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<document id="22A9B0F11C9867C247E2BB7AFAF8B0C1" ID-DOI="10.1649/0010-065X-71.1.159" ID-ISSN="1938-4394" ID-Zenodo-Dep="5362679" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630237506140" checkinUser="diego" docAuthor="Reese, Ellen M. &amp; Swanson, Alexander P." docDate="2017" docId="03C187891E72FF8CF6857C18790CF9C0" docLanguage="en" docName="0010-065X-71.1.159.pdf" docOrigin="The Coleopterists Bulletin 71 (1)" docSource="http://dx.doi.org/10.1649/0010-065x-71.1.159" docStyle="DocumentStyle:F824235CA89CB83023C7D5789122F870.3:TheColeopBull.2017-.journal_article.1cover" docStyleId="F824235CA89CB83023C7D5789122F870" docStyleName="TheColeopBull.2017-.journal_article.1cover" docStyleVersion="3" docTitle="Carcinops wenzeli Reese and Swanson 2017, new species" docType="treatment" docVersion="4" lastPageNumber="184" masterDocId="FFF8FFF11E6AFF96F6437876785DFFF2" masterDocTitle="A Review of the CactophilicCarcinopsMarseul (Coleoptera: Histeridae) of the Sonoran Desert Region, with Descriptions of Six New Species" masterLastPageNumber="190" masterPageNumber="159" pageNumber="182" updateTime="1699194676218" updateUser="plazi">
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<mods:title id="8EBFCCF73A7A54E967599665735D8925">A Review of the CactophilicCarcinopsMarseul (Coleoptera: Histeridae) of the Sonoran Desert Region, with Descriptions of Six New Species</mods:title>
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<mods:namePart id="57F2665E6A999E0F9631D073CCAA7129">Reese, Ellen M.</mods:namePart>
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<mods:namePart id="20C591DFE239D48EE37A9717DEF2652C">Swanson, Alexander P.</mods:namePart>
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<mods:title id="6B7B439192C1ECAD2666D87C54356F3A">The Coleopterists Bulletin</mods:title>
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<mods:date id="DC1453A22874F235B569AE269F81F2A4">2017</mods:date>
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<mods:identifier id="70EDE4258E2E1F56902721B240770793" type="ISSN">1938-4394</mods:identifier>
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<treatment id="03C187891E72FF8CF6857C18790CF9C0" LSID="urn:lsid:plazi:treatment:03C187891E72FF8CF6857C18790CF9C0" httpUri="http://treatment.plazi.org/id/03C187891E72FF8CF6857C18790CF9C0" lastPageId="26" lastPageNumber="184" pageId="24" pageNumber="182">
<subSubSection id="C37265141E72FF8EF6857C187998FB6C" pageId="24" pageNumber="182" type="nomenclature">
<paragraph id="8BD7369F1E72FF8EF6857C187998FB6C" blockId="24.[198,592,1134,1209]" pageId="24" pageNumber="182">
<heading id="D09F81F31E72FF8EF6857C187998FB6C" centered="true" fontSize="9" level="2" pageId="24" pageNumber="182" reason="2">
<emphasis id="B91CEA8D1E72FF8EF6857C187998FB6C" bold="true" pageId="24" pageNumber="182">
<taxonomicName id="4C684D1C1E72FF8EF6857C187A17FB76" ID-CoL="9L68J" authority="Reese and Swanson" authorityName="Reese and Swanson" authorityYear="2017" box="[198,586,1134,1156]" class="Insecta" family="Histeridae" genus="Carcinops" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="182" phylum="Arthropoda" rank="species" species="wenzeli" status="sp. nov.">
<emphasis id="B91CEA8D1E72FF8EF6857C187924FB71" bold="true" box="[198,377,1134,1155]" italics="true" pageId="24" pageNumber="182">Carcinops wenzeli</emphasis>
Reese and Swanson
</taxonomicName>
,
<taxonomicNameLabel id="A22F57F61E72FF8EF7137CFF7998FB6C" box="[336,453,1161,1182]" pageId="24" pageNumber="182" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C37265141E72FF8EF7697CD279B7FB4B" box="[298,490,1188,1209]" pageId="24" pageNumber="182" type="description">
<paragraph id="8BD7369F1E72FF8EF7697CD279B7FB4B" blockId="24.[198,592,1134,1209]" box="[298,490,1188,1209]" pageId="24" pageNumber="182">
(
<figureCitation id="13532A1A1E72FF8EF7727CD279D9FB4B" box="[305,388,1188,1209]" captionStart="Fig" captionStartId="4.[175,206,1662,1681]" captionTargetBox="[151,1171,1072,1640]" captionTargetId="figure-19@4.[151,1171,1072,1640]" captionTargetPageId="4" captionText="Fig. 2. Variation of the lateral disc of the first abdominal ventrite in Carcinops species. A) Single stria present (C. torquatus), B) Single stria with microsculptures present (C. rugulus), C) Bistriate (C. consors), D) Bistriate (C. yaqui), E) Tristriate (C. wenzeli). Photographs by A. Swanson." figureDoi="http://doi.org/10.5281/zenodo.5362683" httpUri="https://zenodo.org/record/5362683/files/figure.png" pageId="24" pageNumber="182">Figs. 2E</figureCitation>
,
<figureCitation id="13532A1A1E72FF8EF7D37CD279C3FB4B" box="[400,414,1188,1209]" captionStart="Figs" captionStartId="5.[197,236,962,981]" captionTargetBox="[173,1193,183,938]" captionTargetId="figure-84@5.[173,1193,183,938]" captionTargetPageId="5" captionText="Figs. 36. Head morphology of Carcinops kumeyaay, with arrows showing key character states: 3) C. kumeyaay; 4) C. corticalis; 5) C. curtus. Photographs by E. Reese. 6) Ground punctation patterns on the pronotum of species in the subgenus Carcinopsida (Carcinops wenzeli). Photograph by A. Swanson." figureDoi="http://doi.org/10.5281/zenodo.5362685" httpUri="https://zenodo.org/record/5362685/files/figure.png" pageId="24" pageNumber="182">6</figureCitation>
,
<figureCitation id="13532A1A1E72FF8EF7EA7CD2799DFB4B" box="[425,448,1188,1209]" captionStart="Fig" captionStartId="6.[175,206,1591,1610]" captionTargetBox="[162,620,718,1549]" captionTargetId="figure-558@6.[151,637,712,1567]" captionTargetPageId="6" captionText="Fig. 9. Aedeagal morphology of Carcinops species, with parameres toward the top of the page and basal pieces toward the bottom. A) C. consors, B) C. gilensis, C) C. corticalis, D) C. papagoanus, E) C. opuntiae, F) C. curtus, G) C. rugulus, H) C. torquatus, I) C. yaqui, J) C. wenzeli. Photographs by A. Swanson." figureDoi="http://doi.org/10.5281/zenodo.5362689" httpUri="https://zenodo.org/record/5362689/files/figure.png" pageId="24" pageNumber="182">9J</figureCitation>
,
<figureCitation id="13532A1A1E72FF8EF7887CD279BEFB4B" box="[459,483,1188,1209]" captionStart="Fig" captionStartId="25.[197,228,962,981]" captionTargetBox="[195,1151,206,925]" captionTargetId="figure-361@25.[173,1193,183,938]" captionTargetPageId="25" captionText="Fig. 21. Carcinops wenzeli morphology. A) Dorsal view, B) Ventral view. Photographs by A. Swanson." figureDoi="http://doi.org/10.5281/zenodo.5362717" httpUri="https://zenodo.org/record/5362717/files/figure.png" pageId="24" pageNumber="182">21</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C37265141E72FF8EF6EC7CB8798FFA71" pageId="24" pageNumber="182" type="materials_examined">
<paragraph id="8BD7369F1E72FF8EF6EC7CB8798FFA71" blockId="24.[151,638,1230,1729]" pageId="24" pageNumber="182">
<emphasis id="B91CEA8D1E72FF8EF6EC7CB87971FB11" bold="true" box="[175,300,1230,1251]" pageId="24" pageNumber="182">Type Series.</emphasis>
<materialsCitation id="3B003CC21E72FF8EF77B7CB879D4FABC" collectingDate="2006-11-11" collectionCode="FMNH" collectorName="A. Swanson" country="United States of America" county="Santa Cruz Co." elevation="1400" latitude="31.40946" location="Santa Cruz Co." longLatPrecision="1" longitude="-111.15423" pageId="24" pageNumber="182" specimenCode="APS173" specimenCount="1" typeStatus="holotype">
<emphasis id="B91CEA8D1E72FF8EF77B7CB87998FB11" bold="true" box="[312,453,1230,1251]" pageId="24" pageNumber="182">
<typeStatus id="54D3883D1E72FF8EF77B7CB8799DFB11" box="[312,448,1230,1251]" pageId="24" pageNumber="182" type="holotype">HOLOTYPE</typeStatus>
.
</emphasis>
Male. “
<collectingCountry id="F37F760F1E72FF8EF45B7CB87A16FB11" box="[536,587,1230,1251]" name="United States of America" pageId="24" pageNumber="182">USA</collectingCountry>
: AZ:
<collectingCounty id="62B64E131E72FF8EF6D47C9F7960FB0C" box="[151,317,1257,1278]" pageId="24" pageNumber="182">Santa Cruz Co.</collectingCounty>
/ Sycamore Cyn.
<geoCoordinate id="EE5C50581E72FF8EF4407C9F7A2BFB0C" box="[515,630,1257,1278]" degrees="31.40946" direction="north" orientation="latitude" pageId="24" pageNumber="182" precision="1" value="31.40946">31.40946°N</geoCoordinate>
/
<geoCoordinate id="EE5C50581E72FF8EF6D47D757942FAEA" box="[151,287,1283,1304]" degrees="111.15423" direction="west" orientation="longitude" pageId="24" pageNumber="182" precision="1" value="-111.15423">111.15423°W</geoCoordinate>
,
<quantity id="4C909B7A1E72FF8EF76D7D75792EFAEA" box="[302,371,1283,1304]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="24" pageNumber="182" unit="m" value="1400.0">
<elevation id="0045D1AC1E72FF8EF76D7D75792EFAEA" box="[302,371,1283,1304]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="24" pageNumber="182" unit="m" value="1400.0">1400m</elevation>
</quantity>
,
<date id="FFD6105F1E72FF8EF7C17D7579B1FAEA" box="[386,492,1283,1304]" pageId="24" pageNumber="182" value="2006-11-11">
<collectingDate id="EF92E9B71E72FF8EF7C17D7579B1FAEA" box="[386,492,1283,1304]" pageId="24" pageNumber="182" value="2006-11-11">11.XI.2006</collectingDate>
</date>
/ ex. decaying
<taxonomicName id="4C684D1C1E72FF8EF6D47D6B7913FAC1" box="[151,334,1309,1331]" class="Liliopsida" family="Asparagaceae" genus="Dasylirion" kingdom="Plantae" order="Asparagales" pageId="24" pageNumber="182" phylum="Tracheophyta" rank="species" species="wheeleri">
<emphasis id="B91CEA8D1E72FF8EF6D47D6B7913FAC1" box="[151,334,1309,1331]" italics="true" pageId="24" pageNumber="182">Dasylirion wheeleri</emphasis>
</taxonomicName>
/
<collectorName id="269D53491E72FF8EF7187D687993FAC1" box="[347,462,1310,1331]" pageId="24" pageNumber="182">A. Swanson</collectorName>
,
<specimenCode id="DBCE9EE41E72FF8EF79A7D687A7BFAC1" box="[473,550,1310,1331]" collectionCode="APS" pageId="24" pageNumber="182">APS173</specimenCode>
(
<collectionCode id="ED79AE5A1E72FF8EF4707D687A27FAC1" box="[563,634,1310,1331]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="24" pageNumber="182" type="Museum">FMNH</collectionCode>
; DNA voucher:
<specimenCode id="DBCE9EE41E72FF8EF7727D4F79DFFABC" box="[305,386,1337,1358]" collectionCode="APS" pageId="24" pageNumber="182">APS173</specimenCode>
)
</materialsCitation>
.
<materialsCitation id="3B003CC21E72FF8EF7D47D4F7993FA71" collectingDate="2006-11-11" collectionCode="APS, MCZ, CASC, UNAM, USNM" collectorName="A. Swanson" country="United States of America" county="Santa Cruz Co." elevation="1400" latitude="31.40946" location="Santa Cruz Co." longLatPrecision="1" longitude="-111.15423" pageId="24" pageNumber="182" specimenCount="5" typeStatus="paratype">
<emphasis id="B91CEA8D1E72FF8EF7D47D4F7A7BFABC" bold="true" box="[407,550,1337,1358]" pageId="24" pageNumber="182">
<typeStatus id="54D3883D1E72FF8EF7D47D4F7A7FFABC" box="[407,546,1337,1358]" pageId="24" pageNumber="182" type="paratype">PARATYPES</typeStatus>
.
</emphasis>
<specimenCount id="9D6EFD161E72FF8EF4737D4F7891FA9A" pageId="24" pageNumber="182" type="generic">5 specimens</specimenCount>
. Same data as holotype (
<collectionCode id="ED79AE5A1E72FF8EF78E7D2579A6FA9A" box="[461,507,1363,1384]" pageId="24" pageNumber="182">APS</collectionCode>
, 1;
<collectionCode id="ED79AE5A1E72FF8EF4647D257A01FA9A" box="[551,604,1363,1384]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="24" pageNumber="182" type="Museum">MCZ</collectionCode>
, 1;
<collectionCode id="ED79AE5A1E72FF8EF6DB7D187885FA71" box="[152,216,1390,1411]" pageId="24" pageNumber="182">CASC</collectionCode>
, 1;
<collectionCode id="ED79AE5A1E72FF8EF6B87D187918FA71" box="[251,325,1390,1411]" country="Mexico" httpUri="http://grbio.org/cool/10ei-x9nj" name="Universidad Nacional Autonoma de Mexico" pageId="24" pageNumber="182">UNAM</collectionCode>
, 1;
<collectionCode id="ED79AE5A1E72FF8EF72A7D1879F2FA71" box="[361,431,1390,1411]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="24" pageNumber="182" type="Museum">USNM</collectionCode>
, 1)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C37265141E72FF8EF6EC7DFE7C51FF00" pageId="24" pageNumber="182" type="diagnosis">
<paragraph id="8BD7369F1E72FF8EF6EC7DFE7A20F933" blockId="24.[151,638,1230,1729]" pageId="24" pageNumber="182">
<emphasis id="B91CEA8D1E72FF8EF6EC7DFE79AAFA6F" bold="true" box="[175,503,1416,1437]" pageId="24" pageNumber="182">Diagnosis. Ground punctures:</emphasis>
Micropunctures clustered in groups of 3 or 4 (
<figureCitation id="13532A1A1E72FF8EF7B27DD57A71FA4A" box="[497,556,1443,1464]" captionStart="Figs" captionStartId="5.[197,236,962,981]" captionTargetBox="[173,1193,183,938]" captionTargetId="figure-84@5.[173,1193,183,938]" captionTargetPageId="5" captionText="Figs. 36. Head morphology of Carcinops kumeyaay, with arrows showing key character states: 3) C. kumeyaay; 4) C. corticalis; 5) C. curtus. Photographs by E. Reese. 6) Ground punctation patterns on the pronotum of species in the subgenus Carcinopsida (Carcinops wenzeli). Photograph by A. Swanson." figureDoi="http://doi.org/10.5281/zenodo.5362685" httpUri="https://zenodo.org/record/5362685/files/figure.png" pageId="24" pageNumber="182">Fig. 6</figureCitation>
).
<emphasis id="B91CEA8D1E72FF8EF4037DD57A20FA4A" bold="true" box="[576,637,1443,1464]" pageId="24" pageNumber="182">Head:</emphasis>
Frontal stria continuous with marginal epistomal stria (
<figureCitation id="13532A1A1E72FF8EF68D7DAE796BFA1F" box="[206,310,1496,1517]" captionStart="Figs" captionStartId="5.[197,236,962,981]" captionTargetBox="[173,1193,183,938]" captionTargetId="figure-84@5.[173,1193,183,938]" captionTargetPageId="5" captionText="Figs. 36. Head morphology of Carcinops kumeyaay, with arrows showing key character states: 3) C. kumeyaay; 4) C. corticalis; 5) C. curtus. Photographs by E. Reese. 6) Ground punctation patterns on the pronotum of species in the subgenus Carcinopsida (Carcinops wenzeli). Photograph by A. Swanson." figureDoi="http://doi.org/10.5281/zenodo.5362685" httpUri="https://zenodo.org/record/5362685/files/figure.png" pageId="24" pageNumber="182">Fig. 4B, C</figureCitation>
).
<emphasis id="B91CEA8D1E72FF8EF70A7DAE79EAFA1F" bold="true" box="[329,439,1496,1517]" pageId="24" pageNumber="182">Pronotum:</emphasis>
Marginal stria moderately divergent from anterior edge along midline (
<figureCitation id="13532A1A1E72FF8EF6DD7E7B78B3F9D0" box="[158,238,1549,1570]" captionStart="Figs" captionStartId="5.[197,236,962,981]" captionTargetBox="[173,1193,183,938]" captionTargetId="figure-84@5.[173,1193,183,938]" captionTargetPageId="5" captionText="Figs. 36. Head morphology of Carcinops kumeyaay, with arrows showing key character states: 3) C. kumeyaay; 4) C. corticalis; 5) C. curtus. Photographs by E. Reese. 6) Ground punctation patterns on the pronotum of species in the subgenus Carcinopsida (Carcinops wenzeli). Photograph by A. Swanson." figureDoi="http://doi.org/10.5281/zenodo.5362685" httpUri="https://zenodo.org/record/5362685/files/figure.png" pageId="24" pageNumber="182">Fig. 4A</figureCitation>
).
<emphasis id="B91CEA8D1E72FF8EF7427E7B7914F9D0" bold="true" box="[257,329,1549,1570]" pageId="24" pageNumber="182">Elytra:</emphasis>
Dorsal stria 5 and sutural stria subequal in length to dorsal striae 14, fading into a row of punctures basally but reaching or nearly reaching proximal base of elytra (
<figureCitation id="13532A1A1E72FF8EF7A77E2B7A7CF980" box="[484,545,1629,1650]" captionStart="Figs" captionStartId="5.[197,236,1686,1705]" captionTargetBox="[189,1150,1335,1652]" captionTargetId="figure-138@5.[173,1193,1320,1663]" captionTargetPageId="5" captionText="Figs. 78. 7) Long elytral striae highlighted on Carcinops kumeyaay; 8) Short fifth and sutural elytral striae highlighted on Carcinops yaqui. Photographs by E. Reese." figureDoi="http://doi.org/10.5281/zenodo.5362687" httpUri="https://zenodo.org/record/5362687/files/figure.png" pageId="24" pageNumber="182">Fig. 7</figureCitation>
); internal subhumeral striae present and strongly impressed; external subhumeral stria present as strong punctures.
<emphasis id="B91CEA8D1E72FF8EF69E7EDA79AFF933" bold="true" box="[221,498,1708,1729]" pageId="24" pageNumber="182">First abdominal ventrite:</emphasis>
Lateral disc
</paragraph>
<paragraph id="8BD7369F1E72FF8EF4EE78B47C51FF00" blockId="24.[685,1172,194,1729]" pageId="24" pageNumber="182">
tristriate (
<figureCitation id="13532A1A1E72FF8EF54F78B47B07FF25" box="[780,858,194,215]" captionStart="Fig" captionStartId="4.[175,206,1662,1681]" captionTargetBox="[151,1171,1072,1640]" captionTargetId="figure-19@4.[151,1171,1072,1640]" captionTargetPageId="4" captionText="Fig. 2. Variation of the lateral disc of the first abdominal ventrite in Carcinops species. A) Single stria present (C. torquatus), B) Single stria with microsculptures present (C. rugulus), C) Bistriate (C. consors), D) Bistriate (C. yaqui), E) Tristriate (C. wenzeli). Photographs by A. Swanson." figureDoi="http://doi.org/10.5281/zenodo.5362683" httpUri="https://zenodo.org/record/5362683/files/figure.png" pageId="24" pageNumber="182">Fig. 2E</figureCitation>
).
<emphasis id="B91CEA8D1E72FF8EF52D78B47B8AFF25" bold="true" box="[878,983,194,215]" pageId="24" pageNumber="182">Aedeagus:</emphasis>
Parameres straight and sharply pointed at tips (
<figureCitation id="13532A1A1E72FF8EF5F978AB7C5DFF00" box="[954,1024,221,242]" captionStart="Fig" captionStartId="6.[175,206,1591,1610]" captionTargetBox="[162,620,718,1549]" captionTargetId="figure-558@6.[151,637,712,1567]" captionTargetPageId="6" captionText="Fig. 9. Aedeagal morphology of Carcinops species, with parameres toward the top of the page and basal pieces toward the bottom. A) C. consors, B) C. gilensis, C) C. corticalis, D) C. papagoanus, E) C. opuntiae, F) C. curtus, G) C. rugulus, H) C. torquatus, I) C. yaqui, J) C. wenzeli. Photographs by A. Swanson." figureDoi="http://doi.org/10.5281/zenodo.5362689" httpUri="https://zenodo.org/record/5362689/files/figure.png" pageId="24" pageNumber="182">Fig. 9J</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C37265141E72FF8CF4867881790CF9C0" lastPageId="26" lastPageNumber="184" pageId="24" pageNumber="182" type="description">
<paragraph id="8BD7369F1E72FF8EF48678817B1EFE13" blockId="24.[685,1172,194,1729]" pageId="24" pageNumber="182">
<emphasis id="B91CEA8D1E72FF8EF48678817B1DFEFE" bold="true" box="[709,832,247,268]" pageId="24" pageNumber="182">Description.</emphasis>
L:
<quantity id="4C909B7A1E72FF8EF52878817B9AFEFE" box="[875,967,247,268]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="24" pageNumber="182" unit="mm" value="2.1">2.10 mm</quantity>
; W:
<quantity id="4C909B7A1E72FF8EF5BF78817C05FEFE" box="[1020,1112,247,268]" metricMagnitude="-3" metricUnit="m" metricValue="1.56" pageId="24" pageNumber="182" unit="mm" value="1.56">1.56 mm</quantity>
; E/Pn L: 1.85; E/Pn W: 1.12; Pn W/L: 1.90; E L/W: 0.87; Sterna: 0.64, 0.15,
<quantity id="4C909B7A1E72FF8EF5F6795A7C48FEB3" box="[949,1045,300,321]" metricMagnitude="-4" metricUnit="m" metricValue="6.1" pageId="24" pageNumber="182" unit="mm" value="0.61">0.61 mm</quantity>
. Form oval, moderately depressed; color black, shiny; ground punctation on all surfaces consisting of a compact cluster or row of 3 micropunctures. Frons slightly convex, moderately coarsely, densely punctate; frontal stria continuous with marginal epistomal stria.
</paragraph>
<paragraph id="8BD7369F1E72FF8EF48679907C6DFCF7" blockId="24.[685,1172,194,1729]" pageId="24" pageNumber="182">Pronotum convex, widest at base and converging anteriorly; marginal stria complete, strongly impressed, subparallel to anterior edge of pronotum— not widely separated from anterior edge behind head; anterior angles acute, projecting; pronotal disc finely, densely punctate in lateral thirds to lateral margins, lateral fields of dense punctation sharply defined with internal boundaries diverging posterad; posterior margin gently rounded, marked by a row of coarse, widely separated punctures; antescutellar puncture weakly impressed.</paragraph>
<paragraph id="8BD7369F1E72FF8EF4867B7C7B00FC9D" blockId="24.[685,1172,194,1729]" pageId="24" pageNumber="182">Prosternal lobe coarsely, densely punctate anteriorly; marginal stria complete; prosternal keel convex; carinal striae slightly inwardly arcuate, united posteriorly.</paragraph>
<paragraph id="8BD7369F1E72FF8EF4867B037C23FABF" blockId="24.[685,1172,194,1729]" pageId="24" pageNumber="182">
Elytra finely, densely punctate; coarse punctures densely distributed along apical margins and extending basad to about apical third along elytral suture; epipleuron strigose-punctate in basal half, coarsely punctate in apical half; epipleural marginal stria slightly less outwardly arcuate than elytral striae, striae gradually converging from elytral base to apex; outer edge of elytral stria cariniform, inner edge lined with coarse punctures; external subhumeral striae represented by 23 coarse punctures; internal subhumeral striae complete, strongly impressed, punctate at regular intervals; 1
<superScript id="7C1D9BD71E72FF8EF55F7CD87B75FB4E" attach="none" box="[796,808,1198,1212]" fontSize="6" pageId="24" pageNumber="182">st</superScript>
4
<superScript id="7C1D9BD71E72FF8EF5037CD87B13FB4E" attach="left" box="[832,846,1198,1212]" fontSize="6" pageId="24" pageNumber="182">th</superScript>
dorsal striae complete, strongly impressed, also punctate at regular intervals, 5
<superScript id="7C1D9BD71E72FF8EF2C67CBF7CCEFB25" attach="none" box="[1157,1171,1225,1239]" fontSize="6" pageId="24" pageNumber="182">th</superScript>
dorsal and sutural striae represented by dense rows of strong punctures apically, fading basad but nearly reaching elytral base where 5
<superScript id="7C1D9BD71E72FF8EF2027D6F7C12FAD5" attach="left" box="[1089,1103,1305,1319]" fontSize="6" pageId="24" pageNumber="182">th</superScript>
dorsal stria curves slightly inward to meet sutural stria.
</paragraph>
<paragraph id="8BD7369F1E72FF8EF4867D257B05F933" blockId="24.[685,1172,194,1729]" pageId="24" pageNumber="182">Mesosternum finely, moderately densely punctate; anterior margin strongly emarginated to receive posterior prosternal projection; marginal stria complete, deeply impressed, continuous with lateral metasternal stria; mesometasternal suture represented by a fine line; intercoxal disc of metasternum finely, moderately densely punctate with coarse punctures distributed along inner edges of lateral metasternal striae; lateral metasternal stria strongly sinuate, strongly impressed, nearly reaching extreme outer edge of metacoxal cavity; posterior mesocoxal stria parallel to internal but distinctly shorter; lateral disc coarsely, densely punctate.</paragraph>
<caption id="DF1766171E73FF8FF6867BB47C0DFC27" ID-DOI="http://doi.org/10.5281/zenodo.5362717" ID-Zenodo-Dep="5362717" box="[197,1104,961,981]" httpUri="https://zenodo.org/record/5362717/files/figure.png" pageId="25" pageNumber="183" startId="25.[197,228,962,981]" targetBox="[195,1151,206,925]" targetPageId="25">
<paragraph id="8BD7369F1E73FF8FF6867BB47C0DFC27" blockId="25.[197,1104,961,981]" box="[197,1104,961,981]" pageId="25" pageNumber="183">
<emphasis id="B91CEA8D1E73FF8FF6867BB47955FC27" bold="true" box="[197,264,962,981]" pageId="25" pageNumber="183">Fig. 21.</emphasis>
<taxonomicName id="4C684D1C1E73FF8FF75E7BB779E7FC26" authorityName="Reese and Swanson" authorityYear="2017" box="[285,442,961,980]" class="Insecta" family="Histeridae" genus="Carcinops" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="183" phylum="Arthropoda" rank="species" species="wenzeli">
<emphasis id="B91CEA8D1E73FF8FF75E7BB779E7FC26" box="[285,442,961,980]" italics="true" pageId="25" pageNumber="183">Carcinops wenzeli</emphasis>
</taxonomicName>
morphology. A) Dorsal view, B) Ventral view. Photographs by A. Swanson.
</paragraph>
</caption>
<paragraph id="8BD7369F1E73FF8FF6867C6279B1FB3A" blockId="25.[173,659,1039,1729]" pageId="25" pageNumber="183">
Intercoxal disc of 1
<superScript id="7C1D9BD71E73FF8FF7C77C7979CDFBEF" attach="left" box="[388,400,1039,1053]" fontSize="6" pageId="25" pageNumber="183">st</superScript>
abdominal ventrite finely, moderately densely punctate with some coarse, dense punctation confined to posterolateral regions; bistriate on each side medial to metacoxae; lateral disc with 2 parallel striae medial to lateral marginal stria, surface bare in anterior half with coarse sparse punctures in posterior half.
</paragraph>
<paragraph id="8BD7369F1E73FF8FF6867CB87ACCFB0C" blockId="25.[173,659,1039,1729]" pageId="25" pageNumber="183">Propygidium hexagonal, coarsely, very densely punctate; punctation of pygidium somewhat less so.</paragraph>
<paragraph id="8BD7369F1E73FF8FF6867D7579FBFA20" blockId="25.[173,659,1039,1729]" pageId="25" pageNumber="183">Protibia explanate, roughly triangular; outer margin bidentate in distal half; denticles rounded, clearly projecting beyond tibial margin. Meso- and metatibia roughly cylindrical; outer margins abruptly expanded at distal sixth, forming lateral flanges from which 2 acute denticles project; mesotibia also bearing a single acute denticle at midpoint of outer margin.</paragraph>
<paragraph id="8BD7369F1E73FF8FF6867DAE7958F9D0" blockId="25.[173,659,1039,1729]" pageId="25" pageNumber="183">
Parameres of aedeagus parallel, not converging, each coming to an extremely sharp, hooked point (
<figureCitation id="13532A1A1E73FF8FF6F77E7B78A4F9D0" box="[180,249,1549,1570]" captionStart="Fig" captionStartId="6.[175,206,1591,1610]" captionTargetBox="[162,620,718,1549]" captionTargetId="figure-558@6.[151,637,712,1567]" captionTargetPageId="6" captionText="Fig. 9. Aedeagal morphology of Carcinops species, with parameres toward the top of the page and basal pieces toward the bottom. A) C. consors, B) C. gilensis, C) C. corticalis, D) C. papagoanus, E) C. opuntiae, F) C. curtus, G) C. rugulus, H) C. torquatus, I) C. yaqui, J) C. wenzeli. Photographs by A. Swanson." figureDoi="http://doi.org/10.5281/zenodo.5362689" httpUri="https://zenodo.org/record/5362689/files/figure.png" pageId="25" pageNumber="183">Fig. 9J</figureCitation>
).
</paragraph>
<paragraph id="8BD7369F1E73FF8FF6867E5E7C50FB0C" blockId="25.[173,659,1039,1729]" lastBlockId="25.[706,1194,1043,1729]" pageId="25" pageNumber="183">
<emphasis id="B91CEA8D1E73FF8FF6867E5E7A79F9CF" bold="true" box="[197,548,1576,1597]" pageId="25" pageNumber="183">Comparison with Related Species.</emphasis>
<taxonomicName id="4C684D1C1E73FF8FF4737E5178A9F9A5" authorityName="Reese and Swanson" authorityYear="2017" class="Insecta" family="Histeridae" genus="Carcinops" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="183" phylum="Arthropoda" rank="species" species="wenzeli">
<emphasis id="B91CEA8D1E73FF8FF4737E5178A9F9A5" italics="true" pageId="25" pageNumber="183">Carcinops wenzeli</emphasis>
</taxonomicName>
is virtually indistinguishable from its close relative
<taxonomicName id="4C684D1C1E73FF8FF7777E2A79FBF980" authorityName="Marseul" authorityYear="1855" baseAuthorityName="LeConte" baseAuthorityYear="1851" box="[308,422,1628,1650]" class="Insecta" family="Histeridae" genus="Carcinops" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="183" phylum="Arthropoda" rank="species" species="opuntiae">
<emphasis id="B91CEA8D1E73FF8FF7777E2A79FBF980" box="[308,422,1628,1650]" italics="true" pageId="25" pageNumber="183">C. opuntiae</emphasis>
</taxonomicName>
on the basis of external morphology alone.
<taxonomicName id="4C684D1C1E73FF8FF7307E007A68F97E" authorityName="Marseul" authorityYear="1855" baseAuthorityName="LeConte" baseAuthorityYear="1851" box="[371,565,1654,1676]" class="Insecta" family="Histeridae" genus="Carcinops" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="183" phylum="Arthropoda" rank="species" species="opuntiae">
<emphasis id="B91CEA8D1E73FF8FF7307E007A68F97E" box="[371,565,1654,1676]" italics="true" pageId="25" pageNumber="183">Carcinops opuntiae</emphasis>
</taxonomicName>
tends to be larger and slightly more elongate, but there is substantial overlap in these dimensions. Where their ranges overlap in
<collectingRegion id="49ACF87D1E73FF8FF5FB7C627C5AFBDB" box="[952,1031,1044,1065]" country="United States of America" name="Arizona" pageId="25" pageNumber="183">Arizona</collectingRegion>
,
<taxonomicName id="4C684D1C1E73FF8FF2547C657CD1FBDB" authorityName="Marseul" authorityYear="1855" baseAuthorityName="LeConte" baseAuthorityYear="1851" box="[1047,1164,1043,1065]" class="Insecta" family="Histeridae" genus="Carcinops" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="183" phylum="Arthropoda" rank="species" species="opuntiae">
<emphasis id="B91CEA8D1E73FF8FF2547C657CD1FBDB" box="[1047,1164,1043,1065]" italics="true" pageId="25" pageNumber="183">C. opuntiae</emphasis>
</taxonomicName>
is restricted to the necroses of various cacti, while
<taxonomicName id="4C684D1C1E73FF8FF4817C3E7B77FBAC" authorityName="Reese and Swanson" authorityYear="2017" box="[706,810,1096,1118]" class="Insecta" family="Histeridae" genus="Carcinops" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="183" phylum="Arthropoda" rank="species" species="wenzeli">
<emphasis id="B91CEA8D1E73FF8FF4817C3E7B77FBAC" box="[706,810,1096,1118]" italics="true" pageId="25" pageNumber="183">C. wenzeli</emphasis>
</taxonomicName>
is found only on decaying yucca and agave.
<taxonomicName id="4C684D1C1E73FF8FF5447C157BE5FB8B" authorityName="Reese and Swanson" authorityYear="2017" box="[775,952,1123,1145]" class="Insecta" family="Histeridae" genus="Carcinops" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="183" phylum="Arthropoda" rank="species" species="wenzeli">
<emphasis id="B91CEA8D1E73FF8FF5447C157BE5FB8B" box="[775,952,1123,1145]" italics="true" pageId="25" pageNumber="183">Carcinops wenzeli</emphasis>
</taxonomicName>
and
<taxonomicName id="4C684D1C1E73FF8FF5A97C157C07FB8B" authorityName="Marseul" authorityYear="1855" baseAuthorityName="LeConte" baseAuthorityYear="1851" box="[1002,1114,1123,1145]" class="Insecta" family="Histeridae" genus="Carcinops" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="183" phylum="Arthropoda" rank="species" species="opuntiae">
<emphasis id="B91CEA8D1E73FF8FF5A97C157C07FB8B" box="[1002,1114,1123,1145]" italics="true" pageId="25" pageNumber="183">C. opuntiae</emphasis>
</taxonomicName>
may be definitively separated only through comparison of the aedeagus. In
<taxonomicName id="4C684D1C1E73FF8FF5347CEE7BBCFB5C" authorityName="Reese and Swanson" authorityYear="2017" box="[887,993,1176,1198]" class="Insecta" family="Histeridae" genus="Carcinops" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="183" phylum="Arthropoda" rank="species" species="wenzeli">
<emphasis id="B91CEA8D1E73FF8FF5347CEE7BBCFB5C" box="[887,993,1176,1198]" italics="true" pageId="25" pageNumber="183">C. wenzeli</emphasis>
</taxonomicName>
, the parameres are straight and sharply pointed at the tips (
<figureCitation id="13532A1A1E73FF8FF2167CC57CC3FB3A" box="[1109,1182,1203,1224]" captionStart="Fig" captionStartId="6.[175,206,1591,1610]" captionTargetBox="[162,620,718,1549]" captionTargetId="figure-558@6.[151,637,712,1567]" captionTargetPageId="6" captionText="Fig. 9. Aedeagal morphology of Carcinops species, with parameres toward the top of the page and basal pieces toward the bottom. A) C. consors, B) C. gilensis, C) C. corticalis, D) C. papagoanus, E) C. opuntiae, F) C. curtus, G) C. rugulus, H) C. torquatus, I) C. yaqui, J) C. wenzeli. Photographs by A. Swanson." figureDoi="http://doi.org/10.5281/zenodo.5362689" httpUri="https://zenodo.org/record/5362689/files/figure.png" pageId="25" pageNumber="183">Fig. 9J</figureCitation>
). In
<taxonomicName id="4C684D1C1E73FF8FF49D7CBB7B12FB11" authorityName="Marseul" authorityYear="1855" baseAuthorityName="LeConte" baseAuthorityYear="1851" box="[734,847,1229,1251]" class="Insecta" family="Histeridae" genus="Carcinops" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="183" phylum="Arthropoda" rank="species" species="opuntiae">
<emphasis id="B91CEA8D1E73FF8FF49D7CBB7B12FB11" box="[734,847,1229,1251]" italics="true" pageId="25" pageNumber="183">C. opuntiae</emphasis>
</taxonomicName>
, the parameres are laterally sinuate and rounded at the tips (
<figureCitation id="13532A1A1E73FF8FF5ED7C9F7C5FFB0C" box="[942,1026,1257,1278]" captionStart="Fig" captionStartId="6.[175,206,1591,1610]" captionTargetBox="[162,620,718,1549]" captionTargetId="figure-558@6.[151,637,712,1567]" captionTargetPageId="6" captionText="Fig. 9. Aedeagal morphology of Carcinops species, with parameres toward the top of the page and basal pieces toward the bottom. A) C. consors, B) C. gilensis, C) C. corticalis, D) C. papagoanus, E) C. opuntiae, F) C. curtus, G) C. rugulus, H) C. torquatus, I) C. yaqui, J) C. wenzeli. Photographs by A. Swanson." figureDoi="http://doi.org/10.5281/zenodo.5362689" httpUri="https://zenodo.org/record/5362689/files/figure.png" pageId="25" pageNumber="183">Figs. 9E</figureCitation>
).
</paragraph>
<paragraph id="8BD7369F1E73FF8FF4987D757B81F9D0" blockId="25.[706,1194,1043,1729]" pageId="25" pageNumber="183">
<emphasis id="B91CEA8D1E73FF8FF4987D757C61FAEA" bold="true" box="[731,1084,1283,1304]" pageId="25" pageNumber="183">Geographic Distribution and Habitat.</emphasis>
<taxonomicName id="4C684D1C1E73FF8FF2057D747B54FAC1" authorityName="Reese and Swanson" authorityYear="2017" class="Insecta" family="Histeridae" genus="Carcinops" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="183" phylum="Arthropoda" rank="species" species="wenzeli">
<emphasis id="B91CEA8D1E73FF8FF2057D747B54FAC1" italics="true" pageId="25" pageNumber="183">Carcinops wenzeli</emphasis>
</taxonomicName>
occurs in the oak-juniper woodland of the lower slopes of the Sky Islands of southeastern Arizona and the Sierra Madre of
<collectingRegion id="49ACF87D1E73FF8FF2487D257C12FA9A" box="[1035,1103,1363,1384]" country="Mexico" name="Sonora" pageId="25" pageNumber="183">Sonora</collectingRegion>
,
<collectingCountry id="F37F760F1E73FF8FF21F7D257CFBFA9A" box="[1116,1190,1363,1384]" name="Mexico" pageId="25" pageNumber="183">Mexico</collectingCountry>
. It is generally associated with decaying basal stalks of plants in the subfamilies
<taxonomicName id="4C684D1C1E73FF8FF5AE7DFE7C01FA6F" box="[1005,1116,1416,1437]" class="Liliopsida" family="Asparagaceae" kingdom="Plantae" order="Asparagales" pageId="25" pageNumber="183" phylum="Tracheophyta" rank="subFamily" subFamily="Agavoideae">Agavoideae</taxonomicName>
(
<taxonomicName id="4C684D1C1E73FF8FF2317DFE7AB1FA4A" class="Liliopsida" family="Asparagaceae" genus="Yucca" kingdom="Plantae" order="Asparagales" pageId="25" pageNumber="183" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B91CEA8D1E73FF8FF2317DFE7CF4FA6F" box="[1138,1193,1416,1437]" italics="true" pageId="25" pageNumber="183">Yucca</emphasis>
spp.
</taxonomicName>
,
<taxonomicName id="4C684D1C1E73FF8FF4B97DD47B3AFA4A" box="[762,871,1442,1464]" class="Liliopsida" family="Asparagaceae" genus="Agave" kingdom="Plantae" order="Asparagales" pageId="25" pageNumber="183" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B91CEA8D1E73FF8FF4B97DD47B68FA4A" box="[762,821,1442,1464]" italics="true" pageId="25" pageNumber="183">Agave</emphasis>
spp.
</taxonomicName>
) and
<taxonomicName id="4C684D1C1E73FF8FF5E67DD57C4BFA4A" box="[933,1046,1443,1464]" class="Liliopsida" family="Asparagaceae" kingdom="Plantae" order="Asparagales" pageId="25" pageNumber="183" phylum="Tracheophyta" rank="subFamily" subFamily="Nolinoideae">Nolinoideae</taxonomicName>
(
<taxonomicName id="4C684D1C1E73FF8FF2697DD57CC1FA4A" box="[1066,1180,1442,1464]" class="Liliopsida" family="Asparagaceae" genus="Dasylirion" kingdom="Plantae" order="Asparagales" pageId="25" pageNumber="183" phylum="Tracheophyta" rank="species" species="wheeleri">
<emphasis id="B91CEA8D1E73FF8FF2697DD57CC1FA4A" box="[1066,1180,1442,1464]" italics="true" pageId="25" pageNumber="183">D. wheeleri</emphasis>
</taxonomicName>
), although APS collected
<specimenCount id="9D6EFD161E73FF8FF5877DCB7C13FA20" box="[964,1102,1469,1490]" pageId="25" pageNumber="183" type="generic">one specimen</specimenCount>
from a necrotic
<taxonomicName id="4C684D1C1E73FF8FF5517DA17BDDFA1F" box="[786,896,1495,1517]" class="Magnoliopsida" family="Cactaceae" genus="Opuntia" kingdom="Plantae" order="Caryophyllales" pageId="25" pageNumber="183" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B91CEA8D1E73FF8FF5517DA17B00FA1F" box="[786,861,1495,1517]" italics="true" pageId="25" pageNumber="183">Opuntia</emphasis>
sp.
</taxonomicName>
cladode (see
<bibRefCitation id="EFF94B6E1E73FF8FF5BC7DAE7CF4FA1F" author="Martin, P. S. &amp; Yetman, M. &amp; Fishbein, P. &amp; Jenkins, T. R. Van" box="[1023,1193,1495,1517]" pageId="25" pageNumber="183" refId="ref20000" refString="Martin, P. S., D. Yetman, M. Fishbein, P. Jenkins, T. R. Van Devender, and R. K. Wilson. 1998. Gentry' s Rio Mayo Plants: The Tropical Deciduous Forest and Environs of Northwest Mexico. University of Arizona Press, Tucson, AZ." type="book" year="1998">
Martin
<emphasis id="B91CEA8D1E73FF8FF2077DAE7C33FA1F" box="[1092,1134,1495,1517]" italics="true" pageId="25" pageNumber="183">et al</emphasis>
. 1998
</bibRefCitation>
for a brief discussion of this unidentified
<taxonomicName id="4C684D1C1E73FF8FF21E7D847B58F9D0" class="Magnoliopsida" family="Cactaceae" genus="Opuntia" kingdom="Plantae" order="Caryophyllales" pageId="25" pageNumber="183" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B91CEA8D1E73FF8FF21E7D847CF5F9FA" box="[1117,1192,1522,1544]" italics="true" pageId="25" pageNumber="183">Opuntia</emphasis>
species
</taxonomicName>
) near Alamos,
<collectingRegion id="49ACF87D1E73FF8FF5D77E7B7B85F9D0" box="[916,984,1549,1570]" country="Mexico" name="Sonora" pageId="25" pageNumber="183">Sonora</collectingRegion>
.
</paragraph>
<paragraph id="8BD7369F1E73FF8CF4987E5E792EFA3A" blockId="25.[706,1194,1043,1729]" lastBlockId="26.[151,639,1326,1586]" lastPageId="26" lastPageNumber="184" pageId="25" pageNumber="183">
<emphasis id="B91CEA8D1E73FF8FF4987E5E7C6AF9CF" bold="true" box="[731,1079,1576,1597]" pageId="25" pageNumber="183">Additional Material Examined.</emphasis>
<specimenCount id="9D6EFD161E73FF8FF2007E5E7AAAF9A5" pageId="25" pageNumber="183" type="generic">28 specimens</specimenCount>
.
<emphasis id="B91CEA8D1E73FF8FF5477E347B94F9A5" bold="true" box="[772,969,1602,1623]" pageId="25" pageNumber="183">
<collectingCountry id="F37F760F1E73FF8FF5477E347B31F9A5" box="[772,876,1602,1623]" name="Mexico" pageId="25" pageNumber="183">MEXICO</collectingCountry>
:
<collectingRegion id="49ACF87D1E73FF8FF53A7E347B98F9A5" box="[889,965,1602,1623]" country="Mexico" name="Sonora" pageId="25" pageNumber="183">Sonora</collectingRegion>
:
</emphasis>
Arroyo Santa Barbara nr. Alamos,
<geoCoordinate id="EE5C50581E73FF8FF57C7E2B7BEEF980" box="[831,947,1629,1650]" degrees="27.11326" direction="north" orientation="latitude" pageId="25" pageNumber="183" precision="1" value="27.11326">27.11326°N</geoCoordinate>
<geoCoordinate id="EE5C50581E73FF8FF5FF7E2B7C19F980" box="[956,1092,1629,1650]" degrees="108.72591" direction="west" orientation="longitude" pageId="25" pageNumber="183" precision="1" value="-108.72591">108.72591°W</geoCoordinate>
,
<quantity id="4C909B7A1E73FF8FF2127E2B7CF8F980" box="[1105,1189,1629,1650]" metricMagnitude="3" metricUnit="m" metricValue="1.35" pageId="25" pageNumber="183" unit="m" value="1350.0">1,350 m</quantity>
,
<date id="FFD6105F1E73FF8FF4817E017B77F97E" box="[706,810,1655,1676]" pageId="25" pageNumber="183" value="2007-05-29">29.V.2007</date>
, ex. decaying
<taxonomicName id="4C684D1C1E73FF8FF5FC7E007C7FF97E" box="[959,1058,1654,1676]" class="Liliopsida" family="Asparagaceae" genus="Agave" kingdom="Plantae" order="Asparagales" pageId="25" pageNumber="183" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B91CEA8D1E73FF8FF5FC7E007BA7F97E" box="[959,1018,1654,1676]" italics="true" pageId="25" pageNumber="183">Agave</emphasis>
sp.
</taxonomicName>
, A. Swanson (APS, 24; DNA vouchers: APS137, APS174); Arroyo Santa Barbara nr. Alamos,
<geoCoordinate id="EE5C50581E73FF8FF2777EDA7CF5F933" box="[1076,1192,1708,1729]" degrees="27.11547" direction="north" orientation="latitude" pageId="25" pageNumber="183" precision="1" value="27.11547">27.11547°N</geoCoordinate>
<geoCoordinate id="EE5C50581E70FF8CF6D47D58797DFAB1" box="[151,288,1326,1347]" degrees="108.70767" direction="west" orientation="longitude" pageId="26" pageNumber="184" precision="1" value="-108.70767">108.70767°W</geoCoordinate>
,
<quantity id="4C909B7A1E70FF8CF7717D58792AFAB1" box="[306,375,1326,1347]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="26" pageNumber="184" unit="m" value="1400.0">1400m</quantity>
,
<date id="FFD6105F1E70FF8CF7C97D5879B0FAB1" box="[394,493,1326,1347]" pageId="26" pageNumber="184" value="2007-06-03">3.VI.2007</date>
, ex. Necrotic
<taxonomicName id="4C684D1C1E70FF8CF6D47D3E7957FAAF" box="[151,266,1352,1374]" class="Magnoliopsida" family="Cactaceae" genus="Opuntia" kingdom="Plantae" order="Caryophyllales" pageId="26" pageNumber="184" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B91CEA8D1E70FF8CF6D47D3E78B9FAAC" box="[151,228,1352,1374]" italics="true" pageId="26" pageNumber="184">Opuntia</emphasis>
sp.
</taxonomicName>
, A. Swanson (APS, 1; DNA voucher: APS141).
<emphasis id="B91CEA8D1E70FF8CF6B97D1579CDFA8A" bold="true" box="[250,400,1379,1400]" pageId="26" pageNumber="184">
<collectingCountry id="F37F760F1E70FF8CF6B97D157971FA8A" box="[250,300,1379,1400]" name="United States of America" pageId="26" pageNumber="184">USA</collectingCountry>
:
<collectingRegion id="49ACF87D1E70FF8CF77B7D1579D1FA8A" box="[312,396,1379,1400]" country="United States of America" name="Arizona" pageId="26" pageNumber="184">Arizona</collectingRegion>
:
</emphasis>
Cochise Co., Chiricahua Mts., Pinery Cyn,
<date id="FFD6105F1E70FF8CF7077D0879E0FA61" box="[324,445,1406,1427]" pageId="26" pageNumber="184" value="2006-08-15">15.VIII.2006</date>
, ex. decaying
<taxonomicName id="4C684D1C1E70FF8CF4077D0B78EFFA5F" class="Liliopsida" family="Asparagaceae" genus="Agave" kingdom="Plantae" order="Asparagales" pageId="26" pageNumber="184" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B91CEA8D1E70FF8CF4077D0B7A20FA61" box="[580,637,1405,1427]" italics="true" pageId="26" pageNumber="184">Agave</emphasis>
sp.
</taxonomicName>
, A. Swanson (APS, 2); Patagonia,
<date id="FFD6105F1E70FF8CF45C7DEE7A27FA5F" box="[543,634,1432,1453]" pageId="26" pageNumber="184" value="1936-07">VII.1936</date>
, E. S. Ross (CASC, 1).
</paragraph>
<caption id="DF1766171E70FF8CF6EC7C9C7B8FFB0F" ID-DOI="http://doi.org/10.5281/zenodo.5362719" ID-Zenodo-Dep="5362719" box="[175,978,1257,1277]" httpUri="https://zenodo.org/record/5362719/files/figure.png" pageId="26" pageNumber="184" startId="26.[175,206,1258,1277]" targetBox="[151,1171,183,1231]" targetPageId="26">
<paragraph id="8BD7369F1E70FF8CF6EC7C9C7B8FFB0F" blockId="26.[175,978,1257,1277]" box="[175,978,1257,1277]" pageId="26" pageNumber="184">
<emphasis id="B91CEA8D1E70FF8CF6EC7C9C78AEFB0F" bold="true" box="[175,243,1258,1277]" pageId="26" pageNumber="184">Fig. 22.</emphasis>
COI gene tree constructed from sequences from this study and
<bibRefCitation id="EFF94B6E1E70FF8CF5677C9C7B90FB0F" author="Pfeiler, E. &amp; Johnson, M. P." box="[804,973,1257,1277]" pageId="26" pageNumber="184" pagination="491 - 501" refId="ref20161" refString="Pfeiler, E., S. Johnson, M. P. Richmond, and T. A. Markow. 2013. Population genetics and phylogenetic relationships of beetles (Coleoptera: Histeridae and Staphylinidae) from the Sonoran Desert associated with rotting columnar cacti. Molecular Phylogenetics and Evolution 69 (3): 491 - 501." type="journal article" year="2013">
Pfeiler
<emphasis id="B91CEA8D1E70FF8CF5227C9C7BDAFB0E" box="[865,903,1257,1276]" italics="true" pageId="26" pageNumber="184">et al</emphasis>
. (2013)
</bibRefCitation>
.
</paragraph>
</caption>
<paragraph id="8BD7369F1E70FF8CF6EC7DBB790CF9C0" blockId="26.[151,639,1326,1586]" pageId="26" pageNumber="184">
<emphasis id="B91CEA8D1E70FF8CF6EC7DBB7A52FA10" bold="true" box="[175,527,1485,1506]" pageId="26" pageNumber="184">Derivation of Specific Epithet.</emphasis>
<taxonomicName id="4C684D1C1E70FF8CF4587DBA7883FA0F" authorityName="Reese and Swanson" authorityYear="2017" class="Insecta" family="Histeridae" genus="Carcinops" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="184" phylum="Arthropoda" rank="species" species="wenzeli">
<emphasis id="B91CEA8D1E70FF8CF4587DBA7883FA0F" italics="true" pageId="26" pageNumber="184">Carcinops wenzeli</emphasis>
</taxonomicName>
is named for the former curator of histerids at the Field Museum of Natural History, the late Dr. Rupert Wenzel.
</paragraph>
</subSubSection>
</treatment>
</document>