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<document id="92C30DC473FD27A786E75359ABCCDADA" ID-DOI="10.1080/00222933.2010.534190" ID-ISSN="1464-5262" ID-Zenodo-Dep="4666850" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1617734185754" checkinUser="carolina" docAuthor="Yang, Zhuo, Zhou, Hong-Zhang &amp; Schillhammer, Harald" docDate="2011" docId="03FF87CDFFEC8632FE5BF9CCFD44D3C6" docLanguage="en" docName="JNATHIST.45.7-8.407-433.pdf" docOrigin="Journal of Natural History 45 (7 - 8)" docSource="http://dx.doi.org/10.1080/00222933.2010.534190" docStyle="DocumentStyle:DE8A86AD4006EFA56F95B2AAEFBF2813.5:JNatHist.2008-2012.journal_article.2cover" docStyleId="DE8A86AD4006EFA56F95B2AAEFBF2813" docStyleName="JNatHist.2008-2012.journal_article.2cover" docStyleVersion="5" docTitle="Thoracostrongylus aduncatus Yang &amp; Zhou &amp; Schillhammer 2011, sp. nov." docType="treatment" docVersion="3" lastPageNumber="414" masterDocId="FFC6FFB5FFE4863BFFBDFFCBFFF7D554" masterDocTitle="Taxonomy of the genus Thoracostrongylus Bernhauer (Coleoptera: Staphylinidae) with descriptions of five new species from China" masterLastPageNumber="433" masterPageNumber="407" pageNumber="413" updateTime="1698901293237" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="655F6ECFBCA83D5A0F66B8D05FD4A6D3" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="348B78103A2C4CC05B834E1B34729E7A">
<mods:title id="AFCB31DEDAEC91DDC425614F727E0C5B">Taxonomy of the genus Thoracostrongylus Bernhauer (Coleoptera: Staphylinidae) with descriptions of five new species from China</mods:title>
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<mods:name id="0B425DE0DC4376970B104A1FEB393A0B" type="personal">
<mods:role id="B3FEB7A29A65103DEEE7A619929E96B3">
<mods:roleTerm id="79C784BEAF10F9C1D80401040BACEF06">Author</mods:roleTerm>
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<mods:namePart id="2DF73E55928EDF4CCC684D5DB9CC827B">Yang, Zhuo</mods:namePart>
</mods:name>
<mods:name id="5E2AD3E60C064FE9B4126AF057DBC599" type="personal">
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<mods:roleTerm id="E1021B520E070AC2C51FB4092141F5F3">Author</mods:roleTerm>
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<mods:namePart id="B5BD88FB599BD5A04A21BB83DD0EAC00">Zhou, Hong-Zhang</mods:namePart>
</mods:name>
<mods:name id="1909ECDF5F899FB94DAAC253A44D728C" type="personal">
<mods:role id="C158AB7B1B93A7376F7817B93C8ED99D">
<mods:roleTerm id="F7DA58A3170CC569BDA42CF1836DDC76">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="051F88E58494974EF4C377D4E357934C">Schillhammer, Harald</mods:namePart>
</mods:name>
<mods:typeOfResource id="34EF9D5EDEE55BD7F0C4C5C76EBD6AB2">text</mods:typeOfResource>
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<mods:title id="6C6E200F19ED9F2F653516C95B9D0C14">Journal of Natural History</mods:title>
</mods:titleInfo>
<mods:part id="E2792DC852206ED7462572243CA356D1">
<mods:date id="A5F80448E6A1F9EA6B6884AD0F2D966D">2011</mods:date>
<mods:detail id="ACE2825B3E0E6F4D0BC37B8EDCC3F081" type="pubDate">
<mods:number id="B819A25E99E3B456923E2A698836D254">2011-02-08</mods:number>
</mods:detail>
<mods:detail id="22D85F926A4A941F660F42FEDB80A9DB" type="volume">
<mods:number id="BEA26919CADD85281444738D169E2D31">45</mods:number>
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<mods:number id="C77285A0BB582EE2C9EBB064CF550A3D">7 - 8</mods:number>
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<mods:url id="A509E8F76478D6C51D137567319B0D67">http://dx.doi.org/10.1080/00222933.2010.534190</mods:url>
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<mods:classification id="16DD43F91C05C78CC321BDF5F40314DF">journal article</mods:classification>
<mods:identifier id="4425A6CEEBEFC1FDA4F469A0D6D00639" type="DOI">10.1080/00222933.2010.534190</mods:identifier>
<mods:identifier id="50B2AED9EF9B5DC5895748627EFF0F94" type="ISSN">1464-5262</mods:identifier>
<mods:identifier id="93866E23CDB369C08536D837D881A95D" type="Zenodo-Dep">4666850</mods:identifier>
</mods:mods>
<treatment id="03FF87CDFFEC8632FE5BF9CCFD44D3C6" LSID="urn:lsid:plazi:treatment:03FF87CDFFEC8632FE5BF9CCFD44D3C6" httpUri="http://treatment.plazi.org/id/03FF87CDFFEC8632FE5BF9CCFD44D3C6" lastPageId="9" lastPageNumber="414" pageId="8" pageNumber="413">
<subSubSection id="C34C6550FFEC8633FE5BF9CCFC75D34B" box="[486,898,1543,1567]" pageId="8" pageNumber="413" type="nomenclature">
<paragraph id="8BE936DBFFEC8633FE5BF9CCFC75D34B" blockId="8.[486,898,1543,1567]" box="[486,898,1543,1567]" pageId="8" pageNumber="413">
<heading id="D0A181B7FFEC8633FE5BF9CCFC75D34B" box="[486,898,1543,1567]" centered="true" fontSize="36" level="2" pageId="8" pageNumber="413" reason="7">
<taxonomicName id="4C564D58FFEC8633FE5BF9CCFCD0D34B" ID-CoL="56KH4" authority="Yang &amp; Zhou &amp; Schillhammer, 2011" authorityName="Yang &amp; Zhou &amp; Schillhammer" authorityYear="2011" box="[486,807,1543,1567]" class="Insecta" family="Staphylinidae" genus="Thoracostrongylus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="413" phylum="Arthropoda" rank="species" species="aduncatus" status="sp. nov.">
<emphasis id="B922EAC9FFEC8633FE5BF9CCFCD0D34B" bold="true" box="[486,807,1543,1567]" italics="true" pageId="8" pageNumber="413">Thoracostrongylus aduncatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21157B2FFEC8633FC93F9C3FC75D34B" box="[814,898,1544,1567]" pageId="8" pageNumber="413" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34C6550FFEC8632FDFAF9FFFD44D3C6" lastPageId="9" lastPageNumber="414" pageId="8" pageNumber="413" type="description">
<paragraph id="8BE936DBFFEC8633FDFAF9FFFCD5D318" blockId="8.[583,802,1588,1612]" box="[583,802,1588,1612]" pageId="8" pageNumber="413">
(
<figureCitation id="136D2A5EFFEC8633FDF3F9FFFD3BD318" box="[590,716,1588,1612]" captionStart="Figure 1" captionStartId="7.[144,211,1692,1714]" captionTargetBox="[143,1099,162,1653]" captionTargetId="figure-5@7.[142,1102,160,1656]" captionTargetPageId="7" captionText="Figure 1. (A) Thoracostrongylus acerosus sp. nov.; (B) Thoracostrongylus aduncatus sp. nov.; (C) Thoracostrongylus baoxingensis sp. nov.; (D) Thoracostrongylus diaoluoensis sp. nov.; (E) Thoracostrongylus fujianensis sp. nov.; (F) Thoracostrongylus birmanus, (Fauvel, 1895)." figureDoi="http://doi.org/10.5281/zenodo.4666852" httpUri="https://zenodo.org/record/4666852/files/figure.png" pageId="8" pageNumber="413">Figures 1B</figureCitation>
,
<figureCitation id="136D2A5EFFEC8633FD65F9FFFD13D318" box="[728,740,1588,1612]" captionStart="Figure 3" captionStartId="10.[213,280,1142,1164]" captionTargetBox="[215,1168,162,1108]" captionTargetId="figure-5@10.[211,1171,161,1110]" captionTargetPageId="10" captionText="Figure 3. Thoracostrongylus aduncatus sp. nov., holotype, male from Yunnan, China. (A) 8th sternite, male; (B) 10th tergite, male; (C) 9th sternite, male; (D) aedeagus, ventral view; (E) aedeagus, lateral view. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4666856" httpUri="https://zenodo.org/record/4666856/files/figure.png" pageId="8" pageNumber="413">3</figureCitation>
A–E)
</paragraph>
<paragraph id="8BE936DBFFEC8633FF68F9B3FE99D3DB" blockId="8.[213,366,1655,1679]" box="[213,366,1655,1679]" pageId="8" pageNumber="413">
<emphasis id="B922EAC9FFEC8633FF68F9B3FE99D3DB" box="[213,366,1655,1679]" italics="true" pageId="8" pageNumber="413">Type material</emphasis>
</paragraph>
<paragraph id="8BE936DBFFEC8633FF68F954FD13D382" blockId="8.[213,1170,1695,1750]" pageId="8" pageNumber="413">
<materialsCitation id="3B3E3C86FFEC8633FF68F954FD13D382" collectorName="Gaoligongshan Nat. Res., E &amp; O. Sausa" country="China" location="Baoshan" pageId="8" pageNumber="413" specimenCount="1" stateProvince="Yunnan" typeStatus="holotype">
<emphasis id="B922EAC9FFEC8633FF68F954FECBD3E3" box="[213,316,1695,1719]" italics="true" pageId="8" pageNumber="413">
<typeStatus id="54ED8879FFEC8633FF68F954FECBD3E3" box="[213,316,1695,1719]" pageId="8" pageNumber="413" type="holotype">Holotype</typeStatus>
</emphasis>
. Male,
<collectingCountry id="F341764BFFEC8633FE30F954FE19D3E3" box="[397,494,1695,1719]" name="China" pageId="8" pageNumber="413">CHINA</collectingCountry>
:
<collectingRegion id="4992F839FFEC8633FE4BF954FDA4D3E2" box="[502,595,1695,1718]" country="China" name="Yunnan" pageId="8" pageNumber="413">Yunnan</collectingRegion>
, 14
<emphasis id="B922EAC9FFEC8633FDCAF954FD73D3E3" bold="true" box="[631,644,1695,1719]" pageId="8" pageNumber="413">–</emphasis>
21.6,
<quantity id="4CAE9B3EFFEC8633FD02F954FCE6D3E3" box="[703,785,1695,1719]" metricMagnitude="5" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="413" unit="km" value="100.0">100 km</quantity>
W
<location id="8E896000FFEC8633FC88F954FC6DD3E3" LSID="urn:lsid:plazi:treatment:03FF87CDFFEC8632FE5BF9CCFD44D3C6:8E896000FFEC8633FC88F954FC6DD3E3" box="[821,922,1695,1719]" country="China" name="Baoshan" pageId="8" pageNumber="413" stateProvince="Yunnan">Baoshan</location>
, 1993,
<collectorName id="26A3530DFFEC8633FC58F954FE94D382" pageId="8" pageNumber="413">Gaoligongshan Nat. Res., E</collectorName>
. Jendek &amp;
<collectorName id="26A3530DFFEC8633FE5EF975FDBFD382" box="[483,584,1726,1750]" pageId="8" pageNumber="413">O. Sausa</collectorName>
leg. (NMW).
</materialsCitation>
</paragraph>
<paragraph id="8BE936DBFFED8632FF2DFF6CFEF8D5EA" blockId="9.[144,271,167,190]" box="[144,271,167,190]" pageId="9" pageNumber="414">
<emphasis id="B922EAC9FFED8632FF2DFF6CFEF8D5EA" box="[144,271,167,190]" italics="true" pageId="9" pageNumber="414">Description</emphasis>
</paragraph>
<paragraph id="8BE936DBFFED8632FF2DFF04FDB5D452" blockId="9.[144,1101,206,262]" pageId="9" pageNumber="414">
<emphasis id="B922EAC9FFED8632FF2DFF04FEC7D5B2" box="[144,304,207,230]" italics="true" pageId="9" pageNumber="414">Measurements</emphasis>
. Body length:
<quantity id="4CAE9B3EFFED8632FE6BFF05FDD8D5B2" box="[470,559,206,230]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" pageId="9" pageNumber="414" unit="mm" value="9.5">9.5 mm</quantity>
.
<specimenCount id="9D50FD52FFED8632FD84FF05FDB9D5B2" box="[569,590,206,230]" pageId="9" pageNumber="414" type="male">♂</specimenCount>
: HL: 1.3; HW: 1.8; CL: 0.9; PO: 0.2; PL: 1.7; PW: 1.5; EL: 2.1; EW: 2.05; ELS: 1.15.
</paragraph>
<paragraph id="8BE936DBFFED8632FF2DFEE5FC83D4D1" blockId="9.[144,1101,302,1282]" pageId="9" pageNumber="414">
<emphasis id="B922EAC9FFED8632FF2DFEE5FF3DD412" box="[144,202,302,326]" italics="true" pageId="9" pageNumber="414">Male</emphasis>
. Dark brown, dull; mandibles reddish black, maxillary palpi, labial palpi and first four segments of antennae reddish brown, remaining segments black brown; legs brown yellow, femora with black ring at midlength and distal end.
</paragraph>
<paragraph id="8BE936DBFFED8632FF06FE46FC99D7D0" blockId="9.[144,1101,302,1282]" pageId="9" pageNumber="414">
Head large, transverse, distinctly wider than pronotum; eyes large and prominent, occupying almost entire sides, longitudinal diameter four times as long as temporal region; vertex with a small longitudinal specular black spot medially, more or less indistinct; surface densely and regularly covered with umbilicate punctures, with short, scarce, brassy pubescence. Antennae with 1st segment longest, 2nd narrower than and almost one-third as long as 1st, 3rd narrower and two times longer than 2nd, 4th longer than wide, 5
<emphasis id="B922EAC9FFED8632FE27FD87FE50D730" bold="true" box="[410,423,588,612]" pageId="9" pageNumber="414">–</emphasis>
10th gradually increasing in breadth, longer than wide, last segment two times as long as wide and subacuminate towards tip.
</paragraph>
<paragraph id="8BE936DBFFED8632FF06FD46FCDAD610" blockId="9.[144,1101,302,1282]" pageId="9" pageNumber="414">Pronotum oblong and dorsal surface well convex, distinctly longer than wide, and much narrower than elytra; widest at level of large lateral seta, slightly narrowed towards rounded anterior angles, distinctly concavely narrowed towards base; anterior margin almost straight, posterior margin moderately convex, hind angles broadly rounded; punctation and pubescence similar to that on head, with brassy pubescence. Scutellum triangular, with dense, black, velvety pubescence.</paragraph>
<paragraph id="8BE936DBFFED8632FF06FC80FCEED6D7" blockId="9.[144,1101,302,1282]" pageId="9" pageNumber="414">Elytra subquadrate, slightly dilated posteriorly; surface densely and finely punctate, with brassy pubescence, mixed with grey pubescence.</paragraph>
<paragraph id="8BE936DBFFED8632FF06FC40FEE1D137" blockId="9.[144,1101,302,1282]" pageId="9" pageNumber="414">
Abdomen gradually narrower towards apex, finely and densely punctate; 3–7th tergites dark, with black pubescence, abdomen lateral plate black; apical margin of 8th sternite (
<figureCitation id="136D2A5EFFED8632FE9FFC00FE6FD6B7" box="[290,408,971,995]" captionStart="Figure 3" captionStartId="10.[213,280,1142,1164]" captionTargetBox="[215,1168,162,1108]" captionTargetId="figure-5@10.[211,1171,161,1110]" captionTargetPageId="10" captionText="Figure 3. Thoracostrongylus aduncatus sp. nov., holotype, male from Yunnan, China. (A) 8th sternite, male; (B) 10th tergite, male; (C) 9th sternite, male; (D) aedeagus, ventral view; (E) aedeagus, lateral view. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4666856" httpUri="https://zenodo.org/record/4666856/files/figure.png" pageId="9" pageNumber="414">Figure 3A</figureCitation>
) shallowly and broadly excised medially, with densely short, golden, erect pubescence; 10th tergite (
<figureCitation id="136D2A5EFFED8632FDE9FC20FD3ED157" box="[596,713,1003,1027]" captionStart="Figure 3" captionStartId="10.[213,280,1142,1164]" captionTargetBox="[215,1168,162,1108]" captionTargetId="figure-5@10.[211,1171,161,1110]" captionTargetPageId="10" captionText="Figure 3. Thoracostrongylus aduncatus sp. nov., holotype, male from Yunnan, China. (A) 8th sternite, male; (B) 10th tergite, male; (C) 9th sternite, male; (D) aedeagus, ventral view; (E) aedeagus, lateral view. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4666856" httpUri="https://zenodo.org/record/4666856/files/figure.png" pageId="9" pageNumber="414">Figure 3B</figureCitation>
) wider than long, posterior margin almost straight with abundant long golden pubescence; apical margin of 9th sternite (
<figureCitation id="136D2A5EFFED8632FF45FBE0FE86D117" box="[248,369,1067,1091]" captionStart="Figure 3" captionStartId="10.[213,280,1142,1164]" captionTargetBox="[215,1168,162,1108]" captionTargetId="figure-5@10.[211,1171,161,1110]" captionTargetPageId="10" captionText="Figure 3. Thoracostrongylus aduncatus sp. nov., holotype, male from Yunnan, China. (A) 8th sternite, male; (B) 10th tergite, male; (C) 9th sternite, male; (D) aedeagus, ventral view; (E) aedeagus, lateral view. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4666856" httpUri="https://zenodo.org/record/4666856/files/figure.png" pageId="9" pageNumber="414">Figure 3C</figureCitation>
) shallowly emarginate medially, with moderately long, yellow pubescence.
</paragraph>
<paragraph id="8BE936DBFFED8632FF06FBA1FBBAD056" blockId="9.[144,1101,302,1282]" pageId="9" pageNumber="414">
Male genitalia (
<figureCitation id="136D2A5EFFED8632FEC4FBA1FDECD1D6" box="[377,539,1130,1154]" captionStart="Figure 3" captionStartId="10.[213,280,1142,1164]" captionTargetBox="[215,1168,162,1108]" captionTargetId="figure-5@10.[211,1171,161,1110]" captionTargetPageId="10" captionText="Figure 3. Thoracostrongylus aduncatus sp. nov., holotype, male from Yunnan, China. (A) 8th sternite, male; (B) 10th tergite, male; (C) 9th sternite, male; (D) aedeagus, ventral view; (E) aedeagus, lateral view. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4666856" httpUri="https://zenodo.org/record/4666856/files/figure.png" pageId="9" pageNumber="414">Figure 3D, E</figureCitation>
) moderately sclerotized, median lobe elongate, in ventral view almost symmetrical, gradually constricted above phallobase, apex distinctly sharply pointed; paramere stout, gradually narrowed towards apex, apex moderately pointed, with eight fine apical setae; in lateral view, apical half bent dorsad, with a small acutely pointed process projecting dorsad at about one-fifth from apex.
</paragraph>
<paragraph id="8BE936DBFFED8632FF2DFAE3FF05D014" blockId="9.[144,242,1320,1344]" box="[144,242,1320,1344]" pageId="9" pageNumber="414">
<emphasis id="B922EAC9FFED8632FF2DFAE3FF05D014" box="[144,242,1320,1344]" italics="true" pageId="9" pageNumber="414">Remarks</emphasis>
</paragraph>
<paragraph id="8BE936DBFFED8632FF2DFA9BFF0ED0F3" blockId="9.[144,1101,1360,1447]" pageId="9" pageNumber="414">
In general appearance, the species is similar to
<taxonomicName id="4C564D58FFED8632FD18FA9BFC1BD03C" authority="Bernhauer, 1943" authorityName="Bernhauer" authorityYear="1943" box="[677,1004,1360,1384]" class="Insecta" family="Staphylinidae" genus="Thoracostrongylus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="414" phylum="Arthropoda" rank="species" species="miyakei">
<emphasis id="B922EAC9FFED8632FD18FA9BFCDED03C" box="[677,809,1360,1384]" italics="true" pageId="9" pageNumber="414">Th. miyakei</emphasis>
<bibRefCitation id="EFC74B2AFFED8632FC8CFA9BFC1BD03C" author="Bernhauer M" box="[817,1004,1360,1384]" pageId="9" pageNumber="414" pagination="169 - 188" refId="ref11732" refString="Bernhauer M. 1943. Neuheiten der palaearktischen Staphylinidenfauna. Mitt munch entomol Ges. 33: 169 - 188." type="journal article" year="1943">Bernhauer, 1943</bibRefCitation>
</taxonomicName>
, but differs from the latter by the colour of abdomen lateral plate and the shape of the male genitalia.
</paragraph>
<paragraph id="8BE936DBFFED8632FF2DFA06FEE2D0B1" blockId="9.[144,277,1485,1509]" box="[144,277,1485,1509]" pageId="9" pageNumber="414">
<emphasis id="B922EAC9FFED8632FF2DFA06FEE2D0B1" box="[144,277,1485,1509]" italics="true" pageId="9" pageNumber="414">Distribution</emphasis>
</paragraph>
<paragraph id="8BE936DBFFED8632FF2DFA3EFC00D359" blockId="9.[144,1015,1525,1550]" box="[144,1015,1525,1550]" pageId="9" pageNumber="414">
<materialsCitation id="3B3E3C86FFED8632FF2DFA3EFC03D359" box="[144,1012,1525,1550]" country="China" location="Yunnan" pageId="9" pageNumber="414" specimenCount="1" stateProvince="Yunnan">
The species is at present known only from the
<typeStatus id="54ED8879FFED8632FD2BFA3DFD31D35A" box="[662,710,1526,1550]" pageId="9" pageNumber="414">type</typeStatus>
locality in
<collectingRegion id="4992F839FFED8632FCFEFA3DFC57D359" box="[835,928,1526,1549]" country="China" name="Yunnan" pageId="9" pageNumber="414">Yunnan</collectingRegion>
,
<collectingCountry id="F341764BFFED8632FC16FA3EFC03D359" box="[939,1012,1525,1549]" name="China" pageId="9" pageNumber="414">China</collectingCountry>
</materialsCitation>
.
</paragraph>
<paragraph id="8BE936DBFFED8632FF2DF998FEFFD33F" blockId="9.[144,264,1619,1643]" box="[144,264,1619,1643]" pageId="9" pageNumber="414">
<emphasis id="B922EAC9FFED8632FF2DF998FEFFD33F" box="[144,264,1619,1643]" italics="true" pageId="9" pageNumber="414">Etymology</emphasis>
</paragraph>
<paragraph id="8BE936DBFFED8632FF2DF9B1FD44D3C6" blockId="9.[144,691,1658,1682]" box="[144,691,1658,1682]" pageId="9" pageNumber="414">The species name (Latin) means “hook-shaped”.</paragraph>
</subSubSection>
</treatment>
</document>
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