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<document id="3E2F31F5A08C7C17F142C07B496440B4" ID-CLB-Dataset="32718" ID-DOI="10.11646/zootaxa.4290.2.1" ID-GBIF-Dataset="43ebda6c-201a-46af-8b54-e7980b8d79e1" ID-ISSN="1175-5326" ID-Zenodo-Dep="828965" ID-ZooBank="52D6E3C3-BA0A-49C7-865C-00F3F5E5C5D4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1500110315309" checkinUser="plazi" docAuthor="Gunter, Nicole L. &amp; Weir, Thomas A." docDate="2017" docId="03C887A8FFDFFFD3FF737FAD5C8EB46A" docLanguage="en" docName="zootaxa.4290.2.1.pdf" docOrigin="Zootaxa 4290 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Matthewsius speewah Storey &amp; Monteith 2000, new combination" docType="treatment" docVersion="7" lastPageNumber="221" masterDocId="FFF1FFD0FFCBFFC7FFE47F3A5F6FB348" masterDocTitle="Two new genera of Australian dung beetles (Coleoptera: Scarabaeidae: Scarabaeinae) with the description of six new species and transfer of six described species" masterLastPageNumber="243" masterPageNumber="201" pageNumber="221" updateTime="1698465402479" updateUser="ExternalLinkService">
<mods:mods id="AEFB46253FBE6C72EFFBB1A6D6144568" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="AD7D2DD16A0CBC2FF0CACEC6811B5490">Two new genera of Australian dung beetles (Coleoptera: Scarabaeidae: Scarabaeinae) with the description of six new species and transfer of six described species</mods:title>
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<mods:namePart id="031BEAB557E088B5D4F3DC4856A80791">Gunter, Nicole L.</mods:namePart>
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<mods:role id="7AD5CB3FF5A90CBD0E2888D6B2BC819F">
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<mods:namePart id="40BE13F228C7182D57A7691F642A492B">Weir, Thomas A.</mods:namePart>
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<mods:typeOfResource id="CC40F67CF8C27542529A2BAF54F6F030">text</mods:typeOfResource>
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<mods:title id="76D8967A85BA878DEF22E8179FCD7585">Zootaxa</mods:title>
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<mods:part id="990487CC4C495B75EB351F0ED0D9ACA9">
<mods:date id="37D27016A36920A9C98E382F50C3716A">2017</mods:date>
<mods:detail id="A82907F62B05FFB361B2A1C7309EC877" type="volume">
<mods:number id="A2167061C7739F664042CCFDFEC4D916">4290</mods:number>
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<mods:detail id="54731CA3A2D6EB8005E182EB426DDB19" type="issue">
<mods:number id="EB6FB4627518EF0D038FB787FD535522">2</mods:number>
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<mods:start id="D410F0A09B72E1C2E059BF0828658223">201</mods:start>
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<mods:classification id="1D6ADC9695DDCE58E6946D568B1924F9">journal article</mods:classification>
<mods:identifier id="B6CA89652090E245E22A2902BA81EF5B" type="CLB-Dataset">32718</mods:identifier>
<mods:identifier id="9CFFA3E151FD8CFB6802484462515736" type="DOI">10.11646/zootaxa.4290.2.1</mods:identifier>
<mods:identifier id="A77FE1599A901E49D0A9CB7B6ED4B569" type="GBIF-Dataset">43ebda6c-201a-46af-8b54-e7980b8d79e1</mods:identifier>
<mods:identifier id="F44032134C702FAC683A5CEC3C36D898" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="584C209A0CF61223067E62B7C08D02A9" type="Zenodo-Dep">828965</mods:identifier>
<mods:identifier id="366B6E4B5520139456F081FB3593F297" type="ZooBank">52D6E3C3-BA0A-49C7-865C-00F3F5E5C5D4</mods:identifier>
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<treatment id="03C887A8FFDFFFD3FF737FAD5C8EB46A" ID-DOI="http://doi.org/10.5281/zenodo.6033004" ID-GBIF-Taxon="132549213" ID-Zenodo-Dep="6033004" LSID="urn:lsid:plazi:treatment:03C887A8FFDFFFD3FF737FAD5C8EB46A" httpUri="http://treatment.plazi.org/id/03C887A8FFDFFFD3FF737FAD5C8EB46A" lastPageNumber="221" pageId="20" pageNumber="221">
<subSubSection id="C37B6535FFDFFFD3FF737FAD5E20B39B" pageId="20" pageNumber="221" type="nomenclature">
<paragraph id="8BDE36BEFFDFFFD3FF737FAD5CA5B3FA" blockId="20.[151,970,151,211]" box="[151,970,151,178]" pageId="20" pageNumber="221">
<heading id="D09681D2FFDFFFD3FF737FAD5CA5B3FA" bold="true" box="[151,970,151,178]" fontSize="11" level="1" pageId="20" pageNumber="221" reason="1">
<emphasis id="B915EAACFFDFFFD3FF737FAD5CA5B3FA" bold="true" box="[151,970,151,178]" pageId="20" pageNumber="221">
<taxonomicName id="4C614D3DFFDFFFD3FF737FAD5D83B3FA" ID-CoL="3YF86" authority="Storey &amp; Monteith, 2000" authorityName="Storey &amp; Monteith" authorityYear="2000" box="[151,748,151,178]" class="Insecta" family="Scarabaeidae" genus="Matthewsius" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="221" phylum="Arthropoda" rank="species" species="speewah" status="comb. nov.">
<emphasis id="B915EAACFFDFFFD3FF737FAD5EF1B3F9" bold="true" box="[151,414,151,177]" italics="true" pageId="20" pageNumber="221">Matthewsius speewah</emphasis>
(
<bibRefCitation id="EFF04B4FFFDFFFD3FE497FA25D8CB3FA" author="Storey" box="[429,739,152,178]" pageId="20" pageNumber="221" refString="Storey, R. I. &amp; Monteith, G. B. (2000) Five new species of Aptenocanthon Matthews (Coleoptera: Scarbaeidae: Scarabaeinae) from tropical Australia, with notes on distribution. Memoirs of the Queensland Museum, 46, 349 - 358." type="journal article" year="2000">Storey &amp; Monteith, 2000</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A22657D7FFDFFFD3FD177FA25CA5B3FA" box="[755,970,152,178]" pageId="20" pageNumber="221">new combination</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BDE36BEFFDFFFD3FF737F805E20B39B" blockId="20.[151,970,151,211]" box="[151,335,186,211]" pageId="20" pageNumber="221">
<figureCitation id="135A2A3BFFDFFFD3FF737F805FB4B39B" box="[151,219,186,211]" captionStart="FIGURE 3" captionStartId="11.[151,250,1894,1916]" captionTargetBox="[164,1410,211,1857]" captionTargetId="figure@11.[163,1424,193,1872]" captionTargetPageId="11" captionText="FIGURE 3. Pygidium and aedeagus (left view, dorsal view, right view) of species of Matthewsius new genus, all scale bars 0.5 mm. A, Matthewsius illawarrensis new combination; B, M. penelopae new combination; C, M. rossi new combination; D, M. speewah new combination; E, M. stricklandensis new species; F, M. vanleeuweni new species." httpUri="https://zenodo.org/record/828971/files/figure.png" pageId="20" pageNumber="221">Figs 3</figureCitation>
D, 6D, 5B
</paragraph>
</subSubSection>
<subSubSection id="C37B6535FFDFFFD3FF737E385DAAB250" box="[151,709,258,280]" pageId="20" pageNumber="221" type="reference_group">
<paragraph id="8BDE36BEFFDFFFD3FF737E385DAAB250" blockId="20.[151,709,258,280]" box="[151,709,258,280]" pageId="20" pageNumber="221">
<treatmentCitationGroup id="AB711190FFDFFFD3FF737E385DAAB250" box="[151,709,258,280]" pageId="20" pageNumber="221">
<taxonomicName id="4C614D3DFFDFFFD3FF737E385DAAB250" ID-CoL="FYD5" authority="Storey &amp; Monteith, 2000: 353" authorityName="Storey &amp; Monteith" authorityPageNumber="353" authorityYear="2000" box="[151,709,258,280]" class="Insecta" family="Scarabaeidae" genus="Aptenocanthon" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="221" phylum="Arthropoda" rank="species" species="speewah">
<emphasis id="B915EAACFFDFFFD3FF737E385EE2B250" box="[151,397,258,280]" italics="true" pageId="20" pageNumber="221">Aptenocanthon speewah</emphasis>
<treatmentCitation id="0AC010AFFFDFFFD3FE707E385DAAB250" author="Storey" box="[404,709,258,280]" page="353" pageId="20" pageNumber="221" year="2000">
<bibRefCitation id="EFF04B4FFFDFFFD3FE707E385DF9B250" author="Storey" box="[404,662,258,280]" pageId="20" pageNumber="221" refString="Storey, R. I. &amp; Monteith, G. B. (2000) Five new species of Aptenocanthon Matthews (Coleoptera: Scarbaeidae: Scarabaeinae) from tropical Australia, with notes on distribution. Memoirs of the Queensland Museum, 46, 349 - 358." type="journal article" year="2000">Storey &amp; Monteith, 2000</bibRefCitation>
: 353
</treatmentCitation>
</taxonomicName>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C37B6535FFDFFFD3FF737E7F5E59B2A6" pageId="20" pageNumber="221" type="materials_examined">
<paragraph id="8BDE36BEFFDFFFD3FF737E7F5E59B2A6" blockId="20.[151,1437,325,1826]" pageId="20" pageNumber="221">
<materialsCitation id="3B093CE3FFDFFFD3FF737E7F5D20B2CA" ID-GBIF-Occurrence="1572325640" collectingDate="1993-12-02" collectingDateMax="1993-12-03" collectingDateMin="1993-12-02" collectionCode="ANIC" county="Janetzki" latitude="-16.916666" location="Material" longLatPrecision="21" longitude="145.66667" municipality="Monteith" pageId="20" pageNumber="221" specimenCode="ANIC 25-053840" specimenCount="1" typeStatus="paratype">
<emphasis id="B915EAACFFDFFFD3FF737E7F5EEBB216" bold="true" box="[151,388,325,350]" pageId="20" pageNumber="221">
<location id="8EBE6065FFDFFFD3FF737E7F5F90B216" LSID="urn:lsid:plazi:treatment:03C887A8FFDFFFD3FF737FAD5C8EB46A:8EBE6065FFDFFFD3FF737E7F5F90B216" box="[151,255,325,350]" county="Janetzki" latitude="-16.916666" longLatPrecision="21" longitude="145.66667" municipality="Monteith" name="Material" pageId="20" pageNumber="221">Material</location>
examined.
</emphasis>
<typeStatus id="54DA881CFFDFFFD3FE6A7E7C5D6FB216" box="[398,512,326,350]" pageId="20" pageNumber="221" type="paratype">Paratypes</typeStatus>
: [
<geoCoordinate id="EE555079FFDFFFD3FDF17E7F5DF7B216" box="[533,664,325,350]" direction="south" orientation="latitude" pageId="20" pageNumber="221" precision="15" value="-16.916666">16°55'00&quot;S</geoCoordinate>
,
<geoCoordinate id="EE555079FFDFFFD3FD4C7E7F5C54B216" box="[680,827,325,350]" direction="east" orientation="longitude" pageId="20" pageNumber="221" precision="15" value="145.66667">145°40'00&quot;E</geoCoordinate>
],
<location id="8EBE6065FFDFFFD3FCB57E7C5B60B216" LSID="urn:lsid:plazi:treatment:03C887A8FFDFFFD3FF737FAD5C8EB46A:8EBE6065FFDFFFD3FCB57E7C5B60B216" box="[849,1039,325,350]" county="Janetzki" latitude="-16.916666" longLatPrecision="21" longitude="145.66667" municipality="Monteith" name="Mount Williams" pageId="20" pageNumber="221">Mount Williams</location>
,
<location id="8EBE6065FFDFFFD3FBFA7E7F5B32B216" LSID="urn:lsid:plazi:treatment:03C887A8FFDFFFD3FF737FAD5C8EB46A:8EBE6065FFDFFFD3FBFA7E7F5B32B216" box="[1054,1117,325,350]" county="Janetzki" latitude="-16.916666" longLatPrecision="21" longitude="145.66667" municipality="Monteith" name="Cook" pageId="20" pageNumber="221">Cook</location>
,
<collectingMunicipality id="6BBAACC4FFDFFFD3FB887E7F5BB9B216" box="[1132,1238,325,350]" pageId="20" pageNumber="221">Monteith</collectingMunicipality>
, &amp;
<collectingCounty id="62BF4E32FFDFFFD3FAE17E7F5A0BB216" box="[1285,1380,325,350]" pageId="20" pageNumber="221">Janetzki</collectingCounty>
,
<collectingDate id="EF9BE996FFDFFFD3FA967E7C5E24B2C9" pageId="20" pageNumber="221" value="1993-12-02" valueMax="1993-12-03" valueMin="1993-12-02">
<date id="FFDF107EFFDFFFD3FA967E7C5E24B2C9" pageId="20" pageNumber="221" value="1993-12-02" valueMax="1993-12-03" valueMin="1993-12-02">2–3 December 1993</date>
</collectingDate>
(in
<specimenCode id="DBC79EC5FFDFFFD3FE9F7E535D28B2CA" box="[379,583,361,386]" collectionCode="ANIC" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="20" pageNumber="221">
<collectionCode id="ED70AE7BFFDFFFD3FE9F7E535ED0B2C9" box="[379,447,361,385]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="20" pageNumber="221">ANIC</collectionCode>
25-053840
</specimenCode>
)
</materialsCitation>
.
<materialsCitation id="3B093CE3FFDFFFD3FDBF7E535A6FB2EE" ID-GBIF-Occurrence="1572325672" collectingDate="2014-11-05" collectingDateMax="2014-11-06" collectingDateMin="2014-11-05" collectionCode="ANIC" country="Aland Islands" county="Monteith" elevation="950" latitude="-16.911943" location="Other" longLatPrecision="21" longitude="145.66306" municipality="Mount Williams" pageId="20" pageNumber="221" specimenCode="ANIC 25-065500-2" specimenCount="1" typeStatus="paratype">
<location id="8EBE6065FFDFFFD3FDBF7E535DF4B2CA" LSID="urn:lsid:plazi:treatment:03C887A8FFDFFFD3FF737FAD5C8EB46A:8EBE6065FFDFFFD3FDBF7E535DF4B2CA" box="[603,667,361,386]" country="Aland Islands" county="Monteith" latitude="-16.911943" longLatPrecision="21" longitude="145.66306" municipality="Mount Williams" name="Other" pageId="20" pageNumber="221">Other</location>
material:
<geoCoordinate id="EE555079FFDFFFD3FCF47E535CFEB2CA" box="[784,913,361,386]" direction="south" orientation="latitude" pageId="20" pageNumber="221" precision="15" value="-16.911943">16°54'43&quot;S</geoCoordinate>
,
<geoCoordinate id="EE555079FFDFFFD3FC7B7E535B5FB2CA" box="[927,1072,361,386]" direction="east" orientation="longitude" pageId="20" pageNumber="221" precision="15" value="145.66306">145°39'47&quot;E</geoCoordinate>
,
<collectingMunicipality id="6BBAACC4FFDFFFD3FBDA7E505B96B2CA" box="[1086,1273,361,386]" pageId="20" pageNumber="221">Mount Williams</collectingMunicipality>
,
<elevation id="004CD18DFFDFFFD3FAE37E535AF8B2C9" box="[1287,1431,361,385]" metricMagnitude="2" metricUnit="m" metricValue="9.5" metricValueMax="10.0" metricValueMin="9.0" pageId="20" pageNumber="221" unit="m" value="950.0" valueMax="1000.0" valueMin="900.0">
<quantity id="4C999B5BFFDFFFD3FAE37E535AF8B2C9" box="[1287,1431,361,385]" metricMagnitude="2" metricUnit="m" metricValue="9.5" metricValueMax="10.0" metricValueMin="9.0" pageId="20" pageNumber="221" unit="m" value="950.0" valueMax="1000.0" valueMin="900.0">900–1000 m</quantity>
</elevation>
,
<collectingCounty id="62BF4E32FFDFFFD3FF737EB75F91B2EE" box="[151,254,397,422]" pageId="20" pageNumber="221">Monteith</collectingCounty>
&amp;
<collectingCountry id="F376762EFFDFFFD3FED27EB75E12B2EE" box="[310,381,397,422]" name="Aland Islands" pageId="20" pageNumber="221">Aland</collectingCountry>
, baited pitfalls rainforest,
<collectingDate id="EF9BE996FFDFFFD3FD2F7EB75CA8B2EE" box="[715,967,397,422]" pageId="20" pageNumber="221" value="2014-11-05" valueMax="2014-11-06" valueMin="2014-11-05">
<date id="FFDF107EFFDFFFD3FD2F7EB75CA8B2EE" box="[715,967,397,422]" pageId="20" pageNumber="221" value="2014-11-05" valueMax="2014-11-06" valueMin="2014-11-05">5–6 November 2014</date>
</collectingDate>
(in
<specimenCode id="DBC79EC5FFDFFFD3FBEC7EB45B96B2EE" box="[1032,1273,397,422]" pageId="20" pageNumber="221">
<collectionCode id="ED70AE7BFFDFFFD3FBEC7EB45B22B2EE" box="[1032,1101,398,422]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="20" pageNumber="221">ANIC</collectionCode>
25-065500–2
</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="3B093CE3FFDFFFD3FAF27EB75E59B2A6" ID-GBIF-Occurrence="1572325655" collectingDate="2014-11-05" collectingDateMax="2014-11-06" collectingDateMin="2014-11-05" collectionCode="ANIC" country="Aland Islands" elevation="950" latitude="-16.911943" location="Mount Williams" longLatPrecision="21" longitude="145.66306" municipality="Monteith" pageId="20" pageNumber="221" specimenCode="ANIC 25-065503-4" specimenCount="1" typeStatus="paratype">
<geoCoordinate id="EE555079FFDFFFD3FAF27EB75AF8B2EE" box="[1302,1431,397,422]" direction="south" orientation="latitude" pageId="20" pageNumber="221" precision="15" value="-16.911943">16°54'43&quot;S</geoCoordinate>
,
<geoCoordinate id="EE555079FFDFFFD3FF737E8B5E47B282" box="[151,296,433,458]" direction="east" orientation="longitude" pageId="20" pageNumber="221" precision="15" value="145.66306">145°39'47&quot;E</geoCoordinate>
,
<location id="8EBE6065FFDFFFD3FED17E885E9FB282" LSID="urn:lsid:plazi:treatment:03C887A8FFDFFFD3FF737FAD5C8EB46A:8EBE6065FFDFFFD3FED17E885E9FB282" box="[309,496,433,458]" country="Aland Islands" latitude="-16.911943" longLatPrecision="21" longitude="145.66306" municipality="Monteith" name="Mount Williams" pageId="20" pageNumber="221">Mount Williams</location>
,
<elevation id="004CD18DFFDFFFD3FE197E8B5DE2B281" box="[509,653,433,457]" metricMagnitude="2" metricUnit="m" metricValue="9.5" metricValueMax="10.0" metricValueMin="9.0" pageId="20" pageNumber="221" unit="m" value="950.0" valueMax="1000.0" valueMin="900.0">
<quantity id="4C999B5BFFDFFFD3FE197E8B5DE2B281" box="[509,653,433,457]" metricMagnitude="2" metricUnit="m" metricValue="9.5" metricValueMax="10.0" metricValueMin="9.0" pageId="20" pageNumber="221" unit="m" value="950.0" valueMax="1000.0" valueMin="900.0">900–1000 m</quantity>
</elevation>
”
<collectingMunicipality id="6BBAACC4FFDFFFD3FD447E8B5C69B282" box="[672,774,433,458]" pageId="20" pageNumber="221">Monteith</collectingMunicipality>
&amp;
<collectingCountry id="F376762EFFDFFFD3FCC97E8B5C1BB282" box="[813,884,433,458]" name="Aland Islands" pageId="20" pageNumber="221">Aland</collectingCountry>
, litter, rainforest,
<collectingDate id="EF9BE996FFDFFFD3FBDB7E8B5A47B281" box="[1087,1320,433,458]" pageId="20" pageNumber="221" value="2014-11-05" valueMax="2014-11-06" valueMin="2014-11-05">
<date id="FFDF107EFFDFFFD3FBDB7E8B5A47B281" box="[1087,1320,433,458]" pageId="20" pageNumber="221" value="2014-11-05" valueMax="2014-11-06" valueMin="2014-11-05">5–6 November 2014</date>
</collectingDate>
(in
<specimenCode id="DBC79EC5FFDFFFD3FABC7E8B5E41B2A6" pageId="20" pageNumber="221">
<collectionCode id="ED70AE7BFFDFFFD3FABC7E8B5AF2B281" box="[1368,1437,433,457]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="20" pageNumber="221">ANIC</collectionCode>
25-065503–4
</specimenCode>
)
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C37B6535FFDFFFD3FF237EC35C0EB591" pageId="20" pageNumber="221" type="description">
<paragraph id="8BDE36BEFFDFFFD3FF237EC35DECB159" blockId="20.[151,1437,325,1826]" box="[199,643,505,530]" pageId="20" pageNumber="221">
<emphasis id="B915EAACFFDFFFD3FF237EC35E36B15A" bold="true" box="[199,345,505,530]" pageId="20" pageNumber="221">Description.</emphasis>
Total Length:
<quantity id="4C999B5BFFDFFFD3FE197EC35DECB159" box="[509,643,505,529]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" metricValueMax="4.2" metricValueMin="3.4" pageId="20" pageNumber="221" unit="mm" value="3.8" valueMax="4.2" valueMin="3.4">3.4–4.2 mm</quantity>
</paragraph>
<paragraph id="8BDE36BEFFDFFFD3FF237D275D58B17E" blockId="20.[151,1437,325,1826]" box="[199,567,541,566]" pageId="20" pageNumber="221">Colour: Black. Antennae yellow.</paragraph>
<paragraph id="8BDE36BEFFDFFFD3FF237D7B5A5BB1A2" blockId="20.[151,1437,325,1826]" pageId="20" pageNumber="221">Male: Head: Broad U-shape between clypeal teeth, which are slightly upturned. Rest of margin of head feebly convex to eye. Mentum almost square, bordered anteriorly, reticulate with setation on posterior corners. Edge of head bordered, wider between eye canthus and suture; genal angle slightly prominent. Dorsal parts of eyes notched; eyes narrow, separated by an interocular space of about 14 times eye width; eye canthus not dividing eye. Surface smooth, nitid, glabrous, with round punctures separated by 2 diameters. Head width to length ratio 40:62.</paragraph>
<paragraph id="8BDE36BEFFDFFFD3FF237DCC5FB1B0D6" blockId="20.[151,1437,325,1826]" pageId="20" pageNumber="221">Pronotum: Anterior angles almost 90°, with a distinct border. Basal angles rounded and obtuse, basal margin rounded, lateral edge finely margined. Hypomeron surface finely reticulate, glabrous with a few punctures. Punctures along basal margin extend to basal edge, with punctures different from disc. All round punctures without setae, separated by 2 diameters, less dense medially. Hypomeral stria present, long. Pronotum width to length ratio 52:27.</paragraph>
<paragraph id="8BDE36BEFFDFFFD3FF237C935CC1B766" blockId="20.[151,1437,325,1826]" pageId="20" pageNumber="221">
<emphasis id="B915EAACFFDFFFD3FF237C935E79B08A" box="[199,278,937,962]" italics="true" pageId="20" pageNumber="221">Elytra:</emphasis>
Surface with extremely fine setae, finely reticulate; sculpturing more evident apically. Intervals flat, finely punctate; punctures masked by fine meshes. Elytral striae narrow, shallowly impressed, with simple punctures. All striae on disc equal width, stria 7 only present in basal 1/3rd; striae 8 confined to groove posteriorly. Ratio of length of elytra along suture to maximum elytral width 48:58.
</paragraph>
<paragraph id="8BDE36BEFFDFFFD3FF237B005CA4B606" blockId="20.[151,1437,325,1826]" pageId="20" pageNumber="221">Legs: Prothoracic legs with trochantofemoral anterior pit and trochanter excavated dorsally for reception of inner apical edge of protibia. Protibiae with 3 teeth on outer edge, which is only serrate between 2nd and 3rd teeth and proximal to them, each tooth with 2 groups of setae dorsally, front edge straight between apical tooth and short, broad apical spur. The tooth on underside near base of tarsus is a sharp projection. Protibiae inner apical angle produced, with an apical comb of short, flattened bristles and on lower surface a brush of long setae extending beyond edge of comb. Claws small and simple. Mesotibiae broad and densely setose on the inner side. Metatibiae are slightly bent with the inner edge of tibia not crenulate. Males with small blunt tooth at inner apical edge of metatibia. Basal tarsomere wider than the other tarsomeres but not lobed.</paragraph>
<paragraph id="8BDE36BEFFDFFFD3FF237A635F85B696" blockId="20.[151,1437,325,1826]" pageId="20" pageNumber="221">Abdomen: Pygidium with a deep, transverse groove at the base reaching basal angle; narrowed medially and terminating in a pit at each end. Narrow, impressed line around the pygidium. Ventrites finely reticulate, with a row of punctures along the anterior edge of ventrites 2–5, ventrite 6 punctate. Suture between abdominal ventrites simple.</paragraph>
<paragraph id="8BDE36BEFFDFFFD3FF237AD35B30B525" blockId="20.[151,1437,325,1826]" pageId="20" pageNumber="221">Pterothorax: Medial lobe of metaventrite nitid, finely punctate, broadly margined between mesocoxae. Mesometaventral suture slightly arched. Lateral lobe of the metaventrite with impressed, close-spaced, horseshoeshaped punctures. Mesoventrite nitid, lateral edges with ocellate punctures and medially finely punctate; not notched anteriorly. Mesepimera punctate. Metanepisterna with rudimentary punctures.</paragraph>
<paragraph id="8BDE36BEFFDFFFD3FF2379435EC0B5DA" blockId="20.[151,1437,325,1826]" box="[199,431,1657,1682]" pageId="20" pageNumber="221">Hind wings: Absent.</paragraph>
<paragraph id="8BDE36BEFFDFFFD3FF2379A75C0EB591" blockId="20.[151,1437,325,1826]" pageId="20" pageNumber="221">Female: Protibiae not expanded at the inner apical angle with longer apical spur. Setae on tarsomeres shorter and less dense than males. Mesotibiae lack inner brush of setae.</paragraph>
</subSubSection>
<subSubSection id="C37B6535FFDFFFD3FF2379DF5C8EB46A" pageId="20" pageNumber="221" type="distribution">
<paragraph id="8BDE36BEFFDFFFD3FF2379DF5C8EB46A" blockId="20.[151,1437,325,1826]" pageId="20" pageNumber="221">
<emphasis id="B915EAACFFDFFFD3FF2379DF5E0FB5B6" bold="true" box="[199,352,1765,1790]" pageId="20" pageNumber="221">Distribution.</emphasis>
<figureCitation id="135A2A3BFFDFFFD3FE8879DF5EDCB5B6" box="[364,435,1765,1790]" captionStart="FIGURE 5" captionStartId="13.[151,250,1554,1576]" captionTargetBox="[170,1385,589,1501]" captionTargetId="figure@13.[151,1436,589,1532]" captionTargetPageId="13" captionText="FIGURE 5. Distributional maps of species belonging to the new genera Matthewsius and Monteithocanthon including close up insets of A, Pilbara region, Western Australia; B, northeastern Queensland; C, southeastern Queensland; D, southwestern Western Australia; E, southeastern South Australia; F, central to southeastern New South Wales." httpUri="https://zenodo.org/record/828975/files/figure.png" pageId="20" pageNumber="221">Fig. 5</figureCitation>
B. Known only from the summit of Mount Williams from
<quantity id="4C999B5BFFDFFFD3FB8979DC5A6FB5B5" box="[1133,1280,1766,1790]" metricMagnitude="2" metricUnit="m" metricValue="9.5" metricValueMax="10.0" metricValueMin="9.0" pageId="20" pageNumber="221" unit="m" value="950.0" valueMax="1000.0" valueMin="900.0">900–1000 m</quantity>
,
<collectingRegion id="49A5F85CFFDFFFD3FAF579DF5AF6B5B6" box="[1297,1433,1765,1790]" country="Australia" name="Queensland" pageId="20" pageNumber="221">Queensland</collectingRegion>
. Collected in rainforest in pitfall traps, dung baited traps, and berlesed litter.
</paragraph>
</subSubSection>
</treatment>
</document>
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