195 lines
23 KiB
XML
195 lines
23 KiB
XML
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<mods:title id="6082BEC7EDA7B50A8ACAB4887D06BED8">A new species of Tychus Leach, 1817 from Lebanon (Coleoptera, Staphylinidae, Pselaphinae)</mods:title>
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<mods:namePart id="34AE897F49F527C4FF0C93F226BFA757">Sabella, Giorgio</mods:namePart>
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<mods:namePart id="9AB7FCB1922FB102494A3028B8D21917">Viglianisi, Fabio Massimo</mods:namePart>
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<mods:namePart id="BA35D23C428AA746046EECC393560BAF">D’Urso, Vera</mods:namePart>
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<mods:title id="FE8B5EEEEFE0E27CCB90F6ED690F30B6">Zootaxa</mods:title>
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<mods:date id="520FDC8C4EF48FB935FEF6B7257038B7">2019</mods:date>
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<emphasis id="B915EAE4FF8BBD60FF1BFFB58FF481E1" bold="true" box="[151,464,153,180]" pageId="1" pageNumber="597">
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<taxonomicName id="4C614D75FF8BBD60FF1BFFB58FA281E1" ID-CoL="8YC53" authority="Sabella & Viglianisi & D’Urso, 2019" authorityName="Sabella & Viglianisi & D’Urso" authorityYear="2019" box="[151,390,153,180]" class="Insecta" family="Staphylinidae" genus="Tychus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="597" phylum="Arthropoda" rank="species" species="meggiolaroi" status="sp. nov.">
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<emphasis id="B915EAE4FF8BBD60FF1BFFB58FA281E1" bold="true" box="[151,390,153,180]" italics="true" pageId="1" pageNumber="597">Tychus meggiolaroi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A226579FFF8BBD60FE02FFB68FF481E1" box="[398,464,154,180]" pageId="1" pageNumber="597" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<subSubSection id="C37B657DFF8BBD60FF1BFF918F2F818D" box="[151,267,189,216]" pageId="1" pageNumber="597" type="description">
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<paragraph id="8BDE36F6FF8BBD60FF1BFF918F2F818D" blockId="1.[151,464,153,216]" box="[151,267,189,216]" pageId="1" pageNumber="597">
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(
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<figureCitation id="135A2A73FF8BBD60FF13FF918F26818D" box="[159,258,189,216]" captionStart-0="FIGURES 1−5" captionStart-1="FIGURE 6" captionStartId-0="1.[151,264,1862,1887]" captionStartId-1="3.[151,250,1673,1698]" captionTargetBox-0="[152,1436,376,1835]" captionTargetBox-1="[151,1436,658,1605]" captionTargetId-0="figure@1.[151,1436,345,1842]" captionTargetId-1="figure@3.[151,1436,658,1605]" captionTargetPageId-0="1" captionTargetPageId-1="3" captionText-0="FIGURES 1−5. Tychus meggiolaroi sp. n., holotype male. 1. Head, dorsal view, schematic draw. 2. Right maxillary palpus. 3. Right antenna. 4. aedeagus, lateral view. 5. aedeagus, ventral view. Scale bar: 0.1 mm." captionText-1="FIGURE 6. Distribution map of the species of Tychus of the cilicicus group. ': T. decens; l: T. dubius;: uT. cilicicus; ▲: T. sodalicius; *: T. meggiolaroi sp. n.; ▲: T. galileus." httpUri-0="https://zenodo.org/record/3368963/files/figure.png" httpUri-1="https://zenodo.org/record/3368965/files/figure.png" pageId="1" pageNumber="597">Figs 1–6</figureCitation>
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)
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</paragraph>
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<subSubSection id="C37B657DFF8BBD63FF1BFE2A8C878592" lastPageId="2" lastPageNumber="598" pageId="1" pageNumber="597" type="materials_examined">
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<paragraph id="8BDE36F6FF8BBD60FF1BFE2A8F268016" blockId="1.[151,1437,261,323]" pageId="1" pageNumber="597">
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<emphasis id="B915EAE4FF8BBD60FF1BFE2A8F668075" bold="true" box="[151,322,261,288]" pageId="1" pageNumber="597">Type material</emphasis>
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.
