metricValue="1.72" pageId="3" pageNumber="160" unit="mm" value="17.2">17.2 mm</quantity>
|
||
, elongate, almost parallel-sided, strongly convex. Dorsal and ventral surfaces weakly shiny, elytra reddish brown, macrosetation pale (
|
||
<figureCitation id="136D2A71E117FFA01C03FA40FB25FA52" box="[1123,1216,1471,1496]" captionStart="FIGURES 2" captionStartId="4.[151,264,909,932]" captionTargetBox="[157,1424,206,882]" captionTargetId="figure@4.[150,1436,193,889]" captionTargetPageId="4" captionText="FIGURES 2 A – D. Entypophana biapicata Moser, 1913 male, lectotype; length 17.2 mm. A, dorsal view; B, detail of head, anterior view; C, head and pronotum, dorsal view; D, labels. Not to scale." httpUri="https://zenodo.org/record/569444/files/figure.png" pageId="3" pageNumber="160">Fig. 2A</figureCitation>
|
||
). Dorsal surface of head, pronotum, scutellum, and elytra covered with short, recumbent, scale-like, white macrosetae; ventral surface of thorax and abdomen with recumbent, long, yellow macrosetae. Head appendages and legs covered with moderately long, yellow macrosetae.
|
||
</paragraph>
|
||
<paragraph id="8BE936F4E117FFA018A7F9B0FC2AF85B" blockId="3.[151,1437,1183,2001]" pageId="3" pageNumber="160">
|
||
Head with labrum transverse, deeply bilobed; lobes rounded, unevenly weakly punctate; lobes covered with long, erect macrosetae. Head including clypeus smooth, glossy, densely coarsely punctate, with each puncture bearing an erect macroseta shorter than puncture diameter. Frontoclypeal carina elevated, medially sinuate forward; base as punctate as clypeus. Frontoclypeal suture undulate, medially arched toward occipital carina. Occipital carina always present, prominent, medially with two rounded summits, laterally diminishing (
|
||
<figureCitation id="136D2A71E117FFA01D55F920FA6AF972" box="[1333,1423,1759,1784]" captionStart="FIGURES 2" captionStartId="4.[151,264,909,932]" captionTargetBox="[157,1424,206,882]" captionTargetId="figure@4.[150,1436,193,889]" captionTargetPageId="4" captionText="FIGURES 2 A – D. Entypophana biapicata Moser, 1913 male, lectotype; length 17.2 mm. A, dorsal view; B, detail of head, anterior view; C, head and pronotum, dorsal view; D, labels. Not to scale." httpUri="https://zenodo.org/record/569444/files/figure.png" pageId="3" pageNumber="160">Fig. 2B</figureCitation>
|
||
). Space between summits and frontoclypeal suture triangular, without punctures or setae. Occiput evenly, moderately punctate. Canthus narrow, short, glabrous. Combined width of both eyes approximately equals maximum width of frons between eyes, eye distinctly extending beyond canthus. Antenna with 10 antennomeres; club with three antennomeres, almost straight, distinctly shorter than antennal shaft (antennomeres 1–7 combined). Antennomeres 1–7 with sparse, long macrosetae; club with sparse, short macrosetae. Terminal maxillary palpomere expanded apically, shorter than palpomeres 2 and 3 combined.
|
||
</paragraph>
|
||
<caption id="DF29667CE110FFA718F7FC72FC64FC49" httpUri="https://zenodo.org/record/569444/files/figure.png" pageId="4" pageNumber="161" targetBox="[157,1424,206,882]" targetPageId="4">
|
||
<paragraph id="8BE936F4E110FFA718F7FC72FC64FC49" blockId="4.[151,1436,909,963]" pageId="4" pageNumber="161">
|
||
<emphasis id="B922EAE6E110FFA718F7FC72FEB1FC29" bold="true" box="[151,340,909,932]" pageId="4" pageNumber="161">FIGURES 2A–D.</emphasis>
|
||
<taxonomicName id="4C564D77E110FFA7193DFC71FD32FC2E" authority="Moser, 1913" authorityName="Moser" authorityYear="1913" box="[349,727,910,932]" class="Insecta" family="Melolonthidae" genus="Entypophana" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="161" phylum="Arthropoda" rank="species" species="biapicata">
|
||
<emphasis id="B922EAE6E110FFA7193DFC71FDA9FC29" box="[349,588,910,931]" italics="true" pageId="4" pageNumber="161">Entypophana biapicata</emphasis>
|
||
Moser, 1913
|
||
</taxonomicName>
|
||
male, lectotype; length 17.2 mm. A, dorsal view; B, detail of head, anterior view; C, head and pronotum, dorsal view; D, labels. Not to scale.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BE936F4E110FFA718A7FC0FFE98FAA2" blockId="4.[151,1437,1007,2004]" pageId="4" pageNumber="161">
|
||
Pronotum transverse, widest behind midlength, with anteromedial semioval depression widest and flat in anterior third and then gradually reclined toward crest. Front margin straight with angles extended forward. Lateral margins crenulate and macrosetose. Base bordered, medially gently arched toward elytra, with smooth margin paralleled by a row of coarse punctures and hind angles broadly rounded. Crest delimiting anteromedial depression prominent, with frontal and top edges impunctate. Anterior part of pronotum deeply punctate, punctures separate, setae no longer than puncture diameters. Remaining parts similarly punctate but punctures shallow; setae yellow, scale-like, semirecumbent, no longer than puncture diameters. Scutellum large, almost equilaterally triangular; sides and apex rounded; sides punctate; each puncture bearing short, narrow, scale-like, semirecumbent, white macroseta (
|
||
<figureCitation id="136D2A71E110FFA71978FAF0FE95FAA2" box="[280,368,1295,1320]" captionStart="FIGURES 2" captionStartId="4.[151,264,909,932]" captionTargetBox="[157,1424,206,882]" captionTargetId="figure@4.[150,1436,193,889]" captionTargetPageId="4" captionText="FIGURES 2 A – D. Entypophana biapicata Moser, 1913 male, lectotype; length 17.2 mm. A, dorsal view; B, detail of head, anterior view; C, head and pronotum, dorsal view; D, labels. Not to scale." httpUri="https://zenodo.org/record/569444/files/figure.png" pageId="4" pageNumber="161">Fig. 2C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BE936F4E110FFA718A7FACCFCDFFA51" blockId="4.[151,1437,1007,2004]" pageId="4" pageNumber="161">Elytra weakly convex, parallel-sided, rounded apically; apical angle approximately rectangular. Striae absent except weakly indicated sutural stria. Humeral umbones present, weakly swollen. Surface not microsculptured, moderately shiny; punctation coarse, almost evenly spaced; punctures separated by twice their diameter. Each puncture bearing narrow, scale-like, semirecumbent, yellow macroseta as long as puncture diameter. Epipleuron distinct, complete, narrow, glabrous laterally. Macropterous.</paragraph>
|
||
<paragraph id="8BE936F4E110FFA718A7FA17FAD7F9E6" blockId="4.[151,1437,1007,2004]" pageId="4" pageNumber="161">
|
||
Legs with femora narrow, shiny, unevenly punctate, macrosetaceous. Protibia narrow, distinctly tridentate; terminal spine inserted against medial tooth. Mesotibiae and metatibiae slightly expanded distally, each with one macrosetiferous, longitudinal carina. Upper terminal spine of metatibia flattened, slightly curved, acute apically; 1/ 3 longer than lower, apically trunctate, chisel-shaped spine. Claws bifid, with ventrobasal teeth (
|
||
<figureCitation id="136D2A71E110FFA71CADF9ACFAC3F9E6" box="[1229,1318,1619,1644]" captionStart="FIGURES 9" captionStartId="13.[151,264,925,948]" captionTargetBox="[159,1429,206,892]" captionTargetId="figure@13.[151,1436,195,904]" captionTargetPageId="13" captionText="FIGURES 9 A – F. Inner mesotarsal claw. A, Entypophana apicata Moser, 1913, holotype; B, Entypophana biapicata Moser, 1913, lectotype; C, Entypophana hulstaerti Burgeon, 1946, lectotype; D, Entypophana lujai Moser, 1917, lectotype; E, Entypophana maynei Burgeon, 1946, lectotype; F, Entypophana njiapanda new species, holotype. Scale bar: 1.0 mm." httpUri="https://zenodo.org/record/569451/files/figure.png" pageId="4" pageNumber="161">Fig. 9B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BE936F4E110FFA718A7F988FA78F939" blockId="4.[151,1437,1007,2004]" pageId="4" pageNumber="161">Ventral surface covered only by long, semierect, yellow macrosetae. Pygidium slightly transverse, convex, bordered completely, apically broadly rounded, unevenly covered by coarse umbilicate, macrosetiferous punctures.</paragraph>
|
||
<paragraph id="8BE936F4E110FFA718A7F940FE95F8AA" blockId="4.[151,1437,1007,2004]" pageId="4" pageNumber="161">
|
||
Male genitalia (
|
||
<figureCitation id="136D2A71E110FFA7191FF940FE00F952" box="[383,485,1727,1752]" captionStart="FIGURES 7" captionStartId="11.[151,264,1701,1724]" captionTargetBox="[185,1411,848,1668]" captionTargetId="figure@11.[177,1414,841,1674]" captionTargetPageId="11" captionText="FIGURES 7 A – F. Aedeagus in dorsal view. A, Entypophana apicata Moser, 1913, holotype; B, Entypophana biapicata Moser, 1913, lectotype; C, Entypophana hulstaerti Burgeon, 1946, lectotype; D, Entypophana lujai Moser, 1917, lectotype; E, Entypophana maynei Burgeon, 1946, lectotype; F, Entypophana njiapanda new species, holotype. Scale bar: 1.0 mm." httpUri="https://zenodo.org/record/569449/files/figure.png" pageId="4" pageNumber="161">Figs. 7B</figureCitation>
|
||
,
|
||
<figureCitation id="136D2A71E110FFA71995F93FFDFDF952" box="[501,536,1728,1752]" captionStart="FIGURES 8" captionStartId="12.[151,264,1326,1349]" captionTargetBox="[170,1427,419,1302]" captionTargetId="figure@12.[151,1437,411,1305]" captionTargetPageId="12" captionText="FIGURES 8 A – F. Aedeagus in lateral view. A, Entypophana apicata Moser, 1913, holotype; B, Entypophana biapicata Moser, 1913, lectotype; C, Entypophana hulstaerti Burgeon, 1946, lectotype; D, Entypophana lujai Moser, 1917, lectotype; E, Entypophana maynei Burgeon, 1946, lectotype; F, Entypophana njiapanda new species, holotype. Scale bar: 1.0 mm." httpUri="https://zenodo.org/record/569450/files/figure.png" pageId="4" pageNumber="161">8B</figureCitation>
|
||
). Parameres symmetrical, longer than phallobase, evenly curved and apically rounded with double hump in lateral aspect, with single apex in dorsal aspect; covered by short, fine, semierect, yellow macrosetae.
|
||
</paragraph>
|
||
<paragraph id="8BE936F4E110FFA718A7F8D4FAB7F8C9" blockId="4.[151,1437,1007,2004]" box="[199,1362,1835,1860]" pageId="4" pageNumber="161">
|
||
<emphasis id="B922EAE6E110FFA718A7F8D4FE4AF8CE" bold="true" box="[199,431,1835,1860]" pageId="4" pageNumber="161">Sexual dimorphism</emphasis>
|
||
. Female differs from male in the following characters: body length
|
||
<quantity id="4CAE9B11E110FFA71CC8F8D4FAABF8C9" box="[1192,1358,1835,1860]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" metricValueMax="1.8399999999999999" metricValueMin="1.56" pageId="4" pageNumber="161" unit="mm" value="17.0" valueMax="18.4" valueMin="15.6">15.6–18.4 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BE936F4E110FFA718A7F8B0FC62F8E2" blockId="4.[151,1437,1007,2004]" box="[199,903,1871,1897]" pageId="4" pageNumber="161">
|
||
<materialsCitation id="3B3E3CA9E110FFA718A7F8B0FC62F8E2" ID-GBIF-Occurrence="1499294134" box="[199,903,1871,1897]" country="Tanzania" location="Tanga Region" pageId="4" pageNumber="161" specimenCount="1" stateProvince="Tanga Region">
|
||
<emphasis id="B922EAE6E110FFA718A7F8B0FE02F8E2" bold="true" box="[199,487,1871,1896]" pageId="4" pageNumber="161">Geographic distribution</emphasis>
|
||
.
|
||
<collectingCountry id="F3417664E110FFA71995F8B0FDB8F8E2" box="[501,605,1871,1896]" name="Tanzania" pageId="4" pageNumber="161">Tanzania</collectingCountry>
|
||
,
|
||
<collectingRegion id="4992F816E110FFA71A08F8AEFCE1F8E2" box="[616,772,1871,1897]" country="Tanzania" name="Tanga" pageId="4" pageNumber="161">Tanga Region</collectingRegion>
|
||
(
|
||
<figureCitation id="136D2A71E110FFA71B73F8B0FC9FF8E2" box="[787,890,1871,1896]" captionStart="FIGURES 10" captionStartId="13.[151,264,1927,1950]" captionTargetBox="[151,1435,1059,1906]" captionTargetId="figure@13.[151,1435,1059,1906]" captionTargetPageId="13" captionText="FIGURES 10 A – F. Map with currently known geographical distribution. A, Entypophana apicata Moser, 1913; B, Entypophana biapicata Moser, 1913; C, Entypophana hulstaerti Burgeon, 1946; D, Entypophana lujai Moser, 1917; E, Entypophana maynei Burgeon, 1946; F, Entypophana njiapanda new species." httpUri="https://zenodo.org/record/569452/files/figure.png" pageId="4" pageNumber="161">Fig. 10B</figureCitation>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34C657FE110FFA718A7F88CFC97F85E" pageId="4" pageNumber="161" type="discussion">
|
||
<paragraph id="8BE936F4E110FFA718A7F88CFC97F85E" blockId="4.[151,1437,1007,2004]" pageId="4" pageNumber="161">
|
||
<emphasis id="B922EAE6E110FFA718A7F88CFED5F806" bold="true" box="[199,304,1907,1932]" pageId="4" pageNumber="161">Remark.</emphasis>
|
||
This species was described based on an unstated number of specimens of both sexes. A
|
||
<typeStatus id="54ED8856E110FFA71D7BF88CFA67F806" box="[1307,1410,1907,1932]" pageId="4" pageNumber="161" type="lectotype">lectotype</typeStatus>
|
||
is designated here to enhance stability of nomenclature (International Commission on
|
||
<bibRefCitation id="EFC74B05E110FFA71C18F868FF37F85E" author="Zoological" pageId="4" pageNumber="161" refString="International Commission on Zoological Nomenclature (1999) International Code of Zoological Nomenclature, 4 th Edition. International Trust for Zoological Nomenclature, London, 306 pp." type="book" year="1999">Zoological Nomenclature 1999</bibRefCitation>
|
||
, Article 74). The location of the
|
||
<typeStatus id="54ED8856E110FFA71A20F844FD07F85E" box="[576,738,1979,2004]" pageId="4" pageNumber="161" type="paralectotype">paralectotypes</typeStatus>
|
||
is unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |