treatments-xml/data/03/C8/87/03C8879CFFB7FFB1FEC7FEB8B37AF984.xml
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<document id="0FF0BB7E4681418C3BDFD76D67CF89BC" ID-CLB-Dataset="50793" ID-DOI="10.5281/zenodo.171647" ID-GBIF-Dataset="d8261f5c-e4dc-4d7e-ad54-dd85e76a43bb" ID-ISSN="1175­5326" ID-Zenodo-Dep="171647" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459755668747" checkinUser="plazi" docAuthor="Provenzano, Francisco &amp; Milani, Nadia" docDate="2006" docId="03C8879CFFB7FFB1FEC7FEB8B37AF984" docLanguage="en" docName="zt01116p041.pdf" docOrigin="Zootaxa 1116" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Cordylancistrus nephelion Provenzano &amp; Milani, 2006, new species" docType="treatment" docVersion="10" lastPageNumber="35" masterDocId="FFF1FFE4FFB5FFB7FFCFFC44B729FFE6" masterDocTitle="Cordylancistrus nephelion (Siluriformes, Loricariidae), a new and endangered species of suckermouth armored catfish from the Tuy River, north­central Venezuela" masterLastPageNumber="41" masterPageNumber="29" pageNumber="31" updateTime="1698217605369" updateUser="ExternalLinkService">
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<mods:title id="36C092FACD8013E1928F47D7B913DEE9">Cordylancistrus nephelion (Siluriformes, Loricariidae), a new and endangered species of suckermouth armored catfish from the Tuy River, north­central Venezuela</mods:title>
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<mods:namePart id="A2E7D59F300868C279472FF6C9FEEA9A">Provenzano, Francisco</mods:namePart>
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<mods:namePart id="1A814A7A44348C0E7338BCBBF004F45C">Milani, Nadia</mods:namePart>
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<mods:date id="485F13390F28434681638624982CC6C0">2006</mods:date>
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<treatment id="03C8879CFFB7FFB1FEC7FEB8B37AF984" ID-DOI="http://doi.org/10.5281/zenodo.5696570" ID-GBIF-Taxon="119343552" ID-Zenodo-Dep="5696570" LSID="urn:lsid:zoobank.org:act:1E0501EB-1364-4851-8E25-D499E73CB754" httpUri="http://treatment.plazi.org/id/03C8879CFFB7FFB1FEC7FEB8B37AF984" lastPageId="6" lastPageNumber="35" pageId="2" pageNumber="31">
<subSubSection id="C37B6501FFB7FFB5FEC7FEB8B6F1FCDF" pageId="2" pageNumber="31" type="nomenclature">
<paragraph id="8BDE368AFFB7FFB5FEC7FEB8B5F6FCF0" blockId="2.[264,735,764,825]" box="[264,735,764,790]" pageId="2" pageNumber="31">
<heading id="D09681E6FFB7FFB5FEC7FEB8B5F6FCF0" bold="true" box="[264,735,764,790]" fontSize="11" level="1" pageId="2" pageNumber="31" reason="1">
<emphasis id="B915EA98FFB7FFB5FEC7FEB8B5F6FCF0" bold="true" box="[264,735,764,790]" pageId="2" pageNumber="31">
<taxonomicName id="4C614D09FFB7FFB5FEC7FEB8B56EFCF0" ID-CoL="YCPQ" box="[264,583,764,790]" class="Actinopterygii" family="Loricariidae" genus="Cordylancistrus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="31" phylum="Chordata" rank="species" species="nephelion" status="sp. nov.">
<emphasis id="B915EA98FFB7FFB5FEC7FEB8B56EFCF0" bold="true" box="[264,583,764,790]" italics="true" pageId="2" pageNumber="31">Cordylancistrus nephelion</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22657E3FFB7FFB5FD80FEB8B5F6FCF0" box="[591,735,764,790]" pageId="2" pageNumber="31" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BDE368AFFB7FFB5FEC7FF5BB6F1FCDF" blockId="2.[264,735,764,825]" box="[264,472,799,825]" pageId="2" pageNumber="31">
<figureCitation id="135A2A0FFFB7FFB5FEC7FF5BB65FFCDF" box="[264,374,799,825]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="6.[264,368,1434,1458]" captionStartId-1="7.[264,367,1442,1466]" captionTargetBox-0="[340,1246,329,1413]" captionTargetBox-1="[340,1246,809,1415]" captionTargetId-0="figure@6.[340,1248,329,1413]" captionTargetId-1="figure@7.[340,1248,809,1416]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionText-0="FIGURE 1. Cordylancistrus nephelion MBUCV­V­ 21800, 129. 1 mm SL, holotype, A: lateral, B: dorsal, and C: ventral views. Photos by N. Milani." captionText-1="FIGURE 2. Cordylancistrus nephelion MBUCV­V­ 21800, 129. 1 mm SL, holotype, Detail of the ventral surface of head. Photo by N. Milani" httpUri-0="https://zenodo.org/record/171648/files/figure.png" httpUri-1="https://zenodo.org/record/171649/files/figure.png" pageId="2" pageNumber="31">Figs. 12</figureCitation>
,
<tableCitation id="C6E30331FFB7FFB5FE4DFF5BB6F1FCDF" box="[386,472,799,825]" captionStart="TABLE 1" captionStartId="4.[264,352,483,507]" captionTargetBox="[276,1293,650,1815]" captionTargetPageId="4" captionText="TABLE 1. Morphometric data of Cordylancistrus nephelion. Morphometric data 1 15 are expressed as percentage of standard length, 16 20 as percentage of the head length, and 21 24 in ratios. MIN: minimum, MAX: maximum, AVE: average, STD: standard deviation, HL: head length, IOW: interorbital width, MRL: mandibular ramus length, OD: orbital diameter." httpUri="http://table.plazi.org/id/DF1E6602FFB1FFB3FEC7FDA7B38CFD85" pageId="2" pageNumber="31" tableUuid="DF1E6602FFB1FFB3FEC7FDA7B38CFD85">Table 1</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="C37B6501FFB7FFB5FEC7FF2AB569F9EF" pageId="2" pageNumber="31" type="materials_examined">
<paragraph id="8BDE368AFFB7FFB5FEC7FF2AB4DAFC3F" blockId="2.[264,1323,878,1905]" pageId="2" pageNumber="31">
<emphasis id="B915EA98FFB7FFB5FEC7FF2AB6AAFC6E" bold="true" box="[264,387,878,904]" pageId="2" pageNumber="31">
<typeStatus id="54DA8828FFB7FFB5FEC7FF2AB657FC6E" box="[264,382,878,904]" pageId="2" pageNumber="31" type="holotype">Holotype</typeStatus>
:
</emphasis>
MBUCV­V­
<quantity id="4C999B6FFFB7FFB5FDD0FF2BB5C4FC6F" box="[543,749,879,905]" metricMagnitude="4" metricUnit="m" metricValue="2.18001291" pageId="2" pageNumber="31" unit="mm" value="2.18001291E7">21800, 129.1 mm</quantity>
<collectionCode id="ED70AE4FFFB7FFB5FD39FF2BB432FC6F" box="[758,795,879,905]" pageId="2" pageNumber="31">SL</collectionCode>
;
<collectingCountry id="F376761AFFB7FFB5FCE7FF2BB48AFC6F" box="[808,931,879,905]" name="Venezuela" pageId="2" pageNumber="31">Venezuela</collectingCountry>
, Caribbean Sea basin, Tuy River system, Mesia River, tributary of the Guare River, near Village Corocito, approximately
<geoCoordinate id="EE55504DFFB7FFB5FEC7FFFBB642FC3F" box="[264,363,959,985]" direction="north" orientation="latitude" pageId="2" pageNumber="31" precision="925" value="10.2">10°12'N</geoCoordinate>
,
<geoCoordinate id="EE55504DFFB7FFB5FEB8FFFBB6C8FC3F" box="[375,481,959,985]" direction="west" orientation="longitude" pageId="2" pageNumber="31" precision="925" value="-67.083336">67°05'W</geoCoordinate>
; N. Padilla and H. Camejo;
<date id="FFDF104AFFB7FFB5FCE5FFFBB4C6FC3F" box="[810,1007,959,985]" pageId="2" pageNumber="31" value="1992-01-26">26 January 1992</date>
.
</paragraph>
<paragraph id="8BDE368AFFB7FFB5FEF7FFA2B569F9EF" blockId="2.[264,1323,878,1905]" pageId="2" pageNumber="31">
<emphasis id="B915EA98FFB7FFB5FEF7FFA2B697FBE6" bold="true" box="[312,446,998,1024]" pageId="2" pageNumber="31">
<typeStatus id="54DA8828FFB7FFB5FEF7FFA2B693FBE6" box="[312,442,998,1024]" pageId="2" pageNumber="31" type="paratype">Paratypes</typeStatus>
:
</emphasis>
All from
<collectingCountry id="F376761AFFB7FFB5FD8DFFA3B594FBE7" box="[578,701,999,1025]" name="Venezuela" pageId="2" pageNumber="31">Venezuela</collectingCountry>
, Caribbean Sea basin, Tuy River system. Miranda State: MBUCV­V­21801, 1 ex.,
<quantity id="4C999B6FFFB7FFB5FD44F84BB42CFBCF" box="[651,773,1039,1065]" metricMagnitude="-1" metricUnit="m" metricValue="1.212" pageId="2" pageNumber="31" unit="mm" value="121.2">121.2 mm</quantity>
<collectionCode id="ED70AE4FFFB7FFB5FCC0F84BB41CFBCF" box="[783,821,1039,1065]" pageId="2" pageNumber="31">SL</collectionCode>
; collected with the
<typeStatus id="54DA8828FFB7FFB5FBEBF84BB3A7FBCF" box="[1060,1166,1039,1065]" pageId="2" pageNumber="31" type="holotype">holotype</typeStatus>
. MBUCV­V­ 12526, 1 ex., 61.0 mm
<collectionCode id="ED70AE4FFFB7FFB5FDD9F873B515FBB7" box="[534,572,1079,1105]" pageId="2" pageNumber="31">SL</collectionCode>
; Grande River approximately
<quantity id="4C999B6FFFB7FFB5FC53F873B4CFFBB7" box="[924,998,1079,1105]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="31" unit="m" value="500.0">500 m</quantity>
upstream from its mouth in Santa Cruz River, a tributary of Taguaza River, Guatopo National Park, approx.
<geoCoordinate id="EE55504DFFB7FFB5FB0DF81BB20CFB9F" box="[1218,1317,1119,1145]" direction="north" orientation="latitude" pageId="2" pageNumber="31" precision="925" value="10.083333">10°05'N</geoCoordinate>
,
<geoCoordinate id="EE55504DFFB7FFB5FEC7F8C3B658FB47" box="[264,369,1159,1185]" direction="west" orientation="longitude" pageId="2" pageNumber="31" precision="925" value="-66.48333">66°29'W</geoCoordinate>
; R. Royero and party; 0
<date id="FFDF104AFFB7FFB5FD54F8C3B46CFB47" box="[667,837,1159,1185]" pageId="2" pageNumber="31" value="1981-03-01">1 March 1981</date>
. MBUCV­V­16382, 1 ex.,
<quantity id="4C999B6FFFB7FFB5FB44F8C3B3DFFB47" box="[1163,1270,1159,1185]" metricMagnitude="-2" metricUnit="m" metricValue="5.52" pageId="2" pageNumber="31" unit="mm" value="55.2">55.2 mm</quantity>
<collectionCode id="ED70AE4FFFB7FFB5FACFF8C3B20FFB47" box="[1280,1318,1159,1185]" pageId="2" pageNumber="31">SL</collectionCode>
; Mesia River, tributary of the Guare River, Paso del Viento, near the road to Tácata, approx.
<geoCoordinate id="EE55504DFFB7FFB5FEC7F893B642FB17" box="[264,363,1239,1265]" direction="north" orientation="latitude" pageId="2" pageNumber="31" precision="925" value="10.2">10°12'N</geoCoordinate>
,
<geoCoordinate id="EE55504DFFB7FFB5FEB7F893B6CBFB17" box="[376,482,1239,1265]" direction="west" orientation="longitude" pageId="2" pageNumber="31" precision="925" value="-67.03333">67°02'W</geoCoordinate>
; N. Padilla and L. Aguana;
<date id="FFDF104AFFB7FFB5FCE1F893B4DCFB17" box="[814,1013,1239,1265]" pageId="2" pageNumber="31" value="1992-01-30">30 January 1992</date>
. MBUCV­V­21806, 1 ex.,
<quantity id="4C999B6FFFB7FFB5FEC7F8BBB65DFAFF" box="[264,372,1279,1305]" metricMagnitude="-2" metricUnit="m" metricValue="3.11" pageId="2" pageNumber="31" unit="mm" value="31.1">31.1 mm</quantity>
<collectionCode id="ED70AE4FFFB7FFB5FEB1F8BBB68DFAFF" box="[382,420,1279,1305]" pageId="2" pageNumber="31">SL</collectionCode>
; Cagua River, Boca de Cagua, approximately
<quantity id="4C999B6FFFB7FFB5FC19F8BBB30AFAFF" box="[982,1059,1279,1305]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="31" unit="km" value="10.0">10 km</quantity>
E of Village Guayas, approx.
<geoCoordinate id="EE55504DFFB7FFB5FEA8F963B6E1FAA7" box="[359,456,1319,1345]" direction="north" orientation="latitude" pageId="2" pageNumber="31" precision="925" value="10.233334">10°14'N</geoCoordinate>
<geoCoordinate id="EE55504DFFB7FFB5FE00F963B511FAA7" box="[463,568,1319,1345]" direction="west" orientation="longitude" pageId="2" pageNumber="31" precision="925" value="-67.11667">67°07'W</geoCoordinate>
; R. Royero, F. Gil and C. Navarrese, 0
<date id="FFDF104AFFB7FFB5FC36F963B3B8FAA7" box="[1017,1169,1319,1345]" pageId="2" pageNumber="31" value="1991-04-06">6 April 1991</date>
. MBUCV­V­ 27872, 1 ex,
<quantity id="4C999B6FFFB7FFB5FE6CF90BB525FA8F" box="[419,524,1359,1385]" metricMagnitude="-2" metricUnit="m" metricValue="7.64" pageId="2" pageNumber="31" unit="mm" value="76.4">76.4 mm</quantity>
<collectionCode id="ED70AE4FFFB7FFB5FDDBF90BB513FA8F" box="[532,570,1359,1385]" pageId="2" pageNumber="31">SL</collectionCode>
; Mesia River, tributary of the Guare River, Pozo El Cirilo, near Village El Salado, approx.
<geoCoordinate id="EE55504DFFB7FFB5FD9EF933B59BFA77" box="[593,690,1399,1425]" direction="north" orientation="latitude" pageId="2" pageNumber="31" precision="925" value="10.216666">10°13'N</geoCoordinate>
<geoCoordinate id="EE55504DFFB7FFB5FD72F933B40FFA77" box="[701,806,1399,1425]" direction="west" orientation="longitude" pageId="2" pageNumber="31" precision="925" value="-67.05">67°03'W</geoCoordinate>
; N. Padilla and E. Camejo;
<date id="FFDF104AFFB7FFB5FB4FF933B20FFA77" box="[1152,1318,1399,1425]" pageId="2" pageNumber="31" value="1994-05-29">29 May 1994</date>
. Aragua State: MBUCV­V­22700, 1 ex.,
<quantity id="4C999B6FFFB7FFB5FD13F9DBB46DFA5F" box="[732,836,1439,1465]" metricMagnitude="-2" metricUnit="m" metricValue="7.59" pageId="2" pageNumber="31" unit="mm" value="75.9">75.9 mm</quantity>
<collectionCode id="ED70AE4FFFB7FFB5FC84F9DBB458FA5F" box="[843,881,1439,1465]" pageId="2" pageNumber="31">SL</collectionCode>
; Quebrada Agua Amarilla, a tributary of the Cagua River, Village Agua Amarilla, SE of Tejerias, approx.
<geoCoordinate id="EE55504DFFB7FFB5FBE7F983B3A2FA07" box="[1064,1163,1479,1505]" direction="north" orientation="latitude" pageId="2" pageNumber="31" precision="925" value="10.2">10°12'N</geoCoordinate>
,
<geoCoordinate id="EE55504DFFB7FFB5FB57F983B228FA07" box="[1176,1281,1479,1505]" direction="west" orientation="longitude" pageId="2" pageNumber="31" precision="925" value="-67.05">67°03'W</geoCoordinate>
; N. Padilla;
<date id="FFDF104AFFB7FFB5FEA6F9ABB515F9EF" box="[361,572,1519,1545]" pageId="2" pageNumber="31" value="1992-02-23">23 February 1992</date>
.
</paragraph>
</subSubSection>
<subSubSection id="C37B6501FFB7FFB5FEF7FA52B6DBF94F" pageId="2" pageNumber="31" type="diagnosis">
<paragraph id="8BDE368AFFB7FFB5FEF7FA52B6DBF94F" blockId="2.[264,1323,878,1905]" pageId="2" pageNumber="31">
<emphasis id="B915EA98FFB7FFB5FEF7FA52B69EF9D6" bold="true" box="[312,439,1558,1584]" pageId="2" pageNumber="31">Diagnosis.</emphasis>
<taxonomicName id="4C614D09FFB7FFB5FE0BFA53B429F9D6" box="[452,768,1559,1584]" class="Actinopterygii" family="Loricariidae" genus="Cordylancistrus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="31" phylum="Chordata" rank="species" species="nephelion">
<emphasis id="B915EA98FFB7FFB5FE0BFA53B429F9D6" box="[452,768,1559,1584]" italics="true" pageId="2" pageNumber="31">Cordylancistrus nephelion</emphasis>
</taxonomicName>
can be distinguished from the other species assigned to its genus by its unique color pattern. Head and body blackish to greenish brown with white irregular spots. Abdomen uniformly white. All fins with whitish spots that simulate bands.
</paragraph>
</subSubSection>
<subSubSection id="C37B6501FFB7FFB3FEF7FAF2B4BBFE60" lastPageId="4" lastPageNumber="33" pageId="2" pageNumber="31" type="description">
<paragraph id="8BDE368AFFB7FFB4FEF7FAF2B227FC68" blockId="2.[264,1323,878,1905]" lastBlockId="3.[264,1323,284,1870]" lastPageId="3" lastPageNumber="32" pageId="2" pageNumber="31">
<emphasis id="B915EA98FFB7FFB5FEF7FAF2B6F9F936" bold="true" box="[312,464,1718,1744]" pageId="2" pageNumber="31">Description.</emphasis>
Morphometric data are given in
<tableCitation id="C6E30331FFB7FFB5FC46FAF3B4C3F937" box="[905,1002,1719,1745]" captionStart="TABLE 1" captionStartId="4.[264,352,483,507]" captionTargetBox="[276,1293,650,1815]" captionTargetPageId="4" captionText="TABLE 1. Morphometric data of Cordylancistrus nephelion. Morphometric data 1 15 are expressed as percentage of standard length, 16 20 as percentage of the head length, and 21 24 in ratios. MIN: minimum, MAX: maximum, AVE: average, STD: standard deviation, HL: head length, IOW: interorbital width, MRL: mandibular ramus length, OD: orbital diameter." httpUri="http://table.plazi.org/id/DF1E6602FFB1FFB3FEC7FDA7B38CFD85" pageId="2" pageNumber="31" tableUuid="DF1E6602FFB1FFB3FEC7FDA7B38CFD85">Table 1</tableCitation>
. Head wide and slightly depressed. Snout edge covered with small plates, its contour oval. Eyes in dorsal position, dorsal edge of the orbits slightly raised, small odontodes found around orbits. Interorbital space flat. Posterior edge of the supraoccipital without a fleshy keel. Mouth wide, lips papillose, papillae of the anterior lip slightly larger. Edge of the posterior lip with marked undulations. Maxillary barbels very short and joined to lower lip by a fleshy flap, leaving tip of barbel free. Premaxilla and dentary enlarged. Inside mouth behind premaxilla with three to five larger papillae. Also behind the dentary three to five larger and smooth papillae present that increase in size toward center of mouth (
<figureCitation id="135A2A0FFFB6FFB4FC2EFDD0B303FE48" box="[993,1066,404,430]" captionStart="FIGURE 2" captionStartId="7.[264,367,1442,1466]" captionTargetBox="[340,1246,809,1415]" captionTargetId="figure@7.[340,1248,809,1416]" captionTargetPageId="7" captionText="FIGURE 2. Cordylancistrus nephelion MBUCV­V­ 21800, 129. 1 mm SL, holotype, Detail of the ventral surface of head. Photo by N. Milani" httpUri="https://zenodo.org/record/171649/files/figure.png" pageId="3" pageNumber="32">Fig. 2</figureCitation>
). Teeth numerous, 49 to 105 teeth on each premaxilla and 53 to 129 on each dentary. Teeth are very thin and elongated, with apex curved toward interior of mouth. Teeth bifurcate, one lobe slightly longer than other, both tips are pointed. Tooth apex yellowish, stalk whitish. Interopercular hypertrophied evertible odontodes vary in size; longest odontode sometimes reaches the cleithrum but surpasses it in only one specimen. Hypertrophied odontodes vary in number from six on smallest specimen to 25 on the largest. Body robust and relatively deep. Caudal peduncle compressed. Lateral plates of the body without spiny keels. Lateral line plates 23 to 24. Post­anal plates 10 to 11. Inter­dorsal plates six, the last with a slight keel. Ventral surface of body naked to origin of anal fin. Dorsal fin
<date id="FFDF104AFFB6FFB4FC24FEB8B323FCF0" box="[1003,1034,764,790]" pageId="3" pageNumber="32">I8.</date>
Pectoral fin
<date id="FFDF104AFFB6FFB4FB6BFEB8B396FCF0" box="[1188,1215,764,790]" pageId="3" pageNumber="32">I6</date>
, pectoral spine relatively short, reaching one­third length of pelvic spine or less. In all examined specimens, pectoral spine with odontodes on posterodorsal surface. Anal fin with four branched rays, three on one specimen. Pelvic fin
<date id="FFDF104AFFB6FFB4FC87FF30B441FC68" box="[840,872,884,910]" pageId="3" pageNumber="32">I5.</date>
Caudal fin
<date id="FFDF104AFFB6FFB4FC3DFF30B333FC68" box="[1010,1050,884,910]" pageId="3" pageNumber="32" value="1914-01">I14</date>
I, obliquely truncate.
</paragraph>
<paragraph id="8BDE368AFFB6FFB4FEF7FFD8B378FA88" blockId="3.[264,1323,284,1870]" pageId="3" pageNumber="32">
<emphasis id="B915EA98FFB6FFB4FEF7FFD8B6ADFC50" bold="true" box="[312,388,924,950]" pageId="3" pageNumber="32">Color.</emphasis>
In specimens preserved in 70% ethanol head and body color varies from blackish to greenish brown with white irregular spots; abdomen uniformly white. Dorsal surface of head with small spots that become larger posteriorly. Spots along body approximately as wide as the eye; in some cases two or more spots unite to form a spot of greater size. Spine and branched rays of dorsal fin with four to five whitish spots. Interradial membrane with irregular translucent spots, more evident towards the distal margin. In largest specimens (121 and
<quantity id="4C999B6FFFB6FFB4FD00F8C8B419FB40" box="[719,816,1164,1190]" metricMagnitude="-1" metricUnit="m" metricValue="1.29" pageId="3" pageNumber="32" unit="mm" value="129.0">129 mm</quantity>
SL), pectoral­fin and pelvic­fin spine have six to seven whitish spots and branched rays have five to six, while in smaller specimens (
<quantity id="4C999B6FFFB6FFB4FEDFF898B6A9FB10" box="[272,384,1244,1270]" metricMagnitude="-2" metricUnit="m" metricValue="7.64" pageId="3" pageNumber="32" unit="mm" value="76.4">76.4 mm</quantity>
SL or less) spines with four to five and branched rays with three to four. Interradial membrane hyaline. Anal fin generally whitish with dark distal border. Adipose fin with whitish band. Caudal­fin spines with six to seven whitish spots and branched rays with four to six. Interradial membrane with pattern similar to dorsal fin.
</paragraph>
<paragraph id="8BDE368AFFB6FFB4FEF7F938B597F930" blockId="3.[264,1323,284,1870]" pageId="3" pageNumber="32">
<emphasis id="B915EA98FFB6FFB4FEF7F938B6A7FA70" bold="true" box="[312,398,1404,1430]" pageId="3" pageNumber="32">Range.</emphasis>
The specimens of
<taxonomicName id="4C614D09FFB6FFB4FDA4F938B48BFA73" box="[619,930,1404,1429]" class="Actinopterygii" family="Loricariidae" genus="Cordylancistrus" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="32" phylum="Chordata" rank="species" species="nephelion">
<emphasis id="B915EA98FFB6FFB4FDA4F938B48BFA73" box="[619,930,1404,1429]" italics="true" pageId="3" pageNumber="32">Cordylancistrus nephelion</emphasis>
</taxonomicName>
were captured in three tributaries of the Tuy River: the Cagua River, the Mesia River (a tributary of the Guare River) and the Santa Cruz River (a tributary of the Taguaza River) at
<quantity id="4C999B6FFFB6FFB4FC5AF988B4CDFA00" box="[917,996,1484,1510]" metricMagnitude="4" metricUnit="m" metricValue="4.0" pageId="3" pageNumber="32" unit="km" value="40.0">40 km</quantity>
,
<quantity id="4C999B6FFFB6FFB4FC3EF988B314FA00" box="[1009,1085,1484,1510]" metricMagnitude="4" metricUnit="m" metricValue="5.5" pageId="3" pageNumber="32" unit="km" value="55.0">55 km</quantity>
and
<quantity id="4C999B6FFFB6FFB4FBB6F988B3FAFA00" box="[1145,1235,1484,1510]" metricMagnitude="5" metricUnit="m" metricValue="1.3" pageId="3" pageNumber="32" unit="km" value="130.0">130 km</quantity>
east of the headwaters of the Tuy River respectively (
<figureCitation id="135A2A0FFFB6FFB4FCFBF9B0B454F9E8" box="[820,893,1524,1550]" captionStart="FIGURE 3" captionStartId="8.[264,367,1825,1849]" captionTargetBox="[390,1213,1170,1753]" captionTargetId="figure@8.[376,1215,1150,1795]" captionTargetPageId="8" captionText="FIGURE 3. Geographical distributions of Cordylancistrus species in Venezuela. C. nephelion circles, white circle is the type locality. C. torbesensis white square. C. perija black square." httpUri="https://zenodo.org/record/171650/files/figure.png" pageId="3" pageNumber="32">Fig. 3</figureCitation>
). This distribution suggests that the species may be present throughout the Tuy River basin. The species seems to inhabit small rivers of the north slope of the Serranía del Interior, Cordillera de La Costa. These rivers have transparent waters, with moderate to strong flow and a temperature of 15° to 20° C. The bottom generally consists of stone, gravel and sand. The predominant vegetation in this mountainous area is cloud forest.
</paragraph>
<paragraph id="8BDE368AFFB6FFB3FEF7FAA0B4BBFE60" blockId="3.[264,1323,284,1870]" lastBlockId="4.[264,1323,284,390]" lastPageId="4" lastPageNumber="33" pageId="3" pageNumber="32">
An exhaustive review of the Venezuelan fish collections shows that the geographic distribution of the three species of
<taxonomicName id="4C614D09FFB6FFB4FD6CFB48B476F8C3" box="[675,863,1804,1829]" class="Actinopterygii" family="Loricariidae" genus="Cordylancistrus" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="32" phylum="Chordata" rank="genus">
<emphasis id="B915EA98FFB6FFB4FD6CFB48B476F8C3" box="[675,863,1804,1829]" italics="true" pageId="3" pageNumber="32">Cordylancistrus</emphasis>
</taxonomicName>
present in the country is restricted and disjunct.
<taxonomicName id="4C614D09FFB6FFB4FEBAFB70B596F8AB" box="[373,703,1844,1869]" class="Actinopterygii" family="Loricariidae" genus="Cordylancistrus" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="32" phylum="Chordata" rank="species" species="torbesensis">
<emphasis id="B915EA98FFB6FFB4FEBAFB70B596F8AB" box="[373,703,1844,1869]" italics="true" pageId="3" pageNumber="32">Cordylancistrus torbesensis</emphasis>
</taxonomicName>
inhabits the foothill rivers of the south slope of the Cordillera de Mérida;
<taxonomicName id="4C614D09FFB1FFB3FDC0FD58B5ACFED3" box="[527,645,284,309]" class="Actinopterygii" family="Loricariidae" genus="Cordylancistrus" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="33" phylum="Chordata" rank="species" species="perijae">
<emphasis id="B915EA98FFB1FFB3FDC0FD58B5ACFED3" box="[527,645,284,309]" italics="true" pageId="4" pageNumber="33">C. perijae</emphasis>
</taxonomicName>
is found in rivers of the eastern slope, north of the Sierra of
<taxonomicName id="4C614D09FFB1FFB3FEE7FD00B645FEB8" box="[296,364,324,350]" class="Actinopterygii" family="Loricariidae" genus="Cordylancistrus" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="37" phylum="Chordata" rank="species" species="perija">Perijá</taxonomicName>
and
<taxonomicName id="4C614D09FFB1FFB3FE67FD00B516FEBB" box="[424,575,324,349]" class="Actinopterygii" family="Loricariidae" genus="Cordylancistrus" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="33" phylum="Chordata" rank="species" species="nephelion">
<emphasis id="B915EA98FFB1FFB3FE67FD00B516FEBB" box="[424,575,324,349]" italics="true" pageId="4" pageNumber="33">C. nephelion</emphasis>
</taxonomicName>
inhabits the Tuy River basin, in rivers of the north slope of the Serranía of the Interior, Cordillera de La Costa (
<figureCitation id="135A2A0FFFB1FFB3FCF4FD28B4ABFE60" box="[827,898,364,390]" captionStart="FIGURE 3" captionStartId="8.[264,367,1825,1849]" captionTargetBox="[390,1213,1170,1753]" captionTargetId="figure@8.[376,1215,1150,1795]" captionTargetPageId="8" captionText="FIGURE 3. Geographical distributions of Cordylancistrus species in Venezuela. C. nephelion circles, white circle is the type locality. C. torbesensis white square. C. perija black square." httpUri="https://zenodo.org/record/171650/files/figure.png" pageId="4" pageNumber="33">Fig. 3</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF1E6602FFB1FFB3FEC7FDA7B38CFD85" ID-Table-UUID="DF1E6602FFB1FFB3FEC7FDA7B38CFD85" httpUri="http://table.plazi.org/id/DF1E6602FFB1FFB3FEC7FDA7B38CFD85" pageId="4" pageNumber="33" targetBox="[276,1293,650,1815]" targetIsTable="true" targetPageId="4">
<paragraph id="8BDE368AFFB1FFB3FEC7FDA7B38CFD85" blockId="4.[264,1323,483,611]" pageId="4" pageNumber="33">
<emphasis id="B915EA98FFB1FFB3FEC7FDA7B6ADFE1D" bold="true" box="[264,388,483,507]" pageId="4" pageNumber="33">TABLE 1.</emphasis>
Morphometric data of
<taxonomicName id="4C614D09FFB1FFB3FD6DFDA0B4EFFE1D" box="[674,966,484,507]" class="Actinopterygii" family="Loricariidae" genus="Cordylancistrus" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="33" phylum="Chordata" rank="species" species="nephelion">
<emphasis id="B915EA98FFB1FFB3FD6DFDA0B4EFFE1D" box="[674,966,484,507]" italics="true" pageId="4" pageNumber="33">Cordylancistrus nephelion</emphasis>
</taxonomicName>
. Morphometric data 115 are expressed as percentage of standard length, 1620 as percentage of the head length, and 2124 in ratios. MIN: minimum, MAX: maximum, AVE: average, STD: standard deviation, HL: head length, IOW: interorbital width, MRL: mandibular ramus length, OD: orbital diameter.
</paragraph>
</caption>
<paragraph id="8BDE368AFFB1FFB3FD05FECEB222F8F1" pageId="4" pageNumber="33">
<table id="F961C42AFFB10048FEDBFECEB224F8F1" box="[276,1293,650,1815]" gridcols="6" gridrows="26" pageId="4" pageNumber="33">
<tr id="355134C8FFB10048FEDBFECEB224FD47" box="[276,1293,650,673]" gridrow="0" pageId="4" pageNumber="33" rowspan-0="1">
<th id="76805DB4FFB10048FD05FECEB410FD47" box="[714,825,650,673]" gridcol="1" gridrow="0" pageId="4" pageNumber="33">Holotype</th>
<th id="76805DB4FFB10048FC94FECEB488FD47" box="[859,929,650,673]" gridcol="2" gridrow="0" pageId="4" pageNumber="33">MIN</th>
<th id="76805DB4FFB10048FC0AFECEB333FD47" box="[965,1050,650,673]" gridcol="3" gridrow="0" pageId="4" pageNumber="33">MAX</th>
<th id="76805DB4FFB10048FB8BFECEB3BAFD47" box="[1092,1171,650,673]" gridcol="4" gridrow="0" pageId="4" pageNumber="33">AVE</th>
<th id="76805DB4FFB10048FB71FECEB224FD47" box="[1214,1293,650,673]" gridcol="5" gridrow="0" pageId="4" pageNumber="33">STD</th>
</tr>
<tr id="355134C8FFB10048FEDBFEFEB224FD37" box="[276,1293,698,721]" gridrow="1" pageId="4" pageNumber="33">
<th id="76805DB4FFB10048FEDBFEFEB5B3FD37" box="[276,666,698,721]" gridcol="0" gridrow="1" pageId="4" pageNumber="33">SL (mm)</th>
<td id="76805DB4FFB10048FD05FEFEB410FD37" box="[714,825,698,721]" gridcol="1" gridrow="1" pageId="4" pageNumber="33">129,10</td>
<td id="76805DB4FFB10048FC94FEFEB488FD37" box="[859,929,698,721]" gridcol="2" gridrow="1" pageId="4" pageNumber="33">31,10</td>
<td id="76805DB4FFB10048FC0AFEFEB333FD37" box="[965,1050,698,721]" gridcol="3" gridrow="1" pageId="4" pageNumber="33">129,10</td>
<td id="76805DB4FFB10048FB8BFEFEB3BAFD37" box="[1092,1171,698,721]" gridcol="4" gridrow="1" pageId="4" pageNumber="33">78,56</td>
<td id="76805DB4FFB10048FB71FEFEB224FD37" box="[1214,1293,698,721]" gridcol="5" gridrow="1" pageId="4" pageNumber="33">35,33</td>
</tr>
<tr id="355134C8FFB10048FEDBFEAEB224FCE7" box="[276,1293,746,769]" gridrow="2" pageId="4" pageNumber="33">
<th id="76805DB4FFB10048FEDBFEAEB5B3FCE7" box="[276,666,746,769]" gridcol="0" gridrow="2" pageId="4" pageNumber="33">1 Head length</th>
<td id="76805DB4FFB10048FD05FEAEB410FCE7" box="[714,825,746,769]" gridcol="1" gridrow="2" pageId="4" pageNumber="33">35,24</td>
<td id="76805DB4FFB10048FC94FEAEB488FCE7" box="[859,929,746,769]" gridcol="2" gridrow="2" pageId="4" pageNumber="33">34,32</td>
<td id="76805DB4FFB10048FC0AFEAEB333FCE7" box="[965,1050,746,769]" gridcol="3" gridrow="2" pageId="4" pageNumber="33">41,16</td>
<td id="76805DB4FFB10048FB8BFEAEB3BAFCE7" box="[1092,1171,746,769]" gridcol="4" gridrow="2" pageId="4" pageNumber="33">36,32</td>
<td id="76805DB4FFB10048FB71FEAEB224FCE7" box="[1214,1293,746,769]" gridcol="5" gridrow="2" pageId="4" pageNumber="33">2,41</td>
</tr>
<tr id="355134C8FFB10048FEDBFF53B224FCC8" box="[276,1293,791,814]" gridrow="3" pageId="4" pageNumber="33">
<th id="76805DB4FFB10048FEDBFF53B5B3FCC8" box="[276,666,791,814]" gridcol="0" gridrow="3" pageId="4" pageNumber="33">2 Predorsal length</th>
<td id="76805DB4FFB10048FD05FF53B410FCC8" box="[714,825,791,814]" gridcol="1" gridrow="3" pageId="4" pageNumber="33">43,14</td>
<td id="76805DB4FFB10048FC94FF53B488FCC8" box="[859,929,791,814]" gridcol="2" gridrow="3" pageId="4" pageNumber="33">42,90</td>
<td id="76805DB4FFB10048FC0AFF53B333FCC8" box="[965,1050,791,814]" gridcol="3" gridrow="3" pageId="4" pageNumber="33">46,62</td>
<td id="76805DB4FFB10048FB8BFF53B3BAFCC8" box="[1092,1171,791,814]" gridcol="4" gridrow="3" pageId="4" pageNumber="33">44,17</td>
<td id="76805DB4FFB10048FB71FF53B224FCC8" box="[1214,1293,791,814]" gridcol="5" gridrow="3" pageId="4" pageNumber="33">1,37</td>
</tr>
<tr id="355134C8FFB10048FEDBFF01B224FCBA" box="[276,1293,837,860]" gridrow="4" pageId="4" pageNumber="33">
<th id="76805DB4FFB10048FEDBFF01B5B3FCBA" box="[276,666,837,860]" gridcol="0" gridrow="4" pageId="4" pageNumber="33">3 Postdorsal length</th>
<td id="76805DB4FFB10048FD05FF01B410FCBA" box="[714,825,837,860]" gridcol="1" gridrow="4" pageId="4" pageNumber="33">36,64</td>
<td id="76805DB4FFB10048FC94FF01B488FCBA" box="[859,929,837,860]" gridcol="2" gridrow="4" pageId="4" pageNumber="33">35,05</td>
<td id="76805DB4FFB10048FC0AFF01B333FCBA" box="[965,1050,837,860]" gridcol="3" gridrow="4" pageId="4" pageNumber="33">37,68</td>
<td id="76805DB4FFB10048FB8BFF01B3BAFCBA" box="[1092,1171,837,860]" gridcol="4" gridrow="4" pageId="4" pageNumber="33">35,98</td>
<td id="76805DB4FFB10048FB71FF01B224FCBA" box="[1214,1293,837,860]" gridcol="5" gridrow="4" pageId="4" pageNumber="33">0,95</td>
</tr>
<tr id="355134C8FFB10048FEDBFF36B224FC6F" box="[276,1293,882,905]" gridrow="5" pageId="4" pageNumber="33">
<th id="76805DB4FFB10048FEDBFF36B5B3FC6F" box="[276,666,882,905]" gridcol="0" gridrow="5" pageId="4" pageNumber="33">4 Interdorsal length</th>
<td id="76805DB4FFB10048FD05FF36B410FC6F" box="[714,825,882,905]" gridcol="1" gridrow="5" pageId="4" pageNumber="33">14,79</td>
<td id="76805DB4FFB10048FC94FF36B488FC6F" box="[859,929,882,905]" gridcol="2" gridrow="5" pageId="4" pageNumber="33">10,13</td>
<td id="76805DB4FFB10048FC0AFF36B333FC6F" box="[965,1050,882,905]" gridcol="3" gridrow="5" pageId="4" pageNumber="33">15,31</td>
<td id="76805DB4FFB10048FB8BFF36B3BAFC6F" box="[1092,1171,882,905]" gridcol="4" gridrow="5" pageId="4" pageNumber="33">13,29</td>
<td id="76805DB4FFB10048FB71FF36B224FC6F" box="[1214,1293,882,905]" gridcol="5" gridrow="5" pageId="4" pageNumber="33">1,93</td>
</tr>
<tr id="355134C8FFB10048FEDBFFE4B224FC51" box="[276,1293,928,951]" gridrow="6" pageId="4" pageNumber="33">
<th id="76805DB4FFB10048FEDBFFE4B5B3FC51" box="[276,666,928,951]" gridcol="0" gridrow="6" pageId="4" pageNumber="33">5 Preanal length</th>
<td id="76805DB4FFB10048FD05FFE4B410FC51" box="[714,825,928,951]" gridcol="1" gridrow="6" pageId="4" pageNumber="33">64,14</td>
<td id="76805DB4FFB10048FC94FFE4B488FC51" box="[859,929,928,951]" gridcol="2" gridrow="6" pageId="4" pageNumber="33">64,14</td>
<td id="76805DB4FFB10048FC0AFFE4B333FC51" box="[965,1050,928,951]" gridcol="3" gridrow="6" pageId="4" pageNumber="33">69,45</td>
<td id="76805DB4FFB10048FB8BFFE4B3BAFC51" box="[1092,1171,928,951]" gridcol="4" gridrow="6" pageId="4" pageNumber="33">67,36</td>
<td id="76805DB4FFB10048FB71FFE4B224FC51" box="[1214,1293,928,951]" gridcol="5" gridrow="6" pageId="4" pageNumber="33">1,84</td>
</tr>
<tr id="355134C8FFB10048FEDBFF89B224FC02" box="[276,1293,973,996]" gridrow="7" pageId="4" pageNumber="33">
<th id="76805DB4FFB10048FEDBFF89B5B3FC02" box="[276,666,973,996]" gridcol="0" gridrow="7" pageId="4" pageNumber="33">6 Postanal length</th>
<td id="76805DB4FFB10048FD05FF89B410FC02" box="[714,825,973,996]" gridcol="1" gridrow="7" pageId="4" pageNumber="33">28,43</td>
<td id="76805DB4FFB10048FC94FF89B488FC02" box="[859,929,973,996]" gridcol="2" gridrow="7" pageId="4" pageNumber="33">28,43</td>
<td id="76805DB4FFB10048FC0AFF89B333FC02" box="[965,1050,973,996]" gridcol="3" gridrow="7" pageId="4" pageNumber="33">35,11</td>
<td id="76805DB4FFB10048FB8BFF89B3BAFC02" box="[1092,1171,973,996]" gridcol="4" gridrow="7" pageId="4" pageNumber="33">30,50</td>
<td id="76805DB4FFB10048FB71FF89B224FC02" box="[1214,1293,973,996]" gridcol="5" gridrow="7" pageId="4" pageNumber="33">2,20</td>
</tr>
<tr id="355134C8FFB10048FEDBFFBFB224FBF4" box="[276,1293,1019,1042]" gridrow="8" pageId="4" pageNumber="33">
<th id="76805DB4FFB10048FEDBFFBFB5B3FBF4" box="[276,666,1019,1042]" gridcol="0" gridrow="8" pageId="4" pageNumber="33">7 Thoracic length</th>
<td id="76805DB4FFB10048FD05FFBFB410FBF4" box="[714,825,1019,1042]" gridcol="1" gridrow="8" pageId="4" pageNumber="33">20,99</td>
<td id="76805DB4FFB10048FC94FFBFB488FBF4" box="[859,929,1019,1042]" gridcol="2" gridrow="8" pageId="4" pageNumber="33">20,58</td>
<td id="76805DB4FFB10048FC0AFFBFB333FBF4" box="[965,1050,1019,1042]" gridcol="3" gridrow="8" pageId="4" pageNumber="33">23,44</td>
<td id="76805DB4FFB10048FB8BFFBFB3BAFBF4" box="[1092,1171,1019,1042]" gridcol="4" gridrow="8" pageId="4" pageNumber="33">21,93</td>
<td id="76805DB4FFB10048FB71FFBFB224FBF4" box="[1214,1293,1019,1042]" gridcol="5" gridrow="8" pageId="4" pageNumber="33">1,00</td>
</tr>
<tr id="355134C8FFB10048FEDBF86CB224FBD9" box="[276,1293,1064,1087]" gridrow="9" pageId="4" pageNumber="33">
<th id="76805DB4FFB10048FEDBF86CB5B3FBD9" box="[276,666,1064,1087]" gridcol="0" gridrow="9" pageId="4" pageNumber="33">8 Abdominal length</th>
<td id="76805DB4FFB10048FD05F86CB410FBD9" box="[714,825,1064,1087]" gridcol="1" gridrow="9" pageId="4" pageNumber="33">22,46</td>
<td id="76805DB4FFB10048FC94F86CB488FBD9" box="[859,929,1064,1087]" gridcol="2" gridrow="9" pageId="4" pageNumber="33">19,61</td>
<td id="76805DB4FFB10048FC0AF86CB333FBD9" box="[965,1050,1064,1087]" gridcol="3" gridrow="9" pageId="4" pageNumber="33">24,01</td>
<td id="76805DB4FFB10048FB8BF86CB3BAFBD9" box="[1092,1171,1064,1087]" gridcol="4" gridrow="9" pageId="4" pageNumber="33">21,69</td>
<td id="76805DB4FFB10048FB71F86CB224FBD9" box="[1214,1293,1064,1087]" gridcol="5" gridrow="9" pageId="4" pageNumber="33">1,40</td>
</tr>
<tr id="355134C8FFB10048FEDBF812B224FB8B" box="[276,1293,1110,1133]" gridrow="10" pageId="4" pageNumber="33">
<th id="76805DB4FFB10048FEDBF812B5B3FB8B" box="[276,666,1110,1133]" gridcol="0" gridrow="10" pageId="4" pageNumber="33">9 Dorsal fin base</th>
<td id="76805DB4FFB10048FD05F812B410FB8B" box="[714,825,1110,1133]" gridcol="1" gridrow="10" pageId="4" pageNumber="33">22,08</td>
<td id="76805DB4FFB10048FC94F812B488FB8B" box="[859,929,1110,1133]" gridcol="2" gridrow="10" pageId="4" pageNumber="33">18,49</td>
<td id="76805DB4FFB10048FC0AF812B333FB8B" box="[965,1050,1110,1133]" gridcol="3" gridrow="10" pageId="4" pageNumber="33">22,53</td>
<td id="76805DB4FFB10048FB8BF812B3BAFB8B" box="[1092,1171,1110,1133]" gridcol="4" gridrow="10" pageId="4" pageNumber="33">20,91</td>
<td id="76805DB4FFB10048FB71F812B224FB8B" box="[1214,1293,1110,1133]" gridcol="5" gridrow="10" pageId="4" pageNumber="33">1,49</td>
</tr>
<tr id="355134C8FFB10048FEDBF8C7B224FB7C" box="[276,1293,1155,1178]" gridrow="11" pageId="4" pageNumber="33">
<th id="76805DB4FFB10048FEDBF8C7B5B3FB7C" box="[276,666,1155,1178]" gridcol="0" gridrow="11" pageId="4" pageNumber="33">10 Dorsal spine length</th>
<td id="76805DB4FFB10048FD05F8C7B410FB7C" box="[714,825,1155,1178]" gridcol="1" gridrow="11" pageId="4" pageNumber="33">24,98</td>
<td id="76805DB4FFB10048FC94F8C7B488FB7C" box="[859,929,1155,1178]" gridcol="2" gridrow="11" pageId="4" pageNumber="33">21,34</td>
<td id="76805DB4FFB10048FC0AF8C7B333FB7C" box="[965,1050,1155,1178]" gridcol="3" gridrow="11" pageId="4" pageNumber="33">25,50</td>
<td id="76805DB4FFB10048FB8BF8C7B3BAFB7C" box="[1092,1171,1155,1178]" gridcol="4" gridrow="11" pageId="4" pageNumber="33">23,50</td>
<td id="76805DB4FFB10048FB71F8C7B224FB7C" box="[1214,1293,1155,1178]" gridcol="5" gridrow="11" pageId="4" pageNumber="33">1,67</td>
</tr>
<tr id="355134C8FFB10048FEDBF8F5B224FB2E" box="[276,1293,1201,1224]" gridrow="12" pageId="4" pageNumber="33">
<th id="76805DB4FFB10048FEDBF8F5B5B3FB2E" box="[276,666,1201,1224]" gridcol="0" gridrow="12" pageId="4" pageNumber="33">11 Pectoral spine length</th>
<td id="76805DB4FFB10048FD05F8F5B410FB2E" box="[714,825,1201,1224]" gridcol="1" gridrow="12" pageId="4" pageNumber="33">32,61</td>
<td id="76805DB4FFB10048FC94F8F5B488FB2E" box="[859,929,1201,1224]" gridcol="2" gridrow="12" pageId="4" pageNumber="33">24,77</td>
<td id="76805DB4FFB10048FC0AF8F5B333FB2E" box="[965,1050,1201,1224]" gridcol="3" gridrow="12" pageId="4" pageNumber="33">33,00</td>
<td id="76805DB4FFB10048FB8BF8F5B3BAFB2E" box="[1092,1171,1201,1224]" gridcol="4" gridrow="12" pageId="4" pageNumber="33">27,77</td>
<td id="76805DB4FFB10048FB71F8F5B224FB2E" box="[1214,1293,1201,1224]" gridcol="5" gridrow="12" pageId="4" pageNumber="33">3,65</td>
</tr>
<tr id="355134C8FFB10048FEDBF89AB224FB13" box="[276,1293,1246,1269]" gridrow="13" pageId="4" pageNumber="33">
<th id="76805DB4FFB10048FEDBF89AB5B3FB13" box="[276,666,1246,1269]" gridcol="0" gridrow="13" pageId="4" pageNumber="33">12 Pelvic spine length</th>
<td id="76805DB4FFB10048FD05F89AB410FB13" box="[714,825,1246,1269]" gridcol="1" gridrow="13" pageId="4" pageNumber="33">25,64</td>
<td id="76805DB4FFB10048FC94F89AB488FB13" box="[859,929,1246,1269]" gridcol="2" gridrow="13" pageId="4" pageNumber="33">25,00</td>
<td id="76805DB4FFB10048FC0AF89AB333FB13" box="[965,1050,1246,1269]" gridcol="3" gridrow="13" pageId="4" pageNumber="33">27,31</td>
<td id="76805DB4FFB10048FB8BF89AB3BAFB13" box="[1092,1171,1246,1269]" gridcol="4" gridrow="13" pageId="4" pageNumber="33">25,94</td>
<td id="76805DB4FFB10048FB71F89AB224FB13" box="[1214,1293,1246,1269]" gridcol="5" gridrow="13" pageId="4" pageNumber="33">0,86</td>
</tr>
<tr id="355134C8FFB10048FEDBF948B224FAC5" box="[276,1293,1292,1315]" gridrow="14" pageId="4" pageNumber="33">
<th id="76805DB4FFB10048FEDBF948B5B3FAC5" box="[276,666,1292,1315]" gridcol="0" gridrow="14" pageId="4" pageNumber="33">13 Cleithral width</th>
<td id="76805DB4FFB10048FD05F948B410FAC5" box="[714,825,1292,1315]" gridcol="1" gridrow="14" pageId="4" pageNumber="33">36,17</td>
<td id="76805DB4FFB10048FC94F948B488FAC5" box="[859,929,1292,1315]" gridcol="2" gridrow="14" pageId="4" pageNumber="33">32,80</td>
<td id="76805DB4FFB10048FC0AF948B333FAC5" box="[965,1050,1292,1315]" gridcol="3" gridrow="14" pageId="4" pageNumber="33">36,17</td>
<td id="76805DB4FFB10048FB8BF948B3BAFAC5" box="[1092,1171,1292,1315]" gridcol="4" gridrow="14" pageId="4" pageNumber="33">34,61</td>
<td id="76805DB4FFB10048FB71F948B224FAC5" box="[1214,1293,1292,1315]" gridcol="5" gridrow="14" pageId="4" pageNumber="33">1,14</td>
</tr>
<tr id="355134C8FFB10048FEDBF97DB224FAB6" box="[276,1293,1337,1360]" gridrow="15" pageId="4" pageNumber="33">
<th id="76805DB4FFB10048FEDBF97DB5B3FAB6" box="[276,666,1337,1360]" gridcol="0" gridrow="15" pageId="4" pageNumber="33">14 Caudal peduncle depth</th>
<td id="76805DB4FFB10048FD05F97DB410FAB6" box="[714,825,1337,1360]" gridcol="1" gridrow="15" pageId="4" pageNumber="33">14,64</td>
<td id="76805DB4FFB10048FC94F97DB488FAB6" box="[859,929,1337,1360]" gridcol="2" gridrow="15" pageId="4" pageNumber="33">12,50</td>
<td id="76805DB4FFB10048FC0AF97DB333FAB6" box="[965,1050,1337,1360]" gridcol="3" gridrow="15" pageId="4" pageNumber="33">14,64</td>
<td id="76805DB4FFB10048FB8BF97DB3BAFAB6" box="[1092,1171,1337,1360]" gridcol="4" gridrow="15" pageId="4" pageNumber="33">13,43</td>
<td id="76805DB4FFB10048FB71F97DB224FAB6" box="[1214,1293,1337,1360]" gridcol="5" gridrow="15" pageId="4" pageNumber="33">0,84</td>
</tr>
<tr id="355134C8FFB10048FEDBF923B224FA98" box="[276,1293,1383,1406]" gridrow="16" pageId="4" pageNumber="33">
<th id="76805DB4FFB10048FEDBF923B5B3FA98" box="[276,666,1383,1406]" gridcol="0" gridrow="16" pageId="4" pageNumber="33">15 Head depth</th>
<td id="76805DB4FFB10048FD05F923B410FA98" box="[714,825,1383,1406]" gridcol="1" gridrow="16" pageId="4" pageNumber="33">19,33</td>
<td id="76805DB4FFB10048FC94F923B488FA98" box="[859,929,1383,1406]" gridcol="2" gridrow="16" pageId="4" pageNumber="33">17,21</td>
<td id="76805DB4FFB10048FC0AF923B333FA98" box="[965,1050,1383,1406]" gridcol="3" gridrow="16" pageId="4" pageNumber="33">20,16</td>
<td id="76805DB4FFB10048FB8BF923B3BAFA98" box="[1092,1171,1383,1406]" gridcol="4" gridrow="16" pageId="4" pageNumber="33">18,81</td>
<td id="76805DB4FFB10048FB71F923B224FA98" box="[1214,1293,1383,1406]" gridcol="5" gridrow="16" pageId="4" pageNumber="33">1,02</td>
</tr>
<tr id="355134C8FFB10048FEDBF9D0B224FA4D" box="[276,1293,1428,1451]" gridrow="17" pageId="4" pageNumber="33">
<th id="76805DB4FFB10048FEDBF9D0B5B3FA4D" box="[276,666,1428,1451]" gridcol="0" gridrow="17" pageId="4" pageNumber="33">16 Snout length</th>
<td id="76805DB4FFB10048FD05F9D0B410FA4D" box="[714,825,1428,1451]" gridcol="1" gridrow="17" pageId="4" pageNumber="33">69,67</td>
<td id="76805DB4FFB10048FC94F9D0B488FA4D" box="[859,929,1428,1451]" gridcol="2" gridrow="17" pageId="4" pageNumber="33">59,38</td>
<td id="76805DB4FFB10048FC0AF9D0B333FA4D" box="[965,1050,1428,1451]" gridcol="3" gridrow="17" pageId="4" pageNumber="33">70,55</td>
<td id="76805DB4FFB10048FB8BF9D0B3BAFA4D" box="[1092,1171,1428,1451]" gridcol="4" gridrow="17" pageId="4" pageNumber="33">66,67</td>
<td id="76805DB4FFB10048FB71F9D0B224FA4D" box="[1214,1293,1428,1451]" gridcol="5" gridrow="17" pageId="4" pageNumber="33">4,29</td>
</tr>
<tr id="355134C8FFB10048FEDBF986B224FA3F" box="[276,1293,1474,1497]" gridrow="18" pageId="4" pageNumber="33">
<th id="76805DB4FFB10048FEDBF986B5B3FA3F" box="[276,666,1474,1497]" gridcol="0" gridrow="18" pageId="4" pageNumber="33">17 Interorbital width</th>
<td id="76805DB4FFB10048FD05F986B410FA3F" box="[714,825,1474,1497]" gridcol="1" gridrow="18" pageId="4" pageNumber="33">29,23</td>
<td id="76805DB4FFB10048FC94F986B488FA3F" box="[859,929,1474,1497]" gridcol="2" gridrow="18" pageId="4" pageNumber="33">27,34</td>
<td id="76805DB4FFB10048FC0AF986B333FA3F" box="[965,1050,1474,1497]" gridcol="3" gridrow="18" pageId="4" pageNumber="33">30,29</td>
<td id="76805DB4FFB10048FB8BF986B3BAFA3F" box="[1092,1171,1474,1497]" gridcol="4" gridrow="18" pageId="4" pageNumber="33">28,45</td>
<td id="76805DB4FFB10048FB71F986B224FA3F" box="[1214,1293,1474,1497]" gridcol="5" gridrow="18" pageId="4" pageNumber="33">1,03</td>
</tr>
<tr id="355134C8FFB10048FEDBF9ABB224F9E0" box="[276,1293,1519,1542]" gridrow="19" pageId="4" pageNumber="33">
<th id="76805DB4FFB10048FEDBF9ABB5B3F9E0" box="[276,666,1519,1542]" gridcol="0" gridrow="19" pageId="4" pageNumber="33">18 Orbital diameter</th>
<td id="76805DB4FFB10048FD05F9ABB410F9E0" box="[714,825,1519,1542]" gridcol="1" gridrow="19" pageId="4" pageNumber="33">13,63</td>
<td id="76805DB4FFB10048FC94F9ABB488F9E0" box="[859,929,1519,1542]" gridcol="2" gridrow="19" pageId="4" pageNumber="33">13,63</td>
<td id="76805DB4FFB10048FC0AF9ABB333F9E0" box="[965,1050,1519,1542]" gridcol="3" gridrow="19" pageId="4" pageNumber="33">16,52</td>
<td id="76805DB4FFB10048FB8BF9ABB3BAF9E0" box="[1092,1171,1519,1542]" gridcol="4" gridrow="19" pageId="4" pageNumber="33">15,37</td>
<td id="76805DB4FFB10048FB71F9ABB224F9E0" box="[1214,1293,1519,1542]" gridcol="5" gridrow="19" pageId="4" pageNumber="33">0,94</td>
</tr>
<tr id="355134C8FFB10048FEDBFA59B224F9D2" box="[276,1293,1565,1588]" gridrow="20" pageId="4" pageNumber="33">
<th id="76805DB4FFB10048FEDBFA59B5B3F9D2" box="[276,666,1565,1588]" gridcol="0" gridrow="20" pageId="4" pageNumber="33">19 Longest interopercular odontode</th>
<td id="76805DB4FFB10048FD05FA59B410F9D2" box="[714,825,1565,1588]" gridcol="1" gridrow="20" pageId="4" pageNumber="33">40,88</td>
<td id="76805DB4FFB10048FC94FA59B488F9D2" box="[859,929,1565,1588]" gridcol="2" gridrow="20" pageId="4" pageNumber="33">11,72</td>
<td id="76805DB4FFB10048FC0AFA59B333F9D2" box="[965,1050,1565,1588]" gridcol="3" gridrow="20" pageId="4" pageNumber="33">40,88</td>
<td id="76805DB4FFB10048FB8BFA59B3BAF9D2" box="[1092,1171,1565,1588]" gridcol="4" gridrow="20" pageId="4" pageNumber="33">21,23</td>
<td id="76805DB4FFB10048FB71FA59B224F9D2" box="[1214,1293,1565,1588]" gridcol="5" gridrow="20" pageId="4" pageNumber="33">9,81</td>
</tr>
<tr id="355134C8FFB10048FEDBFA0FB224F984" box="[276,1293,1611,1634]" gridrow="21" pageId="4" pageNumber="33">
<th id="76805DB4FFB10048FEDBFA0FB5B3F984" box="[276,666,1611,1634]" gridcol="0" gridrow="21" pageId="4" pageNumber="33">20 Mandibular ramus length</th>
<td id="76805DB4FFB10048FD05FA0FB410F984" box="[714,825,1611,1634]" gridcol="1" gridrow="21" pageId="4" pageNumber="33">29,89</td>
<td id="76805DB4FFB10048FC94FA0FB488F984" box="[859,929,1611,1634]" gridcol="2" gridrow="21" pageId="4" pageNumber="33">23,44</td>
<td id="76805DB4FFB10048FC0AFA0FB333F984" box="[965,1050,1611,1634]" gridcol="3" gridrow="21" pageId="4" pageNumber="33">30,77</td>
<td id="76805DB4FFB10048FB8BFA0FB3BAF984" box="[1092,1171,1611,1634]" gridcol="4" gridrow="21" pageId="4" pageNumber="33">28,36</td>
<td id="76805DB4FFB10048FB71FA0FB224F984" box="[1214,1293,1611,1634]" gridcol="5" gridrow="21" pageId="4" pageNumber="33">2,51</td>
</tr>
<tr id="355134C8FFB10048FEDBFA3CB224F969" box="[276,1293,1656,1679]" gridrow="22" pageId="4" pageNumber="33">
<th id="76805DB4FFB10048FEDBFA3CB5B3F969" box="[276,666,1656,1679]" gridcol="0" gridrow="22" pageId="4" pageNumber="33">21 IOW/MRL</th>
<td id="76805DB4FFB10048FD05FA3CB410F969" box="[714,825,1656,1679]" gridcol="1" gridrow="22" pageId="4" pageNumber="33">0,98</td>
<td id="76805DB4FFB10048FC94FA3CB488F969" box="[859,929,1656,1679]" gridcol="2" gridrow="22" pageId="4" pageNumber="33">1.17</td>
<td id="76805DB4FFB10048FC0AFA3CB333F969" box="[965,1050,1656,1679]" gridcol="3" gridrow="22" pageId="4" pageNumber="33">0.93</td>
<td id="76805DB4FFB10048FB8BFA3CB3BAF969" box="[1092,1171,1656,1679]" gridcol="4" gridrow="22" pageId="4" pageNumber="33">1.0 1</td>
<td id="76805DB4FFB10048FB71FA3CB224F969" box="[1214,1293,1656,1679]" gridcol="5" gridrow="22" pageId="4" pageNumber="33">0.0 8</td>
</tr>
<tr id="355134C8FFB10048FEDBFAE1B224F95A" box="[276,1293,1701,1724]" gridrow="23" pageId="4" pageNumber="33">
<th id="76805DB4FFB10048FEDBFAE1B5B3F95A" box="[276,666,1701,1724]" gridcol="0" gridrow="23" pageId="4" pageNumber="33">22 HL/IOW</th>
<td id="76805DB4FFB10048FD05FAE1B410F95A" box="[714,825,1701,1724]" gridcol="1" gridrow="23" pageId="4" pageNumber="33">3,42</td>
<td id="76805DB4FFB10048FC94FAE1B488F95A" box="[859,929,1701,1724]" gridcol="2" gridrow="23" pageId="4" pageNumber="33">3.66</td>
<td id="76805DB4FFB10048FC0AFAE1B333F95A" box="[965,1050,1701,1724]" gridcol="3" gridrow="23" pageId="4" pageNumber="33">3.30</td>
<td id="76805DB4FFB10048FB8BFAE1B3BAF95A" box="[1092,1171,1701,1724]" gridcol="4" gridrow="23" pageId="4" pageNumber="33">3.52</td>
<td id="76805DB4FFB10048FB71FAE1B224F95A" box="[1214,1293,1701,1724]" gridcol="5" gridrow="23" pageId="4" pageNumber="33">0.13</td>
</tr>
<tr id="355134C8FFB10048FEDBFA97B224F90C" box="[276,1293,1747,1770]" gridrow="24" pageId="4" pageNumber="33">
<th id="76805DB4FFB10048FEDBFA97B5B3F90C" box="[276,666,1747,1770]" gridcol="0" gridrow="24" pageId="4" pageNumber="33">23 IOW/OD</th>
<td id="76805DB4FFB10048FD05FA97B410F90C" box="[714,825,1747,1770]" gridcol="1" gridrow="24" pageId="4" pageNumber="33">2,15</td>
<td id="76805DB4FFB10048FC94FA97B488F90C" box="[859,929,1747,1770]" gridcol="2" gridrow="24" pageId="4" pageNumber="33">2.15</td>
<td id="76805DB4FFB10048FC0AFA97B333F90C" box="[965,1050,1747,1770]" gridcol="3" gridrow="24" pageId="4" pageNumber="33">1.73</td>
<td id="76805DB4FFB10048FB8BFA97B3BAF90C" box="[1092,1171,1747,1770]" gridcol="4" gridrow="24" pageId="4" pageNumber="33">1.86</td>
<td id="76805DB4FFB10048FB71FA97B224F90C" box="[1214,1293,1747,1770]" gridcol="5" gridrow="24" pageId="4" pageNumber="33">0.17</td>
</tr>
<tr id="355134C8FFB10048FEDBFB44B224F8F1" box="[276,1293,1792,1815]" gridrow="25" pageId="4" pageNumber="33">
<th id="76805DB4FFB10048FEDBFB44B5B3F8F1" box="[276,666,1792,1815]" gridcol="0" gridrow="25" pageId="4" pageNumber="33">24 HL/OD</th>
<td id="76805DB4FFB10048FD05FB44B410F8F1" box="[714,825,1792,1815]" gridcol="1" gridrow="25" pageId="4" pageNumber="33">7,34</td>
<td id="76805DB4FFB10048FC94FB44B488F8F1" box="[859,929,1792,1815]" gridcol="2" gridrow="25" pageId="4" pageNumber="33">7.34</td>
<td id="76805DB4FFB10048FC0AFB44B333F8F1" box="[965,1050,1792,1815]" gridcol="3" gridrow="25" pageId="4" pageNumber="33">6.0 5</td>
<td id="76805DB4FFB10048FB8BFB44B3BAF8F1" box="[1092,1171,1792,1815]" gridcol="4" gridrow="25" pageId="4" pageNumber="33">6.53</td>
<td id="76805DB4FFB10048FB71FB44B224F8F1" box="[1214,1293,1792,1815]" gridcol="5" gridrow="25" pageId="4" pageNumber="33">0.42</td>
</tr>
</table>
</paragraph>
<caption id="DF1E6602FFB3FFB1FEC7F9DEB436FA37" httpUri="https://zenodo.org/record/171648/files/figure.png" pageId="6" pageNumber="35" targetBox="[340,1246,329,1413]" targetPageId="6">
<paragraph id="8BDE368AFFB3FFB1FEC7F9DEB436FA37" blockId="6.[264,1322,1434,1489]" pageId="6" pageNumber="35">
<emphasis id="B915EA98FFB3FFB1FEC7F9DEB6A5FA54" bold="true" box="[264,396,1434,1458]" pageId="6" pageNumber="35">FIGURE 1.</emphasis>
<taxonomicName id="4C614D09FFB3FFB1FE5BF9DEB598FA57" box="[404,689,1434,1457]" class="Actinopterygii" family="Loricariidae" genus="Cordylancistrus" kingdom="Animalia" order="Siluriformes" pageId="6" pageNumber="35" phylum="Chordata" rank="species" species="nephelion">
<emphasis id="B915EA98FFB3FFB1FE5BF9DEB598FA57" box="[404,689,1434,1457]" italics="true" pageId="6" pageNumber="35">Cordylancistrus nephelion</emphasis>
</taxonomicName>
MBUCV­V­21800, 129.1 mm SL, holotype, A: lateral, B: dorsal, and C: ventral views. Photos by N. Milani.
</paragraph>
</caption>
<subSubSection id="C37B6501FFB3FFB1FEF7FA64B37AF984" pageId="6" pageNumber="35" type="etymology">
<paragraph id="8BDE368AFFB3FFB1FEF7FA64B37AF984" blockId="6.[264,1322,1568,1634]" pageId="6" pageNumber="35">
<emphasis id="B915EA98FFB3FFB1FEF7FA64B694F9DC" bold="true" box="[312,445,1568,1594]" pageId="6" pageNumber="35">Etymology</emphasis>
. The name of the species is taken from the Greek word
<taxonomicName id="4C614D09FFB3FFB1FB7FFA64B20DF9DF" box="[1200,1316,1568,1593]" class="Actinopterygii" family="Loricariidae" genus="Cordylancistrus" kingdom="Animalia" order="Siluriformes" pageId="6" pageNumber="35" phylum="Chordata" rank="species" species="nephelion">
<emphasis id="B915EA98FFB3FFB1FB7FFA64B20DF9DF" box="[1200,1316,1568,1593]" italics="true" pageId="6" pageNumber="35">nephelion</emphasis>
</taxonomicName>
, meaning cloudlike spots, in reference to the color pattern of the species.
</paragraph>
</subSubSection>
</treatment>
</document>