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<materialsCitation id="3B093CABFF8BBD60FED8FE2A8F268016" ID-GBIF-Occurrence="2332606836" collectingDate="1895-04-18" collectionCode="MSNV" collectorName="W. Wittmer" country="Lebanon" elevation="1700" location="Bsharre" pageId="1" pageNumber="597" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
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</emphasis>
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:
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<emphasis id="B915EAE4FF8BBD60FE58FE298D2E8075" bold="true" box="[468,778,261,288]" pageId="1" pageNumber="597">
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NORTHERN
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<collectingCountry id="F3767666FF8BBD60FD0CFE298D2E8075" box="[640,778,261,288]" name="Lebanon" pageId="1" pageNumber="597">LEBANON</collectingCountry>
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</emphasis>
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:
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<specimenCount id="9D67FD7FFF8BBD60FC91FE2A8D118075" box="[797,821,262,288]" pageId="1" pageNumber="597" type="male">♂</specimenCount>
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:
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,
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,
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<collectingDate id="EF9BE9DEFF8BBD60FBD7FE298AF78075" box="[1115,1235,261,288]" pageId="1" pageNumber="597" value="1895-04-18">18.iv.1895</collectingDate>
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,
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<collectorName id="26945320FF8BBD60FB6EFE2A8B4C804A" box="[1250,1384,261,288]" pageId="1" pageNumber="597">W. Wittmer</collectorName>
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leg. (
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<collectionCode id="ED70AE33FF8BBD60FF13FE058ED28016" box="[159,246,297,323]" country="Italy" httpUri="http://grbio.org/cool/vmav-1g60" name="Museo Civico di Storia Naturale di Venezia" pageId="1" pageNumber="597">MSNV</collectionCode>
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).
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</materialsCitation>
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</paragraph>
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<caption id="DF1E667EFF8BBD60FF1BF86A8A2786D7" httpUri="https://zenodo.org/record/3368963/files/figure.png" pageId="1" pageNumber="597" startId="1.[151,264,1862,1887]" targetBox="[152,1436,376,1835]" targetPageId="1">
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<emphasis id="B915EAE4FF8BBD60FF1BF86A8F18860A" bold="true" box="[151,316,1862,1887]" pageId="1" pageNumber="597">FIGURES 1−5.</emphasis>
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<taxonomicName id="4C614D75FF8BBD60FECFF86B8C2E860A" authority="Sabella & Viglianisi & D’Urso, 2019" authorityName="Sabella & Viglianisi & D’Urso" authorityYear="2019" box="[323,522,1863,1887]" class="Insecta" family="Staphylinidae" genus="Tychus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="597" phylum="Arthropoda" rank="species" species="meggiolaroi" status="sp. nov.">
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<emphasis id="B915EAE4FF8BBD60FECFF86B8C2E860A" box="[323,522,1863,1887]" italics="true" pageId="1" pageNumber="597">Tychus meggiolaroi</emphasis>
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</taxonomicName>
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<emphasis id="B915EAE4FF8BBD60FD9DF86B8C6E860A" bold="true" box="[529,586,1863,1887]" pageId="1" pageNumber="597">
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<taxonomicNameLabel id="A226579FFF8BBD60FD9DF86B8C6E860A" box="[529,586,1863,1887]" pageId="1" pageNumber="597" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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, holotype male.
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<emphasis id="B915EAE4FF8BBD60FD78F86A8D25860A" bold="true" box="[756,769,1862,1887]" pageId="1" pageNumber="597">1</emphasis>
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. Head, dorsal view, schematic draw.
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<emphasis id="B915EAE4FF8BBD60FBF6F86A8AA3860A" bold="true" box="[1146,1159,1862,1887]" pageId="1" pageNumber="597">2</emphasis>
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. Right maxillary palpus.
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<emphasis id="B915EAE4FF8BBD60FA05F86A8BB2860A" bold="true" box="[1417,1430,1862,1887]" pageId="1" pageNumber="597">3</emphasis>
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. Right antenna.
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<emphasis id="B915EAE4FF8BBD60FEBDF8468F1A86D6" bold="true" box="[305,318,1898,1923]" pageId="1" pageNumber="597">4</emphasis>
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. aedeagus, lateral view.
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<emphasis id="B915EAE4FF8BBD60FDB4F8478C6186D6" bold="true" box="[568,581,1899,1923]" pageId="1" pageNumber="597">5</emphasis>
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. aedeagus, ventral view. Scale bar: 0.1 mm.
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</paragraph>
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</caption>
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<paragraph id="8BDE36F6FF8BBD63FF4BF89C8F838562" blockId="1.[151,1437,1968,2032]" lastBlockId="2.[151,1437,153,1223]" lastPageId="2" lastPageNumber="598" pageId="1" pageNumber="597">
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<emphasis id="B915EAE4FF8BBD60FF4BF89C8F75869E" bold="true" box="[199,337,1968,1995]" pageId="1" pageNumber="597">Description</emphasis>
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. Length
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, head, pronotum and abdomen dark brown, elytra red, antennae and legs yellowish and yellow palpi. Relatively dense pubescence of long and flattened golden setae on head, pronotum, elytra and abdomen; other setae shorter, yellowish and suberect on antennae, palpi and legs. Tuft of dense setae present behind temples. Tegument smooth and shiny; only elytra with some punctures. Head (
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<figureCitation id="135A2A73FF88BD63FBFEFF918A918182" box="[1138,1205,189,215]" captionStart="FIGURES 1−5" captionStartId="1.[151,264,1862,1887]" captionTargetBox="[152,1436,376,1835]" captionTargetId="figure@1.[151,1436,345,1842]" captionTargetPageId="1" captionText="FIGURES 1−5. Tychus meggiolaroi sp. n., holotype male. 1. Head, dorsal view, schematic draw. 2. Right maxillary palpus. 3. Right antenna. 4. aedeagus, lateral view. 5. aedeagus, ventral view. Scale bar: 0.1 mm." httpUri="https://zenodo.org/record/3368963/files/figure.png" pageId="2" pageNumber="598">Fig. 1</figureCitation>
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) widest across eyes and narrowest posteriorly to antennal tubercles, slightly wider (
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<quantity id="4C999B13FF88BD63FCD3FFCD8DE281AE" box="[863,966,225,251]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="2" pageNumber="598" unit="mm" value="0.3">0.30 mm</quantity>
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) than long (
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<quantity id="4C999B13FF88BD63FBC1FFCD8A9381AE" box="[1101,1207,225,251]" metricMagnitude="-4" metricUnit="m" metricValue="2.9" pageId="2" pageNumber="598" unit="mm" value="0.29">0.29 mm</quantity>
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). Antennal tubercles
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<quantity id="4C999B13FF88BD63FF1BFE288F2D804A" box="[151,265,260,287]" metricMagnitude="-4" metricUnit="m" metricValue="1.45" pageId="2" pageNumber="598" unit="mm" value="0.145">0.145 mm</quantity>
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wide and
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<quantity id="4C999B13FF88BD63FEF0FE298FC4804A" box="[380,480,261,287]" metricMagnitude="-5" metricUnit="m" metricValue="6.0" pageId="2" pageNumber="598" unit="mm" value="0.06">0.06 mm</quantity>
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long, separated by median longitudinal sulcus. Vertex separated from frontal rostrum by transverse depression. Pair of small dorsal vertexal foveae between and close to eyes, the latter prominent, each with 16-18 facets. Small tooth in front of each vertexal fovea. Tempora rounded. Last palpomere (
|
||
<figureCitation id="135A2A73FF88BD63FB69FE618B0D8032" box="[1253,1321,333,359]" captionStart="FIGURES 1−5" captionStartId="1.[151,264,1862,1887]" captionTargetBox="[152,1436,376,1835]" captionTargetId="figure@1.[151,1436,345,1842]" captionTargetPageId="1" captionText="FIGURES 1−5. Tychus meggiolaroi sp. n., holotype male. 1. Head, dorsal view, schematic draw. 2. Right maxillary palpus. 3. Right antenna. 4. aedeagus, lateral view. 5. aedeagus, ventral view. Scale bar: 0.1 mm." httpUri="https://zenodo.org/record/3368963/files/figure.png" pageId="2" pageNumber="598">Fig. 2</figureCitation>
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)
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<quantity id="4C999B13FF88BD63FAB4FE618BB98032" box="[1336,1437,333,359]" metricMagnitude="-4" metricUnit="m" metricValue="1.6" pageId="2" pageNumber="598" unit="mm" value="0.16">0.16 mm</quantity>
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long and
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<quantity id="4C999B13FF88BD63FE88FE5C8F5D80DE" box="[260,377,368,395]" metricMagnitude="-5" metricUnit="m" metricValue="8.5" pageId="2" pageNumber="598" unit="mm" value="0.085">0.085 mm</quantity>
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wide, with slightly rounded lateral margin. Antennae (
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<figureCitation id="135A2A73FF88BD63FC7FFE5D8A1C80DE" box="[1011,1080,369,395]" captionStart="FIGURES 1−5" captionStartId="1.[151,264,1862,1887]" captionTargetBox="[152,1436,376,1835]" captionTargetId="figure@1.[151,1436,345,1842]" captionTargetPageId="1" captionText="FIGURES 1−5. Tychus meggiolaroi sp. n., holotype male. 1. Head, dorsal view, schematic draw. 2. Right maxillary palpus. 3. Right antenna. 4. aedeagus, lateral view. 5. aedeagus, ventral view. Scale bar: 0.1 mm." httpUri="https://zenodo.org/record/3368963/files/figure.png" pageId="2" pageNumber="598">Fig. 3</figureCitation>
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)
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||
<quantity id="4C999B13FF88BD63FBC7FE5D8A9580DE" box="[1099,1201,369,395]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="2" pageNumber="598" unit="mm" value="0.9">0.90 mm</quantity>
|
||
long, club
|
||
<quantity id="4C999B13FF88BD63FABAFE5D8BB980DE" box="[1334,1437,369,395]" metricMagnitude="-4" metricUnit="m" metricValue="3.1" pageId="2" pageNumber="598" unit="mm" value="0.31">0.31 mm</quantity>
|
||
long; scape distinctly longer than wide; antennomeres 2 and 3 longer than wide; antennomere 3 narrow at base, 4 also longer than wide, 5 distinctly larger than other segments of funiculus, longer than wide, with its medial margin enlarged in the apical third, 6-7 wider than long, 7 longer than 6 and 8; 8 smallest and distinctly wider than long. Antennal club with three antennomeres broaden progressively from 9 to 11, antennomeres 9 and 10 transverse, the latter wider than 9, terminal antennomere longer than wide and longer than combined length of antennomeres 9 and 10. Pronotum wider than head, wider (
|
||
<quantity id="4C999B13FF88BD63FDF0FD658CC08336" box="[636,740,585,611]" metricMagnitude="-4" metricUnit="m" metricValue="3.5999999999999996" pageId="2" pageNumber="598" unit="mm" value="0.36">0.36 mm</quantity>
|
||
) than long (
|
||
<quantity id="4C999B13FF88BD63FCE2FD658DFC8336" box="[878,984,585,611]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="2" pageNumber="598" unit="mm" value="0.3">0.30 mm</quantity>
|
||
). It is more clearly tapered and rounded anteriorly than posteriorly with seven small basal pits, median pit larger than lateral ones. Its lateral antebasal foveae large and strongly impressed. Metaventrite with median impression from posterior margin and extending over about 1/2 of its length. Elytra wider (
|
||
<quantity id="4C999B13FF88BD63FE78FD998C7F839A" box="[500,603,693,719]" metricMagnitude="-4" metricUnit="m" metricValue="6.8999999999999995" pageId="2" pageNumber="598" unit="mm" value="0.69">0.69 mm</quantity>
|
||
) than long (
|
||
<quantity id="4C999B13FF88BD63FD68FD988D68839A" box="[740,844,692,719]" metricMagnitude="-4" metricUnit="m" metricValue="5.3" pageId="2" pageNumber="598" unit="mm" value="0.53">0.53 mm</quantity>
|
||
) and longer than pronotum, with protruding humeri. Two basal foveae on each elytron; sutural fovea associated with shallow sutural stria that extends to elytral apex, while discal fovea extended posteriorly in discal stria reaching about half of elytral length. Abdomen with 1
|
||
<superScript id="7C149BBEFF88BD63FAC8FDD08B6B825F" attach="left" box="[1348,1359,764,778]" fontSize="6" pageId="2" pageNumber="598">st</superScript>
|
||
tergite
|
||
<quantity id="4C999B13FF88BD63FF1BFC0D8EDA826E" box="[151,254,801,827]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="598" unit="mm" value="0.2">0.20 mm</quantity>
|
||
long, with short discal carinae detached by pubescent basal impression occupying about 1/2 of tergite width. First tergite with pair of basolateral foveae, 1
|
||
<superScript id="7C149BBEFF88BD63FD6DFC688CC88207" attach="left" box="[737,748,836,850]" fontSize="6" pageId="2" pageNumber="598">st</superScript>
|
||
abdominal paratergite with pair of antebasal impressions, 2
|
||
<superScript id="7C149BBEFF88BD63FA00FC688BB88207" attach="none" box="[1420,1436,836,850]" fontSize="6" pageId="2" pageNumber="598">nd</superScript>
|
||
abdominal tergite with pair of antebasal foveae. All abdominal ventrites not modified. Legs with posterior margin of mesotrochanters extend into short median spine, lateral margin of metatrochanters prolonged into small median spine, femora of all legs slightly thickened. Aedeagus (
|
||
<figureCitation id="135A2A73FF88BD63FC89FC9D8D4F829E" box="[773,875,944,971]" captionStart="FIGURES 1−5" captionStartId="1.[151,264,1862,1887]" captionTargetBox="[152,1436,376,1835]" captionTargetId="figure@1.[151,1436,345,1842]" captionTargetPageId="1" captionText="FIGURES 1−5. Tychus meggiolaroi sp. n., holotype male. 1. Head, dorsal view, schematic draw. 2. Right maxillary palpus. 3. Right antenna. 4. aedeagus, lateral view. 5. aedeagus, ventral view. Scale bar: 0.1 mm." httpUri="https://zenodo.org/record/3368963/files/figure.png" pageId="2" pageNumber="598">Figs 4–5</figureCitation>
|
||
)
|
||
<quantity id="4C999B13FF88BD63FCF6FC9C8DFB829E" box="[890,991,944,971]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="2" pageNumber="598" unit="mm" value="0.25">0.25 mm</quantity>
|
||
long. Dorsal apophysis of median lobe sinuate and canaliculated, and enlarged at apex, bearing in basal third short spine directed medially. Ventral portion of median lobe shorter than dorsal apophysis, larger at base and gradually narrowed towards apex, the latter directed laterally and downward.
|
||
</paragraph>
|
||
<paragraph id="8BDE36F6FF88BD63FF4BFB6D8FBA850E" blockId="2.[151,1437,153,1223]" box="[199,414,1089,1115]" pageId="2" pageNumber="598">Female. Unknown.</paragraph>
|
||
<paragraph id="8BDE36F6FF88BD63FF4BFB488C19852A" blockId="2.[151,1437,153,1223]" box="[199,573,1124,1151]" pageId="2" pageNumber="598">
|
||
<emphasis id="B915EAE4FF88BD63FF4BFB488F45852A" bold="true" box="[199,353,1124,1151]" pageId="2" pageNumber="598">Distribution.</emphasis>
|
||
Northern
|
||
<collectingCountry id="F3767666FF88BD63FE58FB498C1E852A" box="[468,570,1125,1151]" name="Lebanon" pageId="2" pageNumber="598">Lebanon</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BDE36F6FF88BD63FF4BFBA58C878592" blockId="2.[151,1437,153,1223]" pageId="2" pageNumber="598">
|
||
<emphasis id="B915EAE4FF88BD63FF4BFBA58F6A85F6" bold="true" box="[199,334,1161,1187]" pageId="2" pageNumber="598">Etymology.</emphasis>
|
||
<taxonomicName id="4C614D75FF88BD63FEDAFBA58C1785F6" authority="Sabella & Viglianisi & D’Urso, 2019" authorityName="Sabella & Viglianisi & D’Urso" authorityYear="2019" box="[342,563,1161,1187]" class="Insecta" family="Staphylinidae" genus="Tychus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="598" phylum="Arthropoda" rank="species" species="meggiolaroi" status="sp. nov.">
|
||
<emphasis id="B915EAE4FF88BD63FEDAFBA58C1785F6" box="[342,563,1161,1187]" italics="true" pageId="2" pageNumber="598">Tychus meggiolaroi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B915EAE4FF88BD63FDB7FBA58C5F85F6" bold="true" box="[571,635,1161,1187]" pageId="2" pageNumber="598">
|
||
<taxonomicNameLabel id="A226579FFF88BD63FDB7FBA58C5F85F6" box="[571,635,1161,1187]" pageId="2" pageNumber="598" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is dedicated to the memory of the entomologist Giuseppe Meggiolaro, who was the first person to study this material.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |