275 lines
34 KiB
XML
275 lines
34 KiB
XML
<document id="18D6F59C6B50839AC11B09D909FB1235" ID-DOI="10.5281/zenodo.5176513" ID-ISSN="1942-1354" ID-Zenodo-Dep="5176513" ID-ZooBank="068D2C0D-C92D-4E47-93FF-222CA4D5D824" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628329503398" checkinUser="felipe" docAuthor="Canepari, Claudio, Gordon, Robert D. & Hanley, Guy A." docDate="2013" docId="038F8787FFABFFBBFF7C5996FD9DFCE0" docLanguage="en" docName="InsectaMundi.0312.1-91.pdf" docOrigin="Insecta Mundi 2013 (312)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Dilatitibialis Duverger 2001" docType="treatment" docVersion="5" lastPageNumber="4" masterDocId="FFB6FFFFFFADFFBCFFBC5B55FF8DFFFB" masterDocTitle="South American Coccinellidae (Coleoptera), Part XV: systematic revision of Dilatitibialis Duverger (Coccidulinae; Hyperaspidini)" masterLastPageNumber="91" masterPageNumber="1" pageNumber="3" updateTime="1699044533877" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
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<mods:titleInfo id="72A7ADEF372C355144E3826F011AF744">
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<mods:title id="EE3578FF513F1DA4392D6F2BFBCB5A20">South American Coccinellidae (Coleoptera), Part XV: systematic revision of Dilatitibialis Duverger (Coccidulinae; Hyperaspidini)</mods:title>
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<mods:roleTerm id="728C7F3DE96EAC23836342146D1FA66B">Author</mods:roleTerm>
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<mods:namePart id="63BB20AD3242F28541CF294757E5BBBB">Canepari, Claudio</mods:namePart>
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<mods:roleTerm id="0DBD3AEDAD31CD9830754DF405C1662E">Author</mods:roleTerm>
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<mods:namePart id="C71D4D32D716B2A436D3B30A62B67738">Gordon, Robert D.</mods:namePart>
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<mods:name id="083ADE29C1A11D6C31955A4A07F5B94D" type="personal">
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<mods:roleTerm id="FB08EA22D6181E4ED38176A1AAFE2719">Author</mods:roleTerm>
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<mods:namePart id="6218FC6A26A3E0C7B51DF74F7C2E69FB">Hanley, Guy A.</mods:namePart>
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<mods:typeOfResource id="F047C4A254F05878AA269CCF604BA20D">text</mods:typeOfResource>
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<mods:title id="EBFDB1664109482077EF4830B14FD685">Insecta Mundi</mods:title>
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<mods:part id="8A738D5C979E86BBEDA4F156A8A8F310">
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<mods:date id="6348A1F87E1782411C7B5B8A0C8BEBDC">2013</mods:date>
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<mods:number id="C6BC6DB1ECB7CC5F1C926A6DA0EAE3AA">2013-06-28</mods:number>
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<mods:number id="F2047A04552D580F002A99732748DB1E">2013</mods:number>
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<mods:number id="88ED73929889AAC10305F9CA45589CC2">312</mods:number>
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<mods:start id="6AEDEA3951E340F1584E9C34597F7686">1</mods:start>
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<mods:identifier id="93E4CF8F9A02BFFB8BAD37B992649F94" type="DOI">10.5281/zenodo.5176513</mods:identifier>
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<treatment id="038F8787FFABFFBBFF7C5996FD9DFCE0" ID-DOI="http://doi.org/10.5281/zenodo.5188187" ID-Zenodo-Dep="5188187" LSID="urn:lsid:plazi:treatment:038F8787FFABFFBBFF7C5996FD9DFCE0" httpUri="http://treatment.plazi.org/id/038F8787FFABFFBBFF7C5996FD9DFCE0" lastPageId="7" lastPageNumber="4" pageId="6" pageNumber="3">
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<subSubSection id="C33C651AFFABFFBAFF7C5996FE75FD20" box="[192,504,707,731]" pageId="6" pageNumber="3" type="nomenclature">
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<paragraph id="8B993691FFABFFBAFF7C5996FE75FD20" blockId="6.[192,504,707,731]" box="[192,504,707,731]" pageId="6" pageNumber="3">
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<heading id="D0D181FDFFABFFBAFF7C5996FE75FD20" bold="true" box="[192,504,707,731]" fontSize="10" level="2" pageId="6" pageNumber="3" reason="3">
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<taxonomicName id="4C264D12FFABFFBAFF7C5996FE75FD20" authority="Duverger" authorityName="Duverger" authorityYear="2001" box="[192,504,707,731]" class="Insecta" family="Coccinellidae" genus="Dilatitibialis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="3" phylum="Arthropoda" rank="genus">
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||
<emphasis id="B952EA83FFABFFBAFF7C5996FE75FD20" bold="true" box="[192,504,707,731]" pageId="6" pageNumber="3">
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<emphasis id="B952EA83FFABFFBAFF7C5996FEFEFD20" bold="true" box="[192,371,707,731]" italics="true" pageId="6" pageNumber="3">Dilatitibialis</emphasis>
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Duverger
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C33C651AFFABFFBAFF7C5856FCF3FCE0" box="[192,894,771,796]" pageId="6" pageNumber="3" type="reference_group">
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<paragraph id="8B993691FFABFFBAFF7C5856FCF3FCE0" blockId="6.[192,894,771,796]" box="[192,894,771,796]" pageId="6" pageNumber="3">
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<treatmentCitationGroup id="AB3611BFFFABFFBAFF7C5856FCF3FCE0" box="[192,894,771,796]" pageId="6" pageNumber="3">
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<treatmentCitation id="0A871080FFABFFBAFF7C5856FDDAFCE0" author="Duverger, C." box="[192,599,771,796]" page="226" pageId="6" pageNumber="3" year="2001">
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<taxonomicName id="4C264D12FFABFFBAFF7C5856FDDAFCE0" authority="Duverger, 2001: 226" authorityName="Duverger" authorityPageNumber="226" authorityYear="2001" box="[192,599,771,796]" class="Insecta" family="Coccinellidae" genus="Dilatitibialis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="3" phylum="Arthropoda" rank="genus">
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<emphasis id="B952EA83FFABFFBAFF7C5856FED7FCE0" box="[192,346,771,795]" italics="true" pageId="6" pageNumber="3">Dilatitibialis</emphasis>
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<bibRefCitation id="EFB74B60FFABFFBAFEDD5851FDDAFCE0" author="Duverger, C." box="[353,599,771,796]" pageId="6" pageNumber="3" pagination="221 - 228" refId="ref56624" refString="Duverger, C. 2001. Contribution a la connaissance des Hyperaspidinae (2 eme note). Bulletin de la Societe linneene, Bordeaux 29: 221 - 228." type="journal article" year="2001">Duverger, 2001: 226</bibRefCitation>
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</taxonomicName>
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||
</treatmentCitation>
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;
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<treatmentCitation id="0A871080FFABFFBAFDDE5856FCD1FCE0" author="Gordon, R. D. & C. Canepari & G. Hanley" box="[610,860,771,795]" page="7" pageId="6" pageNumber="3" year="2013">
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<bibRefCitation id="EFB74B60FFABFFBAFDDE5856FCD1FCE0" author="Gordon, R. D. & C. Canepari & G. Hanley" box="[610,860,771,795]" pageId="6" pageNumber="3" pagination="1 - 150" refId="ref56862" refString="Gordon, R. D., C. Canepari, and G. Hanley. 2013. South American Coccinellidae (Coleoptera), Part XII: New name for Cyra Mulsant, review of Brachiacantha genera, and systematic revision of Cleothera Mulsant, Hinda Mulsant and Serratitibia Gordon and Canepari, new genus. Insecta Mundi 0278: 1 - 150. [This is actually Part XIV]" type="book chapter" year="2013">Gordon et al. 2013: 7</bibRefCitation>
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</treatmentCitation>
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, 8.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="C33C651AFFABFFBAFF7C5816FBBCFCA0" box="[192,1073,835,859]" pageId="6" pageNumber="3" type="type_taxon">
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<paragraph id="8B993691FFABFFBAFF7C5816FBBCFCA0" blockId="6.[192,1073,835,859]" box="[192,1073,835,859]" pageId="6" pageNumber="3">
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<emphasis id="B952EA83FFABFFBAFF7C5816FEE1FCA0" bold="true" box="[192,364,835,859]" pageId="6" pageNumber="3">
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<typeStatus id="549D8833FFABFFBAFF7C5816FE8EFCA0" box="[192,259,835,859]" pageId="6" pageNumber="3">Type</typeStatus>
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species
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</emphasis>
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.
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<taxonomicName id="4C264D12FFABFFBAFEC75816FC84FCA0" authority="Mulsant 1850" authorityName="Mulsant" authorityYear="1850" box="[379,777,835,859]" class="Insecta" family="Coccinellidae" genus="Cleothera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="3" phylum="Arthropoda" rank="species" species="jucunda">
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<emphasis id="B952EA83FFABFFBAFEC75816FDD4FCA0" box="[379,601,835,859]" italics="true" pageId="6" pageNumber="3">Cleothera jucunda</emphasis>
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<bibRefCitation id="EFB74B60FFABFFBAFDDC5816FC84FCA0" author="Mulsant, M. E." box="[608,777,835,859]" pageId="6" pageNumber="3" pagination="1 - 1104" refId="ref57076" refString="Mulsant, M. E. 1850. Species de Coleopteres trimeres securipalpes. Annales des Sciences Physiques et Naturelles, Lyon 2: 1 - 1104." type="journal article" year="1850">Mulsant 1850</bibRefCitation>
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</taxonomicName>
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, by original designation.
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</paragraph>
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</subSubSection>
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<subSubSection id="C33C651AFFABFFBAFF7C58D6FC49FAA0" pageId="6" pageNumber="3" type="description">
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<paragraph id="8B993691FFABFFBAFF7C58D6FC49FAA0" blockId="6.[192,1441,899,1371]" pageId="6" pageNumber="3">
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<emphasis id="B952EA83FFABFFBAFF7C58D6FEEFFC60" bold="true" box="[192,354,899,923]" pageId="6" pageNumber="3">Description</emphasis>
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.
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<taxonomicName id="4C264D12FFABFFBAFECE58D6FDBBFC60" authorityName="Mulsant" authorityYear="1850" box="[370,566,899,923]" class="Insecta" family="Coccinellidae" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="3" phylum="Arthropoda" rank="tribe" tribe="Brachiacanthini">Brachiacanthini</taxonomicName>
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with form variable, round or oval, convex, occasionally elongate. Head usually yellow in male, at least partially brown or black in female. Elytral color variable, but usually dark with pale maculation, or pale with dark maculation, rarely vittate. Antenna with 11 articles, basal article twice as long as wide, antennal insertion exposed. Clypeus with apical margin nearly truncate or distinctly emarginate. Labrum rectangular. Apical maxillary palpomere securiform with sides slightly diverging. Scutellum large, wider than long. Elytral epipleuron narrow or wide, deeply excavated for reception of tibiae. Prosternal process slightly convex, with two carinae. Protibia with narrow, usually arcuate flange, or flange wide, outwardly arcuate (
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<figureCitation id="131D2A14FFABFFBAFC995F36FCE0FB80" box="[805,877,1123,1147]" captionStart="Figures 1-16. 1-7" captionStartId="75.[192,286,1635,1657]" captionTargetBox="[210,1413,206,1586]" captionTargetId="figure-72@75.[192,1440,192,1613]" captionTargetPageId="75" captionText="Figures 1-16. 1-7) Dilatitibialis semicincta. 1) Habitus. 2) Protibia. 3) Phallobase ventral. 4) Phallobase lateral. 5) Enlarged siphonal apex. 6) Sipho. 7) Female genitalia. 8-12) Dilatitibialis lois. 8) habitus. 9) Phallobase ventral. 10) Phallobase lateral. 11) Enlarged siphonal apex. 12) Sipho. 13-16) Dilatitibialis tina. 13) Habitus. 14) Phallobase ventral. 15) Phallobase lateral. 16) Female genitalia." figureDoi="http://doi.org/10.5281/zenodo.5176517" httpUri="https://zenodo.org/record/5176517/files/figure.png" pageId="6" pageNumber="3">Fig. 2</figureCitation>
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). Abdomen with visible primary pores present between ventrites 4 and 5. Tarsal claw with subquadrate basal tooth. Postcoxal line on basal abdominal ventrite incomplete, of
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<taxonomicName id="4C264D12FFABFFBAFE5D5FF1FD55FB40" authority="(Scymnus)" box="[481,728,1187,1211]" class="Insecta" family="Coccinellidae" genus="Scymnus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="3" phylum="Arthropoda" rank="subGenus" subGenus="Scymnus">
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<emphasis id="B952EA83FFABFFBAFE5D5FF1FDC3FB40" box="[481,590,1188,1211]" italics="true" pageId="6" pageNumber="3">Scymnus</emphasis>
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(
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<emphasis id="B952EA83FFABFFBAFDDE5FF1FD42FB40" box="[610,719,1188,1211]" italics="true" pageId="6" pageNumber="3">Scymnus</emphasis>
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)
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</taxonomicName>
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<typeStatus id="549D8833FFABFFBAFD5E5FF1FC97FB47" box="[738,794,1188,1212]" pageId="6" pageNumber="3">type</typeStatus>
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. Fifth abdominal ventrite nearly always without tubercle on each side of median apical emargination, if tubercle present, then without surrounding clump of setae; 6th ventrite with median emargination, lateral angle bearing setal tuft. Male genitalia with asymmetrical basal lobe, paramere variable,
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<emphasis id="B952EA83FFABFFBAFD6C5E51FC80FAE0" box="[720,781,1284,1307]" italics="true" pageId="6" pageNumber="3">Unm</emphasis>
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(paramere unmodified),
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<emphasis id="B952EA83FFABFFBAFB8F5E51FBD1FAE0" box="[1075,1116,1284,1307]" italics="true" pageId="6" pageNumber="3">Psc</emphasis>
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(paramere similar in shape to a scimitar), or
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<emphasis id="B952EA83FFABFFBAFE375E76FE3EFAC0" box="[395,435,1315,1339]" italics="true" pageId="6" pageNumber="3">Pvl</emphasis>
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(Paramere mostly unmodified, but distinctly lobed ventrally). Female genitalia with spermathecal capsule variable but usually short, wide (
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<figureCitation id="131D2A14FFABFFBAFCE35E16FC3AFAA0" box="[863,951,1347,1371]" captionStart="Figures 17-32. 17-21" captionStartId="76.[192,287,1606,1628]" captionTargetBox="[215,1416,202,1589]" captionTargetId="figure-19@76.[192,1440,192,1618]" captionTargetPageId="76" captionText="Figures 17-32. 17-21) Dilatitibialis phyllis. 17) Habitus. 18) Phallobase ventral. 19) Phallobase lateral. 20) Sipho. 21) Enlarged siphonal apex. 22-26) Dilatitibialis cindy. 22) Habitus. 23) Phallobase ventral. 24) Phallobase lateral. 25) Sipho. 26) Enlarged siphonal apex. 27-32) Dilatitibialis norma. 27) Habitus. 28) Phallobase ventral. 29) Phallobase lateral. 30) Enlarged siphonal apex. 31) Sipho. 32) Female genitalia." figureDoi="http://doi.org/10.5281/zenodo.5176519" httpUri="https://zenodo.org/record/5176519/files/figure.png" pageId="6" pageNumber="3">Fig. 32</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C33C651AFFABFFBBFF7C5ED6FD9DFCE0" lastPageId="7" lastPageNumber="4" pageId="6" pageNumber="3" type="discussion">
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<paragraph id="8B993691FFABFFBAFF7C5ED6FD46F960" blockId="6.[192,1441,1411,1916]" pageId="6" pageNumber="3">
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<emphasis id="B952EA83FFABFFBAFF7C5ED6FEB7FA60" bold="true" box="[192,314,1411,1435]" pageId="6" pageNumber="3">Remarks</emphasis>
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. This genus was originally defined by the presence of a distinct protibial flange, a character that has proved nearly useless as a generic character because all degrees of flange development have been observed between
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<taxonomicName id="4C264D12FFABFFBAFE275E96FDBAFA20" authorityName="Duverger" authorityYear="2001" box="[411,567,1475,1499]" class="Insecta" family="Coccinellidae" genus="Dilatitibialis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="3" phylum="Arthropoda" rank="genus">
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<emphasis id="B952EA83FFABFFBAFE275E96FDBAFA20" box="[411,567,1475,1499]" italics="true" pageId="6" pageNumber="3">Dilatitibialis</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C264D12FFABFFBAFDD35E91FB9CFA20" authority="Gordon and Canepari. There" authorityName="Gordon and Canepari. There" box="[623,1041,1475,1499]" class="Globothalamea" genus="Cyrea" kingdom="Chromista" order="Textulariida" pageId="6" pageNumber="3" phylum="Foraminifera" rank="genus">
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<emphasis id="B952EA83FFABFFBAFDD35E91FD39FA20" box="[623,692,1476,1499]" italics="true" pageId="6" pageNumber="3">Cyrea</emphasis>
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Gordon and Canepari. There
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</taxonomicName>
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are certainly two distinct genera, and we have elected to use the presence or absence of primary abdominal pores to distinguish between them. This character varies in degree of development from extremely large, outstanding primary pores between abdominal ventrites 4-5 to pores that are small and may have to be observed from inside the abdomen, but it is always either present or absent. Both
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<taxonomicName id="4C264D12FFABFFBAFCC15D16FC46F9A0" authorityName="Mulsant" authorityYear="1850" box="[893,971,1603,1627]" class="Insecta" family="Coccinellidae" genus="Hinda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="3" phylum="Arthropoda" rank="genus">
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<emphasis id="B952EA83FFABFFBAFCC15D16FC46F9A0" box="[893,971,1603,1627]" italics="true" pageId="6" pageNumber="3">Hinda</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C264D12FFABFFBAFBB65D16FB79F9A0" box="[1034,1268,1603,1627]" class="Insecta" family="Coccinellidae" genus="Serratitibia" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="3" phylum="Arthropoda" rank="species" species="undetermined">
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<emphasis id="B952EA83FFABFFBAFBB65D16FB1AF9A0" box="[1034,1175,1603,1627]" italics="true" pageId="6" pageNumber="3">Serratitibia</emphasis>
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species
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</taxonomicName>
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have primary pores, but both are distinguished from
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<taxonomicName id="4C264D12FFABFFBAFD2C5D36FCA1F980" authorityName="Duverger" authorityYear="2001" box="[656,812,1635,1659]" class="Insecta" family="Coccinellidae" genus="Dilatitibialis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="3" phylum="Arthropoda" rank="genus">
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<emphasis id="B952EA83FFABFFBAFD2C5D36FCA1F980" box="[656,812,1635,1659]" italics="true" pageId="6" pageNumber="3">Dilatitibialis</emphasis>
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</taxonomicName>
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by different male abdominal ventrites and nearly all species having outwardly serrate protibiae.
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</paragraph>
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<paragraph id="8B993691FFABFFBBFF4C5DF6FC1DFDC7" blockId="6.[192,1441,1411,1916]" lastBlockId="7.[192,1441,195,796]" lastPageId="7" lastPageNumber="4" pageId="6" pageNumber="3">
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Groups are difficult to define within
|
||
<taxonomicName id="4C264D12FFABFFBAFD145DF6FCC9F940" authorityName="Duverger" authorityYear="2001" box="[680,836,1699,1723]" class="Insecta" family="Coccinellidae" genus="Dilatitibialis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="3" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B952EA83FFABFFBAFD145DF6FCC9F940" box="[680,836,1699,1723]" italics="true" pageId="6" pageNumber="3">Dilatitibialis</emphasis>
|
||
</taxonomicName>
|
||
, but species are generally arranged on the basis of male genitalia
|
||
<typeStatus id="549D8833FFABFFBAFECB5D91FE22F927" box="[375,431,1732,1756]" pageId="6" pageNumber="3">type</typeStatus>
|
||
. Those species having genitalia whose parameres are not
|
||
<emphasis id="B952EA83FFABFFBAFBC55D91FB2FF920" box="[1145,1186,1732,1755]" italics="true" pageId="6" pageNumber="3">Psc</emphasis>
|
||
shaped and have no unifying characters are placed first. For convenience of reference, this assemblage is called the semicincta group, and contains those species from
|
||
<taxonomicName id="4C264D12FFABFFBAFD2A5C51FC1BF8E0" authority="(Weise)" baseAuthorityName="Weise" box="[662,918,1795,1819]" class="Insecta" family="Coccinellidae" genus="Cleothera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="3" phylum="Arthropoda" rank="species" species="semicincta">
|
||
<emphasis id="B952EA83FFABFFBAFD2A5C51FD27F8E0" box="[662,682,1796,1819]" italics="true" pageId="6" pageNumber="3">D</emphasis>
|
||
.
|
||
<emphasis id="B952EA83FFABFFBAFD045C51FCB5F8E0" box="[696,824,1796,1819]" italics="true" pageId="6" pageNumber="3">semicincta</emphasis>
|
||
(Weise)
|
||
</taxonomicName>
|
||
through
|
||
<taxonomicName id="4C264D12FFABFFBAFBB85C51FBE7F8E0" authority="Canepari & Gordon & Hanley, 2013" authorityName="Canepari & Gordon & Hanley" authorityYear="2013" box="[1028,1130,1795,1819]" class="Insecta" family="Coccinellidae" genus="Dilatitibialis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="3" phylum="Arthropoda" rank="species" species="paula" status="sp. nov.">
|
||
<emphasis id="B952EA83FFABFFBAFBB85C51FB94F8E0" box="[1028,1049,1796,1819]" italics="true" pageId="6" pageNumber="3">D</emphasis>
|
||
.
|
||
<emphasis id="B952EA83FFABFFBAFB995C56FBE7F8E0" box="[1061,1130,1795,1819]" italics="true" pageId="6" pageNumber="3">paula</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A26157F8FFABFFBAFBC45C51FB35F8E0" box="[1144,1208,1796,1819]" pageId="6" pageNumber="3" rank="species">n. sp.</taxonomicNameLabel>
|
||
These species have mostly dissimilar male parameres that defy attempts to further group them. A large group of species, referred to as the mulsanti group, is defined by having
|
||
<emphasis id="B952EA83FFABFFBAFCF15C11FCFBF8A0" box="[845,886,1860,1883]" italics="true" pageId="6" pageNumber="3">Psc</emphasis>
|
||
shaped male parameres (
|
||
<figureCitation id="131D2A14FFABFFBAFB195C16FB75F8A0" box="[1189,1272,1859,1883]" captionStart="Figures 67-85. 67-73" captionStartId="79.[192,286,1608,1630]" captionTargetBox="[209,1426,206,1600]" captionTargetId="figure-15@79.[192,1440,192,1623]" captionTargetPageId="79" captionText="Figures 67-85. 67-73) Dilatitibialis suzannae. 67) Habitus. 68) protibia. 69) Phallobase ventral. 70) Phallobase lateral. 71) Enlarged siphonal apex. 72) Sipho. 73) Female genitalia. 74-79) Dilatitibialis carolinae. 74) Habitus. 75) Phallobase ventral. 76) Phallobase lateral. 77) Enlarged siphonal apex. 78) Sipho. 79) Female genitalia. 80-85) Dilatitibialis annie. 80) Habitus. 81) protibia. 82) Phallobase ventral. 83) Phallobase lateral. 84) Enlarged siphonal apex. 85) Sipho." figureDoi="http://doi.org/10.5281/zenodo.5176525" httpUri="https://zenodo.org/record/5176525/files/figure.png" pageId="6" pageNumber="3">Fig. 76</figureCitation>
|
||
) and contains those species from
|
||
<taxonomicName id="4C264D12FFABFFBAFE1E5C31FD17F880" authority="(Kirsch)" baseAuthorityName="Kirsch" box="[418,666,1891,1915]" class="Insecta" family="Coccinellidae" genus="Dilatitibialis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="3" phylum="Arthropoda" rank="species" species="mulsanti">
|
||
<emphasis id="B952EA83FFABFFBAFE1E5C31FE3AF880" box="[418,439,1892,1915]" italics="true" pageId="6" pageNumber="3">D</emphasis>
|
||
.
|
||
<emphasis id="B952EA83FFABFFBAFE785C36FDBCF880" box="[452,561,1891,1915]" italics="true" pageId="6" pageNumber="3">mulsanti</emphasis>
|
||
(Kirsch)
|
||
</taxonomicName>
|
||
through
|
||
<taxonomicName id="4C264D12FFABFFBAFCB45C31FC16F880" authority="Canepari & Gordon & Hanley, 2013" authorityName="Canepari & Gordon & Hanley" authorityYear="2013" box="[776,923,1891,1915]" class="Insecta" family="Coccinellidae" genus="Dilatitibialis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="3" phylum="Arthropoda" rank="species" species="josephine" status="sp. nov.">
|
||
<emphasis id="B952EA83FFABFFBAFCB45C31FC90F880" box="[776,797,1892,1915]" italics="true" pageId="6" pageNumber="3">D</emphasis>
|
||
.
|
||
<emphasis id="B952EA83FFABFFBAFC965C36FC16F880" box="[810,923,1891,1915]" italics="true" pageId="6" pageNumber="3">josephine</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A26157F8FFABFFBAFC145C31FC65F880" box="[936,1000,1892,1915]" pageId="6" pageNumber="3" rank="species">n. sp.</taxonomicNameLabel>
|
||
A small group of five species from
|
||
<taxonomicName id="4C264D12FFABFFBBFA385C31FE9EFF20" authority="Canepari & Gordon & Hanley, 2013" authorityName="Canepari & Gordon & Hanley" authorityYear="2013" class="Insecta" family="Coccinellidae" genus="Dilatitibialis" higherTaxonomySource="GBIF" kingdom="Animalia" lastPageId="7" lastPageNumber="4" order="Coleoptera" pageId="6" pageNumber="3" phylum="Arthropoda" rank="species" species="thelma" status="sp. nov.">
|
||
<emphasis id="B952EA83FFABFFBAFA385C31FA14F880" box="[1412,1433,1892,1915]" italics="true" pageId="6" pageNumber="3">D</emphasis>
|
||
.
|
||
<emphasis id="B952EA83FFAAFFBBFF7C5B96FE9EFF20" box="[192,275,195,219]" italics="true" pageId="7" pageNumber="4">thelma</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A26157F8FFAAFFBBFE9F5B91FEE9FF20" box="[291,356,196,219]" pageId="7" pageNumber="4" rank="species">n. sp.</taxonomicNameLabel>
|
||
through
|
||
<taxonomicName id="4C264D12FFAAFFBBFE6A5B91FDBFFF20" authority="Canepari & Gordon & Hanley, 2013" authorityName="Canepari & Gordon & Hanley" authorityYear="2013" box="[470,562,195,219]" class="Insecta" family="Coccinellidae" genus="Dilatitibialis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="4" phylum="Arthropoda" rank="species" species="ellen" status="sp. nov.">
|
||
<emphasis id="B952EA83FFAAFFBBFE6A5B91FE66FF20" box="[470,491,196,219]" italics="true" pageId="7" pageNumber="4">D</emphasis>
|
||
.
|
||
<emphasis id="B952EA83FFAAFFBBFE455B96FDBFFF20" box="[505,562,195,219]" italics="true" pageId="7" pageNumber="4">ellen</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A26157F8FFAAFFBBFDFD5B91FD0EFF20" box="[577,643,196,219]" pageId="7" pageNumber="4" rank="species">n. sp.</taxonomicNameLabel>
|
||
forms a compact, uniform group referred to as the thelma group, defined by a basal abdominal ventrite with median tuft of setae; ventrites 5-6 deeply, widely emarginate apically; apical tergite deeply, broadly emarginate with lateral angle large, projecting; and male genitalia with parameres apically emarginate (
|
||
<figureCitation id="131D2A14FFAAFFBBFD3C5A76FD6AFEC0" box="[640,743,291,315]" captionStart="Figures 284-301. 284-289" captionStartId="92.[192,286,1620,1642]" captionTargetBox="[215,1432,205,1583]" captionTargetId="figure-85@92.[192,1440,192,1615]" captionTargetPageId="92" captionText="Figures 284-301. 284-289) Dilatitibialis shannon. 284) Habitus. 285) Phallobase ventral. 286) Phallobase lateral. 287) Enlarged siphonal apex. 288) Sipho. 289) Female genitalia. 290-295) Dilatitibialis sheila. 290) Habitus. 291) Phallobase ventral. 292) Phallobase lateral. 293) Enlarged siphonal apex. 294) Sipho. 295) Female genitalia. 296- 301) Dilatitibialis ethel. 296) Habitus. 297) Phallobase ventral. 298) Phallobase lateral. 299) Enlarged siphonal apex. 300) Sipho. 301) Female genitalia." figureDoi="http://doi.org/10.5281/zenodo.5176551" httpUri="https://zenodo.org/record/5176551/files/figure.png" pageId="7" pageNumber="4">Fig. 292</figureCitation>
|
||
). Another group of two species,
|
||
<taxonomicName id="4C264D12FFAAFFBBFBD85A71FAC7FEC0" authority="(Crotch)" baseAuthorityName="Crotch" box="[1124,1354,291,315]" class="Insecta" family="Coccinellidae" genus="Dilatitibialis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="4" phylum="Arthropoda" rank="species" species="ceciliae">
|
||
<emphasis id="B952EA83FFAAFFBBFBD85A71FBF4FEC0" box="[1124,1145,292,315]" italics="true" pageId="7" pageNumber="4">D</emphasis>
|
||
.
|
||
<emphasis id="B952EA83FFAAFFBBFB3A5A76FB53FEC0" box="[1158,1246,291,315]" italics="true" pageId="7" pageNumber="4">ceciliae</emphasis>
|
||
(Crotch)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C264D12FFAAFFBBFA385A71FE85FEA0" authority="Canepari & Gordon & Hanley, 2013" authorityName="Canepari & Gordon & Hanley" authorityYear="2013" class="Insecta" family="Coccinellidae" genus="Dilatitibialis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="4" phylum="Arthropoda" rank="species" species="elaine" status="sp. nov.">
|
||
<emphasis id="B952EA83FFAAFFBBFA385A71FA14FEC0" box="[1412,1433,292,315]" italics="true" pageId="7" pageNumber="4">D</emphasis>
|
||
.
|
||
<emphasis id="B952EA83FFAAFFBBFF7C5A16FE85FEA0" box="[192,264,323,347]" italics="true" pageId="7" pageNumber="4">elaine</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A26157F8FFAAFFBBFEAA5A11FED5FEA0" box="[278,344,324,347]" pageId="7" pageNumber="4" rank="species">n. sp.</taxonomicNameLabel>
|
||
, the ceciliae group, is defined by abdominal ventrite 5 with a tubercle on each side of apical emargination, tubercle not partially concealed by clump of setae, and male genitalia with basal lobe long, apically bent to one side (
|
||
<figureCitation id="131D2A14FFAAFFBBFDB95AD6FDE2FE60" box="[517,623,387,411]" captionStart="Figures 302-317. 302-306" captionStartId="93.[192,286,1606,1628]" captionTargetBox="[207,1440,204,1559]" captionTargetId="figure-75@93.[192,1441,192,1565]" captionTargetPageId="93" captionText="Figures 302-317. 302-306). Dilatitibialis ellen. 302) Habitus. 303) Phallobase ventral. 304) Phallobase lateral. 305) Sipho. 306) Enlarged siphonal apex. 307-312) Dilatitibialis ceciliae. 307) Habitus. 308) Phallobase ventral. 309) Phallobase lateral. 310) Enlarged siphonal apex. 311) Sipho. 312) Female genitalia. 313-317) Dilatitibialis elaine. 313) Habitus. 314) Phallobase ventral. 315) Phallobase lateral. 316) Sipho. 317) Enlarged siphonal apex." figureDoi="http://doi.org/10.5281/zenodo.5176553" httpUri="https://zenodo.org/record/5176553/files/figure.png" pageId="7" pageNumber="4">Fig. 308</figureCitation>
|
||
). The tuberculate 5th abdominal ventrite of the latter group is a character that would seem to place them in
|
||
<taxonomicName id="4C264D12FFAAFFBBFD795AF6FCDCFE40" authorityName="Canepari & Gordon & Hanley" authorityYear="2013" box="[709,849,419,443]" class="Insecta" family="Coccinellidae" genus="Serratitibia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B952EA83FFAAFFBBFD795AF6FCDCFE40" box="[709,849,419,443]" italics="true" pageId="7" pageNumber="4">Serratitibia</emphasis>
|
||
</taxonomicName>
|
||
(except that the tubercle in
|
||
<taxonomicName id="4C264D12FFAAFFBBFB225AF6FAA7FE40" authorityName="Canepari & Gordon & Hanley" authorityYear="2013" box="[1182,1322,419,443]" class="Insecta" family="Coccinellidae" genus="Serratitibia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B952EA83FFAAFFBBFB225AF6FAA7FE40" box="[1182,1322,419,443]" italics="true" pageId="7" pageNumber="4">Serratitibia</emphasis>
|
||
</taxonomicName>
|
||
is at least partially concealed by a dense clump of setae), but the unusual male genitalia exclude them from that genus. Therefore they are placed in
|
||
<taxonomicName id="4C264D12FFAAFFBBFD3C5AB6FC91FE00" authorityName="Duverger" authorityYear="2001" box="[640,796,483,507]" class="Insecta" family="Coccinellidae" genus="Dilatitibialis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B952EA83FFAAFFBBFD3C5AB6FC91FE00" box="[640,796,483,507]" italics="true" pageId="7" pageNumber="4">Dilatitibialis</emphasis>
|
||
</taxonomicName>
|
||
, seemingly the next best alternative. Those females distinctive enough to be distinguished from other
|
||
<taxonomicName id="4C264D12FFAAFFBBFC975956FBABFDE0" box="[811,1062,515,539]" class="Insecta" family="Coccinellidae" genus="Dilatitibialis" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="4" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="B952EA83FFAAFFBBFC975956FC4AFDE0" box="[811,967,515,539]" italics="true" pageId="7" pageNumber="4">Dilatitibialis</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
are grouped together because, without male genitalia, they cannot be assigned to a group.
|
||
</paragraph>
|
||
<paragraph id="8B993691FFAAFFBBFF4C5916FE63FD80" blockId="7.[192,1441,195,796]" pageId="7" pageNumber="4">Illustrations consist of color photographs of the habitus of nearly all species as well as pen and ink illustrations of genitalia.</paragraph>
|
||
<paragraph id="8B993691FFAAFFBBFF4C59D6FD9DFCE0" blockId="7.[192,1441,195,796]" pageId="7" pageNumber="4">
|
||
This genus was first recognized by
|
||
<bibRefCitation id="EFB74B60FFAAFFBBFD2459D6FCBEFD60" author="El-Ali, A. A." box="[664,819,643,667]" pageId="7" pageNumber="4" refId="ref56655" refString="El-Ali, A. A. 1972. A biosystematic study of Hyperaspini of California with emphasis on the immature stages. Ph. D. dissertation, University of California; Berkeley. 722 p." type="book" year="1972">El-Ali (1972)</bibRefCitation>
|
||
in an unpublished doctoral dissertation. He named it
|
||
<emphasis id="B952EA83FFAAFFBBFF6759F6FE04FD40" box="[219,393,675,699]" italics="true" pageId="7" pageNumber="4">Dilatitibiaspis</emphasis>
|
||
in recognition of the widely flanged protibia found in many species of the genus, and designated
|
||
<taxonomicName id="4C264D12FFAAFFBBFEF45996FD63FD20" authority="(Mulsant)" baseAuthorityName="Mulsant" box="[328,750,707,731]" class="Insecta" family="Coccinellidae" genus="Cleothera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="4" phylum="Arthropoda" rank="species" species="fuscomaculata">
|
||
<emphasis id="B952EA83FFAAFFBBFEF45996FDFCFD20" box="[328,625,707,731]" italics="true" pageId="7" pageNumber="4">Cleothera fuscomaculata</emphasis>
|
||
(Mulsant)
|
||
</taxonomicName>
|
||
as the
|
||
<typeStatus id="549D8833FFAAFFBBFCFD5991FCF8FD27" box="[833,885,708,732]" pageId="7" pageNumber="4">type</typeStatus>
|
||
species.
|
||
<bibRefCitation id="EFB74B60FFAAFFBBFC615991FB2CFD20" author="Duverger, C." box="[989,1185,707,732]" pageId="7" pageNumber="4" pagination="221 - 228" refId="ref56624" refString="Duverger, C. 2001. Contribution a la connaissance des Hyperaspidinae (2 eme note). Bulletin de la Societe linneene, Bordeaux 29: 221 - 228." type="journal article" year="2001">Duverger (2001)</bibRefCitation>
|
||
recognized the validity of El-Ali’s genus, described it, modified the original name to
|
||
<taxonomicName id="4C264D12FFAAFFBBFC7859B6FBEDFD00" authorityName="Duverger" authorityYear="2001" box="[964,1120,739,763]" class="Insecta" family="Coccinellidae" genus="Dilatitibialis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B952EA83FFAAFFBBFC7859B6FBEDFD00" box="[964,1120,739,763]" italics="true" pageId="7" pageNumber="4">Dilatitibialis</emphasis>
|
||
</taxonomicName>
|
||
, and designated
|
||
<taxonomicName id="4C264D12FFAAFFBBFA9159B6FEA9FCE0" authorityName="Mulsant" authorityYear="1850" class="Insecta" family="Coccinellidae" genus="Cleothera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="4" phylum="Arthropoda" rank="species" species="jucunda">
|
||
<emphasis id="B952EA83FFAAFFBBFA9159B6FEA9FCE0" italics="true" pageId="7" pageNumber="4">Cleothera jucunda</emphasis>
|
||
</taxonomicName>
|
||
as the
|
||
<typeStatus id="549D8833FFAAFFBBFEC55851FE20FCE7" box="[377,429,772,796]" pageId="7" pageNumber="4">type</typeStatus>
|
||
species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |