169 lines
20 KiB
XML
169 lines
20 KiB
XML
<document id="3C3334DCAD8F6B2D9AA60DAEC658888C" ID-DOI="10.24199/j.mmv.2004.61.11" ID-ISSN="1447-2554" ID-Zenodo-Dep="12207880" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1718988272896" checkinUser="felipe" docAuthor="Bock, Philip E. & Cook, Patricia L." docDate="2004" docId="03C18788102DFFC164B44EAEFB2DFE82" docLanguage="en" docName="MemMusVictoria.61.2.135-182.pdf" docOrigin="Memoirs of Museum Victoria 61 (2)" docSource="https://museumsvictoria.com.au/collections-research/journals/memoirs-of-museum-victoria/volume-61-issue-2-2004/pages-135-182/" docStyle="DocumentStyle:5B3B5C052A93BFEEE334AD6097A94E45.2:MemMusVictoria.2003-.journal_article.type1" docStyleId="5B3B5C052A93BFEEE334AD6097A94E45" docStyleName="MemMusVictoria.2003-.journal_article.type1" docStyleVersion="2" docTitle="Trochosodon diommatus Bock & Cook, 2004, sp. nov." docType="treatment" docVersion="2" lastPageNumber="168" masterDocId="FFF8FFF0100CFFE364374B1DFFDFFF91" masterDocTitle="A review of Australian Conescharellinidae (Bryozoa: Cheilostomata)" masterLastPageNumber="182" masterPageNumber="135" pageNumber="168" updateTime="1718989335086" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo id="45F4C0AADA9E4716223037A39563DA06">
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<mods:title id="F7B6F6887940F6F282423FFA574775AD">A review of Australian Conescharellinidae (Bryozoa: Cheilostomata)</mods:title>
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<mods:namePart id="4018F7B50E1C872D7AC374E45024101F">Bock, Philip E.</mods:namePart>
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<mods:namePart id="5DA14BCC7BEC1E55F4C012FF68556BD0">Cook, Patricia L.</mods:namePart>
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<mods:typeOfResource id="7DE554ADF87ACAA03BA807B29499FCDA">text</mods:typeOfResource>
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<mods:title id="66875164B25D7043789AEECDB20DC08B">Memoirs of Museum Victoria</mods:title>
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<mods:part id="F98267B802B4A47E2B70D06236643470">
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<mods:date id="8B0CFBDB34FE7B2194C3221A61FFE8FA">2004</mods:date>
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<mods:number id="07D56AD91F2DEE208BC42373306DFFD9">2004-12-31</mods:number>
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<mods:number id="8331862F96227814378B681B35B6036B">61</mods:number>
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<mods:url id="A86FB6CFA44E9E71E2167EA5E0A7F362">https://museumsvictoria.com.au/collections-research/journals/memoirs-of-museum-victoria/volume-61-issue-2-2004/pages-135-182/</mods:url>
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<mods:classification id="424E6CFABFAA4C4121E8B04064E3480A">journal article</mods:classification>
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<mods:identifier id="5308E39DFE61AE696D4AF22A3649E7E7" type="DOI">10.24199/j.mmv.2004.61.11</mods:identifier>
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<mods:identifier id="3C1401DC2E5543400C946FA52F380763" type="ISSN">1447-2554</mods:identifier>
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<mods:identifier id="13D74AD0882C75F9023A7641C882639B" type="Zenodo-Dep">12207880</mods:identifier>
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</mods:mods>
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<treatment id="03C18788102DFFC164B44EAEFB2DFE82" LSID="urn:lsid:plazi:treatment:03C18788102DFFC164B44EAEFB2DFE82" httpUri="http://treatment.plazi.org/id/03C18788102DFFC164B44EAEFB2DFE82" lastPageId="34" lastPageNumber="168" pageId="33" pageNumber="168">
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<subSubSection id="C3726515102DFFC264B44EAEFE02FA58" box="[131,477,1459,1481]" pageId="33" pageNumber="168" type="nomenclature">
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<paragraph id="8BD7369E102DFFC264B44EAEFE02FA58" blockId="33.[131,477,1459,1481]" box="[131,477,1459,1481]" pageId="33" pageNumber="168">
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<heading id="D09F81F2102DFFC264B44EAEFE02FA58" box="[131,477,1459,1481]" fontSize="9" level="2" pageId="33" pageNumber="168" reason="5">
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<taxonomicName id="4C684D1D102DFFC264B44EAEFE59FA58" ID-CoL="597QH" authorityName="Bock & Cook" authorityYear="2004" box="[131,390,1459,1481]" class="Gymnolaemata" family="Conescharellinidae" genus="Trochosodon" kingdom="Animalia" order="Cheilostomatida" pageId="33" pageNumber="168" phylum="Bryozoa" rank="species" species="diommatus" status="sp. nov.">
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<emphasis id="B91CEA8C102DFFC264B44EAEFE59FA58" bold="true" box="[131,390,1459,1481]" italics="true" pageId="33" pageNumber="168">Trochosodon diommatus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A22F57F7102DFFC265BA4EAEFE02FA58" box="[397,477,1459,1481]" pageId="33" pageNumber="168" rank="species">sp. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3726515102DFFC164B44EFDFB2DFE82" lastPageId="34" lastPageNumber="169" pageId="33" pageNumber="168" type="description">
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<paragraph id="8BD7369E102DFFC264B44EFDFE99FA67" blockId="33.[131,326,1504,1526]" box="[131,326,1504,1526]" pageId="33" pageNumber="168">
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<figureCitation id="13532A1B102DFFC264B44EFDFF2DFA67" box="[131,242,1504,1526]" captionStart="Figure 20" captionStartId="33.[811,870,972,992]" captionTargetBox="[809,1452,221,954]" captionTargetId="figure-747@33.[809,1453,221,955]" captionTargetPageId="33" captionText="Figure 20. Trochosodon diommatus sp. nov. NMV F99012, holotype. adapical view of colony showing lunate root pores, scale = 500 µm." figureDoi="http://doi.org/10.5281/zenodo.12207925" httpUri="https://zenodo.org/record/12207925/files/figure.png" pageId="33" pageNumber="168">Figures 20</figureCitation>
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,
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<figureCitation id="13532A1B102DFFC264CA4EFDFE99FA67" box="[253,326,1504,1526]" captionStart="Figure 21" captionStartId="34.[139,198,1781,1801]" captionTargetBox="[171,762,221,1759]" captionTargetId="figure-463@34.[170,780,220,1763]" captionTargetPageId="34" captionText="Figure 21. Trochosodon diommatus sp. nov. A. NMV F99012, holotype. Detail of orifices and adapical pores, scale = 200 µm. B, NMV F99013, paratype, antapical view of marginal peristomes, showing paired avicularia, scale =200 µm. C. NMV F99014, paratype, detail of peristome, orifice, and lunate root pores, scale = 200 µm." figureDoi="http://doi.org/10.5281/zenodo.12207929" httpUri="https://zenodo.org/record/12207929/files/figure.png" pageId="33" pageNumber="168">21A–C</figureCitation>
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</paragraph>
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<paragraph id="8BD7369E102DFFC264B44D17FD48F98F" blockId="33.[131,772,1546,1673]" box="[131,663,1546,1566]" pageId="33" pageNumber="168">
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<materialsCitation id="3B003CC3102DFFC264B44D17FD48F98F" box="[131,663,1546,1566]" collectionCode="NMV" pageId="33" pageNumber="168" specimenCode="F99012" specimenCount="1" typeStatus="holotype">
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<emphasis id="B91CEA8C102DFFC264B44D17FF09F98F" box="[131,214,1546,1566]" italics="true" pageId="33" pageNumber="168">
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<typeStatus id="54D3883C102DFFC264B44D17FF09F98F" box="[131,214,1546,1566]" pageId="33" pageNumber="168" type="holotype">Holotype</typeStatus>
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</emphasis>
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.
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<collectionCode id="ED79AE5B102DFFC264D54D17FEC8F98F" box="[226,279,1546,1566]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34978" lsid="urn:lsid:biocol.org:col:34978" name="Museum Victoria" pageId="33" pageNumber="168" type="Museum">NMV</collectionCode>
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<specimenCode id="DBCE9EE5102DFFC2652A4D17FEBAF98F" box="[285,357,1546,1566]" collectionCode="NMV" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34978" lsid="urn:lsid:biocol.org:col:34978" name="Museum Victoria" pageId="33" pageNumber="168" type="Museum">F99012</specimenCode>
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, figured specimen, stn SLOPE-7.
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</materialsCitation>
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</paragraph>
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<paragraph id="8BD7369E102DFFC264A94D38FCDBF9A8" blockId="33.[131,772,1546,1673]" box="[158,772,1573,1593]" pageId="33" pageNumber="168">
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<materialsCitation id="3B003CC3102DFFC264A94D38FCDFF9A8" box="[158,768,1573,1593]" collectionCode="NMV" pageId="33" pageNumber="168" specimenCode="F99013, F99014" specimenCount="1" typeStatus="paratype">
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<emphasis id="B91CEA8C102DFFC264A94D38FF30F9A8" box="[158,239,1573,1593]" italics="true" pageId="33" pageNumber="168">
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<typeStatus id="54D3883C102DFFC264A94D38FF30F9A8" box="[158,239,1573,1593]" pageId="33" pageNumber="168" type="paratype">Paratype</typeStatus>
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</emphasis>
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.
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<collectionCode id="ED79AE5B102DFFC264CD4D38FEF0F9A8" box="[250,303,1573,1593]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34978" lsid="urn:lsid:biocol.org:col:34978" name="Museum Victoria" pageId="33" pageNumber="168" type="Museum">NMV</collectionCode>
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<specimenCode id="DBCE9EE5102DFFC265034D38FEA3F9A8" box="[308,380,1573,1593]" collectionCode="NMV" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34978" lsid="urn:lsid:biocol.org:col:34978" name="Museum Victoria" pageId="33" pageNumber="168" type="Museum">F99013</specimenCode>
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,
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<specimenCode id="DBCE9EE5102DFFC265B34D38FE13F9A8" box="[388,460,1573,1593]" collectionCode="NMV" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34978" lsid="urn:lsid:biocol.org:col:34978" name="Museum Victoria" pageId="33" pageNumber="168" type="Museum">F99014</specimenCode>
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, figured specimens, stn SLOPE-7
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="8BD7369E102DFFC264A94D22FDBCF918" blockId="33.[131,772,1546,1673]" pageId="33" pageNumber="168">
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<emphasis id="B91CEA8C102DFFC264A94D22FEE2F9C2" box="[158,317,1599,1619]" italics="true" pageId="33" pageNumber="168">Other specimens.</emphasis>
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NMV F101970, stn SLOPE-6 (4 colonies, 3 very young); NMV F101971, stn SLOPE-7 (22 colonies, 10 with roots); NMV F101972, stn SLOPE-45 (1 colony with root).
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</paragraph>
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<paragraph id="8BD7369E102DFFC264B44D82FDE9F943" blockId="33.[131,771,1695,1746]" pageId="33" pageNumber="168">
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<emphasis id="B91CEA8C102DFFC264B44D82FECEF924" box="[131,273,1695,1717]" italics="true" pageId="33" pageNumber="168">Etymology. di</emphasis>
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– two and
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<emphasis id="B91CEA8C102DFFC265BD4D82FE3AF924" box="[394,485,1695,1717]" italics="true" pageId="33" pageNumber="168">ommatos</emphasis>
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– an eye (Gr.), referring to the paired antapical peristomial avicularia.
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</paragraph>
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<paragraph id="8BD7369E102DFFC264B44DF5FF27F8E2" blockId="33.[131,771,1768,1907]" pageId="33" pageNumber="168">
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<emphasis id="B91CEA8C102DFFC264B44DF5FF36F96F" box="[131,233,1768,1790]" italics="true" pageId="33" pageNumber="168">Diagnosis</emphasis>
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.
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<taxonomicName id="4C684D1D102DFFC264CF4DF5FEA4F96F" authorityName="Canu and Bassler" authorityYear="1927" box="[248,379,1768,1790]" class="Gymnolaemata" family="Conescharellinidae" genus="Trochosodon" kingdom="Animalia" order="Cheilostomatida" pageId="33" pageNumber="168" phylum="Bryozoa" rank="genus">
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<emphasis id="B91CEA8C102DFFC264CF4DF5FEA4F96F" box="[248,379,1768,1790]" italics="true" pageId="33" pageNumber="168">Trochosodon</emphasis>
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</taxonomicName>
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with stellate, radial peristomes, calcification smooth to finely tuberculate. Zooid orifices deeply concealed, with a narrow sinus. Frontal avicularia minute; a prominent pair on the antapical surface of the marginal peristomes.
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</paragraph>
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<paragraph id="8BD7369E102DFFC2671E4F2AFC77FA3D" blockId="33.[809,1450,1078,1511]" pageId="33" pageNumber="168">
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<emphasis id="B91CEA8C102DFFC2671E4F2AFC7EFBDC" box="[809,929,1079,1101]" italics="true" pageId="33" pageNumber="168">Description</emphasis>
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. Colony stellate, fairly flat, distinctly wider than high, with prominent marginal peristomes. Orifices quincuncial at first, becoming radial. Primary orifice at the base of the long, tubular but not prominent peristome, with an elongate, fairly narrow sinus and large, paired condyles. An adapical pore present on the edge of the peristome of some peripheral zooids. Avicularia single, lateral and antapical between the peristomes, rostrum semicircular, with a bar but no ligula; other small avicularia scattered. Lunate root pores frequent in the adapical region, each with a pair of avicularia laterally. Antapical surface with marginal pores and avicularia; a pair of avicularia on the antapical surface of each peristome (cf.
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<taxonomicName id="4C684D1D102DFFC261324E64FAA6FA1E" authorityName="Bock & Cook" authorityYear="2004" box="[1285,1401,1401,1423]" class="Gymnolaemata" family="Conescharellinidae" genus="Conescharellina" kingdom="Animalia" order="Cheilostomatida" pageId="33" pageNumber="168" phylum="Bryozoa" rank="species" species="ocellata">
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<emphasis id="B91CEA8C102DFFC261324E64FAA6FA1E" box="[1285,1401,1401,1423]" italics="true" pageId="33" pageNumber="168">C. ocellata</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C684D1D102DFFC2671E4E8AFC45FA3C" baseAuthorityName="Maplestone" baseAuthorityYear="1909" box="[809,922,1431,1453]" class="Gymnolaemata" family="Conescharellinidae" genus="Conescharellina" kingdom="Animalia" order="Cheilostomatida" pageId="33" pageNumber="168" phylum="Bryozoa" rank="species" species="eburnea">
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<emphasis id="B91CEA8C102DFFC2671E4E8AFC45FA3C" box="[809,922,1431,1453]" italics="true" pageId="33" pageNumber="168">C. eburnea</emphasis>
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</taxonomicName>
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).
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</paragraph>
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<paragraph id="8BD7369E102DFFC2677E4EA9FB24FA76" blockId="33.[809,1450,1078,1511]" pageId="33" pageNumber="168">
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Colony with up to 4 whorls and 4–5 zooids per whorl. Diameter up to
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<quantity id="4C909B7B102DFFC267FE4ECCFBC5FA76" box="[969,1050,1489,1511]" metricMagnitude="-3" metricUnit="m" metricValue="4.7" pageId="33" pageNumber="168" unit="mm" value="4.7">4.7 mm</quantity>
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, height up to
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<quantity id="4C909B7B102DFFC260914ECCFB28FA76" box="[1190,1271,1489,1511]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="33" pageNumber="168" unit="mm" value="1.5">1.5 mm</quantity>
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.
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</paragraph>
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<paragraph id="8BD7369E102DFFC1671E4EE0FB2DFE82" blockId="33.[809,1449,1533,1907]" lastBlockId="34.[817,1457,224,275]" lastPageId="34" lastPageNumber="169" pageId="33" pageNumber="168">
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<emphasis id="B91CEA8C102DFFC2671E4EE0FC56F982" box="[809,905,1533,1555]" italics="true" pageId="33" pageNumber="168">Remarks.</emphasis>
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Ovicells have not been seen in
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<taxonomicName id="4C684D1D102DFFC260E64EE0FA85F982" authorityName="Bock & Cook" authorityYear="2004" box="[1233,1370,1533,1555]" class="Gymnolaemata" family="Conescharellinidae" genus="Trochosodon" kingdom="Animalia" order="Cheilostomatida" pageId="33" pageNumber="168" phylum="Bryozoa" rank="species" species="diommatus">
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<emphasis id="B91CEA8C102DFFC260E64EE0FA85F982" box="[1233,1370,1533,1555]" italics="true" pageId="33" pageNumber="168">T. diommatus</emphasis>
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</taxonomicName>
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but the central position of the adapical pore suggests that they would be symmetrical, like those of
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<taxonomicName id="4C684D1D102DFFC260644D25FB19F9DF" authorityName="Bock & Cook" authorityYear="2004" box="[1107,1222,1592,1614]" class="Gymnolaemata" family="Conescharellinidae" genus="Trochosodon" kingdom="Animalia" order="Cheilostomatida" pageId="33" pageNumber="168" phylum="Bryozoa" rank="species" species="fecundus">
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<emphasis id="B91CEA8C102DFFC260644D25FB19F9DF" box="[1107,1222,1592,1614]" italics="true" pageId="33" pageNumber="168">T. fecundus</emphasis>
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</taxonomicName>
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, rather than asymmetrical, as in
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<taxonomicName id="4C684D1D102DFFC267AF4D48FBE2F9FA" authorityName="Bock & Cook" authorityYear="2004" box="[920,1085,1621,1643]" class="Gymnolaemata" family="Conescharellinidae" genus="Trochosodon" kingdom="Animalia" order="Cheilostomatida" pageId="33" pageNumber="168" phylum="Bryozoa" rank="species" species="asymmetricus">
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<emphasis id="B91CEA8C102DFFC267AF4D48FBE2F9FA" box="[920,1085,1621,1643]" italics="true" pageId="33" pageNumber="168">T. asymmetricus</emphasis>
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</taxonomicName>
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. Several colonies from stn SLOPE- 7 have roots present; these are 0.5–1.0 mm long.
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<taxonomicName id="4C684D1D102DFFC261174D6EFA76F918" authorityName="Bock & Cook" authorityYear="2004" box="[1312,1449,1651,1673]" class="Gymnolaemata" family="Conescharellinidae" genus="Trochosodon" kingdom="Animalia" order="Cheilostomatida" pageId="33" pageNumber="168" phylum="Bryozoa" rank="species" species="diommatus">
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<emphasis id="B91CEA8C102DFFC261174D6EFA76F918" box="[1312,1449,1651,1673]" italics="true" pageId="33" pageNumber="168">T. diommatus</emphasis>
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</taxonomicName>
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is easily distinguished by the presence of the pair of minute avicularia on the antapical side of the marginal zooid peristomes. It resembles two other species in the presence of antapical peristomial avicularia. It differs completely from fossil
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<taxonomicName id="4C684D1D102DFFC2671E4C18FC43F88A" authorityName="Bock & Cook" authorityYear="2004" box="[809,924,1797,1819]" class="Gymnolaemata" family="Conescharellinidae" genus="Conescharellina" kingdom="Animalia" order="Cheilostomatida" pageId="33" pageNumber="168" phylum="Bryozoa" rank="species" species="ocellata">
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<emphasis id="B91CEA8C102DFFC2671E4C18FC43F88A" box="[809,924,1797,1819]" italics="true" pageId="33" pageNumber="168">C. ocellata</emphasis>
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</taxonomicName>
|
||
in dimensions and arrangement of the zooid orifices, that have a longer, more acutely subtriangular sinus. It differs from
|
||
<taxonomicName id="4C684D1D102DFFC2679C4C5DFBC2F8C7" baseAuthorityName="Maplestone" baseAuthorityYear="1909" box="[939,1053,1856,1878]" class="Gymnolaemata" family="Conescharellinidae" genus="Conescharellina" kingdom="Animalia" order="Cheilostomatida" pageId="33" pageNumber="168" phylum="Bryozoa" rank="species" species="eburnea">
|
||
<emphasis id="B91CEA8C102DFFC2679C4C5DFBC2F8C7" box="[939,1053,1856,1878]" italics="true" pageId="33" pageNumber="168">C. eburnea</emphasis>
|
||
</taxonomicName>
|
||
in its long peristomes and narrow orificial sinus, as well as the form of its root pores.
|
||
<taxonomicName id="4C684D1D102DFFC261174C40FA76F8E2" authorityName="Bock & Cook" authorityYear="2004" box="[1312,1449,1885,1907]" class="Gymnolaemata" family="Conescharellinidae" genus="Trochosodon" kingdom="Animalia" order="Cheilostomatida" pageId="33" pageNumber="168" phylum="Bryozoa" rank="species" species="diommatus">
|
||
<emphasis id="B91CEA8C102DFFC261174C40FA76F8E2" box="[1312,1449,1885,1907]" italics="true" pageId="33" pageNumber="168">T. diommatus</emphasis>
|
||
</taxonomicName>
|
||
has a similar distribution to
|
||
<taxonomicName id="4C684D1D102EFFC160664BFDFB29FF67" authorityName="Bock & Cook" authorityYear="2004" box="[1105,1270,224,246]" class="Gymnolaemata" family="Conescharellinidae" genus="Trochosodon" kingdom="Animalia" order="Cheilostomatida" pageId="34" pageNumber="169" phylum="Bryozoa" rank="species" species="asymmetricus">
|
||
<emphasis id="B91CEA8C102EFFC160664BFDFB29FF67" box="[1105,1270,224,246]" italics="true" pageId="34" pageNumber="169">T. asymmetricus</emphasis>
|
||
</taxonomicName>
|
||
, with the addition of a record from
|
||
<quantity id="4C909B7B102EFFC167E84BE0FBFFFE82" box="[991,1056,253,275]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="34" pageNumber="169" unit="m" value="800.0">800 m</quantity>
|
||
depth off
|
||
<collectingRegion id="49ACF87C102EFFC160BD4BE0FB31FE82" box="[1162,1262,253,275]" country="Australia" name="Tasmania" pageId="34" pageNumber="169">Tasmania</collectingRegion>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |