treatments-xml/data/03/8F/87/038F8787C1074356FF6CEFFCFEEBFCCC.xml

<document id="DAB638DA4D86E6F09367AF046C3A100C" ID-CLB-Dataset="5952" ID-DOI="10.11646/zootaxa.3760.2.1" ID-GBIF-Dataset="7ba5f9ac-4317-459f-9829-2975de0a42ff" ID-ISSN="1175-5326" ID-Zenodo-Dep="4908324" ID-ZooBank="C0E4CA6E-6D93-477E-99F6-14FFB652A2A7" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1623095615282" checkinUser="felipe" docAuthor="Michonneau, François &amp; Paulay, Gustav" docDate="2014" docId="038F8787C1074356FF6CEFFCFEEBFCCC" docLanguage="en" docName="zt03760p140.pdf" docOrigin="Zootaxa 3760 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Phyrella thyonoides" docType="treatment" docVersion="7" lastPageNumber="129" masterDocId="FFB6FFFFC110434AFFFBEA5BFFEEFFFB" masterDocTitle="Revision of the genus Phyrella (Holothuroidea: Dendrochirotida) with the description of a new species from Guam" masterLastPageNumber="140" masterPageNumber="101" pageNumber="124" updateTime="1699002177088" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="785C7D33B95DF553D568E5FFCCED3C27">Revision of the genus Phyrella (Holothuroidea: Dendrochirotida) with the description of a new species from Guam</mods:title>
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<heading id="D0D181FDC107435DFF6CEFFCFDDAFA39" bold="true" box="[151,564,1447,1474]" fontSize="11" level="1" pageId="23" pageNumber="124" reason="1">
<taxonomicName id="4C264D12C107435DFF6CEFFCFDDAFA39" ID-CoL="77GG5" authority="(Clark, 1938)" baseAuthorityName="Clark" baseAuthorityYear="1938" box="[151,564,1447,1474]" class="Holothuroidea" family="Phyllophoridae" genus="Phyrella" kingdom="Animalia" order="Dendrochirotida" pageId="23" pageNumber="124" phylum="Echinodermata" rank="species" species="thyonoides">
<emphasis id="B952EA83C107435DFF6CEFFCFDDAFA39" bold="true" box="[151,564,1447,1474]" pageId="23" pageNumber="124">
<emphasis id="B952EA83C107435DFF6CEFFCFE68FA3A" bold="true" box="[151,390,1447,1473]" italics="true" pageId="23" pageNumber="124">Phyrella thyonoides</emphasis>
(
<bibRefCitation id="EFB74B60C107435DFE6DEFF3FDC2FA39" author="Clark, H. L." box="[406,556,1448,1474]" pageId="23" pageNumber="124" refId="ref19549" refString="Clark, H. L. (1938) Memoirs of Comparative Zoology at Harvard College. Vol. LV. Echinoderms from Australia an account of collections made in 1929 and 1938. Museum of Comparative Zoology, Cambridge, MA, USA, 596 pp. + 28 plates." type="book" year="1938">Clark, 1938</bibRefCitation>
)
</emphasis>
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<subSubSection id="C33C651AC107435DFF6CEF91FE69FA18" box="[151,391,1482,1507]" pageId="23" pageNumber="124" type="description">
<paragraph id="8B993691C107435DFF6CEF91FE69FA18" blockId="23.[151,564,1447,1507]" box="[151,391,1482,1507]" pageId="23" pageNumber="124">
<figureCitation id="131D2A14C107435DFF6CEF91FED5FA18" box="[151,315,1482,1507]" captionStart-0="FIGURE 17" captionStart-1="FIGURE 18" captionStart-2="FIGURE 19" captionStart-3="FIGURE 20" captionStart-4="FIGURE 21" captionStart-5="FIGURE 22" captionStartId-0="24.[151,249,1223,1245]" captionStartId-1="24.[151,249,1728,1750]" captionStartId-2="26.[151,249,1941,1963]" captionStartId-3="27.[151,249,887,909]" captionStartId-4="27.[151,249,1690,1712]" captionStartId-5="28.[151,249,624,646]" captionTargetBox-0="[151,1436,393,1202]" captionTargetBox-1="[151,1434,1295,1706]" captionTargetBox-2="[202,1384,198,1910]" captionTargetBox-3="[171,1414,196,863]" captionTargetBox-4="[259,1327,994,1665]" captionTargetBox-5="[151,1436,193,603]" captionTargetId-0="figure-252@24.[151,1436,381,1202]" captionTargetId-1="figure-256@24.[151,1436,1295,1706]" captionTargetId-2="figure-67@26.[195,1396,193,1914]" captionTargetId-3="figure-192@27.[169,1418,193,866]" captionTargetId-4="figure-196@27.[253,1333,989,1668]" captionTargetId-5="figure-505@28.[151,1436,193,603]" captionTargetPageId-0="24" captionTargetPageId-1="24" captionTargetPageId-2="26" captionTargetPageId-3="27" captionTargetPageId-4="27" captionTargetPageId-5="28" captionText-0="FIGURE 17. Holotype of Phyrella thyonoides MCZ HOL1654. Lateral view. Scale bar, 1 cm." captionText-1="FIGURE 18. Live appearance of Phyrella cf. thyonoides A: UF 9598, dorsal view, about 35 mm long; B: UF 9601, lateral view, about 30 mm." captionText-2="FIGURE 19. Ossicle assemblage of Phyrella cf. thyonoides UF 9601. a–e: perforated plates from mid-dorsal body wall; f–i: perforated plates from mid-ventral body wall; j–q: incomplete tables (l) and perforated plates (j,k,m–q) from introvert; r–y: tables (r–w) and perforated plates (x–y) from cloacal region; z: rods from tentacles." captionText-3="FIGURE 20. Ossicle assemblage of Phyrella cf. thyonoides UF 9598. a–e: perforated plates from mid-ventral body wall; f–j: tables (f–i) and perforated plate (j) from cloacal region; k–l: perforated plates from podia that anal teeth." captionText-4="FIGURE 21. Ossicle assemblage of Phyrella cf. thyonoides UF 9414. a–d: perforated plates from podia that flank anal teeth." captionText-5="FIGURE 22. Rosettes of Phyrella cf. thyonoides A: UF 9601, introvert; B: UF 9598, cloacal region. Scale bar, 20 µm." figureDoi-0="http://doi.org/10.5281/zenodo.4908388" figureDoi-1="http://doi.org/10.5281/zenodo.4908392" figureDoi-2="http://doi.org/10.5281/zenodo.4908394" figureDoi-3="http://doi.org/10.5281/zenodo.4908400" figureDoi-4="http://doi.org/10.5281/zenodo.4908404" figureDoi-5="http://doi.org/10.5281/zenodo.4908406" httpUri-0="https://zenodo.org/record/4908388/files/figure.png" httpUri-1="https://zenodo.org/record/4908392/files/figure.png" httpUri-2="https://zenodo.org/record/4908394/files/figure.png" httpUri-3="https://zenodo.org/record/4908400/files/figure.png" httpUri-4="https://zenodo.org/record/4908404/files/figure.png" httpUri-5="https://zenodo.org/record/4908406/files/figure.png" pageId="23" pageNumber="124">Figures 17–22</figureCitation>
; 
<figureCitation id="131D2A14C107435DFEBDEF91FE69FA18" box="[326,391,1482,1507]" captionStart="FIGURE 27" captionStartId="35.[151,249,1914,1936]" captionTargetBox="[177,1410,534,1892]" captionTargetId="figure-173@35.[177,1410,531,1892]" captionTargetPageId="35" captionText="FIGURE 27. Calcareous rings. a–c: Phyrella fragilis (a: UF 4096, b: WAM Z26012, c: WAM Z26013); d–e: Phyrella trapeza, syntypes (d: MCZ HOL1488, thick body wall; e: MCZ HOL1488, thin body wall); f: Phyrella thyonoides, holotype (MCZ HOL1654); g: Phyrella cf. thyonoides (UF 9601); h: Phyrella mookiei, paratype (UF 4770); i: Phyllophorus aculeatus, holotype (USNM 1871)." figureDoi="http://doi.org/10.5281/zenodo.4908418" httpUri="https://zenodo.org/record/4908418/files/figure.png" pageId="23" pageNumber="124">27 f,g</figureCitation>
</paragraph>
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<treatmentCitationGroup id="AB3611BFC107435DFF6CEC49FEBDF9BC" pageId="23" pageNumber="124">
<treatmentCitation id="0A871080C107435DFF6CEC49FD95F9D3" author="Clark, H. L." box="[151,635,1554,1576]" page="494" pageId="23" pageNumber="124" year="1938">
<taxonomicName id="4C264D12C107435DFF6CEC49FD95F9D3" ID-CoL="4H9TH" authority=", Clark, 1938: 492 - 494" authorityName=", Clark" authorityPageNumber="492 - 494" authorityYear="1938" box="[151,635,1554,1576]" class="Holothuroidea" family="Phyllophoridae" genus="Phyllophorus" kingdom="Animalia" order="Dendrochirotida" pageId="23" pageNumber="124" phylum="Echinodermata" rank="species" species="thyonoides">
<emphasis id="B952EA83C107435DFF6CEC49FE7DF9D3" box="[151,403,1554,1576]" italics="true" pageId="23" pageNumber="124">Phyllophorus thyonoides</emphasis>
, 
<materialsCitation id="3B4E3CCCC107435DFE5BEC49FD95F9D3" ID-GBIF-Occurrence="3311708310" box="[416,635,1554,1576]" collectionCode="MCZ" location="Cottesloe Beach" pageId="23" pageNumber="124" specimenCode="HOL1654" specimenCount="1" stateProvince="Western Australia" typeStatus="holotype">
<bibRefCitation id="EFB74B60C107435DFE5BEC49FD95F9D3" author="Clark, H. L." box="[416,635,1554,1576]" pageId="23" pageNumber="124" refId="ref19549" refString="Clark, H. L. (1938) Memoirs of Comparative Zoology at Harvard College. Vol. LV. Echinoderms from Australia an account of collections made in 1929 and 1938. Museum of Comparative Zoology, Cambridge, MA, USA, 596 pp. + 28 plates." type="book" year="1938">Clark, 1938:492–494</bibRefCitation>
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. fig. 48. 
<typeStatus id="549D8833C107435DFD29EC49FCDDF9D3" box="[722,819,1554,1576]" pageId="23" pageNumber="124" type="holotype">Holotype</typeStatus>
, MCZ 
<specimenCode id="DB809EEAC107435DFC81EC49FC0AF9D3" box="[890,996,1554,1576]" collectionCode="HOL" pageId="23" pageNumber="124">HOL1654</specimenCode>
; type locality: 
<collectingRegion id="49E2F873C107435DFB82EC48FADFF9D3" box="[1145,1329,1554,1576]" country="Australia" name="Western Australia" pageId="23" pageNumber="124">Western Australia</collectingRegion>
, 
<location id="8EF9604AC107435DFAC7EC49FEE8F9BC" LSID="urn:lsid:plazi:treatment:038F8787C1074356FF6CEFFCFEEBFCCC:8EF9604AC107435DFAC7EC49FEE8F9BC" name="Cottesloe Beach" pageId="23" pageNumber="124" stateProvince="Western Australia">Cottesloe Beach</location>
(Perth)
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</paragraph>
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<treatmentCitationGroup id="AB3611BFC107435DFF6CEC0BFCF5F99D" box="[151,795,1616,1638]" pageId="23" pageNumber="124">
<treatmentCitation id="0A871080C107435DFF6CEC0BFD20F99D" author="Heding, S. G. &amp; Panning, A." box="[151,718,1616,1638]" page="184" pageId="23" pageNumber="124" year="1954">
<taxonomicName id="4C264D12C107435DFF6CEC0BFD20F99D" ID-CoL="77GG5" authority=": Heding &amp; Panning, 1954: 183 - 184" authorityName="Heding &amp; Panning" authorityPageNumber="183 - 184" authorityYear="1954" box="[151,718,1616,1638]" class="Holothuroidea" family="Phyllophoridae" genus="Phyrella" kingdom="Animalia" order="Dendrochirotida" pageId="23" pageNumber="124" phylum="Echinodermata" rank="species" species="thyonoides">
<emphasis id="B952EA83C107435DFF6CEC0BFE84F99D" box="[151,362,1616,1638]" italics="true" pageId="23" pageNumber="124">Phyrella thyonoides:</emphasis>
<bibRefCitation id="EFB74B60C107435DFE94EC0BFD20F99D" author="Heding, S. G. &amp; Panning, A." box="[367,718,1616,1638]" pageId="23" pageNumber="124" pagination="1 - 209" refId="ref19970" refString="Heding, S. G. &amp; Panning, A. (1954) Phyllophoridae. Eine bearbeitung der polytentaculaten dendrochiroten holothurien des zoologischen museums. in Kopenhagen. Spolia Zoologica Musei Hauniensis, 13, 1 - 209." type="journal article" year="1954">Heding &amp; Panning, 1954:183–184</bibRefCitation>
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, fig. 88
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<emphasis id="B952EA83C107435DFF6CECC9FE68F950" bold="true" box="[151,390,1682,1707]" inLineHeading="true" pageId="23" pageNumber="124" reason="1">Material examined.</emphasis>
<materialsCitation id="3B4E3CCCC107435DFE69ECC9FCF0F934" ID-GBIF-Occurrence="3311708326" collectingDate="1926-07" collectionCode="MCZ" collectorName="L. Glauert." location="Cottesloe Beach" pageId="23" pageNumber="124" specimenCode="HOL1654" specimenCount="1" stateProvince="Western Australia" typeStatus="holotype">
<collectionCode id="ED37AE54C107435DFE69ECC9FE21F950" box="[402,463,1682,1707]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="23" pageNumber="124" type="Museum">MCZ</collectionCode>
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, (
<typeStatus id="549D8833C107435DFD99ECC9FD3EF950" box="[610,720,1682,1707]" pageId="23" pageNumber="124" type="holotype">Holotype</typeStatus>
), 
<collectingRegion id="49E2F873C107435DFD13ECC8FC57F950" box="[744,953,1682,1707]" country="Australia" name="Western Australia" pageId="23" pageNumber="124">Western Australia</collectingRegion>
, 
<location id="8EF9604AC107435DFC31ECC9FB67F950" LSID="urn:lsid:plazi:treatment:038F8787C1074356FF6CEFFCFEEBFCCC:8EF9604AC107435DFC31ECC9FB67F950" box="[970,1161,1682,1707]" name="Cottesloe Beach" pageId="23" pageNumber="124" stateProvince="Western Australia">Cottesloe Beach</location>
, from beach drift, 
<date id="FF981051C107435DFA96ECC9FF3CF934" pageId="23" pageNumber="124" value="1926-07">
<collectingDate id="EFDCE9B9C107435DFA96ECC9FF3CF934" pageId="23" pageNumber="124" value="1926-07">July 1926</collectingDate>
</date>
, coll. 
<collectorName id="26D35347C107435DFEE1ECECFE7BF934" box="[282,405,1718,1743]" pageId="23" pageNumber="124">L. Glauert.</collectorName>
<taxonomicName id="4C264D12C107435DFE65ECEDFD32F935" authority=": UF" authorityName="UF" box="[414,732,1718,1743]" class="Holothuroidea" family="Phyllophoridae" genus="Phyrella" kingdom="Animalia" order="Dendrochirotida" pageId="23" pageNumber="124" phylum="Echinodermata" rank="species" species="thyonoides">
<emphasis id="B952EA83C107435DFE65ECEDFE13F934" box="[414,509,1718,1743]" italics="true" pageId="23" pageNumber="124">Phyrella</emphasis>
cf. 
<emphasis id="B952EA83C107435DFDD6ECEDFD48F934" box="[557,678,1718,1743]" italics="true" pageId="23" pageNumber="124">thyonoides</emphasis>
: UF
</taxonomicName>
9414
</materialsCitation>
, 
<materialsCitation id="3B4E3CCCC107435DFCD0ECECFC7BF934" ID-GBIF-Occurrence="3311708307" box="[811,917,1718,1743]" collectionCode="UF" pageId="23" pageNumber="124" specimenCode="UF 9598" specimenCount="1" stateProvince="Western Australia" typeStatus="holotype">
<specimenCode id="DB809EEAC107435DFCD0ECECFC7FF934" box="[811,913,1718,1743]" collectionCode="UF" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="23" pageNumber="124" type="Museum">UF 9598</specimenCode>
,
</materialsCitation>
<materialsCitation id="3B4E3CCCC107435DFC64ECECFAC7F908" ID-GBIF-Occurrence="3311708323" collectingDate="2009-05" collectionCode="UF" collectorName="Francois Michonneau." elevation="3" latitude="-22.6753" location="Ningaloo station" longLatPrecision="51" longitude="113.684" municipality="Ningaloo Reef" pageId="23" pageNumber="124" specimenCode="UF 9601" specimenCount="1" stateProvince="Western Australia" typeStatus="holotype">
<specimenCode id="DB809EEAC107435DFC64ECECFBEBF934" box="[927,1029,1718,1743]" collectionCode="UF" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="23" pageNumber="124" type="Museum">UF 9601</specimenCode>
, 
<collectingRegion id="49E2F873C107435DFBE9ECECFB0EF934" box="[1042,1248,1718,1743]" country="Australia" name="Western Australia" pageId="23" pageNumber="124">Western Australia</collectingRegion>
, 
<collectingMunicipality id="6BFDACEBC107435DFB15ECEDFA76F934" box="[1262,1432,1718,1743]" pageId="23" pageNumber="124">Ningaloo Reef</collectingMunicipality>
, 
<location id="8EF9604AC107435DFF6CEC81FEBDF908" LSID="urn:lsid:plazi:treatment:038F8787C1074356FF6CEFFCFEEBFCCC:8EF9604AC107435DFF6CEC81FEBDF908" box="[151,339,1754,1779]" latitude="-22.6753" longLatPrecision="51" longitude="113.684" municipality="Ningaloo Reef" name="Ningaloo station" pageId="23" pageNumber="124" stateProvince="Western Australia">Ningaloo station</location>
, 
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<geoCoordinate id="EE125056C107435DFE31EC81FDC9F908" box="[458,551,1754,1779]" degrees="113.684" direction="east" orientation="longitude" pageId="23" pageNumber="124" precision="55" value="113.684">113.684</geoCoordinate>
, 1– 
<quantity id="4CDE9B74C107435DFDB5EC81FD95F909" box="[590,635,1754,1779]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="23" pageNumber="124" unit="m" value="3.0">
<elevation id="000BD1A2C107435DFDB5EC81FD95F909" box="[590,635,1754,1779]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="23" pageNumber="124" unit="m" value="3.0">3 m</elevation>
</quantity>
, under rocks on sand, 
<date id="FF981051C107435DFC8DEC80FC02F908" box="[886,1004,1754,1779]" pageId="23" pageNumber="124" value="2009-05">
<collectingDate id="EFDCE9B9C107435DFC8DEC80FC02F908" box="[886,1004,1754,1779]" pageId="23" pageNumber="124" value="2009-05">May 2009</collectingDate>
</date>
. coll. 
<collectorName id="26D35347C107435DFBD5EC81FAC7F908" box="[1070,1321,1754,1779]" pageId="23" pageNumber="124">François Michonneau.</collectorName>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C33C651AC1074353FF3CECA5FF26FA95" lastPageId="25" lastPageNumber="126" pageId="23" pageNumber="124" type="description">
<paragraph id="8B993691C1074352FF3CECA5FCD5FE9E" blockId="23.[151,1436,1682,2031]" lastBlockId="24.[151,1436,151,357]" lastPageId="24" lastPageNumber="125" pageId="23" pageNumber="124">
<emphasis id="B952EA83C107435DFF3CECA5FE00F8ED" box="[199,494,1790,1815]" inLineHeading="true" pageId="23" pageNumber="124" reason="1">
<emphasis id="B952EA83C107435DFF3CECA5FE09F8EC" bold="true" box="[199,487,1790,1815]" pageId="23" pageNumber="124">
Description of 
<typeStatus id="549D8833C107435DFE7AECA5FE09F8EC" box="[385,487,1790,1815]" pageId="23" pageNumber="124" type="holotype">holotype</typeStatus>
</emphasis>
.
</emphasis>
Specimen in poor condition; 
<quantity id="4CDE9B74C107435DFCAEECA5FC4DF8ED" box="[853,931,1790,1815]" metricMagnitude="-2" metricUnit="m" metricValue="3.3" pageId="23" pageNumber="124" unit="mm" value="33.0">33 mm</quantity>
along dorsum, about 
<quantity id="4CDE9B74C107435DFB50ECA5FB14F8ED" box="[1195,1274,1790,1815]" metricMagnitude="-2" metricUnit="m" metricValue="4.5" pageId="23" pageNumber="124" unit="mm" value="45.0">45 mm</quantity>
along venter, 
<quantity id="4CDE9B74C107435DFF6CED79FF08F8C1" box="[151,230,1826,1851]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="23" pageNumber="124" unit="mm" value="13.0">13 mm</quantity>
wide at mid-body (
<figureCitation id="131D2A14C107435DFE39ED79FDFBF8C0" box="[450,533,1826,1851]" captionStart="FIGURE 17" captionStartId="24.[151,249,1223,1245]" captionTargetBox="[151,1436,393,1202]" captionTargetId="figure-252@24.[151,1436,381,1202]" captionTargetPageId="24" captionText="FIGURE 17. Holotype of Phyrella thyonoides MCZ HOL1654. Lateral view. Scale bar, 1 cm." figureDoi="http://doi.org/10.5281/zenodo.4908388" httpUri="https://zenodo.org/record/4908388/files/figure.png" pageId="23" pageNumber="124">Fig. 17</figureCitation>
). Body wall dried out, very stiff, leathery. Body wall and most organs uniformly dark brown. Introvert retracted; only a small piece remains; 
<quantity id="4CDE9B74C107435DFCC9ED1DFC6FF8A5" box="[818,897,1862,1887]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="23" pageNumber="124" unit="mm" value="12.0">12 mm</quantity>
long. With 16 tentacles of varying sizes, longest about 
<quantity id="4CDE9B74C107435DFF19ED31FEDDF879" box="[226,307,1898,1923]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="23" pageNumber="124" unit="mm" value="13.0">13 mm</quantity>
, shortest about 
<quantity id="4CDE9B74C107435DFE0FED31FDD6F879" box="[500,568,1898,1923]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="23" pageNumber="124" unit="mm" value="1.0">1 mm</quantity>
; arrangement difficult to discern because of the state of contraction and preservation of specimen. Cloacal region dissected out, missing. Calcareous ring 
<quantity id="4CDE9B74C107435DFB97EDD5FB55F85D" box="[1132,1211,1934,1959]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="23" pageNumber="124" unit="mm" value="17.0">17 mm</quantity>
long, 
<quantity id="4CDE9B74C107435DFAEBEDD5FABFF85D" box="[1296,1361,1934,1959]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="23" pageNumber="124" unit="mm" value="8.0">8 mm</quantity>
wide, embedded in a fairly thin and transparent membrane (
<figureCitation id="131D2A14C107435DFCDAEDE9FC91F830" box="[801,895,1970,1995]" captionStart="FIGURE 27" captionStartId="35.[151,249,1914,1936]" captionTargetBox="[177,1410,534,1892]" captionTargetId="figure-173@35.[177,1410,531,1892]" captionTargetPageId="35" captionText="FIGURE 27. Calcareous rings. a–c: Phyrella fragilis (a: UF 4096, b: WAM Z26012, c: WAM Z26013); d–e: Phyrella trapeza, syntypes (d: MCZ HOL1488, thick body wall; e: MCZ HOL1488, thin body wall); f: Phyrella thyonoides, holotype (MCZ HOL1654); g: Phyrella cf. thyonoides (UF 9601); h: Phyrella mookiei, paratype (UF 4770); i: Phyllophorus aculeatus, holotype (USNM 1871)." figureDoi="http://doi.org/10.5281/zenodo.4908418" httpUri="https://zenodo.org/record/4908418/files/figure.png" pageId="23" pageNumber="124">Fig. 27f</figureCitation>
). Anterior margins of radials divided slightly unequally by deep notch. Anterior margins of the inter-radials arrow-head shaped. Inter-radials each composed of 11–13 elements. Radials each composed of 14–18 elements, forming long posterior tails. Radials and inter-radials closely associated along most of their lengths. Single Polian vesicle, 
<quantity id="4CDE9B74C1084352FC4EEAE7FC18FF28" box="[949,1014,188,213]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="24" pageNumber="125" unit="mm" value="4.0">4 mm</quantity>
long, brown and pear-shaped. Stone canal and madreporite embedded in membrane surrounding the calcareous ring. Stone canal 
<quantity id="4CDE9B74C1084352FAE9EA84FABDFF0C" box="[1298,1363,223,248]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="24" pageNumber="125" unit="mm" value="2.0">2 mm</quantity>
long, madreporite about 
<quantity id="4CDE9B74C1084352FE89EB5FFE5DFEE0" box="[370,435,260,285]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="24" pageNumber="125" unit="mm" value="1.0">1 mm</quantity>
in diameter, spherical with convolutions. After multiple attempts in different parts of specimen, no ossicles were found in the body wall or in the tentacles, likely because of corrosion. Clark illustrated and described body wall tables and tentacle supporting rods.
</paragraph>
<caption id="DF596619C1084352FF6CEE9CFBBDFB26" ID-DOI="http://doi.org/10.5281/zenodo.4908388" ID-Zenodo-Dep="4908388" box="[151,1107,1223,1245]" httpUri="https://zenodo.org/record/4908388/files/figure.png" pageId="24" pageNumber="125" startId="24.[151,249,1223,1245]" targetBox="[151,1436,393,1202]" targetPageId="24">
<paragraph id="8B993691C1084352FF6CEE9CFBBDFB26" blockId="24.[151,1107,1223,1245]" box="[151,1107,1223,1245]" pageId="24" pageNumber="125">
<emphasis id="B952EA83C1084352FF6CEE9CFEF1FB27" bold="true" box="[151,287,1223,1245]" pageId="24" pageNumber="125">FIGURE 17.</emphasis>
Holotype of 
<taxonomicName id="4C264D12C1084352FE5EEE9CFCF3FB26" authority="MCZ HOL 1654" authorityName="MCZ HOL" authorityYear="1654" box="[421,797,1223,1245]" class="Holothuroidea" family="Phyllophoridae" genus="Phyrella" kingdom="Animalia" order="Dendrochirotida" pageId="24" pageNumber="125" phylum="Echinodermata" rank="species" species="thyonoides">
<emphasis id="B952EA83C1084352FE5EEE9CFD9EFB26" box="[421,624,1223,1245]" italics="true" pageId="24" pageNumber="125">Phyrella thyonoides</emphasis>
MCZ HOL1654
</taxonomicName>
. Lateral view. Scale bar, 1 cm.
</paragraph>
</caption>
<caption id="DF596619C1084352FF6CEC9BFE8EF90F" ID-DOI="http://doi.org/10.5281/zenodo.4908392" ID-Zenodo-Dep="4908392" httpUri="https://zenodo.org/record/4908392/files/figure.png" pageId="24" pageNumber="125" startId="24.[151,249,1728,1750]" targetBox="[151,1434,1295,1706]" targetPageId="24">
<paragraph id="8B993691C1084352FF6CEC9BFE8EF90F" blockId="24.[151,1435,1728,1780]" pageId="24" pageNumber="125">
<emphasis id="B952EA83C1084352FF6CEC9BFECCF92E" bold="true" box="[151,290,1728,1750]" pageId="24" pageNumber="125">FIGURE 18.</emphasis>
Live appearance of 
<taxonomicName id="4C264D12C1084352FE00EC9BFD05F92D" authorityName="MCZ HOL" authorityYear="1654" box="[507,747,1728,1750]" class="Holothuroidea" family="Phyllophoridae" genus="Phyrella" kingdom="Animalia" order="Dendrochirotida" pageId="24" pageNumber="125" phylum="Echinodermata" rank="species" species="thyonoides">
<emphasis id="B952EA83C1084352FE00EC9BFDBFF92D" box="[507,593,1728,1750]" italics="true" pageId="24" pageNumber="125">Phyrella</emphasis>
cf. 
<emphasis id="B952EA83C1084352FD86EC9BFD05F92D" box="[637,747,1728,1750]" italics="true" pageId="24" pageNumber="125">thyonoides</emphasis>
</taxonomicName>
A: UF 9598, dorsal view, about 35 mm long; B: UF 9601, lateral view, about 30 mm.
</paragraph>
</caption>
<paragraph id="8B993691C1084353FF3CED47FEC8FF0C" blockId="24.[151,1436,1820,2023]" lastBlockId="25.[151,1437,151,2035]" lastPageId="25" lastPageNumber="126" pageId="24" pageNumber="125">
<emphasis id="B952EA83C1084352FF3CED47FBA7F8CE" bold="true" box="[199,1097,1820,1845]" pageId="24" pageNumber="125">
Description of 
<taxonomicName id="4C264D12C1084352FE8CED47FD6FF8CE" authorityName="MCZ HOL" authorityYear="1654" box="[375,641,1820,1845]" class="Holothuroidea" family="Phyllophoridae" genus="Phyrella" kingdom="Animalia" order="Dendrochirotida" pageId="24" pageNumber="125" phylum="Echinodermata" rank="species" species="thyonoides">
<emphasis id="B952EA83C1084352FE8CED47FE37F8CE" bold="true" box="[375,473,1820,1845]" italics="true" pageId="24" pageNumber="125">Phyrella</emphasis>
cf. 
<emphasis id="B952EA83C1084352FDFFED47FD6FF8CE" bold="true" box="[516,641,1820,1845]" italics="true" pageId="24" pageNumber="125">thyonoides</emphasis>
</taxonomicName>
from Ningaloo. External morphology.
</emphasis>
Body wall slightly firmer and more leathery than in other 
<taxonomicName id="4C264D12C1084352FE28ED64FDDCF8A3" authorityName="Heding &amp; Panning" authorityYear="1954" box="[467,562,1855,1880]" class="Holothuroidea" family="Phyllophoridae" genus="Phyrella" kingdom="Animalia" order="Dendrochirotida" pageId="24" pageNumber="125" phylum="Echinodermata" rank="genus">
<emphasis id="B952EA83C1084352FE28ED64FDDCF8A3" box="[467,562,1855,1880]" italics="true" pageId="24" pageNumber="125">Phyrella</emphasis>
</taxonomicName>
. Living animal dark brown, densely covered with tube feet with yellow base color overlain with dark brown transverse lines and with red ring around sucker margin (
<figureCitation id="131D2A14C1084352FB3BED38FAFAF887" box="[1216,1300,1891,1916]" captionStart="FIGURE 18" captionStartId="24.[151,249,1728,1750]" captionTargetBox="[151,1434,1295,1706]" captionTargetId="figure-256@24.[151,1436,1295,1706]" captionTargetPageId="24" captionText="FIGURE 18. Live appearance of Phyrella cf. thyonoides A: UF 9598, dorsal view, about 35 mm long; B: UF 9601, lateral view, about 30 mm." figureDoi="http://doi.org/10.5281/zenodo.4908392" httpUri="https://zenodo.org/record/4908392/files/figure.png" pageId="24" pageNumber="125">Fig. 18</figureCitation>
). Preserved specimens relatively straight 
<quantity id="4CDE9B74C1084352FE14EDDCFD86F864" box="[495,616,1927,1952]" metricMagnitude="-2" metricUnit="m" metricValue="3.25" metricValueMax="3.5" metricValueMin="3.0" pageId="24" pageNumber="125" unit="mm" value="32.5" valueMax="35.0" valueMin="30.0">30–35 mm</quantity>
(excluding introvert and tentacles when extended) and 
<quantity id="4CDE9B74C1084352FAF8EDDCFABCF864" box="[1283,1362,1927,1952]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="24" pageNumber="125" unit="mm" value="13.0">13 mm</quantity>
wide. Introvert short (
<quantity id="4CDE9B74C1084352FEBDEDF0FE63F83F" box="[326,397,1963,1988]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="24" pageNumber="125" unit="mm" value="4.0">4 mm</quantity>
). Introvert and tentacles white, peppered with small, dark brown flecks; base of tentacles also have small light brown blotches. With 15 tentacles in outer circle (about 
<quantity id="4CDE9B74C1084352FC15ED95FBDEF81C" box="[1006,1072,1998,2023]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="24" pageNumber="125" unit="mm" value="3.0">3 mm</quantity>
) and 5 tentacles in inner circle (&lt;
<quantity id="4CDE9B74C1094353FF4CEACCFF12FF4B" box="[183,252,151,176]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="25" pageNumber="126" unit="mm" value="1.0">1 mm</quantity>
), latter aligned with ambulacra (UF 9414 and UF 9601). Oral disc with light brown line around mouth. Cloacal membrane white, with 5 lightly calcified anal teeth, each associated with a few tube feet with minute brown spots.
</paragraph>
<paragraph id="8B993691C1094353FF3CEB58FE8FFE09" blockId="25.[151,1437,151,2035]" pageId="25" pageNumber="126">
<emphasis id="B952EA83C1094353FF3CEB58FD69FEE7" box="[199,647,259,284]" inLineHeading="true" pageId="25" pageNumber="126" reason="1">
<emphasis id="B952EA83C1094353FF3CEB58FE76FEE7" bold="true" box="[199,408,259,284]" pageId="25" pageNumber="126">Internal anatomy</emphasis>
(based on UF 9601).
</emphasis>
Polian vesicle single, light brown with very sparse minute dark spots, 
<quantity id="4CDE9B74C1094353FF6CEB7DFF36FEC5" box="[151,216,294,319]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="25" pageNumber="126" unit="mm" value="5.0">5 mm</quantity>
long. Madreporite and stone canal inconspicuous, embedded in the membrane surrounding the calcareous ring. Stone canal short (&lt;
<quantity id="4CDE9B74C1094353FE3EEB11FDE9FE98" box="[453,519,330,355]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="25" pageNumber="126" unit="mm" value="2.0">2 mm</quantity>
) and narrow (&lt;
<quantity id="4CDE9B74C1094353FD3EEB11FCE9FE98" box="[709,775,330,355]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="25" pageNumber="126" unit="mm" value="1.0">1 mm</quantity>
) embedded in the membrane surrounding the pharyngeal complex. Gonads with two well-developed tufts of unbranched, semi-transparent tubes, with small dark purple spots, up to 
<quantity id="4CDE9B74C1094353FEDFEBC9FE8BFE52" box="[292,357,402,427]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="25" pageNumber="126" unit="mm" value="5.0">5 mm</quantity>
long. Respiratory trees light gray with minute dark purple spots, both extending almost entire length of animal. Specimen eviscerated, but retaining pharyngeal complex, gonads, respiratory trees and proximal end of esophagus.
</paragraph>
<paragraph id="8B993691C1094353FF3CEBA6FD9BFD5E" blockId="25.[151,1437,151,2035]" pageId="25" pageNumber="126">
<emphasis id="B952EA83C1094353FF3CEBA6FD91FDED" box="[199,639,509,534]" inLineHeading="true" pageId="25" pageNumber="126" reason="1">
<emphasis id="B952EA83C1094353FF3CEBA6FE66FDED" bold="true" box="[199,392,509,534]" pageId="25" pageNumber="126">Calcareous ring</emphasis>
(based on UF 9601).
</emphasis>
Calcareous ring embedded in a relatively thin membrane (
<figureCitation id="131D2A14C1094353FAD6EBA6FA61FDED" box="[1325,1423,509,534]" captionStart="FIGURE 27" captionStartId="35.[151,249,1914,1936]" captionTargetBox="[177,1410,534,1892]" captionTargetId="figure-173@35.[177,1410,531,1892]" captionTargetPageId="35" captionText="FIGURE 27. Calcareous rings. a–c: Phyrella fragilis (a: UF 4096, b: WAM Z26012, c: WAM Z26013); d–e: Phyrella trapeza, syntypes (d: MCZ HOL1488, thick body wall; e: MCZ HOL1488, thin body wall); f: Phyrella thyonoides, holotype (MCZ HOL1654); g: Phyrella cf. thyonoides (UF 9601); h: Phyrella mookiei, paratype (UF 4770); i: Phyllophorus aculeatus, holotype (USNM 1871)." figureDoi="http://doi.org/10.5281/zenodo.4908418" httpUri="https://zenodo.org/record/4908418/files/figure.png" pageId="25" pageNumber="126">
Fig. 
<quantity id="4CDE9B74C1094353FA9AEBA6FA61FDED" box="[1377,1423,509,534]" metricMagnitude="-2" metricUnit="kg" metricValue="2.7" pageId="25" pageNumber="126" unit="g" value="27.0">27g</quantity>
</figureCitation>
). Anterior margins of radials divided unequally by a well-formed notch. Anterior margins of inter-radials arrow-head shaped. Radial and inter-radial elements are closely associated along most of their respective lengths. Inter-radials each composed of 5–8 elements. Radials each composed of 14–18 elements. Radials project posteriorly forming long tails that curve anteriorly at their tips.
</paragraph>
<paragraph id="8B993691C1094353FF3CE8F4FF26FA95" blockId="25.[151,1437,151,2035]" pageId="25" pageNumber="126">
<emphasis id="B952EA83C1094353FF3CE8F4FD7BFD33" bold="true" box="[199,661,687,712]" pageId="25" pageNumber="126">
<emphasis id="B952EA83C1094353FF3CE8F4FEC3FD33" box="[199,301,687,712]" inLineHeading="true" pageId="25" pageNumber="126" reason="1">Ossicles.</emphasis>
Dorsal and ventral body wall
</emphasis>
lacking ossicles except in podia. Podia with perforated plates (
<quantity id="4CDE9B74C1094353FA99E8F4FF39FD10" metricMagnitude="2" metricUnit="m" metricValue="1.7" metricValueMax="2.2" metricValueMin="1.2" pageId="25" pageNumber="126" unit="m" value="170.0" valueMax="220.0" valueMin="120.0">
120– 220 
<emphasis id="B952EA83C1094353FF3CE88FFF39FD10" box="[199,215,724,747]" italics="true" pageId="25" pageNumber="126">µ</emphasis>
</quantity>
m long, 
<quantity id="4CDE9B74C1094353FEC8E888FE61FD10" box="[307,399,723,748]" metricMagnitude="1" metricUnit="m" metricValue="4.25" metricValueMax="6.5" metricValueMin="2.0" pageId="25" pageNumber="126" unit="m" value="42.5" valueMax="65.0" valueMin="20.0">
20–65 
<emphasis id="B952EA83C1094353FE84E88FFE61FD10" box="[383,399,724,747]" italics="true" pageId="25" pageNumber="126">µ</emphasis>
</quantity>
m wide), more elongated and with typically larger holes, in particular towards the ends of the plates than in the other species (
<figureCitation id="131D2A14C1094353FDF5E8ACFD6FFCEB" box="[526,641,759,784]" captionStart="FIGURE 19" captionStartId="26.[151,249,1941,1963]" captionTargetBox="[202,1384,198,1910]" captionTargetId="figure-67@26.[195,1396,193,1914]" captionTargetPageId="26" captionText="FIGURE 19. Ossicle assemblage of Phyrella cf. thyonoides UF 9601. a–e: perforated plates from mid-dorsal body wall; f–i: perforated plates from mid-ventral body wall; j–q: incomplete tables (l) and perforated plates (j,k,m–q) from introvert; r–y: tables (r–w) and perforated plates (x–y) from cloacal region; z: rods from tentacles." figureDoi="http://doi.org/10.5281/zenodo.4908394" httpUri="https://zenodo.org/record/4908394/files/figure.png" pageId="25" pageNumber="126">Fig. 19a–i</figureCitation>
; 
<figureCitation id="131D2A14C1094353FD6AE8ACFCE6FCEB" box="[657,776,759,784]" captionStart="FIGURE 20" captionStartId="27.[151,249,887,909]" captionTargetBox="[171,1414,196,863]" captionTargetId="figure-192@27.[169,1418,193,866]" captionTargetPageId="27" captionText="FIGURE 20. Ossicle assemblage of Phyrella cf. thyonoides UF 9598. a–e: perforated plates from mid-ventral body wall; f–j: tables (f–i) and perforated plate (j) from cloacal region; k–l: perforated plates from podia that anal teeth." figureDoi="http://doi.org/10.5281/zenodo.4908400" httpUri="https://zenodo.org/record/4908400/files/figure.png" pageId="25" pageNumber="126">Fig. 20a–e</figureCitation>
). 
<emphasis id="B952EA83C1094353FCDBE8ACFC15FCEB" bold="true" box="[800,1019,759,784]" pageId="25" pageNumber="126">Cloacal body wall</emphasis>
with tables and rosettes (
<quantity id="4CDE9B74C1094353FADFE8ACFA68FCF4" box="[1316,1414,759,784]" metricMagnitude="1" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="25" pageNumber="126" unit="m" value="30.0" valueMax="40.0" valueMin="20.0">
20–40 
<emphasis id="B952EA83C1094353FA8DE8A3FA68FCF4" box="[1398,1414,760,783]" italics="true" pageId="25" pageNumber="126">µ</emphasis>
</quantity>
m long, 
<figureCitation id="131D2A14C1094353FF2DE940FED2FCCF" box="[214,316,795,820]" captionStart="FIGURE 22" captionStartId="28.[151,249,624,646]" captionTargetBox="[151,1436,193,603]" captionTargetId="figure-505@28.[151,1436,193,603]" captionTargetPageId="28" captionText="FIGURE 22. Rosettes of Phyrella cf. thyonoides A: UF 9601, introvert; B: UF 9598, cloacal region. Scale bar, 20 µm." figureDoi="http://doi.org/10.5281/zenodo.4908406" httpUri="https://zenodo.org/record/4908406/files/figure.png" pageId="25" pageNumber="126">Fig. 22B</figureCitation>
). Tables variables; disc 
<quantity id="4CDE9B74C1094353FDBEE940FD40FCC8" box="[581,686,795,820]" metricMagnitude="2" metricUnit="m" metricValue="1.025" metricValueMax="1.15" metricValueMin="0.9" pageId="25" pageNumber="126" unit="m" value="102.5" valueMax="115.0" valueMin="90.0">
90–115 
<emphasis id="B952EA83C1094353FD65E947FD40FCC8" box="[670,686,796,819]" italics="true" pageId="25" pageNumber="126">µ</emphasis>
</quantity>
m in diameter, with either a single circle of 8–11 holes (
<figureCitation id="131D2A14C1094353FAE3E940FA61FCCF" box="[1304,1423,795,820]" captionStart="FIGURE 19" captionStartId="26.[151,249,1941,1963]" captionTargetBox="[202,1384,198,1910]" captionTargetId="figure-67@26.[195,1396,193,1914]" captionTargetPageId="26" captionText="FIGURE 19. Ossicle assemblage of Phyrella cf. thyonoides UF 9601. a–e: perforated plates from mid-dorsal body wall; f–i: perforated plates from mid-ventral body wall; j–q: incomplete tables (l) and perforated plates (j,k,m–q) from introvert; r–y: tables (r–w) and perforated plates (x–y) from cloacal region; z: rods from tentacles." figureDoi="http://doi.org/10.5281/zenodo.4908394" httpUri="https://zenodo.org/record/4908394/files/figure.png" pageId="25" pageNumber="126">Fig. 19u,v</figureCitation>
), sometimes with a few extra holes at the periphery (
<figureCitation id="131D2A14C1094353FD21E965FCDAFCAC" box="[730,820,830,855]" captionStart="FIGURE 19" captionStartId="26.[151,249,1941,1963]" captionTargetBox="[202,1384,198,1910]" captionTargetId="figure-67@26.[195,1396,193,1914]" captionTargetPageId="26" captionText="FIGURE 19. Ossicle assemblage of Phyrella cf. thyonoides UF 9601. a–e: perforated plates from mid-dorsal body wall; f–i: perforated plates from mid-ventral body wall; j–q: incomplete tables (l) and perforated plates (j,k,m–q) from introvert; r–y: tables (r–w) and perforated plates (x–y) from cloacal region; z: rods from tentacles." figureDoi="http://doi.org/10.5281/zenodo.4908394" httpUri="https://zenodo.org/record/4908394/files/figure.png" pageId="25" pageNumber="126">
Fig. 
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</figureCitation>
), or with very numerous holes that are not arranged in any particular order (
<figureCitation id="131D2A14C1094353FE76E939FE07FC80" box="[397,489,866,891]" captionStart="FIGURE 20" captionStartId="27.[151,249,887,909]" captionTargetBox="[171,1414,196,863]" captionTargetId="figure-192@27.[169,1418,193,866]" captionTargetPageId="27" captionText="FIGURE 20. Ossicle assemblage of Phyrella cf. thyonoides UF 9598. a–e: perforated plates from mid-ventral body wall; f–j: tables (f–i) and perforated plate (j) from cloacal region; k–l: perforated plates from podia that anal teeth." figureDoi="http://doi.org/10.5281/zenodo.4908400" httpUri="https://zenodo.org/record/4908400/files/figure.png" pageId="25" pageNumber="126">Fig. 20i</figureCitation>
), with a highly serrated rim; spire (sometimes wanting, 
<figureCitation id="131D2A14C1094353FB81E939FB36FC80" box="[1146,1240,866,891]" captionStart="FIGURE 19" captionStartId="26.[151,249,1941,1963]" captionTargetBox="[202,1384,198,1910]" captionTargetId="figure-67@26.[195,1396,193,1914]" captionTargetPageId="26" captionText="FIGURE 19. Ossicle assemblage of Phyrella cf. thyonoides UF 9601. a–e: perforated plates from mid-dorsal body wall; f–i: perforated plates from mid-ventral body wall; j–q: incomplete tables (l) and perforated plates (j,k,m–q) from introvert; r–y: tables (r–w) and perforated plates (x–y) from cloacal region; z: rods from tentacles." figureDoi="http://doi.org/10.5281/zenodo.4908394" httpUri="https://zenodo.org/record/4908394/files/figure.png" pageId="25" pageNumber="126">Fig. 19x</figureCitation>
) formed by four (rarely five) pillars united by a single cross-beam, surmounted with a crown; crown often incomplete (
<figureCitation id="131D2A14C1094353FAC9E9DDFA60FC64" box="[1330,1422,902,927]" captionStart="FIGURE 19" captionStartId="26.[151,249,1941,1963]" captionTargetBox="[202,1384,198,1910]" captionTargetId="figure-67@26.[195,1396,193,1914]" captionTargetPageId="26" captionText="FIGURE 19. Ossicle assemblage of Phyrella cf. thyonoides UF 9601. a–e: perforated plates from mid-dorsal body wall; f–i: perforated plates from mid-ventral body wall; j–q: incomplete tables (l) and perforated plates (j,k,m–q) from introvert; r–y: tables (r–w) and perforated plates (x–y) from cloacal region; z: rods from tentacles." figureDoi="http://doi.org/10.5281/zenodo.4908394" httpUri="https://zenodo.org/record/4908394/files/figure.png" pageId="25" pageNumber="126">Fig. 19r</figureCitation>
), when complete sometimes with spines (
<figureCitation id="131D2A14C1094353FD99E9F1FD24FC38" box="[610,714,938,963]" captionStart="FIGURE 19" captionStartId="26.[151,249,1941,1963]" captionTargetBox="[202,1384,198,1910]" captionTargetId="figure-67@26.[195,1396,193,1914]" captionTargetPageId="26" captionText="FIGURE 19. Ossicle assemblage of Phyrella cf. thyonoides UF 9601. a–e: perforated plates from mid-dorsal body wall; f–i: perforated plates from mid-ventral body wall; j–q: incomplete tables (l) and perforated plates (j,k,m–q) from introvert; r–y: tables (r–w) and perforated plates (x–y) from cloacal region; z: rods from tentacles." figureDoi="http://doi.org/10.5281/zenodo.4908394" httpUri="https://zenodo.org/record/4908394/files/figure.png" pageId="25" pageNumber="126">Fig. 19w</figureCitation>
), which in some tables that have numerous holes in their disc, extend and connect to the disc margin (
<figureCitation id="131D2A14C1094353FDABE996FD42FC1D" box="[592,684,973,998]" captionStart="FIGURE 20" captionStartId="27.[151,249,887,909]" captionTargetBox="[171,1414,196,863]" captionTargetId="figure-192@27.[169,1418,193,866]" captionTargetPageId="27" captionText="FIGURE 20. Ossicle assemblage of Phyrella cf. thyonoides UF 9598. a–e: perforated plates from mid-ventral body wall; f–j: tables (f–i) and perforated plate (j) from cloacal region; k–l: perforated plates from podia that anal teeth." figureDoi="http://doi.org/10.5281/zenodo.4908400" httpUri="https://zenodo.org/record/4908400/files/figure.png" pageId="25" pageNumber="126">Fig. 20i</figureCitation>
). Podia with perforated plates; less abundant and generally smaller than in mid body wall, 
<quantity id="4CDE9B74C1094353FE5BE9AAFDF7FBF2" box="[416,537,1009,1034]" metricMagnitude="2" metricUnit="m" metricValue="1.375" metricValueMax="1.65" metricValueMin="1.1" pageId="25" pageNumber="126" unit="m" value="137.5" valueMax="165.0" valueMin="110.0">
110–165 
<emphasis id="B952EA83C1094353FDF2E9A9FDF7FBF2" box="[521,537,1010,1033]" italics="true" pageId="25" pageNumber="126">µ</emphasis>
</quantity>
m long, 
<quantity id="4CDE9B74C1094353FD83E9AAFD38FBF2" box="[632,726,1009,1034]" metricMagnitude="1" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="25" pageNumber="126" unit="m" value="50.0" valueMax="60.0" valueMin="40.0">
40–60 
<emphasis id="B952EA83C1094353FD3DE9A9FD38FBF2" box="[710,726,1010,1033]" italics="true" pageId="25" pageNumber="126">µ</emphasis>
</quantity>
m wide (
<figureCitation id="131D2A14C1094353FCC0E9AAFC79FBF1" box="[827,919,1009,1034]" captionStart="FIGURE 20" captionStartId="27.[151,249,887,909]" captionTargetBox="[171,1414,196,863]" captionTargetId="figure-192@27.[169,1418,193,866]" captionTargetPageId="27" captionText="FIGURE 20. Ossicle assemblage of Phyrella cf. thyonoides UF 9598. a–e: perforated plates from mid-ventral body wall; f–j: tables (f–i) and perforated plate (j) from cloacal region; k–l: perforated plates from podia that anal teeth." figureDoi="http://doi.org/10.5281/zenodo.4908400" httpUri="https://zenodo.org/record/4908400/files/figure.png" pageId="25" pageNumber="126">Fig. 20j</figureCitation>
). Podia that flank anal teeth with rosettes and perforated plates (
<figureCitation id="131D2A14C1094353FE9EEE4EFE2EFBD5" box="[357,448,1045,1070]" captionStart="FIGURE 20" captionStartId="27.[151,249,887,909]" captionTargetBox="[171,1414,196,863]" captionTargetId="figure-192@27.[169,1418,193,866]" captionTargetPageId="27" captionText="FIGURE 20. Ossicle assemblage of Phyrella cf. thyonoides UF 9598. a–e: perforated plates from mid-ventral body wall; f–j: tables (f–i) and perforated plate (j) from cloacal region; k–l: perforated plates from podia that anal teeth." figureDoi="http://doi.org/10.5281/zenodo.4908400" httpUri="https://zenodo.org/record/4908400/files/figure.png" pageId="25" pageNumber="126">Fig. 20l</figureCitation>
, 
<figureCitation id="131D2A14C1094353FE37EE4EFDA8FBD5" box="[460,582,1045,1070]" captionStart="FIGURE 21" captionStartId="27.[151,249,1690,1712]" captionTargetBox="[259,1327,994,1665]" captionTargetId="figure-196@27.[253,1333,989,1668]" captionTargetPageId="27" captionText="FIGURE 21. Ossicle assemblage of Phyrella cf. thyonoides UF 9414. a–d: perforated plates from podia that flank anal teeth." figureDoi="http://doi.org/10.5281/zenodo.4908404" httpUri="https://zenodo.org/record/4908404/files/figure.png" pageId="25" pageNumber="126">Fig. 21a–c</figureCitation>
), some of the latter with unusual lateral extensions (
<figureCitation id="131D2A14C1094353FB67EE4EFB11FBD5" box="[1180,1279,1045,1070]" captionStart="FIGURE 20" captionStartId="27.[151,249,887,909]" captionTargetBox="[171,1414,196,863]" captionTargetId="figure-192@27.[169,1418,193,866]" captionTargetPageId="27" captionText="FIGURE 20. Ossicle assemblage of Phyrella cf. thyonoides UF 9598. a–e: perforated plates from mid-ventral body wall; f–j: tables (f–i) and perforated plate (j) from cloacal region; k–l: perforated plates from podia that anal teeth." figureDoi="http://doi.org/10.5281/zenodo.4908400" httpUri="https://zenodo.org/record/4908400/files/figure.png" pageId="25" pageNumber="126">Fig. 20k</figureCitation>
), many plates incomplete with no or few large perforations and only a few small perforations at the extremities (
<figureCitation id="131D2A14C1094353FAD6EE62FA61FBA9" box="[1325,1423,1081,1106]" captionStart="FIGURE 21" captionStartId="27.[151,249,1690,1712]" captionTargetBox="[259,1327,994,1665]" captionTargetId="figure-196@27.[253,1333,989,1668]" captionTargetPageId="27" captionText="FIGURE 21. Ossicle assemblage of Phyrella cf. thyonoides UF 9414. a–d: perforated plates from podia that flank anal teeth." figureDoi="http://doi.org/10.5281/zenodo.4908404" httpUri="https://zenodo.org/record/4908404/files/figure.png" pageId="25" pageNumber="126">Fig. 21d</figureCitation>
). 
<emphasis id="B952EA83C1094353FF6CEE07FEEBFB8E" bold="true" box="[151,261,1116,1141]" pageId="25" pageNumber="126">Introvert</emphasis>
body wall with variable tables, disc 
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60–80 
<emphasis id="B952EA83C1094353FCF2EE06FCF7FB8F" box="[777,793,1117,1140]" italics="true" pageId="25" pageNumber="126">µ</emphasis>
</quantity>
m in diameter, with smooth rim margins, complete and regular with 10–12 holes in the rim, to irregular either with incomplete rim or with additional scattered holes; spire poorly developed or missing for most (
<figureCitation id="131D2A14C1094353FD8FEEFFFD3EFB46" box="[628,720,1188,1213]" captionStart="FIGURE 19" captionStartId="26.[151,249,1941,1963]" captionTargetBox="[202,1384,198,1910]" captionTargetId="figure-67@26.[195,1396,193,1914]" captionTargetPageId="26" captionText="FIGURE 19. Ossicle assemblage of Phyrella cf. thyonoides UF 9601. a–e: perforated plates from mid-dorsal body wall; f–i: perforated plates from mid-ventral body wall; j–q: incomplete tables (l) and perforated plates (j,k,m–q) from introvert; r–y: tables (r–w) and perforated plates (x–y) from cloacal region; z: rods from tentacles." figureDoi="http://doi.org/10.5281/zenodo.4908394" httpUri="https://zenodo.org/record/4908394/files/figure.png" pageId="25" pageNumber="126">Fig. 19l</figureCitation>
). Podia with abundant rosettes, 
<quantity id="4CDE9B74C1094353FBACEEFFFB55FB47" box="[1111,1211,1188,1213]" metricMagnitude="1" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="25" pageNumber="126" unit="m" value="40.0" valueMax="50.0" valueMin="30.0">
30–50 
<emphasis id="B952EA83C1094353FB50EEFEFB55FB47" box="[1195,1211,1189,1212]" italics="true" pageId="25" pageNumber="126">µ</emphasis>
</quantity>
m long (
<figureCitation id="131D2A14C1094353FADDEEFFFA61FB46" box="[1318,1423,1188,1213]" captionStart="FIGURE 22" captionStartId="28.[151,249,624,646]" captionTargetBox="[151,1436,193,603]" captionTargetId="figure-505@28.[151,1436,193,603]" captionTargetPageId="28" captionText="FIGURE 22. Rosettes of Phyrella cf. thyonoides A: UF 9601, introvert; B: UF 9598, cloacal region. Scale bar, 20 µm." figureDoi="http://doi.org/10.5281/zenodo.4908406" httpUri="https://zenodo.org/record/4908406/files/figure.png" pageId="25" pageNumber="126">Fig. 22A</figureCitation>
); perforated plates that range from typical to small, more or less circular (
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60–80 
<emphasis id="B952EA83C1094353FBE7EE92FBC2FB1B" box="[1052,1068,1225,1248]" italics="true" pageId="25" pageNumber="126">µ</emphasis>
</quantity>
m in diameter), with a few large holes in the center and smaller holes toward the periphery (
<figureCitation id="131D2A14C1094353FCC6EEB0FC2AFAFF" box="[829,964,1259,1284]" captionStart="FIGURE 19" captionStartId="26.[151,249,1941,1963]" captionTargetBox="[202,1384,198,1910]" captionTargetId="figure-67@26.[195,1396,193,1914]" captionTargetPageId="26" captionText="FIGURE 19. Ossicle assemblage of Phyrella cf. thyonoides UF 9601. a–e: perforated plates from mid-dorsal body wall; f–i: perforated plates from mid-ventral body wall; j–q: incomplete tables (l) and perforated plates (j,k,m–q) from introvert; r–y: tables (r–w) and perforated plates (x–y) from cloacal region; z: rods from tentacles." figureDoi="http://doi.org/10.5281/zenodo.4908394" httpUri="https://zenodo.org/record/4908394/files/figure.png" pageId="25" pageNumber="126">
Fig. 
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–q
</figureCitation>
). 
<emphasis id="B952EA83C1094353FC23EEB0FBA9FAFF" bold="true" box="[984,1095,1259,1284]" pageId="25" pageNumber="126">Tentacles</emphasis>
with two sizes of rods. Large rods (
<quantity id="4CDE9B74C1094353FF2EEF54FEA0FADC" box="[213,334,1295,1320]" metricMagnitude="2" metricUnit="m" metricValue="1.9" metricValueMax="2.7" metricValueMin="1.1" pageId="25" pageNumber="126" unit="m" value="190.0" valueMax="270.0" valueMin="110.0">
110–270 
<emphasis id="B952EA83C1094353FEC5EF4BFEA0FADC" box="[318,334,1296,1319]" italics="true" pageId="25" pageNumber="126">µ</emphasis>
</quantity>
m) with one or a few holes at the ends. Small rods (
<quantity id="4CDE9B74C1094353FC73EF54FC1AFADC" box="[904,1012,1295,1320]" metricMagnitude="1" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="25" pageNumber="126" unit="m" value="75.0" valueMax="100.0" valueMin="50.0">
50–100 
<emphasis id="B952EA83C1094353FC1FEF4BFC1AFADC" box="[996,1012,1296,1319]" italics="true" pageId="25" pageNumber="126">µ</emphasis>
</quantity>
m) generally with no holes at the ends (
<figureCitation id="131D2A14C1094353FF64EF68FF11FAB7" box="[159,255,1331,1356]" captionStart="FIGURE 19" captionStartId="26.[151,249,1941,1963]" captionTargetBox="[202,1384,198,1910]" captionTargetId="figure-67@26.[195,1396,193,1914]" captionTargetPageId="26" captionText="FIGURE 19. Ossicle assemblage of Phyrella cf. thyonoides UF 9601. a–e: perforated plates from mid-dorsal body wall; f–i: perforated plates from mid-ventral body wall; j–q: incomplete tables (l) and perforated plates (j,k,m–q) from introvert; r–y: tables (r–w) and perforated plates (x–y) from cloacal region; z: rods from tentacles." figureDoi="http://doi.org/10.5281/zenodo.4908394" httpUri="https://zenodo.org/record/4908394/files/figure.png" pageId="25" pageNumber="126">Fig. 19z</figureCitation>
). No ossicles observed in cloacal, longitudinal and retractor muscles; cloacal wall; gonads; or respiratory tree.
</paragraph>
</subSubSection>
<subSubSection id="C33C651AC1094351FF3CEF21FCD2F833" lastPageId="27" lastPageNumber="128" pageId="25" pageNumber="126" type="discussion">
<paragraph id="8B993691C1094353FF3CEF21FDE9F89F" blockId="25.[151,1437,151,2035]" pageId="25" pageNumber="126">
<emphasis id="B952EA83C1094353FF3CEF21FED5FA68" bold="true" box="[199,315,1402,1427]" inLineHeading="true" pageId="25" pageNumber="126" reason="1">Remarks.</emphasis>
In the original description of 
<taxonomicName id="4C264D12C1094353FD62EF21FC24FA68" authority=", Clark" authorityName="Clark" box="[665,970,1402,1427]" class="Holothuroidea" family="Phyllophoridae" genus="Phyrella" kingdom="Animalia" order="Dendrochirotida" pageId="25" pageNumber="126" phylum="Echinodermata" rank="species" species="thyonoides">
<emphasis id="B952EA83C1094353FD62EF21FC95FA68" box="[665,891,1402,1427]" italics="true" pageId="25" pageNumber="126">Phyrella thyonoides</emphasis>
, Clark
</taxonomicName>
only illustrates one table from the body wall and three rods from the tentacles. Overall, the description he provides and the 
<typeStatus id="549D8833C1094353FBA7EFC5FB50FA4C" box="[1116,1214,1438,1463]" pageId="25" pageNumber="126" type="holotype">holotype</typeStatus>
fit reasonably well with our new material from Ningaloo, but there are some differences. Thus, in Clark's description, the ends of the large rods in the tentacles have numerous perforations and are spiny; in our specimens, the large rods have only a few holes at their ends and are not spiny. In our specimens, we did not observe any tables in the mid-dorsal or midventral body wall, whereas Clark noted that the tables were “rather scattered, but by no means rare” but did not indicate in which part of the animal he observed them. The table he illustrates is relatively similar to the ones we observed in the anal region of our specimens. Unfortunately, the poor condition of the 
<typeStatus id="549D8833C1094353FB8BEC2FFB3CF976" box="[1136,1234,1652,1677]" pageId="25" pageNumber="126" type="holotype">holotype</typeStatus>
and the complete corrosion of the ossicles do not allow for an adequate comparison with the Ningaloo material. A full evaluation of 
<taxonomicName id="4C264D12C1094353FF6CECE6FEDFF92E" box="[151,305,1724,1749]" class="Holothuroidea" family="Phyllophoridae" genus="Phyrella" kingdom="Animalia" order="Dendrochirotida" pageId="25" pageNumber="126" phylum="Echinodermata" rank="species" species="thyonoides">
<emphasis id="B952EA83C1094353FF6CECE6FEDFF92E" box="[151,305,1724,1749]" italics="true" pageId="25" pageNumber="126">P. thyonoides</emphasis>
</taxonomicName>
and its relation to the Ningaloo population must await collection of new topotypical material. Surprisingly, none of the specimens loaned to us by the Western Australian Museum identified as 
<taxonomicName id="4C264D12C1094353FB05ECBAFA7BF903" box="[1278,1429,1759,1784]" class="Holothuroidea" family="Phyllophoridae" genus="Phyrella" kingdom="Animalia" order="Dendrochirotida" pageId="25" pageNumber="126" phylum="Echinodermata" rank="species" species="thyonoides">
<emphasis id="B952EA83C1094353FB05ECBAFA7BF903" box="[1278,1429,1759,1784]" italics="true" pageId="25" pageNumber="126">P. thyonoides</emphasis>
</taxonomicName>
, turned out to belong to this species or to 
<emphasis id="B952EA83C1094353FD8AED58FC99F8E7" box="[625,887,1795,1820]" italics="true" pageId="25" pageNumber="126">
<taxonomicName id="4C264D12C1094353FD8AED58FD3AF8E7" authorityName="Heding &amp; Panning" authorityYear="1954" box="[625,724,1795,1820]" class="Holothuroidea" family="Phyllophoridae" genus="Phyrella" kingdom="Animalia" order="Dendrochirotida" pageId="25" pageNumber="126" phylum="Echinodermata" rank="genus">Phyrella</taxonomicName>
. 
<taxonomicName id="4C264D12C1094353FD1AED5FFC99F8E7" box="[737,887,1795,1820]" class="Holothuroidea" family="Phyllophoridae" genus="Phyrella" kingdom="Animalia" order="Dendrochirotida" pageId="25" pageNumber="126" phylum="Echinodermata" rank="species" species="thyonoides">P. thyonoides</taxonomicName>
</emphasis>
can be distinguished from other 
<taxonomicName id="4C264D12C1094353FB0FED58FABDF8E7" authorityName="Heding &amp; Panning" authorityYear="1954" box="[1268,1363,1795,1820]" class="Holothuroidea" family="Phyllophoridae" genus="Phyrella" kingdom="Animalia" order="Dendrochirotida" pageId="25" pageNumber="126" phylum="Echinodermata" rank="genus">
<emphasis id="B952EA83C1094353FB0FED58FABDF8E7" box="[1268,1363,1795,1820]" italics="true" pageId="25" pageNumber="126">Phyrella</emphasis>
</taxonomicName>
by its thick, leathery, dark brown body wall; the elongated perforated plates with large perforations, and the circular perforated plates in the introvert.
</paragraph>
<paragraph id="8B993691C1094351FF3CED35FCD2F833" blockId="25.[151,1437,151,2035]" lastBlockId="27.[151,1436,1751,1992]" lastPageId="27" lastPageNumber="128" pageId="25" pageNumber="126">
Recently, 
<bibRefCitation id="EFB74B60C1094353FEC1ED35FDBDF87C" author="O'Loughlin, P. M. &amp; Barmos, S. &amp; VandenSpiegel, D." box="[314,595,1902,1927]" pageId="25" pageNumber="126" pagination="269 - 308" refId="ref20475" refString="O'Loughlin, P. M., Barmos, S. &amp; VandenSpiegel, D. (2012) The phyllophorid sea cucumbers of southern Australia (Echinodermata: Holothuroidea: Dendrochirotida: Phyllophoridae). Memoirs of Museum Victoria, 308, 269 - 308." type="journal article" year="2012">
O'Loughlin 
<emphasis id="B952EA83C1094353FE38ED2BFE11F87C" box="[451,511,1902,1927]" italics="true" pageId="25" pageNumber="126">et al.</emphasis>
(2012)
</bibRefCitation>
proposed to transfer 
<taxonomicName id="4C264D12C1094353FCB6ED35FBC1F87C" box="[845,1071,1902,1927]" class="Holothuroidea" family="Phyllophoridae" genus="Phyrella" kingdom="Animalia" order="Dendrochirotida" pageId="25" pageNumber="126" phylum="Echinodermata" rank="species" species="thyonoides">
<emphasis id="B952EA83C1094353FCB6ED35FBC1F87C" box="[845,1071,1902,1927]" italics="true" pageId="25" pageNumber="126">Phyrella thyonoides</emphasis>
</taxonomicName>
to 
<emphasis id="B952EA83C1094353FBA2ED35FAEEF87C" box="[1113,1280,1902,1927]" italics="true" pageId="25" pageNumber="126">Phyllophorella</emphasis>
based on the number of tentacles and the shape of the calcareous ring for the specimens they examined, however most of the specimens they report upon do not appear to pertain to 
<taxonomicName id="4C264D12C1094353FCADEDECFC1AF834" box="[854,1012,1974,1999]" class="Holothuroidea" family="Phyllophoridae" genus="Phyrella" kingdom="Animalia" order="Dendrochirotida" pageId="25" pageNumber="126" phylum="Echinodermata" rank="species" species="thyonoides">
<emphasis id="B952EA83C1094353FCADEDECFC1AF834" box="[854,1012,1974,1999]" italics="true" pageId="25" pageNumber="126">P. thyonoides</emphasis>
</taxonomicName>
, but to an undescribed species of 
<emphasis id="B952EA83C1094353FF6CED81FED0F808" box="[151,318,2010,2035]" italics="true" pageId="25" pageNumber="126">Phyllophorella</emphasis>
. The tables illustrated (
<bibRefCitation id="EFB74B60C1094353FDBEED81FCBFF808" author="O'Loughlin, P. M. &amp; Barmos, S. &amp; VandenSpiegel, D." box="[581,849,2010,2035]" pageId="25" pageNumber="126" pagination="269 - 308" refId="ref20475" refString="O'Loughlin, P. M., Barmos, S. &amp; VandenSpiegel, D. (2012) The phyllophorid sea cucumbers of southern Australia (Echinodermata: Holothuroidea: Dendrochirotida: Phyllophoridae). Memoirs of Museum Victoria, 308, 269 - 308." type="journal article" year="2012">
O'Loughlin 
<emphasis id="B952EA83C1094353FD35ED80FCE8F808" box="[718,774,2010,2035]" italics="true" pageId="25" pageNumber="126">et al.</emphasis>
, 2012
</bibRefCitation>
: 
<figureCitation id="131D2A14C1094353FCAFED81FC4AF808" box="[852,932,2010,2035]" captionStart="FIGURE 10" captionStartId="18.[151,249,1358,1380]" captionTargetBox="[151,1436,373,1337]" captionTargetId="figure-360@18.[151,1436,373,1337]" captionTargetPageId="18" captionText="FIGURE 10. Phyrella mookiei in situ, Guam. The animal is buried in sediment; the oral disc with extended tentacles is visible. Oral disc diameter approximately 1 cm in diameter. Photograph by David Burdick." figureDoi="http://doi.org/10.5281/zenodo.4908360" httpUri="https://zenodo.org/record/4908360/files/figure.png" pageId="25" pageNumber="126">Fig. 10</figureCitation>
) with marginal teeth are unlike those found in any 
<taxonomicName id="4C264D12C10B4351FF1EEC8CFEAAF90B" authorityName="Heding &amp; Panning" authorityYear="1954" box="[229,324,1751,1776]" class="Holothuroidea" family="Phyllophoridae" genus="Phyrella" kingdom="Animalia" order="Dendrochirotida" pageId="27" pageNumber="128" phylum="Echinodermata" rank="genus">
<emphasis id="B952EA83C10B4351FF1EEC8CFEAAF90B" box="[229,324,1751,1776]" italics="true" pageId="27" pageNumber="128">Phyrella</emphasis>
</taxonomicName>
. The general appearance of the animal and shape of the calcareous ring illustrated (
<bibRefCitation id="EFB74B60C10B4351FB06EC8CFF10F8EF" author="O'Loughlin, P. M. &amp; Barmos, S. &amp; VandenSpiegel, D." pageId="27" pageNumber="128" pagination="269 - 308" refId="ref20475" refString="O'Loughlin, P. M., Barmos, S. &amp; VandenSpiegel, D. (2012) The phyllophorid sea cucumbers of southern Australia (Echinodermata: Holothuroidea: Dendrochirotida: Phyllophoridae). Memoirs of Museum Victoria, 308, 269 - 308." type="journal article" year="2012">
O'Loughlin 
<emphasis id="B952EA83C10B4351FA73EC83FF5AF8EF" italics="true" pageId="27" pageNumber="128">et al.</emphasis>
, 2012
</bibRefCitation>
: 
<figureCitation id="131D2A14C10B4351FEFAECA0FEADF8EF" box="[257,323,1787,1812]" captionStart="FIGURE 9" captionStartId="16.[151,249,1890,1912]" captionTargetBox="[340,1247,1531,1869]" captionTargetId="figure-112@16.[340,1247,1531,1869]" captionTargetPageId="16" captionText="FIGURE 9. Rosettes from cloacal region (A) and introvert (B) of Phyrella fragilis UF 4096. Scale bar, 20 µm." figureDoi="http://doi.org/10.5281/zenodo.4908358" httpUri="https://zenodo.org/record/4908358/files/figure.png" pageId="27" pageNumber="128">Fig. 9</figureCitation>
) also do not match 
<taxonomicName id="4C264D12C10B4351FDE6ECA0FD92F8EF" authorityName="Heding &amp; Panning" authorityYear="1954" box="[541,636,1787,1812]" class="Holothuroidea" family="Phyllophoridae" genus="Phyrella" kingdom="Animalia" order="Dendrochirotida" pageId="27" pageNumber="128" phylum="Echinodermata" rank="genus">
<emphasis id="B952EA83C10B4351FDE6ECA0FD92F8EF" box="[541,636,1787,1812]" italics="true" pageId="27" pageNumber="128">Phyrella</emphasis>
</taxonomicName>
. 
<bibRefCitation id="EFB74B60C10B4351FD71ECA0FC73F8EF" author="O'Loughlin, P. M. &amp; Barmos, S. &amp; VandenSpiegel, D." box="[650,925,1787,1812]" pageId="27" pageNumber="128" pagination="269 - 308" refId="ref20475" refString="O'Loughlin, P. M., Barmos, S. &amp; VandenSpiegel, D. (2012) The phyllophorid sea cucumbers of southern Australia (Echinodermata: Holothuroidea: Dendrochirotida: Phyllophoridae). Memoirs of Museum Victoria, 308, 269 - 308." type="journal article" year="2012">
O’Loughlin 
<emphasis id="B952EA83C10B4351FCEFECA7FCA2F8EF" box="[788,844,1787,1812]" italics="true" pageId="27" pageNumber="128">et al.</emphasis>
(2012)
</bibRefCitation>
note that there are single posterior extensions on the calcareous ring in this species (and other 
<emphasis id="B952EA83C10B4351FD55ED44FCBBF8C3" box="[686,853,1823,1848]" italics="true" pageId="27" pageNumber="128">Phyllophorella</emphasis>
) arising either from radial or jointly from radial and inter-radial elements; this contrasts with the clearly separate series of plates that arise from the fragmentation of both inter-radial and radial elements of all 
<taxonomicName id="4C264D12C10B4351FD61ED3CFCB1F87B" box="[666,863,1895,1920]" class="Holothuroidea" family="Phyllophoridae" genus="Phyrella" kingdom="Animalia" order="Dendrochirotida" pageId="27" pageNumber="128" phylum="Echinodermata" rank="species" species="undetermined">
<emphasis id="B952EA83C10B4351FD61ED3CFD17F87B" box="[666,761,1895,1920]" italics="true" pageId="27" pageNumber="128">Phyrella</emphasis>
species.
</taxonomicName>
One of the specimens examined by the authors (WAM Z31837) was sequenced and matches a 
<taxonomicName id="4C264D12C10B4351FD46EDD0FC5EF85F" box="[701,944,1931,1956]" class="Holothuroidea" family="Phyllophoridae" genus="Phyllophorus" kingdom="Animalia" order="Dendrochirotida" pageId="27" pageNumber="128" phylum="Echinodermata" rank="species" species="undetermined">
<emphasis id="B952EA83C10B4351FD46EDD0FCBAF85F" box="[701,852,1931,1956]" italics="true" pageId="27" pageNumber="128">Phyllophorus</emphasis>
species
</taxonomicName>
we collected along with 
<taxonomicName id="4C264D12C10B4351FB27EDD7FA9AF85F" box="[1244,1396,1931,1956]" class="Holothuroidea" family="Phyllophoridae" genus="Phyrella" kingdom="Animalia" order="Dendrochirotida" pageId="27" pageNumber="128" phylum="Echinodermata" rank="species" species="thyonoides">
<emphasis id="B952EA83C10B4351FB27EDD7FA9AF85F" box="[1244,1396,1931,1956]" italics="true" pageId="27" pageNumber="128">P. thyonoides</emphasis>
</taxonomicName>
, in Ningaloo, 
<collectingRegion id="49E2F873C10B4351FEF6EDEBFE38F833" box="[269,470,1967,1992]" country="Australia" name="Western Australia" pageId="27" pageNumber="128">Western Australia</collectingRegion>
(
<taxonomicName id="4C264D12C10B4351FE1DEDF4FD57F833" box="[486,697,1967,1992]" class="Holothuroidea" family="Phyllophoridae" genus="Phyllophorus" kingdom="Animalia" order="Dendrochirotida" pageId="27" pageNumber="128" phylum="Echinodermata" rank="species" species="undefined-1">
<emphasis id="B952EA83C10B4351FE1DEDF4FD93F833" box="[486,637,1967,1992]" italics="true" pageId="27" pageNumber="128">Phyllophorus</emphasis>
sp. 
<quantity id="4CDE9B74C10B4351FD50EDF4FD57F833" box="[683,697,1967,1992]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="27" pageNumber="128" unit="in" value="1.0">1</quantity>
</taxonomicName>
in 
<figureCitation id="131D2A14C10B4351FD26EDF4FCDEF833" box="[733,816,1967,1992]" captionStart="FIGURE 28" captionStartId="37.[151,249,1672,1694]" captionTargetBox="[196,1410,344,1609]" captionTargetId="graphics-127@37.[198,981,352,1598]" captionTargetPageId="37" captionText="FIGURE 28. Maximum-likelihood phylogram of some of the species of the genus Phyrella (estimated using RAxML 8.0.1). Bootstrap values based on 500 rapid replicates are reported above the edges. Bootstrap values within species have been removed for clarity. Tree rooted using Holothuria impatiens (not shown). Tree deposited in TreeBASE http://purl.org/phylo/treebase/phylows/study/TB2:S15218." figureDoi="http://doi.org/10.5281/zenodo.4908420" httpUri="https://zenodo.org/record/4908420/files/figure.png" pageId="27" pageNumber="128">Fig. 28</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF596619C10A4350FF6CEDCEFC0AF813" ID-DOI="http://doi.org/10.5281/zenodo.4908394" ID-Zenodo-Dep="4908394" httpUri="https://zenodo.org/record/4908394/files/figure.png" pageId="26" pageNumber="127" startId="26.[151,249,1941,1963]" targetBox="[202,1384,198,1910]" targetPageId="26">
<paragraph id="8B993691C10A4350FF6CEDCEFC0AF813" blockId="26.[151,1436,1940,2024]" pageId="26" pageNumber="127">
<emphasis id="B952EA83C10A4350FF6CEDCEFECEF851" bold="true" box="[151,288,1941,1963]" pageId="26" pageNumber="127">FIGURE 19.</emphasis>
Ossicle assemblage of 
<taxonomicName id="4C264D12C10A4350FDEFEDCFFCC7F851" authority="UF" authorityName="UF" box="[532,809,1940,1962]" class="Holothuroidea" family="Phyllophoridae" genus="Phyrella" kingdom="Animalia" order="Dendrochirotida" pageId="26" pageNumber="127" phylum="Echinodermata" rank="species" species="thyonoides">
<emphasis id="B952EA83C10A4350FDEFEDCFFD84F851" box="[532,618,1940,1962]" italics="true" pageId="26" pageNumber="127">Phyrella</emphasis>
cf. 
<emphasis id="B952EA83C10A4350FD6FEDCFFCECF851" box="[660,770,1940,1962]" italics="true" pageId="26" pageNumber="127">thyonoides</emphasis>
UF
</taxonomicName>
9601. a–e: perforated plates from mid-dorsal body wall; f–i: perforated plates from mid-ventral body wall; j–q: incomplete tables (l) and perforated plates (j,k,m–q) from introvert; r–y: tables (r–w) and perforated plates (x–y) from cloacal region; z: rods from tentacles.
</paragraph>
</caption>
<caption id="DF596619C10B4351FF6CE92CFB58FC50" ID-DOI="http://doi.org/10.5281/zenodo.4908400" ID-Zenodo-Dep="4908400" httpUri="https://zenodo.org/record/4908400/files/figure.png" pageId="27" pageNumber="128" startId="27.[151,249,887,909]" targetBox="[171,1414,196,863]" targetPageId="27">
<paragraph id="8B993691C10B4351FF6CE92CFB58FC50" blockId="27.[151,1436,887,939]" pageId="27" pageNumber="128">
<emphasis id="B952EA83C10B4351FF6CE92CFECEFC77" bold="true" box="[151,288,887,909]" pageId="27" pageNumber="128">FIGURE 20.</emphasis>
Ossicle assemblage of 
<taxonomicName id="4C264D12C10B4351FDEAE92CFCCBFC77" authority="UF" authorityName="UF" box="[529,805,887,909]" class="Holothuroidea" family="Phyllophoridae" genus="Phyrella" kingdom="Animalia" order="Dendrochirotida" pageId="27" pageNumber="128" phylum="Echinodermata" rank="species" species="thyonoides">
<emphasis id="B952EA83C10B4351FDEAE92CFD89FC76" box="[529,615,887,909]" italics="true" pageId="27" pageNumber="128">Phyrella</emphasis>
cf. 
<emphasis id="B952EA83C10B4351FD6BE92CFD10FC76" box="[656,766,887,909]" italics="true" pageId="27" pageNumber="128">thyonoides</emphasis>
UF
</taxonomicName>
9598. a–e: perforated plates from mid-ventral body wall; f–j: tables (f–i) and perforated plate (j) from cloacal region; k–l: perforated plates from podia that anal teeth.
</paragraph>
</caption>
<caption id="DF596619C10B4351FF6CECC1FA64F94B" ID-DOI="http://doi.org/10.5281/zenodo.4908404" ID-Zenodo-Dep="4908404" box="[151,1418,1690,1712]" httpUri="https://zenodo.org/record/4908404/files/figure.png" pageId="27" pageNumber="128" startId="27.[151,249,1690,1712]" targetBox="[259,1327,994,1665]" targetPageId="27">
<paragraph id="8B993691C10B4351FF6CECC1FA64F94B" blockId="27.[151,1418,1690,1712]" box="[151,1418,1690,1712]" pageId="27" pageNumber="128">
<emphasis id="B952EA83C10B4351FF6CECC1FEF1F954" bold="true" box="[151,287,1690,1712]" pageId="27" pageNumber="128">FIGURE 21.</emphasis>
Ossicle assemblage of 
<taxonomicName id="4C264D12C10B4351FDF6ECC1FCF0F954" authority="UF" authorityName="UF" box="[525,798,1690,1712]" class="Holothuroidea" family="Phyllophoridae" genus="Phyrella" kingdom="Animalia" order="Dendrochirotida" pageId="27" pageNumber="128" phylum="Echinodermata" rank="species" species="thyonoides">
<emphasis id="B952EA83C10B4351FDF6ECC1FD8DF94B" box="[525,611,1690,1712]" italics="true" pageId="27" pageNumber="128">Phyrella</emphasis>
cf. 
<emphasis id="B952EA83C10B4351FD71ECC1FD16F94B" box="[650,760,1690,1712]" italics="true" pageId="27" pageNumber="128">thyonoides</emphasis>
UF
</taxonomicName>
9414. a–d: perforated plates from podia that flank anal teeth.
</paragraph>
</caption>
<caption id="DF596619C10C4356FF6CE82BFAA4FD7D" ID-DOI="http://doi.org/10.5281/zenodo.4908406" ID-Zenodo-Dep="4908406" box="[151,1354,624,646]" httpUri="https://zenodo.org/record/4908406/files/figure.png" pageId="28" pageNumber="129" startId="28.[151,249,624,646]" targetBox="[151,1436,193,603]" targetPageId="28">
<paragraph id="8B993691C10C4356FF6CE82BFAA4FD7D" blockId="28.[151,1354,624,646]" box="[151,1354,624,646]" pageId="28" pageNumber="129">
<emphasis id="B952EA83C10C4356FF6CE82BFECEFD7E" bold="true" box="[151,288,624,646]" pageId="28" pageNumber="129">FIGURE 22.</emphasis>
Rosettes of 
<taxonomicName id="4C264D12C10C4356FE66E82BFD66FD7D" box="[413,648,624,646]" class="Holothuroidea" family="Phyllophoridae" genus="Phyrella" kingdom="Animalia" order="Dendrochirotida" pageId="28" pageNumber="129" phylum="Echinodermata" rank="species" species="thyonoides">
<emphasis id="B952EA83C10C4356FE66E82BFE1DFD7D" box="[413,499,624,646]" italics="true" pageId="28" pageNumber="129">Phyrella</emphasis>
cf. 
<emphasis id="B952EA83C10C4356FDE1E82BFD66FD7D" box="[538,648,624,646]" italics="true" pageId="28" pageNumber="129">thyonoides</emphasis>
</taxonomicName>
A: UF 9601, introvert; B: UF 9598, cloacal region. Scale bar, 20 
<emphasis id="B952EA83C10C4356FADBE82AFAC0FD7D" box="[1312,1326,625,646]" italics="true" pageId="28" pageNumber="129">µ</emphasis>
m.
</paragraph>
</caption>
<subSubSection id="C33C651AC10C4356FF3CE8E8FEEBFCCC" pageId="28" pageNumber="129" type="biology_ecology">
<paragraph id="8B993691C10C4356FF3CE8E8FEEBFCCC" blockId="28.[151,1436,691,823]" pageId="28" pageNumber="129">
<emphasis id="B952EA83C10C4356FF3CE8E8FE34FD37" bold="true" box="[199,474,691,716]" pageId="28" pageNumber="129">Ecology &amp; distribution</emphasis>
. 
<materialsCitation id="3B4E3CCCC10C4356FE12E8E8FEEFFCCC" ID-GBIF-Occurrence="3311708333" county="Known" elevation="2" location="Ningaloo" municipality="Perth" pageId="28" pageNumber="129" specimenCount="1" stateProvince="Western Australia">
In Ningaloo, animals were adhering to dead shells or rocks that were buried in sand and on which algae was growing, close to shore at 
<elevation id="000BD1A2C10C4356FD19E88DFC9DFD14" box="[738,883,726,751]" metricMagnitude="0" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="28" pageNumber="129" unit="m" value="2.0" valueMax="3.0" valueMin="1.0">
<quantity id="4CDE9B74C10C4356FD19E88DFCC7FD15" box="[738,809,726,751]" metricMagnitude="0" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="28" pageNumber="129" unit="m" value="2.0" valueMax="3.0" valueMin="1.0">1–3 m</quantity>
depth
</elevation>
. They co-occurred with 
<taxonomicName id="4C264D12C10C4356FB71E88DFA7BFD14" authorityName="Erwe" authorityYear="1913" box="[1162,1429,726,751]" class="Holothuroidea" family="Holothuriidae" genus="Holothuria" kingdom="Animalia" order="Aspidochirotida" pageId="28" pageNumber="129" phylum="Echinodermata" rank="species" species="michaelseni">
<emphasis id="B952EA83C10C4356FB71E88DFA7BFD14" box="[1162,1429,726,751]" italics="true" pageId="28" pageNumber="129">Holothuria michaelseni</emphasis>
</taxonomicName>
. The 
<typeStatus id="549D8833C10C4356FF34E8A0FEE4FCE8" box="[207,266,763,787]" pageId="28" pageNumber="129">types</typeStatus>
were cast up on the beach, suggesting shallow habits. 
<collectingCounty id="62F84E1DC10C4356FC5DE8A0FC16FCE9" box="[934,1016,763,786]" pageId="28" pageNumber="129">Known</collectingCounty>
from 
<collectingMunicipality id="6BFDACEBC10C4356FBB3E8A0FB6DFCEF" box="[1096,1155,763,788]" pageId="28" pageNumber="129">Perth</collectingMunicipality>
and 
<location id="8EF9604AC10C4356FB3EE8A0FADEFCEF" LSID="urn:lsid:plazi:treatment:038F8787C1074356FF6CEFFCFEEBFCCC:8EF9604AC10C4356FB3EE8A0FADEFCEF" box="[1221,1328,763,788]" county="Known" municipality="Perth" name="Ningaloo" pageId="28" pageNumber="129" stateProvince="Western Australia">Ningaloo</location>
, 
<collectingRegion id="49E2F873C10C4356FABAE8A0FEEFFCCC" country="Australia" name="Western Australia" pageId="28" pageNumber="129">Western Australia</collectingRegion>
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>