295 lines
32 KiB
XML
295 lines
32 KiB
XML
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<mods:title id="D6BEB046F011FE968F749B7D0B9AA534">A cybertaxonomic revision of the new dung beetle tribe Parachoriini (Coleoptera: Scarabaeidae: Scarabaeinae) and its phylogenetic assessment using molecular and morphological data</mods:title>
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<heading id="D09F819BAE44B629DD9E43E4FC9B06B5" bold="true" box="[151,832,1195,1221]" fontSize="11" level="1" pageId="25" pageNumber="126" reason="1">
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<emphasis id="B91CEAE5AE44B629DD9E43E4FC9B06B5" bold="true" box="[151,832,1195,1221]" pageId="25" pageNumber="126">
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<taxonomicName id="4C684D74AE44B629DD9E43E4FDB906B5" ID-CoL="75NM5" authority="(Arrow, 1907)" authorityName="Tarasov" authorityYear="2017" baseAuthorityName="Arrow" baseAuthorityYear="1907" box="[151,610,1195,1221]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="126" phylum="Arthropoda" rank="species" species="humeralis" status="comb. nov.">
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<emphasis id="B91CEAE5AE44B629DD9E43E4FE7506B5" bold="true" box="[151,430,1195,1221]" italics="true" pageId="25" pageNumber="126">Parachorius humeralis</emphasis>
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(
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<bibRefCitation id="EFF94B06AE44B629DCB443E4FD8106B5" author="Arrow" box="[445,602,1195,1221]" pageId="25" pageNumber="126" refString="Arrow, G. J. (1907) Some new species and genera of lamellicorn Coleoptera from the Indian Empire (part 2). The Annals and Magazine of Natural History, including Zoology, Botany and Geology, 7 (19), 416 - 439." type="journal article" year="1907">Arrow, 1907</bibRefCitation>
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)
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</taxonomicName>
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<taxonomicNameLabel id="A22F579EAE44B629DF6343E4FC9B06B5" box="[618,832,1195,1221]" pageId="25" pageNumber="126">new combination</taxonomicNameLabel>
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<paragraph id="8BD736F7AE44B629DD9E4382FE9B0696" blockId="25.[151,832,1195,1254]" box="[151,320,1229,1254]" pageId="25" pageNumber="126">
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(
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<figureCitation id="13532A72AE44B629DD964382FF270696" box="[159,252,1229,1254]" captionStart="FIGURE 13" captionStartId="26.[151,250,1259,1281]" captionTargetBox="[151,1416,193,1211]" captionTargetId="figure@26.[151,1436,193,1237]" captionTargetPageId="26" captionText="FIGURE 13. Morphology of Parachorius humeralis. (A – B) aedeagus (Darjeeling). (C) aedeagus. (D) habitus (male, holotype). (E) habitus (male). (F – H) right metafemur (male). (I) protibia (female). (J) protibia (male)." httpUri="https://zenodo.org/record/1001943/files/figure.png" pageId="25" pageNumber="126">Figs. 13</figureCitation>
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,
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<figureCitation id="13532A72AE44B629DC014382FEE30696" box="[264,312,1229,1254]" captionStart="FIGURE 26" captionStartId="43.[151,250,1905,1927]" captionTargetBox="[151,1436,265,1825]" captionTargetId="figure@43.[151,1436,265,1825]" captionTargetPageId="43" captionText="FIGURE 26. Distribution maps of Parachorius species. (A) Parachorius hookeri. (B) Parachorius humeralis. (C) Parachorius javanus. (D) Parachorius longipenis. (E) Parachorius maruyamai. (F) Parachorius newthayerae. Type locality — type locality of species (holotype or lectotype); Type. Collection data — localities of type specimens associated with species (paratypes or paralectotypes); Collection data — localities of other specimens examined in this study." httpUri="https://zenodo.org/record/1001969/files/figure.png" pageId="25" pageNumber="126">26B</figureCitation>
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)
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</paragraph>
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<subSubSection id="C372657CAE44B629DD9E4259FDD8073A" pageId="25" pageNumber="126" type="reference_group">
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<paragraph id="8BD736F7AE44B629DD9E4259FDCC075C" blockId="25.[151,760,1302,1508]" box="[151,535,1302,1324]" pageId="25" pageNumber="126">
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<taxonomicName id="4C684D74AE44B629DD9E4259FDCC075C" ID-CoL="RNHS" authority="Arrow, 1907: 416" authorityName="Arrow" authorityPageNumber="416" authorityYear="1907" box="[151,535,1302,1324]" class="Insecta" family="Scarabaeidae" genus="Cassolus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="126" phylum="Arthropoda" rank="species" species="humeralis">
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<emphasis id="B91CEAE5AE44B629DD9E4259FE85075C" box="[151,350,1302,1324]" italics="true" pageId="25" pageNumber="126">Cassolus humeralis</emphasis>
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<treatmentCitation id="0AC910E6AE44B629DC6D4259FDCC075C" author="Arrow" box="[356,535,1302,1324]" page="416" pageId="25" pageNumber="126" year="1907">
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: 416
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</taxonomicName>
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</paragraph>
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<paragraph id="8BD736F7AE44B629DD9E427BFDD8073A" blockId="25.[151,760,1302,1508]" box="[151,515,1332,1354]" pageId="25" pageNumber="126">
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<treatmentCitationGroup id="AB7811D9AE44B629DD9E427BFDD8073A" box="[151,515,1332,1354]" pageId="25" pageNumber="126">
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<taxonomicName id="4C684D74AE44B629DD9E427BFE85073A" ID-CoL="RNHS" authorityName="Arrow" authorityYear="1907" box="[151,350,1332,1354]" class="Insecta" family="Scarabaeidae" genus="Cassolus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="126" phylum="Arthropoda" rank="species" species="humeralis">
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<emphasis id="B91CEAE5AE44B629DD9E427BFE85073A" box="[151,350,1332,1354]" italics="true" pageId="25" pageNumber="126">Cassolus humeralis</emphasis>
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</taxonomicName>
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;
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<treatmentCitation id="0AC910E6AE44B629DC62427BFDD8073A" author="Gillet" box="[363,515,1332,1354]" page="39" pageId="25" pageNumber="126" year="1911">
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<bibRefCitation id="EFF94B06AE44B629DC62427BFE3B073A" author="Gillet" box="[363,480,1332,1354]" pageId="25" pageNumber="126" refString="Gillet, J. J. E. (1911) Scarabaeidae: Coprinae I. Coleopterorum Catologus, 38, 1 - 100." type="journal article" year="1911">Gillet 1911</bibRefCitation>
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: 39
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</treatmentCitation>
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</paragraph>
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<paragraph id="8BD736F7AE44B629DD9E421CFDC00719" blockId="25.[151,760,1302,1508]" box="[151,539,1363,1385]" pageId="25" pageNumber="126">
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<treatmentCitationGroup id="AB7811D9AE44B629DD9E421CFDC00719" box="[151,539,1363,1385]" pageId="25" pageNumber="126">
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<emphasis id="B91CEAE5AE44B629DD9E421CFEBD0719" box="[151,358,1363,1385]" italics="true" pageId="25" pageNumber="126">
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<taxonomicName id="4C684D74AE44B629DD9E421CFEBB0719" ID-CoL="RNHS" authorityName="Arrow" authorityYear="1907" box="[151,352,1363,1385]" class="Insecta" family="Scarabaeidae" genus="Cassolus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="126" phylum="Arthropoda" rank="species" species="humeralis">Cassolus humeralis</taxonomicName>
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;
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</emphasis>
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<treatmentCitation id="0AC910E6AE44B629DC65421BFDC00719" author="Arrow" box="[364,539,1363,1385]" page="361" pageId="25" pageNumber="126" year="1931">
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<bibRefCitation id="EFF94B06AE44B629DC65421BFE370719" author="Arrow" box="[364,492,1363,1385]" pageId="25" pageNumber="126" refString="Arrow, G. J. (1931) Coleoptera Lamellicornia part. III. (Coprinae). Taylor and Francis, London, 428 pp., 13 pls." type="book" year="1931">Arrow 1931</bibRefCitation>
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: 361
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</treatmentCitation>
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="C372657CAE44B629DD9E423DFDCD0794" pageId="25" pageNumber="126" type="reference_group">
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<paragraph id="8BD736F7AE44B629DD9E423DFDF407B5" blockId="25.[151,760,1302,1508]" pageId="25" pageNumber="126">
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<treatmentCitationGroup id="AB7811D9AE44B629DD9E423DFDF407B5" pageId="25" pageNumber="126">
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<emphasis id="B91CEAE5AE44B629DD9E423DFEBD07F8" box="[151,358,1394,1416]" italics="true" pageId="25" pageNumber="126">
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<taxonomicName id="4C684D74AE44B629DD9E423DFEBB07F8" ID-CoL="RNHS" authorityName="Arrow" authorityYear="1907" box="[151,352,1394,1416]" class="Insecta" family="Scarabaeidae" genus="Cassolus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="126" phylum="Arthropoda" rank="species" species="humeralis">Cassolus humeralis</taxonomicName>
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||
;
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</emphasis>
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||
<treatmentCitation id="0AC910E6AE44B629DC65423DFDE207F8" author="Balthasar" box="[364,569,1394,1416]" page="262" pageId="25" pageNumber="126" year="1963">
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<bibRefCitation id="EFF94B06AE44B629DC65423DFDD207F8" author="Balthasar" box="[364,521,1394,1416]" pageId="25" pageNumber="126" refString="Balthasar, V. (1963) Monographie der Scarabaeidae und Aphodiidae der palaearktischen und orientalischen Region. Folume 1. Tschechoslowakische Akademie der Wissenschaften, Prague, 391 pp., XXIV pls." type="book" year="1963">Balthasar 1963</bibRefCitation>
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: 262
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</treatmentCitation>
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<emphasis id="B91CEAE5AE44B629DD9E42DFFEBD07D6" box="[151,358,1424,1446]" italics="true" pageId="25" pageNumber="126">
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<taxonomicName id="4C684D74AE44B629DD9E42DFFEBB07D6" ID-CoL="RNHS" authorityName="Arrow" authorityYear="1907" box="[151,352,1424,1446]" class="Insecta" family="Scarabaeidae" genus="Cassolus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="126" phylum="Arthropoda" rank="species" species="humeralis">Cassolus humeralis</taxonomicName>
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;
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</emphasis>
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<treatmentCitation id="0AC910E6AE44B629DC6542DFFD2307D6" author="Kryzhanovsky" box="[364,760,1424,1446]" page="394" pageId="25" pageNumber="126" year="1966">
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<bibRefCitation id="EFF94B06AE44B629DC6542DFFD1307D6" author="Kryzhanovsky" box="[364,712,1424,1446]" pageId="25" pageNumber="126" refString="Kryzhanovsky, O. L. & Medvedev, S. I. (1966) Dung beetles of the subfamily Coprinae in Yunnan province of South-Western China. Part 1. Entomologicheskoe Obozrenie, Revue d'entomologie de l'URSS, 45, 387 - 400." type="journal article" year="1966">Kryzhanovsky & Medvedev 1966</bibRefCitation>
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: 394
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</treatmentCitation>
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<emphasis id="B91CEAE5AE44B629DD9E42E0FEBD07B5" box="[151,358,1455,1477]" italics="true" pageId="25" pageNumber="126">
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<taxonomicName id="4C684D74AE44B629DD9E42E0FEBB07B5" ID-CoL="RNHS" authorityName="Arrow" authorityYear="1907" box="[151,352,1455,1477]" class="Insecta" family="Scarabaeidae" genus="Cassolus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="126" phylum="Arthropoda" rank="species" species="humeralis">Cassolus humeralis</taxonomicName>
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;
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</emphasis>
|
||
<treatmentCitation id="0AC910E6AE44B629DC6542E0FDF407B5" author="Bacchus" box="[364,559,1455,1477]" page="104" pageId="25" pageNumber="126" year="1978">
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<bibRefCitation id="EFF94B06AE44B629DC6542E0FE2407B5" author="Bacchus" box="[364,511,1455,1477]" pageId="25" pageNumber="126" refString="Bacchus, M. E. (1978) A catalogue of the type-specimens of the Scarabaeinae (Scarabaeidae) and the smaller lamellicorn families (Coleoptera) described by G. J. Arrow. Bulletin of the British Museum (Natural History) Entomology Supplement, 37, 97 - 115." type="journal article" year="1978">Bacchus 1978</bibRefCitation>
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: 104
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</treatmentCitation>
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8BD736F7AE44B629DD9E4281FDCD0794" blockId="25.[151,760,1302,1508]" box="[151,534,1486,1508]" pageId="25" pageNumber="126">
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<treatmentCitationGroup id="AB7811D9AE44B629DD9E4281FDCD0794" box="[151,534,1486,1508]" pageId="25" pageNumber="126">
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<taxonomicName id="4C684D74AE44B629DD9E4281FE850794" ID-CoL="RNHS" authorityName="Arrow" authorityYear="1907" box="[151,350,1486,1508]" class="Insecta" family="Scarabaeidae" genus="Cassolus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="126" phylum="Arthropoda" rank="species" species="humeralis">
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<emphasis id="B91CEAE5AE44B629DD9E4281FE850794" box="[151,350,1486,1508]" italics="true" pageId="25" pageNumber="126">Cassolus humeralis</emphasis>
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</taxonomicName>
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;
|
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<treatmentCitation id="0AC910E6AE44B629DC624281FDCD0794" author="Paulian" box="[363,534,1486,1508]" page="57" pageId="25" pageNumber="126" year="1980">
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<bibRefCitation id="EFF94B06AE44B629DC624281FE280794" author="Paulian" box="[363,499,1486,1508]" pageId="25" pageNumber="126" refString="Paulian, R. (1980) Coleopteres Scarabaeidae Canthoninae d'Inde du Sud. Revue Suisse de Zoologie, 87, 57 - 65. https: // doi. org / 10.5962 / bhl. part. 85506" type="journal article" year="1980">Paulian 1980</bibRefCitation>
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: 57
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<paragraph id="8BD736F7AE44B629DD9E415FFCD60459" blockId="25.[151,1436,1552,2009]" box="[151,781,1552,1577]" pageId="25" pageNumber="126">
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<materialsCitation id="3B003CAAAE44B629DD9E415FFCD60459" ID-GBIF-Occurrence="1658411501" box="[151,781,1552,1577]" country="India" location="Assa" pageId="25" pageNumber="126" specimenCount="1" stateProvince="Assam" typeStatus="holotype">
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<emphasis id="B91CEAE5AE44B629DD9E415FFEF60459" bold="true" box="[151,301,1552,1577]" pageId="25" pageNumber="126">
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<typeStatus id="54D38855AE44B629DD9E415FFF090459" box="[151,210,1552,1577]" pageId="25" pageNumber="126">Type</typeStatus>
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locality
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</emphasis>
|
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.
|
||
<collectingRegion id="49ACF815AE44B629DC35415EFE520459" box="[316,393,1553,1577]" country="India" name="Assam" pageId="25" pageNumber="126">Assam</collectingRegion>
|
||
Sudiya [
|
||
<location id="8EB7602CAE44B629DCE3415EFDFD0459" LSID="urn:lsid:plazi:treatment:03C187E1AE44B62BDD9E43E4FC190288:8EB7602CAE44B629DCE3415EFDFD0459" box="[490,550,1553,1577]" country="India" name="Assa" pageId="25" pageNumber="126" stateProvince="Assam">Assa</location>
|
||
, northeastern
|
||
<collectingCountry id="F37F7667AE44B629DFCA415FFCD80459" box="[707,771,1552,1577]" name="India" pageId="25" pageNumber="126">India</collectingCountry>
|
||
].
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C372657CAE44B629DDCE417BFB2D043D" box="[199,1270,1588,1613]" pageId="25" pageNumber="126" type="distribution">
|
||
<paragraph id="8BD736F7AE44B629DDCE417BFB2D043D" blockId="25.[151,1436,1552,2009]" box="[199,1270,1588,1613]" pageId="25" pageNumber="126">
|
||
<emphasis id="B91CEAE5AE44B629DDCE417BFEBB043D" bold="true" box="[199,352,1588,1613]" pageId="25" pageNumber="126">Distribution.</emphasis>
|
||
The species is known only from northeastern
|
||
<collectingCountry id="F37F7667AE44B629DE6F417BFC7B043D" box="[870,928,1588,1613]" name="India" pageId="25" pageNumber="126">India</collectingCountry>
|
||
(
|
||
<collectingRegion id="49ACF815AE44B629DEA7417BFBD8043D" box="[942,1027,1588,1613]" country="India" name="Sikkim" pageId="25" pageNumber="126">Sikkim</collectingRegion>
|
||
,
|
||
<collectingRegion id="49ACF815AE44B629D906417AFB85043D" box="[1039,1118,1589,1613]" country="India" name="Assam" pageId="25" pageNumber="126">Assam</collectingRegion>
|
||
,
|
||
<collectingRegion id="49ACF815AE44B629D960417BFB31043D" box="[1129,1258,1588,1613]" country="India" name="Meghalaya" pageId="25" pageNumber="126">Meghalaya</collectingRegion>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C372657CAE44B629DDCE4117FC120570" pageId="25" pageNumber="126" type="diagnosis">
|
||
<paragraph id="8BD736F7AE44B629DDCE4117FC120570" blockId="25.[151,1436,1552,2009]" pageId="25" pageNumber="126">
|
||
<emphasis id="B91CEAE5AE44B629DDCE4117FE9A0401" bold="true" box="[199,321,1624,1649]" pageId="25" pageNumber="126">Diagnosis.</emphasis>
|
||
This species is most similar to
|
||
<taxonomicName id="4C684D74AE44B629DFAF4116FC870400" authorityName="S. Tarasov" authorityYear="2017" box="[678,860,1625,1648]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="126" phylum="Arthropoda" rank="species" species="asymmetricus">
|
||
<emphasis id="B91CEAE5AE44B629DFAF4116FC870400" box="[678,860,1625,1648]" italics="true" pageId="25" pageNumber="126">P. asymmetricus</emphasis>
|
||
</taxonomicName>
|
||
from which it differs by the symmetrical aedeagus as well as first and second teeth of protibia being distinctly separated from each other. From other
|
||
<taxonomicName id="4C684D74AE44B629D8194133FA4704E5" box="[1296,1436,1660,1685]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="126" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91CEAE5AE44B629D8194133FA4704E5" box="[1296,1436,1660,1685]" italics="true" pageId="25" pageNumber="126">Parachorius</emphasis>
|
||
</taxonomicName>
|
||
species
|
||
<taxonomicName id="4C684D74AE44B629DDE641EEFEA104C9" authorityName="Arrow" authorityYear="1907" box="[239,378,1696,1721]" class="Insecta" family="Scarabaeidae" genus="Cassolus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="126" phylum="Arthropoda" rank="species" species="humeralis">
|
||
<emphasis id="B91CEAE5AE44B629DDE641EEFEA104C9" box="[239,378,1696,1721]" italics="true" pageId="25" pageNumber="126">P. humeralis</emphasis>
|
||
</taxonomicName>
|
||
can be separated by (1) elytron usually bearing yellow humeral spot (rarely not), (2) aedeagus shape, (3) apical half of metafemoral posterior margin from slightly to largely dilated and produced in sharp or rounded angle, and (4) dilation of metafemur steeply reducing near apex.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C372657CAE44B62ADDCE4043FC3E04D4" lastPageId="26" lastPageNumber="127" pageId="25" pageNumber="126" type="description">
|
||
<paragraph id="8BD736F7AE44B629DDCE4043FEB20539" blockId="25.[151,1436,1552,2009]" pageId="25" pageNumber="126">
|
||
<emphasis id="B91CEAE5AE44B629DDCE4043FE820555" bold="true" box="[199,345,1804,1829]" pageId="25" pageNumber="126">Description.</emphasis>
|
||
<emphasis id="B91CEAE5AE44B629DC6D4043FE7F0555" box="[356,420,1804,1829]" italics="true" pageId="25" pageNumber="126">Body.</emphasis>
|
||
Body, black or brown, dark, length 3.8–5.0 mm. Elytra with yellow humeral spot or uniformly colored.
|
||
</paragraph>
|
||
<paragraph id="8BD736F7AE44B629DDCE401BFBAD051D" blockId="25.[151,1436,1552,2009]" box="[199,1142,1876,1901]" pageId="25" pageNumber="126">
|
||
<emphasis id="B91CEAE5AE44B629DDCE401BFED0051D" box="[199,267,1876,1901]" italics="true" pageId="25" pageNumber="126">Head.</emphasis>
|
||
Clypeus with two teeth, outer margin of each tooth distinctly notched basally.
|
||
</paragraph>
|
||
<paragraph id="8BD736F7AE44B629DDCE4036FB7605C5" blockId="25.[151,1436,1552,2009]" pageId="25" pageNumber="126">
|
||
<emphasis id="B91CEAE5AE44B629DDCE4036FEE605E0" box="[199,317,1913,1936]" italics="true" pageId="25" pageNumber="126">Pronotum.</emphasis>
|
||
Pronotum rounded laterally. Frontal angles rounded. Disc covered with punctures separated by 0.5– 2.0 puncture diameter(s). Pronotum in anterior-posterior direction notably shorter than elytra.
|
||
</paragraph>
|
||
<paragraph id="8BD736F7AE44B629DDCE408FFD1B05A8" blockId="25.[151,1436,1552,2009]" box="[199,704,1984,2009]" pageId="25" pageNumber="126">
|
||
<emphasis id="B91CEAE5AE44B629DDCE408FFECE05A9" box="[199,277,1984,2009]" italics="true" pageId="25" pageNumber="126">Elytra.</emphasis>
|
||
Elytra covered with sparse punctures.
|
||
</paragraph>
|
||
<caption id="DF17667FAE47B62ADD9E43A4FB470750" httpUri="https://zenodo.org/record/1001943/files/figure.png" pageId="26" pageNumber="127" targetBox="[151,1416,193,1211]" targetPageId="26">
|
||
<paragraph id="8BD736F7AE47B62ADD9E43A4FB470750" blockId="26.[151,1436,1259,1312]" pageId="26" pageNumber="127">
|
||
<emphasis id="B91CEAE5AE47B62ADD9E43A4FEFC0770" bold="true" box="[151,295,1259,1281]" pageId="26" pageNumber="127">FIGURE 13.</emphasis>
|
||
Morphology of
|
||
<taxonomicName id="4C684D74AE47B62ADCED43A4FD0D0771" authorityName="Tarasov" authorityYear="2017" baseAuthorityName="Arrow" baseAuthorityYear="1907" box="[484,726,1259,1281]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="127" phylum="Arthropoda" rank="species" species="humeralis">
|
||
<emphasis id="B91CEAE5AE47B62ADCED43A4FD0D0771" box="[484,726,1259,1281]" italics="true" pageId="26" pageNumber="127">Parachorius humeralis</emphasis>
|
||
</taxonomicName>
|
||
. (A–B) aedeagus (Darjeeling). (C) aedeagus. (D) habitus (male, holotype). (E) habitus (male). (F–H) right metafemur (male). (I) protibia (female). (J) protibia (male).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BD736F7AE47B62ADDCE4207FD940710" blockId="26.[151,1437,1351,2024]" box="[199,591,1351,1376]" pageId="26" pageNumber="127">
|
||
<emphasis id="B91CEAE5AE47B62ADDCE4207FEC9072F" box="[199,274,1352,1375]" italics="true" pageId="26" pageNumber="127">Wings.</emphasis>
|
||
Wings normally developed.
|
||
</paragraph>
|
||
<paragraph id="8BD736F7AE47B62ADDCE4223FDD2042C" blockId="26.[151,1437,1351,2024]" pageId="26" pageNumber="127">
|
||
<emphasis id="B91CEAE5AE47B62ADDCE4223FED807F3" box="[199,259,1388,1411]" italics="true" pageId="26" pageNumber="127">Legs.</emphasis>
|
||
Profemur with almost straight or sinuate anterior margin. Protibia with apical spur approximately reaching middle of protarsomere 3. First tooth notably modified in males, approximately twice as wide as second tooth; first and second teeth distinctly separated from each other; first tooth distinctly separated from denticles located between first and second teeth. Metafemoral posterior margin not serrate, largely dilated in apical half of metafemur and produced in sharp angle or slightly dilated in apical forth of metafemur and produced in rounded angle preapically; dilation gradually reducing preapically. Metatibia slightly sinuate, inner margin not denticulate, produced in small angle apically.
|
||
</paragraph>
|
||
<paragraph id="8BD736F7AE47B62ADDCE4127FBBD04F0" blockId="26.[151,1437,1351,2024]" box="[199,1126,1639,1664]" pageId="26" pageNumber="127">
|
||
<emphasis id="B91CEAE5AE47B62ADDCE4127FEB9040F" box="[199,354,1640,1663]" italics="true" pageId="26" pageNumber="127">Metasternum.</emphasis>
|
||
Transverse basal ridge of metasternum reduced and extremely short.
|
||
</paragraph>
|
||
<paragraph id="8BD736F7AE47B62ADDCE41C4FC3E04D4" blockId="26.[151,1437,1351,2024]" box="[199,997,1675,1700]" pageId="26" pageNumber="127">
|
||
<emphasis id="B91CEAE5AE47B62ADDCE41C4FEE004D4" box="[199,315,1675,1700]" italics="true" pageId="26" pageNumber="127">Aedeagus.</emphasis>
|
||
Parameres symmetrical, with horn-like upper lobes dorsally.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C372657CAE47B62BDDCE41E0FC190288" lastPageId="27" lastPageNumber="128" pageId="26" pageNumber="127" type="materials_examined">
|
||
<paragraph id="8BD736F7AE47B62BDDCE41E0FC190288" blockId="26.[151,1437,1351,2024]" lastBlockId="27.[151,1436,151,248]" lastPageId="27" lastPageNumber="128" pageId="26" pageNumber="127">
|
||
<emphasis id="B91CEAE5AE47B62ADDCE41E0FDE904B8" bold="true" box="[199,562,1711,1736]" pageId="26" pageNumber="127">Material examined. Lectotype</emphasis>
|
||
:
|
||
<emphasis id="B91CEAE5AE47B62ADF4A41E0FD5804B8" bold="true" box="[579,643,1711,1736]" pageId="26" pageNumber="127">India</emphasis>
|
||
:
|
||
<emphasis id="B91CEAE5AE47B62ADF9D41E0FD3F04B8" bold="true" box="[660,740,1711,1736]" pageId="26" pageNumber="127">1 male</emphasis>
|
||
, "? / LECTOTYPE / Doherty / Assam Sudiya / 60468 / Fry Coll. 1905-100 /
|
||
<taxonomicName id="4C684D74AE47B62ADC5F419CFDF5049C" authorityName="Arrow" authorityYear="1907" box="[342,558,1747,1772]" class="Insecta" family="Scarabaeidae" genus="Cassolus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="127" phylum="Arthropoda" rank="species" species="humeralis">Cassolus humeralis</taxonomicName>
|
||
/ Arrow type? A. a. m., 1907 /
|
||
<taxonomicName id="4C684D74AE47B62ADE9B419CFB60049B" authority="Arrow" authorityName="Arrow" authorityYear="1907" box="[914,1211,1747,1772]" class="Insecta" family="Scarabaeidae" genus="Cassolus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="127" phylum="Arthropoda" rank="species" species="humeralis">Cassolus humeralis Arrow</taxonomicName>
|
||
? M.E. Bacchus det 1975. LECTOTYPE", 27°50'N, 95°45'E (BMNH).
|
||
<emphasis id="B91CEAE5AE47B62ADFF541B8FC770560" bold="true" box="[764,940,1783,1808]" pageId="26" pageNumber="127">Paralectotypes</emphasis>
|
||
:
|
||
<emphasis id="B91CEAE5AE47B62ADECB41B8FBC30560" bold="true" box="[962,1048,1783,1808]" pageId="26" pageNumber="127">1 male</emphasis>
|
||
,
|
||
<emphasis id="B91CEAE5AE47B62AD92441B8FB420560" bold="true" box="[1069,1177,1783,1808]" pageId="26" pageNumber="127">1 female</emphasis>
|
||
, Assam, Patkai Hills (BMNH).
|
||
<emphasis id="B91CEAE5AE47B62ADC1C4054FEB60544" bold="true" box="[277,365,1819,1844]" pageId="26" pageNumber="127">1 male</emphasis>
|
||
, Darjiling, Gopaldhara, Runbong Valley, Sikkim, British Museum 1922-307, 27°2.1'N, 88°15.8'E, on
|
||
<date id="FFD61037AE47B62ADC314070FE4E0528" box="[312,405,1855,1880]" bridgedPair="/" pageId="26" pageNumber="127" value="1920-07">vii/1920</date>
|
||
(H. Stevens) (BMNH).
|
||
<emphasis id="B91CEAE5AE47B62ADFAF4070FD2E0528" bold="true" box="[678,757,1855,1880]" pageId="26" pageNumber="127">1 male</emphasis>
|
||
, Gopaldhara, Runbong Valley, Sikkim, 1916-218, 27°2.1'N, 88°15.8'E (H. Stevens) (ISNB).
|
||
<emphasis id="B91CEAE5AE47B62ADF0B402CFD8A050C" bold="true" box="[514,593,1891,1916]" pageId="26" pageNumber="127">1 male</emphasis>
|
||
, Gopaldhara, Runbong Valley, Sikkim, 1916-218, 27°2.1'N, 88°15.8'E (H. Stevens) (BMNH).
|
||
<emphasis id="B91CEAE5AE47B62ADC7D40C8FE1805D0" bold="true" box="[372,451,1927,1952]" pageId="26" pageNumber="127">1 male</emphasis>
|
||
, Gopaldhara, Runbong Valley, Sikkim, British Museum 1922-307, 27°2.1'N, 88°15.8'E (H. Stevens) (BMNH).
|
||
<emphasis id="B91CEAE5AE47B62ADCAA40E4FE2F05B4" bold="true" box="[419,500,1963,1988]" pageId="26" pageNumber="127">1 male</emphasis>
|
||
, Assam, Bhalukpong, 27°2'N, 92°35'E,
|
||
<quantity id="4C909B12AE47B62ADEB640E4FBD205B3" box="[959,1033,1963,1988]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="26" pageNumber="127" unit="m" value="150.0">150 m</quantity>
|
||
, on
|
||
<date id="FFD61037AE47B62AD93240E4FB3C05B4" box="[1083,1255,1963,1988]" pageId="26" pageNumber="127" value="2009-05-26" valueMax="2009-06-03" valueMin="2009-05-26">26/v–3/vi/2009</date>
|
||
(P. Pocholatko) (BMNH).
|
||
<emphasis id="B91CEAE5AE47B62ADC074080FEBB0598" bold="true" box="[270,352,1999,2024]" pageId="26" pageNumber="127">1 male</emphasis>
|
||
, Darjeeling, Mungpo, B.M. 1931-452, 26°58.3'N, 88°22.3'E (M. Cameron) (BMNH).
|
||
<emphasis id="B91CEAE5AE47B62AD84D4080FA4E0598" bold="true" box="[1348,1429,1999,2024]" pageId="26" pageNumber="127">1 male</emphasis>
|
||
, Meghalaya, SW of Cherrapunjee, 25°13.5'N, 91°40'E,
|
||
<quantity id="4C909B12AE46B62BDE0F47D7FC8A02DF" box="[774,849,152,176]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="27" pageNumber="128" unit="m" value="900.0">900 m</quantity>
|
||
, on
|
||
<date id="FFD61037AE46B62BDE8A47D8FBCB02C0" box="[899,1040,151,176]" pageId="27" pageNumber="128" value="2005-05-05" valueMax="2005-05-24" valueMin="2005-05-05">5–24/v/2005</date>
|
||
(P. Pocholatko) (BMNH).
|
||
<emphasis id="B91CEAE5AE46B62BD84D47D8FA4F02C0" bold="true" box="[1348,1428,151,176]" pageId="27" pageNumber="128">1 male</emphasis>
|
||
, Shilong, 25°33.9'N, 91°53' E,
|
||
<date id="FFD61037AE46B62BDCE547F3FD8102A5" box="[492,602,188,213]" pageId="27" pageNumber="128" value="1918-04-06">6/iv/1918</date>
|
||
, (BMNH).
|
||
<emphasis id="B91CEAE5AE46B62BDFD247F3FCF202A5" bold="true" box="[731,809,188,213]" pageId="27" pageNumber="128">1 male</emphasis>
|
||
, W Bengal, Darjeeling, North Point, No 890, 27°2.1'N, 88°15.8'E,
|
||
<quantity id="4C909B12AE46B62BDC1A4790FEB10287" box="[275,362,223,248]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="27" pageNumber="128" unit="m" value="1500.0">1500 m</quantity>
|
||
, from traps with carcass,
|
||
<date id="FFD61037AE46B62BDF814790FD210288" box="[648,762,223,248]" pageId="27" pageNumber="128" value="1967-10-23">23/x/1967</date>
|
||
(G. Topal) (ZIN).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |