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<document id="0FAD952B7D4125E010C784341AC53826" ID="10.11646/zootaxa.4258.2.5SLASH10907" ID-CLB-Dataset="33155" ID-DOI="10.11646/zootaxa.4258.2.5" ID-GBIF-Dataset="ac8dfde6-c270-402e-835d-cd37633fdfca" ID-ISSN="1175-5326" ID-Zenodo-Dep="569442" ID-ZooBank="312CBC35-CBB7-438C-AF7D-F0B373A460BC" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1493281446952" checkinUser="plazi" docAuthor="Sehnal, Richard" docDate="2017" docId="03FF87E2E111FFA518F7FF68FE2CFB62" docLanguage="en" docName="zootaxa.4258.2.5SLASH10907.pdf" docOrigin="Zootaxa 4258 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Entypophana hulstaerti Burgeon 1946" docType="treatment" docVersion="7" lastPageNumber="163" masterDocId="FFC6FF9AE114FFA31860FFFFFFE5FF8A" masterDocTitle="Review of the central African leaf chafer genus Entypophana Moser, 1913 (Coleoptera: Scarabaeidae: Melolonthinae: Schizonychini)" masterLastPageNumber="171" masterPageNumber="157" pageNumber="162" updateTime="1698439120373" updateUser="ExternalLinkService">
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<mods:title id="C5ED5D7DE465DA472234E5159420B5BF">Review of the central African leaf chafer genus Entypophana Moser, 1913 (Coleoptera: Scarabaeidae: Melolonthinae: Schizonychini)</mods:title>
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<mods:namePart id="9D62B019D455D5BEC04272E033F4536D">Sehnal, Richard</mods:namePart>
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<mods:title id="BD360379CAD1C0EDDC26B6F967E43772">Zootaxa</mods:title>
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<mods:date id="585033DCF3F6C5BD5ED67CBD1B400913">2017</mods:date>
<mods:detail id="568A0B66819A79D98B16714307B34EEA" type="volume">
<mods:number id="E564DC1F962241A111DF7197FEE57754">4258</mods:number>
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<mods:classification id="DA6EE1470E33E6A20D11AD86E9B8DF4E">journal article</mods:classification>
<mods:identifier id="5D2F72A118303A72C9A0C062774AF1FF" type="CLB-Dataset">33155</mods:identifier>
<mods:identifier id="C2E7C32EA16745FE8825CC00A3EFF679" type="DOI">10.11646/zootaxa.4258.2.5</mods:identifier>
<mods:identifier id="46912857DCB3E48703F4DCD9E53891F3" type="GBIF-Dataset">ac8dfde6-c270-402e-835d-cd37633fdfca</mods:identifier>
<mods:identifier id="713C957ABA79A2D1507869E8CADD8520" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="3E5373E15951B1490A506206EDB2E078" type="Zenodo-Dep">569442</mods:identifier>
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<treatment id="03FF87E2E111FFA518F7FF68FE2CFB62" ID-DOI="http://doi.org/10.5281/zenodo.6034954" ID-GBIF-Taxon="128810302" ID-Zenodo-Dep="6034954" LSID="urn:lsid:plazi:treatment:03FF87E2E111FFA518F7FF68FE2CFB62" httpUri="http://treatment.plazi.org/id/03FF87E2E111FFA518F7FF68FE2CFB62" lastPageId="6" lastPageNumber="163" pageId="5" pageNumber="162">
<subSubSection id="C34C657FE111FFA618F7FF68FE06FF59" pageId="5" pageNumber="162" type="nomenclature">
<paragraph id="8BE936F4E111FFA618F7FF68FD95FF38" blockId="5.[151,624,151,211]" box="[151,624,151,178]" pageId="5" pageNumber="162">
<heading id="D0A18198E111FFA618F7FF68FD95FF38" bold="true" box="[151,624,151,178]" fontSize="11" level="1" pageId="5" pageNumber="162" reason="1">
<taxonomicName id="4C564D77E111FFA618F7FF68FD95FF38" authority="Burgeon, 1946" authorityName="Burgeon" authorityYear="1946" box="[151,624,151,178]" class="Insecta" family="Melolonthidae" genus="Entypophana" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="162" phylum="Arthropoda" rank="species" species="hulstaerti">
<emphasis id="B922EAE6E111FFA618F7FF68FD95FF38" bold="true" box="[151,624,151,178]" pageId="5" pageNumber="162">
<emphasis id="B922EAE6E111FFA618F7FF68FE51FF3B" bold="true" box="[151,436,151,177]" italics="true" pageId="5" pageNumber="162">Entypophana hulstaerti</emphasis>
<bibRefCitation id="EFC74B05E111FFA619DCFF67FD95FF38" author="Burgeon" box="[444,624,152,178]" pageId="5" pageNumber="162" refString="Burgeon, L. (1946) Melolonthini et Pachydemini du Congo belge. Revue de Zoologie et de Botanique Africains, 39, 230 - 273." type="journal article" year="1946">Burgeon, 1946</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BE936F4E111FFA618F7FF45FE06FF59" blockId="5.[151,624,151,211]" box="[151,483,186,211]" pageId="5" pageNumber="162">
<figureCitation id="136D2A71E111FFA618F7FF45FEF0FF59" box="[151,277,186,211]" captionStart="FIGURES 3" captionStartId="5.[151,264,1507,1530]" captionTargetBox="[159,1429,778,1479]" captionTargetId="figure@5.[150,1437,762,1486]" captionTargetPageId="5" captionText="FIGURES 3 A – D. Entypophana hulstaerti Burgeon, 1946 male, lectotype; length 18.4 mm. A, dorsal view; B, detail of head, anterior view; C, head and pronotum, dorsal view; D, labels. Not to scale." httpUri="https://zenodo.org/record/569445/files/figure.png" pageId="5" pageNumber="162">Figs 3A–D</figureCitation>
,
<figureCitation id="136D2A71E111FFA61940FF44FEA1FF59" box="[288,324,187,211]" captionStart="FIGURES 7" captionStartId="11.[151,264,1701,1724]" captionTargetBox="[185,1411,848,1668]" captionTargetId="figure@11.[177,1414,841,1674]" captionTargetPageId="11" captionText="FIGURES 7 A – F. Aedeagus in dorsal view. A, Entypophana apicata Moser, 1913, holotype; B, Entypophana biapicata Moser, 1913, lectotype; C, Entypophana hulstaerti Burgeon, 1946, lectotype; D, Entypophana lujai Moser, 1917, lectotype; E, Entypophana maynei Burgeon, 1946, lectotype; F, Entypophana njiapanda new species, holotype. Scale bar: 1.0 mm." httpUri="https://zenodo.org/record/569449/files/figure.png" pageId="5" pageNumber="162">7C</figureCitation>
,
<figureCitation id="136D2A71E111FFA6192FFF44FE97FF59" box="[335,370,187,211]" captionStart="FIGURES 8" captionStartId="12.[151,264,1326,1349]" captionTargetBox="[170,1427,419,1302]" captionTargetId="figure@12.[151,1437,411,1305]" captionTargetPageId="12" captionText="FIGURES 8 A – F. Aedeagus in lateral view. A, Entypophana apicata Moser, 1913, holotype; B, Entypophana biapicata Moser, 1913, lectotype; C, Entypophana hulstaerti Burgeon, 1946, lectotype; D, Entypophana lujai Moser, 1917, lectotype; E, Entypophana maynei Burgeon, 1946, lectotype; F, Entypophana njiapanda new species, holotype. Scale bar: 1.0 mm." httpUri="https://zenodo.org/record/569450/files/figure.png" pageId="5" pageNumber="162">8C</figureCitation>
,
<figureCitation id="136D2A71E111FFA6191EFF44FE45FF59" box="[382,416,187,211]" captionStart="FIGURES 9" captionStartId="13.[151,264,925,948]" captionTargetBox="[159,1429,206,892]" captionTargetId="figure@13.[151,1436,195,904]" captionTargetPageId="13" captionText="FIGURES 9 A – F. Inner mesotarsal claw. A, Entypophana apicata Moser, 1913, holotype; B, Entypophana biapicata Moser, 1913, lectotype; C, Entypophana hulstaerti Burgeon, 1946, lectotype; D, Entypophana lujai Moser, 1917, lectotype; E, Entypophana maynei Burgeon, 1946, lectotype; F, Entypophana njiapanda new species, holotype. Scale bar: 1.0 mm." httpUri="https://zenodo.org/record/569451/files/figure.png" pageId="5" pageNumber="162">9C</figureCitation>
,
<figureCitation id="136D2A71E111FFA619CDFF44FE3AFF59" box="[429,479,187,211]" captionStart="FIGURES 10" captionStartId="13.[151,264,1927,1950]" captionTargetBox="[151,1435,1059,1906]" captionTargetId="figure@13.[151,1435,1059,1906]" captionTargetPageId="13" captionText="FIGURES 10 A – F. Map with currently known geographical distribution. A, Entypophana apicata Moser, 1913; B, Entypophana biapicata Moser, 1913; C, Entypophana hulstaerti Burgeon, 1946; D, Entypophana lujai Moser, 1917; E, Entypophana maynei Burgeon, 1946; F, Entypophana njiapanda new species." httpUri="https://zenodo.org/record/569452/files/figure.png" pageId="5" pageNumber="162">10C</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34C657FE111FFA618F7FEFCFBEAFE92" box="[151,1039,258,280]" pageId="5" pageNumber="162" type="reference_group">
<paragraph id="8BE936F4E111FFA618F7FEFCFBEAFE92" blockId="5.[151,1039,258,280]" box="[151,1039,258,280]" pageId="5" pageNumber="162">
<treatmentCitationGroup id="AB4611DAE111FFA618F7FEFCFBEAFE92" box="[151,1039,258,280]" pageId="5" pageNumber="162">
<taxonomicName id="4C564D77E111FFA618F7FEFCFDB0FE92" authority="Burgeon, 1946: 263" authorityName="Burgeon" authorityPageNumber="263" authorityYear="1946" box="[151,597,258,280]" class="Insecta" family="Melolonthidae" genus="Entypophana" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="162" phylum="Arthropoda" rank="species" species="hulstaerti">
<emphasis id="B922EAE6E111FFA618F7FEFCFE60FE92" box="[151,389,259,280]" italics="true" pageId="5" pageNumber="162">Entypophana hulstaerti</emphasis>
<treatmentCitation id="0AF710E5E111FFA619ECFEFCFDB0FE92" author="Burgeon" box="[396,597,258,280]" page="263" pageId="5" pageNumber="162" year="1946">
<bibRefCitation id="EFC74B05E111FFA619ECFEFCFDC3FE92" author="Burgeon" box="[396,550,258,280]" pageId="5" pageNumber="162" refString="Burgeon, L. (1946) Melolonthini et Pachydemini du Congo belge. Revue de Zoologie et de Botanique Africains, 39, 230 - 273." type="journal article" year="1946">Burgeon, 1946</bibRefCitation>
: 263
</treatmentCitation>
</taxonomicName>
(description and comparison with
<taxonomicName id="4C564D77E111FFA61BD5FEFCFC1AFE92" box="[949,1023,259,280]" class="Insecta" family="Melolonthidae" genus="Entypophana" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="162" phylum="Arthropoda" rank="species" species="lujai">
<emphasis id="B922EAE6E111FFA61BD5FEFCFC1AFE92" box="[949,1023,259,280]" italics="true" pageId="5" pageNumber="162">E. lujai</emphasis>
</taxonomicName>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C34C657FE111FFA618F7FEBAFE08FD9B" pageId="5" pageNumber="162" type="materials_examined">
<paragraph id="8BE936F4E111FFA618F7FEBAFD16FE08" blockId="5.[151,1437,325,710]" pageId="5" pageNumber="162">
<materialsCitation id="3B3E3CA9E111FFA618F7FEBAFD16FE08" ID-GBIF-Occurrence="1499294089" country="Democratic Republic of the Congo" county="Flandria" location="Flandria" municipality="Equator Region" pageId="5" pageNumber="162" specimenCount="1" typeStatus="holotype">
<emphasis id="B922EAE6E111FFA618F7FEBAFEDDFED4" bold="true" box="[151,312,325,350]" pageId="5" pageNumber="162">
<typeStatus id="54ED8856E111FFA618F7FEBAFF37FED4" box="[151,210,325,350]" pageId="5" pageNumber="162">Type</typeStatus>
locality.
</emphasis>
“
<collectingCounty id="62884E78E111FFA6192BFEBAFE4EFED4" box="[331,427,325,350]" pageId="5" pageNumber="162">Flandria</collectingCounty>
[=
<collectingCountry id="F3417664E111FFA619B7FEBAFC89FED4" box="[471,876,325,350]" name="Democratic Republic of the Congo" pageId="5" pageNumber="162">Democratic Republic of the Congo</collectingCountry>
,
<collectingMunicipality id="6B8DAC8EE111FFA61B1AFEB9FBD5FED4" box="[890,1072,325,350]" pageId="5" pageNumber="162">Equator Region</collectingMunicipality>
,
<location id="8E89602FE111FFA61C5EFEBAFB7BFED4" LSID="urn:lsid:plazi:treatment:03FF87E2E111FFA518F7FF68FE2CFB62:8E89602FE111FFA61C5EFEBAFB7BFED4" box="[1086,1182,325,350]" country="Democratic Republic of the Congo" county="Flandria" municipality="Equator Region" name="Flandria" pageId="5" pageNumber="162">Flandria</location>
, near
<location id="8E89602FE111FFA61C85FEBAFA7DFED4" LSID="urn:lsid:plazi:treatment:03FF87E2E111FFA518F7FF68FE2CFB62:8E89602FE111FFA61C85FEBAFA7DFED4" box="[1253,1432,325,350]" country="Democratic Republic of the Congo" county="Flandria" municipality="Equator Region" name="Ingende village" pageId="5" pageNumber="162">Ingende village</location>
, between confluence of
<location id="8E89602FE111FFA619FBFE96FE01FE08" LSID="urn:lsid:plazi:treatment:03FF87E2E111FFA518F7FF68FE2CFB62:8E89602FE111FFA619FBFE96FE01FE08" box="[411,484,361,386]" country="Democratic Republic of the Congo" county="Flandria" municipality="Equator Region" name="Busira" pageId="5" pageNumber="162">Busira</location>
and
<location id="8E89602FE111FFA61A7BFE96FD75FE08" LSID="urn:lsid:plazi:treatment:03FF87E2E111FFA518F7FF68FE2CFB62:8E89602FE111FFA61A7BFE96FD75FE08" box="[539,656,361,386]" country="Democratic Republic of the Congo" county="Flandria" municipality="Equator Region" name="Momboyo" pageId="5" pageNumber="162">Momboyo</location>
rivers]”.
</materialsCitation>
</paragraph>
<paragraph id="8BE936F4E111FFA618A7FE72FC5AFE64" blockId="5.[151,1437,325,710]" pageId="5" pageNumber="162">
<materialsCitation id="3B3E3CA9E111FFA618A7FE72FD5FFE43" ID-GBIF-Occurrence="1499294085" collectionCode="MUSEE" country="Democratic Republic of the Congo" location="Equater" municipality="Burg" pageId="5" pageNumber="162" specimenCount="1" specimenCount-male="1" typeStatus="lectotype">
<emphasis id="B922EAE6E111FFA618A7FE72FE9DFE2C" bold="true" box="[199,376,397,422]" pageId="5" pageNumber="162">
<typeStatus id="54ED8856E111FFA618A7FE72FEE7FE2C" box="[199,258,397,422]" pageId="5" pageNumber="162">Type</typeStatus>
material.
</emphasis>
<typeStatus id="54ED8856E111FFA619E1FE71FE14FE2C" box="[385,497,398,422]" pageId="5" pageNumber="162" type="lectotype">Lectotype</typeStatus>
(by present designation),
<specimenCount id="9D50FD7DE111FFA61B40FE71FCD2FE2F" box="[800,823,398,421]" pageId="5" pageNumber="162" type="male">♂</specimenCount>
, labelled: “
<typeStatus id="54ED8856E111FFA61BDAFE71FBF0FE2C" box="[954,1045,398,422]" pageId="5" pageNumber="162">TYPUS</typeStatus>
[printed] /
<taxonomicName id="4C564D77E111FFA61CFCFE72FAEFFE2C" box="[1180,1290,397,422]" class="Insecta" family="Melolonthidae" genus="Entypophana" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="170" phylum="Arthropoda" rank="species" species="hulstaerti">Hulstaerti</taxonomicName>
/
<collectingMunicipality id="6B8DAC8EE111FFA61D46FE71FABBFE2C" box="[1318,1374,398,422]" pageId="5" pageNumber="162">Burg</collectingMunicipality>
[red, handwritten] //
<collectionCode id="ED47AE31E111FFA61928FE4EFE42FE40" box="[328,423,433,458]" pageId="5" pageNumber="162">MUSÉE</collectionCode>
DU
<collectingCountry id="F3417664E111FFA61982FE4EFDA3FE43" box="[482,582,433,457]" name="Democratic Republic of the Congo" pageId="5" pageNumber="162">CONGO</collectingCountry>
/
<location id="8E89602FE111FFA61A3FFE4DFD5FFE43" LSID="urn:lsid:plazi:treatment:03FF87E2E111FFA518F7FF68FE2CFB62:8E89602FE111FFA61A3FFE4DFD5FFE43" box="[607,698,434,457]" country="Democratic Republic of the Congo" municipality="Burg" name="Equater" pageId="5" pageNumber="162">Equater</location>
</materialsCitation>
:
<materialsCitation id="3B3E3CA9E111FFA61AA8FE4EFC5AFE64" ID-GBIF-Occurrence="1499294122" collectorName="R. P. Hulstaert" country="Democratic Republic of the Congo" location="Flandria" pageId="5" pageNumber="162" specimenCount="1" stateProvince="Flandria" typeStatus="holotype">
<collectingRegion id="4992F816E111FFA61AA8FE4EFCC0FE40" box="[712,805,433,458]" country="Belgium" name="Vlaanderen" pageId="5" pageNumber="162">Flandria</collectingRegion>
[printed] / 1929 [handwritten] /
<collectorName id="26A35322E111FFA61CFEFE4DFAA0FE40" box="[1182,1349,433,458]" pageId="5" pageNumber="162">R. P. Hulstaert</collectorName>
[white, printed] //
<taxonomicName id="4C564D77E111FFA6196DFE2AFE44FE64" box="[269,417,469,494]" class="Insecta" family="Melolonthidae" genus="Entypophana" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="162" phylum="Arthropoda" rank="genus">Entypophana</taxonomicName>
/
<taxonomicName id="4C564D77E111FFA619D6FE2AFDC1FE64" box="[438,548,469,494]" class="Insecta" family="Melolonthidae" genus="Entypophana" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="170" phylum="Arthropoda" rank="species" species="hulstaerti">Hulstaerti</taxonomicName>
/
<typeStatus id="54ED8856E111FFA61A5BFE29FD96FE64" box="[571,627,470,494]" pageId="5" pageNumber="162">Type</typeStatus>
/ Burg [white, handwritten]”.
</materialsCitation>
</paragraph>
<paragraph id="8BE936F4E111FFA618A7FE06FE08FD9B" blockId="5.[151,1437,325,710]" box="[199,493,505,530]" pageId="5" pageNumber="162">
<emphasis id="B922EAE6E111FFA618A7FE06FE69FD98" bold="true" box="[199,396,505,530]" pageId="5" pageNumber="162">Type depository.</emphasis>
MRAC.
</paragraph>
</subSubSection>
<subSubSection id="C34C657FE111FFA518A7FDE2FB50FBF6" lastPageId="6" lastPageNumber="163" pageId="5" pageNumber="162" type="description">
<paragraph id="8BE936F4E111FFA618A7FDE2FD75FD4F" blockId="5.[151,1437,325,710]" pageId="5" pageNumber="162">
<emphasis id="B922EAE6E111FFA618A7FDE2FDEDFDBC" bold="true" box="[199,520,541,566]" pageId="5" pageNumber="162">
Redescription of
<typeStatus id="54ED8856E111FFA619FDFDE2FDEDFDBC" box="[413,520,541,566]" pageId="5" pageNumber="162" type="lectotype">lectotype</typeStatus>
</emphasis>
(
<specimenCount id="9D50FD7DE111FFA61A7EFDE1FDD0FDBC" box="[542,565,542,566]" pageId="5" pageNumber="162" type="male">♂</specimenCount>
). Body length
<quantity id="4CAE9B11E111FFA61A8BFDE1FCB3FDBF" box="[747,854,542,566]" metricMagnitude="-2" metricUnit="m" metricValue="1.8399999999999999" pageId="5" pageNumber="162" unit="mm" value="18.4">18.4 mm</quantity>
, elongate, almost parallel-sided, strongly convex. Dorsal and ventral surfaces weakly shiny, elytra reddish brown, macrosetation pale (
<figureCitation id="136D2A71E111FFA61C03FDBEFB25FDD3" box="[1123,1216,577,602]" captionStart="FIGURES 3" captionStartId="5.[151,264,1507,1530]" captionTargetBox="[159,1429,778,1479]" captionTargetId="figure@5.[150,1437,762,1486]" captionTargetPageId="5" captionText="FIGURES 3 A – D. Entypophana hulstaerti Burgeon, 1946 male, lectotype; length 18.4 mm. A, dorsal view; B, detail of head, anterior view; C, head and pronotum, dorsal view; D, labels. Not to scale." httpUri="https://zenodo.org/record/569445/files/figure.png" pageId="5" pageNumber="162">Fig. 3A</figureCitation>
). Dorsal surface of head, pronotum, scutellum, and elytra covered with short, recumbent, scale-like, white macrosetae; ventral surface of thorax and abdomen with long, recumbent, yellow macrosetae. Head appendages and legs covered with moderately long, reddish-yellow macrosetae.
</paragraph>
<caption id="DF29667CE111FFA618F7FA1CFC64F992" httpUri="https://zenodo.org/record/569445/files/figure.png" pageId="5" pageNumber="162" targetBox="[159,1429,778,1479]" targetPageId="5">
<paragraph id="8BE936F4E111FFA618F7FA1CFC64F992" blockId="5.[151,1436,1507,1560]" pageId="5" pageNumber="162">
<emphasis id="B922EAE6E111FFA618F7FA1CFEB6FA73" bold="true" box="[151,339,1507,1530]" pageId="5" pageNumber="162">FIGURES 3A–D.</emphasis>
<taxonomicName id="4C564D77E111FFA6193AFA1BFD02FA70" authority="Burgeon, 1946" authorityName="Burgeon" authorityYear="1946" box="[346,743,1508,1530]" class="Insecta" family="Melolonthidae" genus="Entypophana" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="162" phylum="Arthropoda" rank="species" species="hulstaerti">
<emphasis id="B922EAE6E111FFA6193AFA1BFDACFA73" box="[346,585,1508,1529]" italics="true" pageId="5" pageNumber="162">Entypophana hulstaerti</emphasis>
Burgeon, 1946
</taxonomicName>
male, lectotype; length 18.4 mm. A, dorsal view; B, detail of head, anterior view; C, head and pronotum, dorsal view; D, labels. Not to scale.
</paragraph>
</caption>
<paragraph id="8BE936F4E111FFA618A7F9BFFEE8F86F" blockId="5.[151,1437,1600,2021]" pageId="5" pageNumber="162">
Head with labrum transverse, deeply bilobed; lobes rounded, unevenly, weakly punctate; lobes covered with long, erect macrosetae. Clypeus transverse, anterior margin weakly upturned and shallowly sinuate, anterior angles rounded. Head including clypeus smooth, matte, densely coarsely punctate; each puncture with an erect macroseta one-half longer than puncture diameter. Frontoclypeal carina elevated, medially sinuate forward; base as punctate as clypeus. Frontoclypeal suture straight. Occipital carina always present, prominent; medially with a flat, broad, glossy, impunctate summit, laterally diminishing (
<figureCitation id="136D2A71E111FFA61AB3F90BFCC8F886" box="[723,813,1780,1805]" captionStart="FIGURES 3" captionStartId="5.[151,264,1507,1530]" captionTargetBox="[159,1429,778,1479]" captionTargetId="figure@5.[150,1437,762,1486]" captionTargetPageId="5" captionText="FIGURES 3 A – D. Entypophana hulstaerti Burgeon, 1946 male, lectotype; length 18.4 mm. A, dorsal view; B, detail of head, anterior view; C, head and pronotum, dorsal view; D, labels. Not to scale." httpUri="https://zenodo.org/record/569445/files/figure.png" pageId="5" pageNumber="162">Fig. 3B</figureCitation>
). Space posterior of frontoclypeal suture with a small, triangular facet devoid of punctures and setae. Occiput evenly, moderately punctate. Canthus narrow, short, glabrous. Combined width of both eyes approximately equals width of frons between eyes, eye distinctly extending beyond canthus. Antenna with 10 antennomeres; club with three antennomeres, almost straight, distinctly shorter than antennal shaft (antennomeres 1–7 combined). Antennomeres 1–7 with sparse, long macrosetae; club with sparse, short macrosetae. Terminal maxillary palpomere expanded apically, shorter than palpomeres 2 and 3 combined.
</paragraph>
<paragraph id="8BE936F4E112FFA518A7FF67FB52FE26" blockId="6.[151,1437,151,1256]" pageId="6" pageNumber="163">
Pronotum transverse, widest behind midlength; with anteromedial, oval depression narrower and flat in anterior third, then gradually widening and reclining toward crest. Front margin straight with angles extended forward. Lateral margins crenulate and macrosetose. Base bordered, medially gently arched toward elytra, with smooth margin paralleled by a row of coarse punctures and hind angles broadly rounded. Crest delimiting anteromedial depression prominent, with frontal and top edges toward base impunctate except a narrow medial strip at base with punctures as those present laterally. Anterior part of pronotum distinctly punctate, setae from within punctures one-half longer than puncture diameters. Remaining parts similarly punctate but punctures shallow, setae yellow, scale-like, semirecumbent, no longer than puncture diameters (
<figureCitation id="136D2A71E112FFA51C32FE6BFB4FFE26" box="[1106,1194,404,429]" captionStart="FIGURES 3" captionStartId="5.[151,264,1507,1530]" captionTargetBox="[159,1429,778,1479]" captionTargetId="figure@5.[150,1437,762,1486]" captionTargetPageId="5" captionText="FIGURES 3 A – D. Entypophana hulstaerti Burgeon, 1946 male, lectotype; length 18.4 mm. A, dorsal view; B, detail of head, anterior view; C, head and pronotum, dorsal view; D, labels. Not to scale." httpUri="https://zenodo.org/record/569445/files/figure.png" pageId="6" pageNumber="163">Fig. 3C</figureCitation>
).
</paragraph>
<paragraph id="8BE936F4E112FFA518A7FE48FC1DFE7E" blockId="6.[151,1437,151,1256]" pageId="6" pageNumber="163">Scutellum large, almost equilaterally triangular, sides and apex rounded; sides punctate, each with four large punctures bearing short, narrow, scale-like, semirecumbent, white macroseta.</paragraph>
<paragraph id="8BE936F4E112FFA518A7FE00FC5AFD2D" blockId="6.[151,1437,151,1256]" pageId="6" pageNumber="163">
Elytra weakly convex, parallel-sided, rounded apically; apical angle approximately rectangular. Striae absent except for weakly indicated sutural stria. Humeral umbones present, weakly swollen. Surface not microsculptured, moderately shiny; punctation coarse, almost evenly spaced; punctures separated by
<date id="FFE81034E112FFA51CE4FDB7FB45FDEA" box="[1156,1184,584,608]" pageId="6" pageNumber="163">1x</date>
their diameter. Each puncture bearing narrow, scale-like, semirecumbent macroseta as long or slightly longer than puncture diameter. Epipleuron distinct, complete, narrow, laterally glabrous. Macropterous.
</paragraph>
<paragraph id="8BE936F4E112FFA518A7FD4AFF2DFCD6" blockId="6.[151,1437,151,1256]" pageId="6" pageNumber="163">
Legs with femora narrow, shiny, unevenly punctate; macrosetaceous. Protibia narrow, distinctly tridentate; terminal spine inserted over medial tooth. Mesotibiae and metatibiae slightly expanded distally, each with one macrosetiferous, longitudinal carina. Upper terminal spine of metatibia flattened, slightly curved, acute apically; 1/ 3 longer than lower, apically trunctate, chisel-shaped metatibial spine. Claws bifid, with ventrobasal teeth (
<figureCitation id="136D2A71E112FFA51D0FFCE0FF5EFCD6" captionStart="FIGURES 9" captionStartId="13.[151,264,925,948]" captionTargetBox="[159,1429,206,892]" captionTargetId="figure@13.[151,1436,195,904]" captionTargetPageId="13" captionText="FIGURES 9 A – F. Inner mesotarsal claw. A, Entypophana apicata Moser, 1913, holotype; B, Entypophana biapicata Moser, 1913, lectotype; C, Entypophana hulstaerti Burgeon, 1946, lectotype; D, Entypophana lujai Moser, 1917, lectotype; E, Entypophana maynei Burgeon, 1946, lectotype; F, Entypophana njiapanda new species, holotype. Scale bar: 1.0 mm." httpUri="https://zenodo.org/record/569451/files/figure.png" pageId="6" pageNumber="163">Fig. 9C</figureCitation>
).
</paragraph>
<paragraph id="8BE936F4E112FFA518A7FC98FEEEFC4D" blockId="6.[151,1437,151,1256]" pageId="6" pageNumber="163">Ventral surface covered only by long, semierect, yellow macrosetae. Pygidium slightly transverse, convex, bordered completely; apically broadly rounded; unevenly covered by coarse, umbilicate, macrosetiferous punctures.</paragraph>
<paragraph id="8BE936F4E112FFA518A7FC2BFB15FB85" blockId="6.[151,1437,151,1256]" pageId="6" pageNumber="163">
Male genitalia (
<figureCitation id="136D2A71E112FFA51916FC2BFE3DFC66" box="[374,472,980,1005]" captionStart="FIGURES 7" captionStartId="11.[151,264,1701,1724]" captionTargetBox="[185,1411,848,1668]" captionTargetId="figure@11.[177,1414,841,1674]" captionTargetPageId="11" captionText="FIGURES 7 A – F. Aedeagus in dorsal view. A, Entypophana apicata Moser, 1913, holotype; B, Entypophana biapicata Moser, 1913, lectotype; C, Entypophana hulstaerti Burgeon, 1946, lectotype; D, Entypophana lujai Moser, 1917, lectotype; E, Entypophana maynei Burgeon, 1946, lectotype; F, Entypophana njiapanda new species, holotype. Scale bar: 1.0 mm." httpUri="https://zenodo.org/record/569449/files/figure.png" pageId="6" pageNumber="163">Figs. 7C</figureCitation>
,
<figureCitation id="136D2A71E112FFA51982FC2BFDE3FC66" box="[482,518,980,1004]" captionStart="FIGURES 8" captionStartId="12.[151,264,1326,1349]" captionTargetBox="[170,1427,419,1302]" captionTargetId="figure@12.[151,1437,411,1305]" captionTargetPageId="12" captionText="FIGURES 8 A – F. Aedeagus in lateral view. A, Entypophana apicata Moser, 1913, holotype; B, Entypophana biapicata Moser, 1913, lectotype; C, Entypophana hulstaerti Burgeon, 1946, lectotype; D, Entypophana lujai Moser, 1917, lectotype; E, Entypophana maynei Burgeon, 1946, lectotype; F, Entypophana njiapanda new species, holotype. Scale bar: 1.0 mm." httpUri="https://zenodo.org/record/569450/files/figure.png" pageId="6" pageNumber="163">8C</figureCitation>
). Parameres symmetrical, longer than phallobase, evenly curved and rounded with hump and deflection apically in lateral aspect; covered by short, fine, semierect yellow macrosetae.
</paragraph>
<paragraph id="8BE936F4E112FFA518A7FBE3FC27FBD2" blockId="6.[151,1437,151,1256]" pageId="6" pageNumber="163">
<emphasis id="B922EAE6E112FFA518A7FBE3FE4BFBBF" bold="true" box="[199,430,1052,1077]" pageId="6" pageNumber="163">Sexual dimorphism</emphasis>
. Female differs from male in the following characters: vertex of head with small swelling, pronotum narrower with weak anteromedial depression (
<bibRefCitation id="EFC74B05E112FFA51B71FBBFFC53FBD2" author="Burgeon" box="[785,950,1087,1112]" pageId="6" pageNumber="163" refString="Burgeon, L. (1946) Melolonthini et Pachydemini du Congo belge. Revue de Zoologie et de Botanique Africains, 39, 230 - 273." type="journal article" year="1946">Burgeon 1946</bibRefCitation>
).
</paragraph>
<paragraph id="8BE936F4E112FFA518A7FB9BFB50FBF6" blockId="6.[151,1437,151,1256]" box="[199,1205,1124,1149]" pageId="6" pageNumber="163">
<emphasis id="B922EAE6E112FFA518A7FB9BFE02FBF7" bold="true" box="[199,487,1124,1149]" pageId="6" pageNumber="163">Geographic distribution</emphasis>
. Democratic Republic of the Congo, Equator region (
<figureCitation id="136D2A71E112FFA51C22FB9BFB4CFBF6" box="[1090,1193,1124,1149]" captionStart="FIGURES 10" captionStartId="13.[151,264,1927,1950]" captionTargetBox="[151,1435,1059,1906]" captionTargetId="figure@13.[151,1435,1059,1906]" captionTargetPageId="13" captionText="FIGURES 10 A – F. Map with currently known geographical distribution. A, Entypophana apicata Moser, 1913; B, Entypophana biapicata Moser, 1913; C, Entypophana hulstaerti Burgeon, 1946; D, Entypophana lujai Moser, 1917; E, Entypophana maynei Burgeon, 1946; F, Entypophana njiapanda new species." httpUri="https://zenodo.org/record/569452/files/figure.png" pageId="6" pageNumber="163">Fig. 10C</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C34C657FE112FFA518A7FB78FE2CFB62" pageId="6" pageNumber="163" type="discussion">
<paragraph id="8BE936F4E112FFA518A7FB78FE2CFB62" blockId="6.[151,1437,151,1256]" pageId="6" pageNumber="163">
<emphasis id="B922EAE6E112FFA518A7FB78FED5FB2A" bold="true" box="[199,304,1159,1184]" pageId="6" pageNumber="163">Remark.</emphasis>
This species was described based on two males. A
<typeStatus id="54ED8856E112FFA51B0BFB78FC34FB2A" box="[875,977,1159,1184]" pageId="6" pageNumber="163" type="lectotype">lectotype</typeStatus>
is designated here to enhance stability of nomenclature (International Commission on Zoological Nomenclature, Article 74). The location of the
<typeStatus id="54ED8856E112FFA518F7FB30FEDCFB62" box="[151,313,1231,1256]" pageId="6" pageNumber="163" type="paralectotype">paralectotypes</typeStatus>
is unknown.
</paragraph>
</subSubSection>
</treatment>
</document>
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