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<document id="8F043AEAD6A549D91A0E0EA4B0C86D6A" ID-CLB-Dataset="25263" ID-DOI="10.11646/zootaxa.4683.4.1" ID-GBIF-Dataset="48496621-de1a-4490-b866-1ae69d81ecbc" ID-ISSN="1175-5326" ID-Zenodo-Dep="3479609" ID-ZooBank="982FE3FB-5610-44A1-AAE3-659B28F45307" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1570688900916" checkinUser="plazi" docAuthor="Lee, Chi-Feng &amp; Bezdĕk, Jan" docDate="2019" docId="03FF87D6FFB5446CFF6BA827FDC3FAF6" docLanguage="en" docName="zootaxa.4683.4.1.pdf" docOrigin="Zootaxa 4683 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Theopea houjayi Lee &amp; Bezdĕk 2019, sp. nov." docType="treatment" docVersion="15" lastPageNumber="487" masterDocId="FFC6FFAEFF974448FFFCAF12FFAEFFA1" masterDocTitle="Revision of the genus Theopea Baly (Coleoptera: Chrysomelidae: Galerucinae) in Sundaland and the Philippines: Redefinition of the genus" masterLastPageNumber="507" masterPageNumber="451" pageNumber="485" updateTime="1698758705207" updateUser="ExternalLinkService">
<mods:mods id="B3B53CA8D5D7D4306A87D7036BAB2448" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="155BEBC2EB9FE1A4ECFAAA0C8ACD9DC2">
<mods:title id="A0B266ADD6E4A35EAAC2B5A7C943F53A">Revision of the genus Theopea Baly (Coleoptera: Chrysomelidae: Galerucinae) in Sundaland and the Philippines: Redefinition of the genus</mods:title>
</mods:titleInfo>
<mods:name id="E111F6AA093275DDD6091FE923221DB6" type="personal">
<mods:role id="3DFDD5FA08EEEFDC34244C2F9E03F19E">
<mods:roleTerm id="78C657E9747E790EFD422E6E3DCB481C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C402908704D4D7D30EE495B3C0BC0F5B">Lee, Chi-Feng</mods:namePart>
</mods:name>
<mods:name id="700BC21190F8273EC3CFC2FA55155128" type="personal">
<mods:role id="B0D1C678DF80B124A18755D5B9D075BA">
<mods:roleTerm id="72A06376062939FD50769493234B0E9E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8CACF81E32F066CA014959D5CBDFF20E">Bezdĕk, Jan</mods:namePart>
</mods:name>
<mods:typeOfResource id="E18A3AEDFD54075D77327DD34FAC752E">text</mods:typeOfResource>
<mods:relatedItem id="26E2CFAFC9BC39A16C1665A5D23621FD" type="host">
<mods:titleInfo id="1EFC54CC5C118CCEE8D37F6773865BED">
<mods:title id="080427CC720E859A791DD608C9EA0470">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="E614892C20E026DFD7B9940CEC9256F6">
<mods:date id="166DF7C7BD0B6C44CCF240AE1AA5E222">2019</mods:date>
<mods:detail id="4B6A947646B1F6DB7EA5BF0D317CBA2F" type="pubDate">
<mods:number id="AA171EADAFB585C75E449B5DF42EBED8">2019-10-10</mods:number>
</mods:detail>
<mods:detail id="4376A32CD45937583697FE4D9D9E1530" type="volume">
<mods:number id="8CD44615AA3DF93A982DB31355A0C43D">4683</mods:number>
</mods:detail>
<mods:detail id="CBCAA9FBFD00009A08C408CDE8FBB94F" type="issue">
<mods:number id="A3C555B50ACA9D61E70FFC349356575A">4</mods:number>
</mods:detail>
<mods:extent id="141E231076873DE9E9A402BDC15B1AD0" unit="page">
<mods:start id="3F3A30B250B302CE7D0804BA708CC9BF">451</mods:start>
<mods:end id="442F09B06AD8D4BF0A0B900A39AD1E3A">507</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="4DD9B5C3C53388675AFB116821B3D745">journal article</mods:classification>
<mods:identifier id="BB762D14A92F3EE2EB5DEE760492A2CF" type="CLB-Dataset">25263</mods:identifier>
<mods:identifier id="F01D3B07EE87DB4DEB2BAF02CC776178" type="DOI">10.11646/zootaxa.4683.4.1</mods:identifier>
<mods:identifier id="E7B13388FC5C185F23D30031AF20DB26" type="GBIF-Dataset">48496621-de1a-4490-b866-1ae69d81ecbc</mods:identifier>
<mods:identifier id="2929FEB05288832ABA6A40875A8019A8" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="0F3EE6D6CAF26266FFD66FF650D744EF" type="Zenodo-Dep">3479609</mods:identifier>
<mods:identifier id="A9EE45634DB75BE31CDFD372B3E00A98" type="ZooBank">982FE3FB-5610-44A1-AAE3-659B28F45307</mods:identifier>
</mods:mods>
<treatment id="03FF87D6FFB5446CFF6BA827FDC3FAF6" ID-DOI="http://doi.org/10.5281/zenodo.4505264" ID-GBIF-Taxon="159690569" ID-Zenodo-Dep="4505264" LSID="urn:lsid:plazi:treatment:03FF87D6FFB5446CFF6BA827FDC3FAF6" httpUri="http://treatment.plazi.org/id/03FF87D6FFB5446CFF6BA827FDC3FAF6" lastPageId="36" lastPageNumber="487" pageId="34" pageNumber="485">
<subSubSection id="C34C654BFFB5446AFF6BA827FE69F8EE" box="[151,455,1845,1872]" pageId="34" pageNumber="485" type="nomenclature">
<paragraph id="8BE936C0FFB5446AFF6BA827FE69F8EE" blockId="34.[151,455,1845,1907]" box="[151,455,1845,1872]" pageId="34" pageNumber="485">
<heading id="D0A181ACFFB5446AFF6BA827FE69F8EE" bold="true" box="[151,455,1845,1872]" fontSize="11" level="1" pageId="34" pageNumber="485" reason="1">
<emphasis id="B922EAD2FFB5446AFF6BA827FE69F8EE" bold="true" box="[151,455,1845,1872]" pageId="34" pageNumber="485">
<taxonomicName id="4C564D43FFB5446AFF6BA827FECCF8F1" authority="Lee &amp; Bezdĕk, 2019" authorityName="Lee &amp; Bezdĕk" authorityYear="2019" box="[151,354,1845,1872]" class="Insecta" family="Chrysomelidae" genus="Theopea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="485" phylum="Arthropoda" rank="species" species="houjayi" status="sp. nov.">
<emphasis id="B922EAD2FFB5446AFF6BA827FECCF8F1" bold="true" box="[151,354,1845,1872]" italics="true" pageId="34" pageNumber="485">Theopea houjayi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21157A9FFB5446AFE95A827FE69F8EE" box="[361,455,1845,1871]" pageId="34" pageNumber="485" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34C654BFFB5446CFF6BA84BFE7AFBB2" lastPageId="36" lastPageNumber="487" pageId="34" pageNumber="485" type="description">
<paragraph id="8BE936C0FFB5446AFF6BA84BFEDBF8D2" blockId="34.[151,455,1845,1907]" box="[151,373,1881,1907]" pageId="34" pageNumber="485">
(
<figureCitation id="136D2A45FFB5446AFF63A84BFF5DF8D2" box="[159,243,1881,1907]" captionStart="FIGURE 14" captionStartId="29.[151,250,1897,1922]" captionTargetBox="[238,1338,183,1873]" captionTargetId="figure@29.[232,1356,174,1878]" captionTargetPageId="29" captionText="FIGURE 14. Habitus of Theopea elegantula Baly, T. houjayi sp. nov., and T. yuae sp. nov. 14A. T. elegantula, male, dorsal view; 14B. Same, ventral view; 14C. T. houjayi sp. nov., male, dorsal view; 14D. Same, ventral view; 14E. T. houjayi sp. nov., female, dorsal view; 14F. T. yuae sp. nov., male, dorsal view; 14G. Same, ventral view; 14H. T. yuae sp. nov., male, dorsal view, color variation; 14I. T. yuae sp. nov., male, dorsal view, color variation." figureDoi="http://doi.org/10.5281/zenodo.3479637" httpUri="https://zenodo.org/record/3479637/files/figure.png" pageId="34" pageNumber="485">Figs 14</figureCitation>
C–14E, 18)
</paragraph>
<paragraph id="8BE936C0FFB5446CFF6BA8B3FA84FEBE" blockId="34.[151,1437,1952,2015]" lastBlockId="36.[151,1437,152,1367]" lastPageId="36" lastPageNumber="487" pageId="34" pageNumber="485">
<emphasis id="B922EAD2FFB5446AFF6BA8B3FEECF81A" bold="true" box="[151,322,1952,1979]" pageId="34" pageNumber="485">Types (n= 15).</emphasis>
<materialsCitation id="3B3E3C9DFFB5446AFEB6A8B3FAB1F87E" ID-GBIF-Occurrence="2425474194" collectingDate="2011-12-08" collectingDateMax="2011-12-11" collectingDateMin="2011-12-08" collectionCode="NMPC" collectorName="J. Hajek &amp; Hajek, J &amp; P. Votruba." elevation="35" latitude="-1.135" location="Forest" longLatPrecision="130" longitude="116.831665" municipality="Sungai Wain" pageId="34" pageNumber="485" specimenCount="1" specimenCount-male="1" stateProvince="Kalimantan Timur" typeStatus="holotype">
<typeStatus id="54ED8862FFB5446AFEB6A8B3FE1CF81A" box="[330,434,1953,1979]" pageId="34" pageNumber="485" type="holotype">Holotype</typeStatus>
<specimenCount id="9D50FD49FFB5446AFE46A8B3FE61F81A" box="[442,463,1953,1979]" pageId="34" pageNumber="485" type="male">♂</specimenCount>
(
<collectionCode id="ED47AE05FFB5446AFE23A8B3FD9AF81A" box="[479,564,1953,1979]" country="Czech Republic" name="National Museum Prague" pageId="34" pageNumber="485">NMPC</collectionCode>
), INDONESIA.
<collectingRegion id="4992F822FFB5446AFD0CA8B3FC00F81A" box="[752,942,1953,1979]" country="Indonesia" name="Kalimantan Timur" pageId="34" pageNumber="485">East Kalimantan</collectingRegion>
.
<locationDeviation id="9A5B2FE2FFB5446AFC46A8B2FB66F81A" box="[954,1224,1952,1979]" location="Balikpapan" pageId="34" pageNumber="485">
<quantity id="4CAE9B25FFB5446AFC46A8B2FBACF81A" box="[954,1026,1952,1979]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="34" pageNumber="485" unit="km" value="15.0">15 km</quantity>
N of Balikpapan
</locationDeviation>
,
<collectingMunicipality id="6B8DACBAFFB5446AFB28A8B3FACDF81A" box="[1236,1379,1953,1979]" pageId="34" pageNumber="485">Sungai Wain</collectingMunicipality>
Pro- tection
<location id="8E89601BFFB5446AFF12A8D7FE99F87E" LSID="urn:lsid:plazi:treatment:03FF87D6FFB5446CFF6BA827FDC3FAF6:8E89601BFFB5446AFF12A8D7FE99F87E" box="[238,311,1989,2015]" latitude="-1.135" longLatPrecision="130" longitude="116.831665" municipality="Sungai Wain" name="Forest" pageId="34" pageNumber="485" stateProvince="Kalimantan Timur">Forest</location>
,
<geoCoordinate id="EE625007FFB5446AFEB9A8D7FE06F87E" box="[325,424,1989,2015]" degrees="1" direction="south" minutes="08.1" orientation="latitude" pageId="34" pageNumber="485" precision="92" value="-1.135">1°08.1’S</geoCoordinate>
<geoCoordinate id="EE625007FFB5446AFE4FA8D7FD9BF87E" box="[435,565,1989,2015]" degrees="116" direction="east" minutes="49.9" orientation="longitude" pageId="34" pageNumber="485" precision="92" value="116.831665">116°49.9’E</geoCoordinate>
,
<quantity id="4CAE9B25FFB5446AFDB8A8D6FD2CF87E" box="[580,642,1988,2015]" metricMagnitude="1" metricUnit="m" metricValue="3.5" pageId="34" pageNumber="485" unit="m" value="35.0">
<elevation id="007BD1F3FFB5446AFDB8A8D6FD2CF87E" box="[580,642,1988,2015]" metricMagnitude="1" metricUnit="m" metricValue="3.5" pageId="34" pageNumber="485" unit="m" value="35.0">35 m</elevation>
</quantity>
,
<date id="FFE81000FFB5446AFD6EA8D7FC99F87E" box="[658,823,1989,2015]" pageId="34" pageNumber="485" value="2011-12-08" valueMax="2011-12-11" valueMin="2011-12-08">
<collectingDate id="EFACE9E8FFB5446AFD6EA8D7FC99F87E" box="[658,823,1989,2015]" pageId="34" pageNumber="485" value="2011-12-08" valueMax="2011-12-11" valueMin="2011-12-08">8–11.XII.2011</collectingDate>
</date>
, leg.
<collectorName id="26A35316FFB5446AFC86A8D7FC72F87E" box="[890,988,1989,2015]" pageId="34" pageNumber="485">J. Hájek</collectorName>
, J. Schneider &amp;
<collectorName id="26A35316FFB5446AFB5DA8D7FAB1F87E" box="[1185,1311,1989,2015]" pageId="34" pageNumber="485">P. Votruba.</collectorName>
</materialsCitation>
<materialsCitation id="3B3E3C9DFFB5446CFAD4A8D7FDDAFF12" ID-GBIF-Occurrence="2425474204" collectingDate="2011-12-08" collectingDateMax="2011-12-11" collectingDateMin="2011-12-08" collectionCode="NMPC" collectorName="J. Hajek &amp; Hajek, J &amp; P. Votruba." elevation="35" lastPageId="36" lastPageNumber="487" latitude="-1.135" location="Forest" longLatPrecision="130" longitude="116.831665" municipality="Sungai Wain" pageId="34" pageNumber="485" specimenCount="4" specimenCount-female="1" specimenCount-male="3" stateProvince="Kalimantan Timur" typeStatus="paratype">
<typeStatus id="54ED8862FFB5446AFAD4A8D7FA37F87E" box="[1320,1433,1989,2015]" pageId="34" pageNumber="485" type="paratype">Paratypes</typeStatus>
.
<specimenCount id="9D50FD49FFB3446CFF6BAF8BFF7CFF12" box="[151,210,153,179]" pageId="36" pageNumber="487" type="male">3♂♂</specimenCount>
,
<specimenCount id="9D50FD49FFB3446CFF22AF8BFEAEFF12" box="[222,256,153,179]" pageId="36" pageNumber="487" type="female">1♀</specimenCount>
(
<collectionCode id="ED47AE05FFB3446CFEECAF8BFECBFF12" box="[272,357,153,179]" country="Czech Republic" name="National Museum Prague" pageId="36" pageNumber="487">NMPC</collectionCode>
), same data as holotype
</materialsCitation>
;
<materialsCitation id="3B3E3C9DFFB3446CFD83AF8BFBA7FF5A" ID-GBIF-Occurrence="2425474202" collectingDate="2001-08" collectionCode="MNHUB, BMNH" collectorName="J. Weise &amp; Brendell &amp; Mendel" country="Indonesia" location="Central Kalimantan" longitude="113.98333" pageId="36" pageNumber="487" specimenCount="3" specimenCount-male="3" stateProvince="Central Kalimantan" typeStatus="paratype">
<specimenCount id="9D50FD49FFB3446CFD83AF8BFD0CFF12" box="[639,674,153,179]" pageId="36" pageNumber="487" type="male">1♂</specimenCount>
(
<collectionCode id="ED47AE05FFB3446CFD4DAF8BFCB1FF12" box="[689,799,153,179]" pageId="36" pageNumber="487">MNHUB</collectionCode>
), labeled: “Borneo / Nonfried [h, w] // ex coll. /
<collectorName id="26A35316FFB3446CFABCAF8BFA33FF12" box="[1344,1437,153,179]" pageId="36" pageNumber="487">J. Weise</collectorName>
[p, w] // Theopea / elegantula / Baly [h, w]”;
<emphasis id="B922EAD2FFB3446CFD60AFAEFC95FF76" bold="true" box="[668,827,188,215]" pageId="36" pageNumber="487">
<collectingCountry id="F3417650FFB3446CFD60AFAEFC95FF76" box="[668,827,188,215]" name="Indonesia" pageId="36" pageNumber="487">INDONESIA</collectingCountry>
</emphasis>
.
<collectingRegion id="4992F822FFB3446CFCB6AFAFFB82FF76" box="[842,1068,189,215]" country="Indonesia" name="Kalimantan Tengah" pageId="36" pageNumber="487">Central Kalimantan</collectingRegion>
:
<specimenCount id="9D50FD49FFB3446CFBCBAFAFFBC1FF76" box="[1079,1135,189,215]" pageId="36" pageNumber="487" type="male">2♂♂</specimenCount>
(
<collectionCode id="ED47AE05FFB3446CFB83AFAFFB77FF76" box="[1151,1241,189,215]" pageId="36" pageNumber="487">BMNH</collectionCode>
), Busang / Rekut confl.,
<geoCoordinate id="EE625007FFB3446CFF19AFF3FE98FF5A" box="[229,310,225,251]" pageId="36" pageNumber="487">0°03”S</geoCoordinate>
<geoCoordinate id="EE625007FFB3446CFEC0AFF2FE0EFF5A" box="[316,416,224,251]" degrees="113" direction="east" minutes="59" orientation="longitude" pageId="36" pageNumber="487" precision="925" value="113.98333">113°59E</geoCoordinate>
,
<date id="FFE81000FFB3446CFE56AFF3FDB5FF5A" box="[426,539,225,251]" pageId="36" pageNumber="487" value="2001-08">
<collectingDate id="EFACE9E8FFB3446CFE56AFF3FDB5FF5A" box="[426,539,225,251]" pageId="36" pageNumber="487" value="2001-08">VIII.2001</collectingDate>
</date>
, leg.
<collectorName id="26A35316FFB3446CFDA9AFF3FD1BFF5A" box="[597,693,225,251]" pageId="36" pageNumber="487">Brendell</collectorName>
&amp;
<collectorName id="26A35316FFB3446CFD2BAFF3FC9EFF5A" box="[727,816,225,251]" pageId="36" pageNumber="487">Mendel</collectorName>
;
<collectingRegion id="4992F822FFB3446CFCC7AFF3FBA7FF5A" box="[827,1033,225,251]" country="Indonesia" name="Kalimantan Selatan" pageId="36" pageNumber="487">South Kalimantan</collectingRegion>
</materialsCitation>
:
<materialsCitation id="3B3E3C9DFFB3446CFBEEAFF3FDF9FEBE" ID-GBIF-Occurrence="2425473989" collectionCode="USNM" collectorName="Martapura &amp; Doherty" country="Indonesia" pageId="36" pageNumber="487" specimenCount="10" specimenCount-female="2" specimenCount-male="8" stateProvince="Kalimantan Tengah" typeStatus="paratype">
<specimenCount id="9D50FD49FFB3446CFBEEAFF3FBE2FF5A" box="[1042,1100,225,251]" pageId="36" pageNumber="487" type="male">4♂♂</specimenCount>
,
<specimenCount id="9D50FD49FFB3446CFBABAFF3FBD4FF5A" box="[1111,1146,225,251]" pageId="36" pageNumber="487" type="female">1♀</specimenCount>
(
<specimenCount id="9D50FD49FFB3446CFB75AFF3FB6BFF5A" box="[1161,1221,225,251]" pageId="36" pageNumber="487" type="male">2♂♂</specimenCount>
: NMPC;
<specimenCount id="9D50FD49FFB3446CFAD0AFF3FAC9FF5A" box="[1324,1383,225,251]" pageId="36" pageNumber="487" type="male">2♂♂</specimenCount>
,
<specimenCount id="9D50FD49FFB3446CFA8DAFF3FA38FF5A" box="[1393,1430,225,251]" pageId="36" pageNumber="487" type="female">1♀</specimenCount>
:
<collectionCode id="ED47AE05FFB3446CFF6BAE17FF42FEBE" box="[151,236,261,287]" country="USA" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="36" pageNumber="487" type="Museum">USNM</collectionCode>
),
<collectorName id="26A35316FFB3446CFF03AE17FED9FEBE" box="[255,375,261,287]" pageId="36" pageNumber="487">Martapura</collectorName>
, 1891, leg.
<collectorName id="26A35316FFB3446CFE04AE17FDF9FEBE" box="[504,599,261,287]" pageId="36" pageNumber="487">Doherty</collectorName>
</materialsCitation>
;
<materialsCitation id="3B3E3C9DFFB3446CFD9FAE17FA88FEBE" ID-GBIF-Occurrence="2425474200" box="[611,1318,260,287]" collectionCode="BMNH" collectorName="Doherty &amp; Coll." country="Indonesia" location="Pangaron" pageId="36" pageNumber="487" specimenCount="2" specimenCount-male="2" stateProvince="Kalimantan Tengah" typeStatus="paratype">
<specimenCount id="9D50FD49FFB3446CFD9FAE17FD35FEBE" box="[611,667,261,287]" pageId="36" pageNumber="487" type="male">2♂♂</specimenCount>
(
<collectionCode id="ED47AE05FFB3446CFD56AE17FCAAFEBE" box="[682,772,261,287]" pageId="36" pageNumber="487">BMNH</collectionCode>
),
<location id="8E89601BFFB3446CFCEAAE17FC2DFEBE" LSID="urn:lsid:plazi:treatment:03FF87D6FFB5446CFF6BA827FDC3FAF6:8E89601BFFB3446CFCEAAE17FC2DFEBE" box="[790,899,261,287]" country="Indonesia" name="Pangaron" pageId="36" pageNumber="487" stateProvince="Kalimantan Tengah">Pangaron</location>
, leg.
<collectorName id="26A35316FFB3446CFC42AE17FBB6FEBE" box="[958,1048,261,287]" pageId="36" pageNumber="487">Doherty</collectorName>
, FREY
<collectorName id="26A35316FFB3446CFB88AE17FB00FEBE" box="[1140,1198,261,287]" pageId="36" pageNumber="487">Coll.</collectorName>
, 1905.100
</materialsCitation>
.
</paragraph>
<caption id="DF296648FFB4446BFF6BA89EFE8FF84C" ID-DOI="http://doi.org/10.5281/zenodo.3479645" ID-Zenodo-Dep="3479645" httpUri="https://zenodo.org/record/3479645/files/figure.png" pageId="35" pageNumber="486" startId="35.[151,250,1932,1957]" targetBox="[160,1426,193,1870]" targetPageId="35">
<paragraph id="8BE936C0FFB4446BFF6BA89EFE8FF84C" blockId="35.[151,1437,1932,2029]" pageId="35" pageNumber="486">
<emphasis id="B922EAD2FFB4446BFF6BA89EFE8EF804" bold="true" box="[151,288,1932,1957]" pageId="35" pageNumber="486">FIGURE 18.</emphasis>
Diagnostic characters of
<taxonomicName id="4C564D43FFB4446BFDDAA89FFD7FF804" authority="Lee &amp; Bezdĕk, 2019" authorityName="Lee &amp; Bezdĕk" authorityYear="2019" box="[550,721,1933,1957]" class="Insecta" family="Chrysomelidae" genus="Theopea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="486" phylum="Arthropoda" rank="species" species="houjayi" status="sp. nov.">
<emphasis id="B922EAD2FFB4446BFDDAA89FFD7FF804" box="[550,721,1933,1957]" italics="true" pageId="35" pageNumber="486">Theopea houjayi</emphasis>
</taxonomicName>
<emphasis id="B922EAD2FFB4446BFD24A89FFC87F804" bold="true" box="[728,809,1933,1957]" pageId="35" pageNumber="486">
<taxonomicNameLabel id="A21157A9FFB4446BFD24A89FFC87F804" box="[728,809,1933,1957]" pageId="35" pageNumber="486" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
18A. Antenna, male; 18B. Antenna, female; 18C. Aedeagus, dorsal view; 18D. Aedeagus, lateral view; 18E. Aedeagus, ventral view; 18F. Abdominal ventrite VIII; 18G. Gonocoxae; 18H. Spermatheca.
</paragraph>
</caption>
<paragraph id="8BE936C0FFB3446CFF3BAE3AFE7AFBB2" blockId="36.[151,1437,152,1367]" pageId="36" pageNumber="487">
<emphasis id="B922EAD2FFB3446CFF3BAE3AFEF7FEE2" bold="true" box="[199,345,296,323]" pageId="36" pageNumber="487">Description.</emphasis>
Length 6.4–7.0 mm, width
<quantity id="4CAE9B25FFB3446CFD5DAE3BFC80FEE2" box="[673,814,297,323]" metricMagnitude="-3" metricUnit="m" metricValue="2.55" metricValueMax="2.7" metricValueMin="2.4" pageId="36" pageNumber="487" unit="mm" value="2.55" valueMax="2.7" valueMin="2.4">2.4–2.7 mm</quantity>
. General color (
<figureCitation id="136D2A45FFB3446CFC16AE3BFBEEFEE2" box="[1002,1088,297,323]" captionStart="FIGURE 14" captionStartId="29.[151,250,1897,1922]" captionTargetBox="[238,1338,183,1873]" captionTargetId="figure@29.[232,1356,174,1878]" captionTargetPageId="29" captionText="FIGURE 14. Habitus of Theopea elegantula Baly, T. houjayi sp. nov., and T. yuae sp. nov. 14A. T. elegantula, male, dorsal view; 14B. Same, ventral view; 14C. T. houjayi sp. nov., male, dorsal view; 14D. Same, ventral view; 14E. T. houjayi sp. nov., female, dorsal view; 14F. T. yuae sp. nov., male, dorsal view; 14G. Same, ventral view; 14H. T. yuae sp. nov., male, dorsal view, color variation; 14I. T. yuae sp. nov., male, dorsal view, color variation." figureDoi="http://doi.org/10.5281/zenodo.3479637" httpUri="https://zenodo.org/record/3479637/files/figure.png" pageId="36" pageNumber="487">Figs 14</figureCitation>
C–14E) metallic blue or grey; antenna black except antennomeres IX–XI white or pale yellow. Antennae filiform in males, but antennomere VII distinctly swollen, X with a broad lateral process at middle (
<figureCitation id="136D2A45FFB3446CFCCCAE63FC39FE2A" box="[816,919,369,395]" captionStart="FIGURE 18" captionStartId="35.[151,250,1932,1957]" captionTargetBox="[160,1426,193,1870]" captionTargetId="figure@35.[151,1436,174,1879]" captionTargetPageId="35" captionText="FIGURE 18. Diagnostic characters of Theopea houjayi sp. nov. 18A. Antenna, male; 18B. Antenna, female; 18C. Aedeagus, dorsal view; 18D. Aedeagus, lateral view; 18E. Aedeagus, ventral view; 18F. Abdominal ventrite VIII; 18G. Gonocoxae; 18H. Spermatheca." figureDoi="http://doi.org/10.5281/zenodo.3479645" httpUri="https://zenodo.org/record/3479645/files/figure.png" pageId="36" pageNumber="487">Fig. 18A</figureCitation>
), length ratios of antennomeres I–XI 1.0: 0.3: 0.8: 0.9: 0.9: 0.9: 1.0: 0.9: 0.9: 0.9: 1.1, length to width ratios of antennomeres I–XI 2.9: 1.3: 2.9: 3.2: 3.0: 2.9: 2.4: 2.6: 2.3: 2.4: 4.6; filiform in females (
<figureCitation id="136D2A45FFB3446CFD69AEABFD54FE72" box="[661,762,441,467]" captionStart="FIGURE 18" captionStartId="35.[151,250,1932,1957]" captionTargetBox="[160,1426,193,1870]" captionTargetId="figure@35.[151,1436,174,1879]" captionTargetPageId="35" captionText="FIGURE 18. Diagnostic characters of Theopea houjayi sp. nov. 18A. Antenna, male; 18B. Antenna, female; 18C. Aedeagus, dorsal view; 18D. Aedeagus, lateral view; 18E. Aedeagus, ventral view; 18F. Abdominal ventrite VIII; 18G. Gonocoxae; 18H. Spermatheca." figureDoi="http://doi.org/10.5281/zenodo.3479645" httpUri="https://zenodo.org/record/3479645/files/figure.png" pageId="36" pageNumber="487">Fig. 18B</figureCitation>
), length ratios of antennomeres I–XI 1.0: 0.3: 0.7: 0.8: 0.8: 0.8: 0.9: 0.9: 0.8: 0.8: 1.0, length to width ratios of antennomeres I–XI 3.3: 1.8: 2.8: 3.2: 2.9: 2.7: 3.2: 3.3: 3.4: 3.5: 4.3. Elytra elongate and parallel-sided, 1.9x longer than wide; disc with dense, coarse punctures, arranged into longitudinal rows, with one weak longitudinal ridge between two longitudinal rows of punctures, ridges near suture abbreviated. Tarsomeres I of front legs swollen in males. Aedeagus (
<figureCitation id="136D2A45FFB3446CFC24AD5BFB84FDC2" box="[984,1066,585,611]" captionStart="FIGURE 18" captionStartId="35.[151,250,1932,1957]" captionTargetBox="[160,1426,193,1870]" captionTargetId="figure@35.[151,1436,174,1879]" captionTargetPageId="35" captionText="FIGURE 18. Diagnostic characters of Theopea houjayi sp. nov. 18A. Antenna, male; 18B. Antenna, female; 18C. Aedeagus, dorsal view; 18D. Aedeagus, lateral view; 18E. Aedeagus, ventral view; 18F. Abdominal ventrite VIII; 18G. Gonocoxae; 18H. Spermatheca." figureDoi="http://doi.org/10.5281/zenodo.3479645" httpUri="https://zenodo.org/record/3479645/files/figure.png" pageId="36" pageNumber="487">Figs 18</figureCitation>
C–18E) slender, 5.9x longer than wide; sides widest at apical 1/3, abruptly narrowed and then gradually broadened towards apex, apex with median, shallow and narrow notch; gradually narrowed towards base; tectum well sclerotized, basally broadened, broader than aedeagus at base; almost straight from apical 2/
<quantity id="4CAE9B25FFB3446CFD16ADA6FCBBFD6E" box="[746,789,692,719]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="36" pageNumber="487" unit="in" value="5.0">5 in</quantity>
lateral view, and then slightly curved, straight from apex to apical 1/6; ventral surface with deep notch from apex to apical 2/5, mesally extending and approximate at apical 1/6; triangular sclerites small; internal sac with one median, elongate sclerite, gradually broadened towards base, about 0.5x as long as aedeagus, with two pairs of hook-like sclerites locate subbasally and subapically. Gonocoxae (
<figureCitation id="136D2A45FFB3446CFF5CAC57FEA9FCFE" box="[160,263,837,863]" captionStart="FIGURE 18" captionStartId="35.[151,250,1932,1957]" captionTargetBox="[160,1426,193,1870]" captionTargetId="figure@35.[151,1436,174,1879]" captionTargetPageId="35" captionText="FIGURE 18. Diagnostic characters of Theopea houjayi sp. nov. 18A. Antenna, male; 18B. Antenna, female; 18C. Aedeagus, dorsal view; 18D. Aedeagus, lateral view; 18E. Aedeagus, ventral view; 18F. Abdominal ventrite VIII; 18G. Gonocoxae; 18H. Spermatheca." figureDoi="http://doi.org/10.5281/zenodo.3479645" httpUri="https://zenodo.org/record/3479645/files/figure.png" pageId="36" pageNumber="487">Fig. 18G</figureCitation>
) elongate, widest at apical 1/6, both gonocoxae combined together from basal 1/7 to apical 1/6; apices narrowly rounded, each gonocoxa with eight setae along outer margin from apex to apical 1/6; with one pair of short lateral processes at basal 2/5. Ventrite VIII (
<figureCitation id="136D2A45FFB3446CFD82AC9FFD70FC06" box="[638,734,909,935]" captionStart="FIGURE 18" captionStartId="35.[151,250,1932,1957]" captionTargetBox="[160,1426,193,1870]" captionTargetId="figure@35.[151,1436,174,1879]" captionTargetPageId="35" captionText="FIGURE 18. Diagnostic characters of Theopea houjayi sp. nov. 18A. Antenna, male; 18B. Antenna, female; 18C. Aedeagus, dorsal view; 18D. Aedeagus, lateral view; 18E. Aedeagus, ventral view; 18F. Abdominal ventrite VIII; 18G. Gonocoxae; 18H. Spermatheca." figureDoi="http://doi.org/10.5281/zenodo.3479645" httpUri="https://zenodo.org/record/3479645/files/figure.png" pageId="36" pageNumber="487">Fig. 18F</figureCitation>
) elongate; disc with several long setae laterally and near apical margin, and with dense, short setae along apical margin; spiculum extremely slender. Receptacle of spermatheca (
<figureCitation id="136D2A45FFB3446CFF5CACC7FEA9FC4E" box="[160,263,981,1007]" captionStart="FIGURE 18" captionStartId="35.[151,250,1932,1957]" captionTargetBox="[160,1426,193,1870]" captionTargetId="figure@35.[151,1436,174,1879]" captionTargetPageId="35" captionText="FIGURE 18. Diagnostic characters of Theopea houjayi sp. nov. 18A. Antenna, male; 18B. Antenna, female; 18C. Aedeagus, dorsal view; 18D. Aedeagus, lateral view; 18E. Aedeagus, ventral view; 18F. Abdominal ventrite VIII; 18G. Gonocoxae; 18H. Spermatheca." figureDoi="http://doi.org/10.5281/zenodo.3479645" httpUri="https://zenodo.org/record/3479645/files/figure.png" pageId="36" pageNumber="487">Fig. 18H</figureCitation>
) strongly swollen; pump slender and strongly curved; proximal spermathecal duct deeply inserted into receptacle, narrow and short
</paragraph>
</subSubSection>
<subSubSection id="C34C654BFFB3446CFF3BAB0EFCB7FB4A" pageId="36" pageNumber="487" type="diagnosis">
<paragraph id="8BE936C0FFB3446CFF3BAB0EFCB7FB4A" blockId="36.[151,1437,152,1367]" pageId="36" pageNumber="487">
<emphasis id="B922EAD2FFB3446CFF3BAB0EFEECFB96" bold="true" box="[199,322,1052,1079]" pageId="36" pageNumber="487">Diagnosis.</emphasis>
Specimens of
<taxonomicName id="4C564D43FFB3446CFE15AB0CFDF7FB96" authority="Lee &amp; Bezdĕk, 2019" authorityName="Lee &amp; Bezdĕk" authorityYear="2019" box="[489,601,1053,1079]" class="Insecta" family="Chrysomelidae" genus="Theopea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="487" phylum="Arthropoda" rank="species" species="houjayi" status="sp. nov.">
<emphasis id="B922EAD2FFB3446CFE15AB0CFDF7FB96" box="[489,601,1053,1079]" italics="true" pageId="36" pageNumber="487">T. houjayi</emphasis>
</taxonomicName>
<emphasis id="B922EAD2FFB3446CFD9DAB0FFD15FB96" bold="true" box="[609,699,1053,1079]" pageId="36" pageNumber="487">
<taxonomicNameLabel id="A21157A9FFB3446CFD9DAB0FFD15FB96" box="[609,699,1053,1079]" pageId="36" pageNumber="487" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
are similar to those of
<taxonomicName id="4C564D43FFB3446CFC3EAB0CFBBDFB96" authority="Lee &amp; Bezdĕk, 2019" authorityName="Lee &amp; Bezdĕk" authorityYear="2019" box="[962,1043,1054,1079]" class="Insecta" family="Chrysomelidae" genus="Theopea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="487" phylum="Arthropoda" rank="species" species="yuae" status="sp. nov.">
<emphasis id="B922EAD2FFB3446CFC3EAB0CFBBDFB96" box="[962,1043,1054,1079]" italics="true" pageId="36" pageNumber="487">T. yuae</emphasis>
</taxonomicName>
<emphasis id="B922EAD2FFB3446CFBE7AB0FFBDBFB96" bold="true" box="[1051,1141,1053,1079]" pageId="36" pageNumber="487">
<taxonomicNameLabel id="A21157A9FFB3446CFBE7AB0FFBDBFB96" box="[1051,1141,1053,1079]" pageId="36" pageNumber="487" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in possessing three apical white antenomeres (two apical white antennomeres in
<taxonomicName id="4C564D43FFB3446CFD0BAB50FCE6FBFA" authority="Lee &amp; Bezdĕk, 2019" authorityName="Lee &amp; Bezdĕk" authorityYear="2019" box="[759,840,1090,1115]" class="Insecta" family="Chrysomelidae" genus="Theopea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="487" phylum="Arthropoda" rank="species" species="tsoui" status="sp. nov.">
<emphasis id="B922EAD2FFB3446CFD0BAB50FCE6FBFA" box="[759,840,1090,1115]" italics="true" pageId="36" pageNumber="487">T. tsoui</emphasis>
</taxonomicName>
<emphasis id="B922EAD2FFB3446CFCACAB53FC07FBFA" bold="true" box="[848,937,1089,1115]" pageId="36" pageNumber="487">
<taxonomicNameLabel id="A21157A9FFB3446CFCACAB53FC07FBFA" box="[848,937,1089,1115]" pageId="36" pageNumber="487" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C564D43FFB3446CFC22AB50FBD9FBFA" box="[990,1143,1089,1115]" class="Insecta" family="Chrysomelidae" genus="Theopea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="487" phylum="Arthropoda" rank="species" species="kedenburgi">
<emphasis id="B922EAD2FFB3446CFC22AB50FBD9FBFA" box="[990,1143,1089,1115]" italics="true" pageId="36" pageNumber="487">T. kedenburgi</emphasis>
</taxonomicName>
) and reduced longitudinal ridges near the suture of the elytra (distinct longitudinal ridges near suture of elytra in
<taxonomicName id="4C564D43FFB3446CFBB0AB74FB72FBDE" box="[1100,1244,1125,1151]" class="Insecta" family="Chrysomelidae" genus="Theopea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="487" phylum="Arthropoda" rank="species" species="elegantula">
<emphasis id="B922EAD2FFB3446CFBB0AB74FB72FBDE" box="[1100,1244,1125,1151]" italics="true" pageId="36" pageNumber="487">T. elegantula</emphasis>
</taxonomicName>
). They cannot be separated from each other based on external morphology. However,
<taxonomicName id="4C564D43FFB3446CFC73AB98FC50FB02" authority="Lee &amp; Bezdĕk, 2019" authorityName="Lee &amp; Bezdĕk" authorityYear="2019" box="[911,1022,1161,1187]" class="Insecta" family="Chrysomelidae" genus="Theopea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="487" phylum="Arthropoda" rank="species" species="houjayi" status="sp. nov.">
<emphasis id="B922EAD2FFB3446CFC73AB98FC50FB02" box="[911,1022,1161,1187]" italics="true" pageId="36" pageNumber="487">T. houjayi</emphasis>
</taxonomicName>
<emphasis id="B922EAD2FFB3446CFBF9AB9BFBF0FB02" bold="true" box="[1029,1118,1161,1187]" pageId="36" pageNumber="487">
<taxonomicNameLabel id="A21157A9FFB3446CFBF9AB9BFBF0FB02" box="[1029,1118,1161,1187]" pageId="36" pageNumber="487" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is only found in Kalimantan and males have slender and notched apices of the aedeagi, in contrast to those of
<taxonomicName id="4C564D43FFB3446CFBE6ABBCFBC4FB66" authority="Lee &amp; Bezdĕk, 2019" authorityName="Lee &amp; Bezdĕk" authorityYear="2019" box="[1050,1130,1198,1223]" class="Insecta" family="Chrysomelidae" genus="Theopea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="487" phylum="Arthropoda" rank="species" species="yuae" status="sp. nov.">
<emphasis id="B922EAD2FFB3446CFBE6ABBCFBC4FB66" box="[1050,1130,1198,1223]" italics="true" pageId="36" pageNumber="487">T. yuae</emphasis>
</taxonomicName>
<emphasis id="B922EAD2FFB3446CFB8CABBFFB67FB66" bold="true" box="[1136,1225,1197,1223]" pageId="36" pageNumber="487">
<taxonomicNameLabel id="A21157A9FFB3446CFB8CABBFFB67FB66" box="[1136,1225,1197,1223]" pageId="36" pageNumber="487" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, which is confined to
<collectingCountry id="F3417650FFB3446CFF48ABC3FEBAFB4A" box="[180,276,1233,1259]" name="Malaysia" pageId="36" pageNumber="487">Sarawak</collectingCountry>
and has broad and rounded apices of aedeagi.
</paragraph>
</subSubSection>
<subSubSection id="C34C654BFFB3446CFF3BABE7FE0BFA92" pageId="36" pageNumber="487" type="etymology">
<paragraph id="8BE936C0FFB3446CFF3BABE7FE0BFA92" blockId="36.[151,1437,152,1367]" pageId="36" pageNumber="487">
<emphasis id="B922EAD2FFB3446CFF3BABE7FEE0FAAE" bold="true" box="[199,334,1269,1295]" pageId="36" pageNumber="487">Etymology.</emphasis>
The new species is dedicated to Mr. Hou-Jay Chen, a member of TCRT for his contribution to diversity of leaf beetles.
</paragraph>
</subSubSection>
<subSubSection id="C34C654BFFB3446CFF3BAA2EFDC3FAF6" box="[199,621,1340,1367]" pageId="36" pageNumber="487" type="distribution">
<paragraph id="8BE936C0FFB3446CFF3BAA2EFDC3FAF6" blockId="36.[151,1437,152,1367]" box="[199,621,1340,1367]" pageId="36" pageNumber="487">
<emphasis id="B922EAD2FFB3446CFF3BAA2EFECFFAF6" bold="true" box="[199,353,1340,1367]" pageId="36" pageNumber="487">Distribution.</emphasis>
<collectingCountry id="F3417650FFB3446CFE9BAA2FFE76FAF6" box="[359,472,1341,1367]" name="Indonesia" pageId="36" pageNumber="487">Indonesia</collectingCountry>
:
<collectingRegion id="4992F822FFB3446CFE1FAA2FFDC7FAF6" box="[483,617,1341,1367]" country="Indonesia" name="Kalimantan Utara" pageId="36" pageNumber="487">Kalimantan</collectingRegion>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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