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<document id="DD091921C63867A5705042C44B66F9D6" ID-CLB-Dataset="54097" ID-DOI="10.11646/megataxa.4.1.1" ID-GBIF-Dataset="7c9bc39d-9910-46f1-9558-c0647b9cbbdb" ID-ISSN="2703-3090" ID-Zenodo-Dep="4591138" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1614033145335" checkinUser="plazi" docAuthor="Kim, Il-Hoi &amp; Boxshall, Geoff A." docDate="2020" docId="03C487CBEF593A39FF4DF949FDE9FAAC" docLanguage="en" docName="megataxa.4.1.1.pdf.imf" docOrigin="Megataxa 4 (1)" docSource="http://zoobank.org/3fdd970e-62f1-4f67-8cce-10870bdb3c01" docStyle="DocumentStyle:50B6B96D96A55C21793B52C23E95B310.5:Megataxa.2020-.journal_article" docStyleId="50B6B96D96A55C21793B52C23E95B310" docStyleName="Megataxa.2020-.journal_article" docStyleVersion="5" docTitle="Bathynotodelphys defayeae Kim &amp; Boxshall 2020, gen. et sp.nov." docType="treatment" docVersion="11" lastPageNumber="13" masterDocId="FFFDFFB3EF513A35FFC5FFCEFFC5FFC1" masterDocTitle="Untold diversity: the astonishing species richness of the Notodelphyidae (Copepoda: Cyclopoida), a family of symbiotic copepods associated with ascidians (Tunicata)" masterLastPageNumber="6" masterPageNumber="1" pageNumber="9" updateTime="1698930853666" updateUser="ExternalLinkService">
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<mods:title id="1878EAB8FB51DFA31DD4CBA4A1072A83">Untold diversity: the astonishing species richness of the Notodelphyidae (Copepoda: Cyclopoida), a family of symbiotic copepods associated with ascidians (Tunicata)</mods:title>
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<mods:namePart id="A5C80B4E8653F36EF910F7743F768760">Kim, Il-Hoi</mods:namePart>
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<mods:affiliation id="C64F1DDFB120A7E077520A1CC2135644">Korea Institute of Coastal Ecology, Inc., 802 - ho, 302 - dong, 397 Seokcheon-ro, Ojeong-gu, Bucheon, Gyeonggi-do 14449, Republic of Korea ® ihkim @ gwnu. ac. kr; https: // orcid. org / 0000 - 0002 - 7332 - 0043</mods:affiliation>
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<mods:namePart id="73737046A6A059A25F65C2C735086B5D">Boxshall, Geoff A.</mods:namePart>
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<mods:affiliation id="A67BEC9FAFCDA5D50B814DD132F362DB">Korea Institute of Coastal Ecology, Inc., 802 - ho, 302 - dong, 397 Seokcheon-ro, Ojeong-gu, Bucheon, Gyeonggi-do 14449, Republic of Korea ® ihkim @ gwnu. ac. kr; https: // orcid. org / 0000 - 0002 - 7332 - 0043</mods:affiliation>
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<mods:title id="B022D55C920BBCC409916BD8A43EF89B">Megataxa</mods:title>
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<treatment id="03C487CBEF593A39FF4DF949FDE9FAAC" ID-DOI="http://doi.org/10.5281/zenodo.5661610" ID-GBIF-Taxon="194474943" ID-Zenodo-Dep="5661610" LSID="urn:lsid:plazi:treatment:03C487CBEF593A39FF4DF949FDE9FAAC" httpUri="http://treatment.plazi.org/id/03C487CBEF593A39FF4DF949FDE9FAAC" lastPageId="12" lastPageNumber="13" pageId="8" pageNumber="9">
<subSubSection id="C3776556EF593A3DFF4DF949FD4DF963" box="[136,648,1671,1698]" pageId="8" pageNumber="9" type="nomenclature">
<paragraph id="8BD236DDEF593A3DFF4DF949FD4DF963" blockId="8.[136,648,1671,1730]" box="[136,648,1671,1698]" pageId="8" pageNumber="9">
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<emphasis id="B919EACFEF593A3DFF4DF949FD4DF963" bold="true" box="[136,648,1671,1698]" pageId="8" pageNumber="9">
<taxonomicName id="4C6D4D5EEF593A3DFF4DF949FE17F963" authority="Kim &amp; Boxshall, 2020" authorityName="Kim &amp; Boxshall" authorityYear="2020" box="[136,466,1671,1698]" class="Maxillopoda" family="Ascidicolidae" genus="Bathynotodelphys" kingdom="Animalia" order="Cyclopoida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="defayeae" status="gen. et sp.nov.">
<emphasis id="B919EACFEF593A3DFF4DF949FE17F963" bold="true" box="[136,466,1671,1698]" italics="true" pageId="8" pageNumber="9">Bathynotodelphys defayeae</emphasis>
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<taxonomicNameLabel id="A22A57B4EF593A3DFE1CF946FD4DF963" box="[473,648,1672,1698]" pageId="8" pageNumber="9" rank="species">gen. et sp.nov.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="C3776556EF593A3DFF4DF969FEC6F903" box="[136,259,1703,1730]" pageId="8" pageNumber="9" type="description">
<paragraph id="8BD236DDEF593A3DFF4DF969FEC6F903" blockId="8.[136,648,1671,1730]" box="[136,259,1703,1730]" pageId="8" pageNumber="9">
(
<figureCitation id="13562A58EF593A3DFF55F969FF3FF903" box="[144,250,1703,1730]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="9.[136,245,1907,1933]" captionStartId-1="10.[136,245,1907,1933]" captionStartId-2="11.[136,245,1567,1593]" captionTargetBox-0="[234,1352,203,1854]" captionTargetBox-1="[238,1342,193,1853]" captionTargetBox-2="[146,1444,210,1521]" captionTargetId-0="figure-18@9.[221,1366,185,1870]" captionTargetId-1="figure-19@10.[230,1356,181,1866]" captionTargetId-2="figure-211@11.[136,1451,200,1532]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionTargetPageId-2="11" captionText-0="FIGURE 1. Bathynotodelphys defayeae gen. et sp. nov., female. A, habitus, lateral; B, urosome, ventral; C, right caudal ramus, dorsal; D, rostrum; E, antennule; F, antenna; G, labrum; H, mandible; I, paragnath; J, maxillule. Scale bars: A, 0.5 mm; B, 0.2 mm; C–H, J, 0.1 mm; I, 0.05 mm." captionText-1="FIGURE 2. Bathynotodelphys defayeae gen. et sp. nov., female. A, maxilla; B, maxilliped; C, leg 1; D, leg 2; E, leg 4; F, leg 5. Scale bars: 0.1 mm." captionText-2="FIGURE 3. Bathynotodelphys defayeae gen. et sp. nov., male. A, habitus, dorsal; B, urosome, ventral; C, antennule. Scale bars: A, 0.5 mm; B, C, 0.1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4591140" figureDoi-1="http://doi.org/10.5281/zenodo.4591142" figureDoi-2="http://doi.org/10.5281/zenodo.4591144" httpUri-0="https://zenodo.org/record/4591140/files/figure.png" httpUri-1="https://zenodo.org/record/4591142/files/figure.png" httpUri-2="https://zenodo.org/record/4591144/files/figure.png" pageId="8" pageNumber="9">Figs. 1–3</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3776556EF593A3DFF4DF922FEC7F813" pageId="8" pageNumber="9" type="materials_examined">
<paragraph id="8BD236DDEF593A3DFF4DF922FEC7F813" blockId="8.[136,778,1771,2070]" pageId="8" pageNumber="9">
<emphasis id="B919EACFEF593A3DFF4DF922FEF3F8C4" bold="true" box="[136,310,1771,1798]" pageId="8" pageNumber="9">Typematerial</emphasis>
.
<materialsCitation id="3B053C80EF593A3DFE89F925FF1BF8E9" ID-GBIF-Occurrence="3748446451" pageId="8" pageNumber="9" specimenCode="MNHN-IU-2014- 21212" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<typeStatus id="54D6887FEF593A3DFE89F925FE71F8C4" box="[332,436,1771,1797]" pageId="8" pageNumber="9" type="holotype">Holotype</typeStatus>
(intact
<specimenCount id="9D6BFD54EF593A3DFDDDF922FDE8F8C7" box="[536,557,1772,1798]" pageId="8" pageNumber="9" type="female">♀</specimenCount>
,
<specimenCode id="DBCB9EA6EF593A3DFD86F922FF17F8E9" pageId="8" pageNumber="9">MNHN-IU-2014- 21212</specimenCode>
),
</materialsCitation>
<materialsCitation id="3B053C80EF593A3DFF2CF8C0FED9F88B" ID-GBIF-Occurrence="3748446508" pageId="8" pageNumber="9" typeStatus="paratype">
<typeStatus id="54D6887FEF593A3DFF2CF8C0FE91F8E9" box="[233,340,1806,1832]" pageId="8" pageNumber="9" type="paratype">paratypes</typeStatus>
(intact
<specimenCount id="9D6BFD54EF593A3DFE75F8C3FE36F8E9" box="[432,499,1805,1832]" pageId="8" pageNumber="9" type="female">6 ♀♀</specimenCount>
and 2 ♂♂, MNHN-IU-2014-21213)
</materialsCitation>
, and dissected
<materialsCitation id="3B053C80EF593A3DFE04F8FEFEC7F813" ID-GBIF-Occurrence="3748446450" latitude="-44.55" longitude="-49.316666" pageId="8" pageNumber="9" typeStatus="paratype">
<typeStatus id="54D6887FEF593A3DFE04F8FEFDE9F88B" box="[449,556,1840,1866]" pageId="8" pageNumber="9" type="paratype">paratypes</typeStatus>
(
<specimenCount id="9D6BFD54EF593A3DFDFBF8FEFDA3F88B" box="[574,614,1840,1866]" pageId="8" pageNumber="9" type="female">1 ♀</specimenCount>
and 1 ♂) from
<taxonomicName id="4C6D4D5EEF593A3DFF4DF89CFCC3F8AA" authority="Monniot F. &amp; Monniot C., 1976" authorityName="Monniot F. &amp; Monniot C." authorityYear="1976" box="[136,774,1873,1900]" class="Ascidiacea" family="Pyuridae" genus="Culeolus" kingdom="Animalia" order="Pleurogona" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="anonymus">
<emphasis id="B919EACFEF593A3DFF4DF89CFEA9F8AA" box="[136,364,1874,1900]" italics="true" pageId="8" pageNumber="9">Culeolus anonymus</emphasis>
Monniot F. &amp; Monniot C., 1976
</taxonomicName>
, Vema Cruise,
<collectingCountry id="F37A764DEF593A3DFEECF8BAFE5DF84F" box="[297,408,1908,1934]" name="Argentina" pageId="8" pageNumber="9">Argentine</collectingCountry>
Basin, South Atlantic (
<geoCoordinate id="EE59501AEF593A3DFD63F8BAFCC0F84F" box="[678,773,1908,1934]" degrees="44" direction="south" minutes="33" orientation="latitude" pageId="8" pageNumber="9" precision="925" value="-44.55">44°33’S</geoCoordinate>
,
<geoCoordinate id="EE59501AEF593A3DFF4DF858FF37F871" box="[136,242,1942,1968]" degrees="49" direction="west" minutes="19" orientation="longitude" pageId="8" pageNumber="9" precision="925" value="-49.316666">49°19’W</geoCoordinate>
), “Vema”, Stn
<collectionCode id="ED7CAE18EF593A3DFE53F858FE6FF871" box="[406,426,1942,1968]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="8" pageNumber="9" type="Museum">V</collectionCode>
17–81, depth
<quantity id="4C959B38EF593A3DFD82F85BFD26F871" box="[583,739,1941,1968]" metricMagnitude="3" metricUnit="m" metricValue="5.330499999999999" metricValueMax="5.332" metricValueMin="5.329" pageId="8" pageNumber="9" unit="m" value="5330.5" valueMax="5332.0" valueMin="5329.0">5329–5332 m</quantity>
,
<date id="FFD3101DEF593A3DFD2BF858FF38F813" pageId="8" pageNumber="9" value="1961-05-27">27 May 1961</date>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3776556EF593A3DFF78F814FAF9FF28" pageId="8" pageNumber="9" type="etymology">
<paragraph id="8BD236DDEF593A3DFF78F814FAF9FF28" blockId="8.[136,778,1771,2070]" lastBlockId="8.[810,1452,173,2070]" pageId="8" pageNumber="9">
<emphasis id="B919EACFEF593A3DFF78F814FEFBF835" bold="true" box="[189,318,2010,2036]" pageId="8" pageNumber="9">Etymology</emphasis>
. The new species is named in honour of Dr. Danielle Defaye who made the Monniot collection of copepods associated with ascidians available to the authors for study, resulting in the present work.
</paragraph>
</subSubSection>
<subSubSection id="C3776556EF593A39FC9AFF3FFDB7FC8C" lastPageId="12" lastPageNumber="13" pageId="8" pageNumber="9" type="description">
<paragraph id="8BD236DDEF593A3DFC9AFF3FFBDEFC8C" blockId="8.[810,1452,173,2070]" pageId="8" pageNumber="9">
<emphasis id="B919EACFEF593A3DFC9AFF3FFBABFECA" bold="true" box="[863,1134,241,267]" pageId="8" pageNumber="9">Descriptionoffemale</emphasis>
. Body (
<figureCitation id="13562A58EF593A3DFB1DFF3FFAF2FECA" box="[1240,1335,241,267]" captionStart="FIGURE 1" captionStartId="9.[136,245,1907,1933]" captionTargetBox="[234,1352,203,1854]" captionTargetId="figure-18@9.[221,1366,185,1870]" captionTargetPageId="9" captionText="FIGURE 1. Bathynotodelphys defayeae gen. et sp. nov., female. A, habitus, lateral; B, urosome, ventral; C, right caudal ramus, dorsal; D, rostrum; E, antennule; F, antenna; G, labrum; H, mandible; I, paragnath; J, maxillule. Scale bars: A, 0.5 mm; B, 0.2 mm; C–H, J, 0.1 mm; I, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4591140" httpUri="https://zenodo.org/record/4591140/files/figure.png" pageId="8" pageNumber="9">Fig. 1A</figureCitation>
) laterally compressed: body length 4.40 mm in dissected specimen. Cephalosome broader than first to third pedigerous somites. Second and third pedigerous somites each with well-developed dorsal tergite. Fourth pedigerous somite forming small brood pouch, much shorter than anterior part of prosome, incorporating fused fifth pedigerous somite. Freeurosome (
<figureCitation id="13562A58EF593A3DFB86FE11FB64FE38" box="[1091,1185,479,505]" captionStart="FIGURE 1" captionStartId="9.[136,245,1907,1933]" captionTargetBox="[234,1352,203,1854]" captionTargetId="figure-18@9.[221,1366,185,1870]" captionTargetPageId="9" captionText="FIGURE 1. Bathynotodelphys defayeae gen. et sp. nov., female. A, habitus, lateral; B, urosome, ventral; C, right caudal ramus, dorsal; D, rostrum; E, antennule; F, antenna; G, labrum; H, mandible; I, paragnath; J, maxillule. Scale bars: A, 0.5 mm; B, 0.2 mm; C–H, J, 0.1 mm; I, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4591140" httpUri="https://zenodo.org/record/4591140/files/figure.png" pageId="8" pageNumber="9">Fig. 1B</figureCitation>
) 5-segmented: genital somite short; suture between genital and first abdominal somitesdistinct on dorsal surface but obscure ventrally. First to fourth free abdominal somites 364×345, 284×320, 218×287, and 255×313 μm, respectively. Caudal rami elongate and slightly divergent; each ramus (
<figureCitation id="13562A58EF593A3DFA82FD47FA67FD62" box="[1351,1442,649,675]" captionStart="FIGURE 1" captionStartId="9.[136,245,1907,1933]" captionTargetBox="[234,1352,203,1854]" captionTargetId="figure-18@9.[221,1366,185,1870]" captionTargetPageId="9" captionText="FIGURE 1. Bathynotodelphys defayeae gen. et sp. nov., female. A, habitus, lateral; B, urosome, ventral; C, right caudal ramus, dorsal; D, rostrum; E, antennule; F, antenna; G, labrum; H, mandible; I, paragnath; J, maxillule. Scale bars: A, 0.5 mm; B, 0.2 mm; C–H, J, 0.1 mm; I, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4591140" httpUri="https://zenodo.org/record/4591140/files/figure.png" pageId="8" pageNumber="9">Fig. 1C</figureCitation>
) constrictedbasallyandtaperingdistally, widestatproximal one-fifth, about 4.1 times as long as wide (502×124 μm) and armed with 6 small setae; 4 distal setae pinnate and remaining 2 setae naked; outer lateral seta positioned at 42% of ramus length.
</paragraph>
<paragraph id="8BD236DDEF593A3DFC9AFC98FBB7FA73" blockId="8.[810,1452,173,2070]" pageId="8" pageNumber="9">
Rostrum (
<figureCitation id="13562A58EF593A3DFC12FC9BFBF0FCAE" box="[983,1077,853,879]" captionStart="FIGURE 1" captionStartId="9.[136,245,1907,1933]" captionTargetBox="[234,1352,203,1854]" captionTargetId="figure-18@9.[221,1366,185,1870]" captionTargetPageId="9" captionText="FIGURE 1. Bathynotodelphys defayeae gen. et sp. nov., female. A, habitus, lateral; B, urosome, ventral; C, right caudal ramus, dorsal; D, rostrum; E, antennule; F, antenna; G, labrum; H, mandible; I, paragnath; J, maxillule. Scale bars: A, 0.5 mm; B, 0.2 mm; C–H, J, 0.1 mm; I, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4591140" httpUri="https://zenodo.org/record/4591140/files/figure.png" pageId="8" pageNumber="9">Fig. 1D</figureCitation>
) 255×215 μm, well-defined at base, tapering towards rounded apex. Antennule (
<figureCitation id="13562A58EF593A3DFA8AFCB9FA67FC50" box="[1359,1442,887,913]" captionStart="FIGURE 1" captionStartId="9.[136,245,1907,1933]" captionTargetBox="[234,1352,203,1854]" captionTargetId="figure-18@9.[221,1366,185,1870]" captionTargetPageId="9" captionText="FIGURE 1. Bathynotodelphys defayeae gen. et sp. nov., female. A, habitus, lateral; B, urosome, ventral; C, right caudal ramus, dorsal; D, rostrum; E, antennule; F, antenna; G, labrum; H, mandible; I, paragnath; J, maxillule. Scale bars: A, 0.5 mm; B, 0.2 mm; C–H, J, 0.1 mm; I, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4591140" httpUri="https://zenodo.org/record/4591140/files/figure.png" pageId="8" pageNumber="9">Fig. 1E</figureCitation>
) gradually narrowing distally, 9-segmented with segmental fusion pattern as in generic diagnosis; armature formula 3, 17, 6, 4+aesthetasc, 6, 4+aesthetasc, 2, 2+aesthetasc, and 7+aesthetasc; about half of setae pinnate (as figured). Antenna (
<figureCitation id="13562A58EF593A3DFC5DFBEFFC2FFBFA" box="[920,1002,1057,1083]" captionStart="FIGURE 1" captionStartId="9.[136,245,1907,1933]" captionTargetBox="[234,1352,203,1854]" captionTargetId="figure-18@9.[221,1366,185,1870]" captionTargetPageId="9" captionText="FIGURE 1. Bathynotodelphys defayeae gen. et sp. nov., female. A, habitus, lateral; B, urosome, ventral; C, right caudal ramus, dorsal; D, rostrum; E, antennule; F, antenna; G, labrum; H, mandible; I, paragnath; J, maxillule. Scale bars: A, 0.5 mm; B, 0.2 mm; C–H, J, 0.1 mm; I, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4591140" httpUri="https://zenodo.org/record/4591140/files/figure.png" pageId="8" pageNumber="9">Fig. 1F</figureCitation>
) 5-segmented, consistingof coxa, basis and 3-segmented endopod: coxashortand unarmed: basis with 1 long pinnate seta and 1 minute naked seta at distal outer corner representing exopod; ornamented with tuft of setules proximally on inner margin: endopod 3-segmented; first segment with 1 seta on inner margin; second segment about 2.7 times as long as wide, armed with 5 setae (1 on middle of inner margin and 4 at inner distal corner); third segment about 0.6 times as long as second, 1.8 times as long as wide, armed with 6 setae (3 distal setae blunt at tip) plus small terminal claw, about 0.6 times as long as segment.
</paragraph>
<paragraph id="8BD236DDEF593A3EFC9AFA77FD49F84F" blockId="8.[810,1452,173,2070]" lastBlockId="11.[136,778,1703,2070]" lastPageId="11" lastPageNumber="12" pageId="8" pageNumber="9">
Labrum (
<figureCitation id="13562A58EF593A3DFC24FA77FB94FA15" box="[993,1105,1465,1492]" captionStart="FIGURE 1" captionStartId="9.[136,245,1907,1933]" captionTargetBox="[234,1352,203,1854]" captionTargetId="figure-18@9.[221,1366,185,1870]" captionTargetPageId="9" captionText="FIGURE 1. Bathynotodelphys defayeae gen. et sp. nov., female. A, habitus, lateral; B, urosome, ventral; C, right caudal ramus, dorsal; D, rostrum; E, antennule; F, antenna; G, labrum; H, mandible; I, paragnath; J, maxillule. Scale bars: A, 0.5 mm; B, 0.2 mm; C–H, J, 0.1 mm; I, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4591140" httpUri="https://zenodo.org/record/4591140/files/figure.png" pageId="8" pageNumber="9">Fig. 1G</figureCitation>
) with rounded, setulose posterolateral corners and spinulose posteromedian lobe. Mandible (
<figureCitation id="13562A58EF593A3DFC6CFA33FBC1F9D9" box="[937,1028,1533,1560]" captionStart="FIGURE 1" captionStartId="9.[136,245,1907,1933]" captionTargetBox="[234,1352,203,1854]" captionTargetId="figure-18@9.[221,1366,185,1870]" captionTargetPageId="9" captionText="FIGURE 1. Bathynotodelphys defayeae gen. et sp. nov., female. A, habitus, lateral; B, urosome, ventral; C, right caudal ramus, dorsal; D, rostrum; E, antennule; F, antenna; G, labrum; H, mandible; I, paragnath; J, maxillule. Scale bars: A, 0.5 mm; B, 0.2 mm; C–H, J, 0.1 mm; I, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4591140" httpUri="https://zenodo.org/record/4591140/files/figure.png" pageId="8" pageNumber="9">Fig. 1H</figureCitation>
) with 6 teethand 2 smallproximal setae on cutting margin of coxal gnathobase; basis with 1 seta subdistally onmedial margin; exopod 4-segmented and armed with 1, 1, 1, and 2 setae on first to fourth segments, respectively (all 5 setae equal in length): endopod with 4 and 10 setae on first and second segments, respectively. Paragnath (
<figureCitation id="13562A58EF593A3DFB86F907FB57F925" box="[1091,1170,1737,1764]" captionStart="FIGURE 1" captionStartId="9.[136,245,1907,1933]" captionTargetBox="[234,1352,203,1854]" captionTargetId="figure-18@9.[221,1366,185,1870]" captionTargetPageId="9" captionText="FIGURE 1. Bathynotodelphys defayeae gen. et sp. nov., female. A, habitus, lateral; B, urosome, ventral; C, right caudal ramus, dorsal; D, rostrum; E, antennule; F, antenna; G, labrum; H, mandible; I, paragnath; J, maxillule. Scale bars: A, 0.5 mm; B, 0.2 mm; C–H, J, 0.1 mm; I, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4591140" httpUri="https://zenodo.org/record/4591140/files/figure.png" pageId="8" pageNumber="9">Fig. 1I</figureCitation>
) as simple lobe bearing setules on medial surface. Maxillule (
<figureCitation id="13562A58EF593A3DFB37F925FA83F8C7" box="[1266,1350,1771,1798]" captionStart="FIGURE 1" captionStartId="9.[136,245,1907,1933]" captionTargetBox="[234,1352,203,1854]" captionTargetId="figure-18@9.[221,1366,185,1870]" captionTargetPageId="9" captionText="FIGURE 1. Bathynotodelphys defayeae gen. et sp. nov., female. A, habitus, lateral; B, urosome, ventral; C, right caudal ramus, dorsal; D, rostrum; E, antennule; F, antenna; G, labrum; H, mandible; I, paragnath; J, maxillule. Scale bars: A, 0.5 mm; B, 0.2 mm; C–H, J, 0.1 mm; I, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4591140" httpUri="https://zenodo.org/record/4591140/files/figure.png" pageId="8" pageNumber="9">Fig. 1J</figureCitation>
) with 9 setaeon arthrite, 1 on coxal endite, 2 on epipodite, and 3 on medial margin of basis; exopod unsegmented with 4 setae distally; endopod 2-segmented with 2 or 3 setae on medial margin of first segment and 4 setae on second segment. Maxilla (
<figureCitation id="13562A58EF593A3DFBC8F858FBAAF871" box="[1037,1135,1942,1968]" captionStart="FIGURE 2" captionStartId="10.[136,245,1907,1933]" captionTargetBox="[238,1342,193,1853]" captionTargetId="figure-19@10.[230,1356,181,1866]" captionTargetPageId="10" captionText="FIGURE 2. Bathynotodelphys defayeae gen. et sp. nov., female. A, maxilla; B, maxilliped; C, leg 1; D, leg 2; E, leg 4; F, leg 5. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4591142" httpUri="https://zenodo.org/record/4591142/files/figure.png" pageId="8" pageNumber="9">Fig. 2A</figureCitation>
) indistinctly 6-segmented; first segment (precoxa) with 4 and 1 setae on proximal and distal endites, respectively; secondsegment (coxa) incompletely articulated from first, armed with 2 and 3 setae on proximal and distal endites, respectively: basis with strong claw bearing spinules along concave margin, plus 2 setae: endopod slender, 3-segmentedwith 1, 1, and 4 setae on first to third segments, respectively; one of 4 setae on third segment small. Maxilliped (
<figureCitation id="13562A58EF5A3A3EFD6CF8E1FCC4F88B" box="[681,769,1839,1866]" captionStart="FIGURE 2" captionStartId="10.[136,245,1907,1933]" captionTargetBox="[238,1342,193,1853]" captionTargetId="figure-19@10.[230,1356,181,1866]" captionTargetPageId="10" captionText="FIGURE 2. Bathynotodelphys defayeae gen. et sp. nov., female. A, maxilla; B, maxilliped; C, leg 1; D, leg 2; E, leg 4; F, leg 5. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4591142" httpUri="https://zenodo.org/record/4591142/files/figure.png" pageId="11" pageNumber="12">Fig. 2B</figureCitation>
) 3-segmented and armed with 10, 1, and 4 setae on first to third segments, respectively; all setae pinnate.
</paragraph>
<caption id="DF126655EF583A3CFF4DF8BDFCD7F813" ID-DOI="http://doi.org/10.5281/zenodo.4591140" ID-Zenodo-Dep="4591140" httpUri="https://zenodo.org/record/4591140/files/figure.png" pageId="9" pageNumber="10" startId="9.[136,245,1907,1933]" targetBox="[234,1352,203,1854]" targetPageId="9">
<paragraph id="8BD236DDEF583A3CFF4DF8BDFCD7F813" blockId="9.[136,1452,1907,2002]" pageId="9" pageNumber="10">
<emphasis id="B919EACFEF583A3CFF4DF8BDFED1F84F" bold="true" box="[136,276,1907,1934]" pageId="9" pageNumber="10">FIGURE 1.</emphasis>
<taxonomicName id="4C6D4D5EEF583A3CFEDBF8BAFD91F84F" authority="Kim &amp; Boxshall, 2020" authorityName="Kim &amp; Boxshall" authorityYear="2020" box="[286,596,1908,1934]" class="Maxillopoda" family="Ascidicolidae" genus="Bathynotodelphys" kingdom="Animalia" order="Cyclopoida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="defayeae" status="gen. et sp. nov.">
<emphasis id="B919EACFEF583A3CFEDBF8BAFD91F84F" box="[286,596,1908,1934]" italics="true" pageId="9" pageNumber="10">Bathynotodelphys defayeae</emphasis>
</taxonomicName>
<emphasis id="B919EACFEF583A3CFD98F8BAFCD1F84F" bold="true" box="[605,788,1908,1934]" pageId="9" pageNumber="10">
<taxonomicNameLabel id="A22A57B4EF583A3CFD98F8BAFCD1F84F" box="[605,788,1908,1934]" pageId="9" pageNumber="10" rank="species">gen. et sp. nov.</taxonomicNameLabel>
</emphasis>
, female. A, habitus, lateral; B, urosome, ventral; C, right caudalramus, dorsal; D, rostrum; E, antennule; F, antenna; G, labrum; H, mandible; I, paragnath; J, maxillule. Scale bars: A, 0.5 mm; B, 0.2 mm; C–H, J, 0.1 mm; I, 0.05 mm.
</paragraph>
</caption>
<caption id="DF126655EF5B3A3FFF4DF8BDFE21F871" ID-DOI="http://doi.org/10.5281/zenodo.4591142" ID-Zenodo-Dep="4591142" httpUri="https://zenodo.org/record/4591142/files/figure.png" pageId="10" pageNumber="11" startId="10.[136,245,1907,1933]" targetBox="[238,1342,193,1853]" targetPageId="10">
<paragraph id="8BD236DDEF5B3A3FFF4DF8BDFE21F871" blockId="10.[136,1451,1907,1968]" pageId="10" pageNumber="11">
<emphasis id="B919EACFEF5B3A3FFF4DF8BDFED4F84F" bold="true" box="[136,273,1907,1934]" pageId="10" pageNumber="11">FIGURE 2.</emphasis>
<taxonomicName id="4C6D4D5EEF5B3A3FFEDCF8BAFD88F84F" authority="Kim &amp; Boxshall, 2020" authorityName="Kim &amp; Boxshall" authorityYear="2020" box="[281,589,1908,1934]" class="Maxillopoda" family="Ascidicolidae" genus="Bathynotodelphys" kingdom="Animalia" order="Cyclopoida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="defayeae" status="gen. et sp. nov.">
<emphasis id="B919EACFEF5B3A3FFEDCF8BAFD88F84F" box="[281,589,1908,1934]" italics="true" pageId="10" pageNumber="11">Bathynotodelphys defayeae</emphasis>
</taxonomicName>
<emphasis id="B919EACFEF5B3A3FFD91F8BAFCC6F84F" bold="true" box="[596,771,1908,1934]" pageId="10" pageNumber="11">
<taxonomicNameLabel id="A22A57B4EF5B3A3FFD91F8BAFCC6F84F" box="[596,771,1908,1934]" pageId="10" pageNumber="11" rank="species">gen. et sp. nov.</taxonomicNameLabel>
</emphasis>
, female. A, maxilla; B, maxilliped; C, leg 1; D, leg 2; E, leg 4; F, leg 5. Scale bars: 0.1 mm.
</paragraph>
</caption>
<caption id="DF126655EF5A3A3EFF4DF9D1FDE8F99D" ID-DOI="http://doi.org/10.5281/zenodo.4591144" ID-Zenodo-Dep="4591144" httpUri="https://zenodo.org/record/4591144/files/figure.png" pageId="11" pageNumber="12" startId="11.[136,245,1567,1593]" targetBox="[146,1444,210,1521]" targetPageId="11">
<paragraph id="8BD236DDEF5A3A3EFF4DF9D1FDE8F99D" blockId="11.[136,1452,1567,1628]" pageId="11" pageNumber="12">
<emphasis id="B919EACFEF5A3A3EFF4DF9D1FED4F9FB" bold="true" box="[136,273,1567,1594]" pageId="11" pageNumber="12">FIGURE 3.</emphasis>
<taxonomicName id="4C6D4D5EEF5A3A3EFEDCF9D1FD88F9F8" authority="Kim &amp; Boxshall, 2020" authorityName="Kim &amp; Boxshall" authorityYear="2020" box="[281,589,1567,1593]" class="Maxillopoda" family="Ascidicolidae" genus="Bathynotodelphys" kingdom="Animalia" order="Cyclopoida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="defayeae" status="gen. et sp. nov.">
<emphasis id="B919EACFEF5A3A3EFEDCF9D1FD88F9F8" box="[281,589,1567,1593]" italics="true" pageId="11" pageNumber="12">Bathynotodelphys defayeae</emphasis>
</taxonomicName>
<emphasis id="B919EACFEF5A3A3EFD96F9EEFCC6F9FB" bold="true" box="[595,771,1568,1594]" pageId="11" pageNumber="12">
<taxonomicNameLabel id="A22A57B4EF5A3A3EFD96F9EEFCC6F9FB" box="[595,771,1568,1594]" pageId="11" pageNumber="12" rank="species">gen. et sp. nov.</taxonomicNameLabel>
</emphasis>
, male. A, habitus, dorsal; B, urosome, ventral; C, antennule. Scale bars: A, 0.5 mm; B, C, 0.1 mm.
</paragraph>
</caption>
<paragraph id="8BD236DDEF5A3A3EFF78F858FAA2F812" blockId="11.[136,778,1703,2070]" lastBlockId="11.[810,1452,1703,2070]" pageId="11" pageNumber="12">
Legs 1–4 biramous with 3-segmented rami (
<figureCitation id="13562A58EF5A3A3EFD19F858FF53F813" captionStart="FIGURE 2" captionStartId="10.[136,245,1907,1933]" captionTargetBox="[238,1342,193,1853]" captionTargetId="figure-19@10.[230,1356,181,1866]" captionTargetPageId="10" captionText="FIGURE 2. Bathynotodelphys defayeae gen. et sp. nov., female. A, maxilla; B, maxilliped; C, leg 1; D, leg 2; E, leg 4; F, leg 5. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4591142" httpUri="https://zenodo.org/record/4591142/files/figure.png" pageId="11" pageNumber="12">Fig. 2</figureCitation>
C–E). Spines on outer margin of exopods slender and fringed with membrane bilaterally in distal half. Inner distal spine on basis of leg 1 fringed with spinules, 106 μm long, slightly longerthan first endopodal segment. Inner coxal seta of legs 2–4 stiff and spiniform. Armature formula as in generic diagnosis. Leg 5 (
<figureCitation id="13562A58EF5A3A3EFB1EF925FAE9F8C7" box="[1243,1324,1771,1798]" captionStart="FIGURE 2" captionStartId="10.[136,245,1907,1933]" captionTargetBox="[238,1342,193,1853]" captionTargetId="figure-19@10.[230,1356,181,1866]" captionTargetPageId="10" captionText="FIGURE 2. Bathynotodelphys defayeae gen. et sp. nov., female. A, maxilla; B, maxilliped; C, leg 1; D, leg 2; E, leg 4; F, leg 5. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4591142" httpUri="https://zenodo.org/record/4591142/files/figure.png" pageId="11" pageNumber="12">Fig. 2F</figureCitation>
) consisting of protopod and free exopod; protopod incorporated into fifth pedigerous somite, armed with naked outer distal seta; exopodal segment elongate, about 4.3 times as long as wide (167×39 μm), ornamented with spinules along inner margin and armed distally with 1 slender spine (82 μm long) and 1 weakly pinnate seta (118 μm long).
</paragraph>
<paragraph id="8BD236DDEF5A3A39FC9AF817FE50FE74" blockId="11.[810,1452,1703,2070]" lastBlockId="12.[136,778,173,1390]" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
<emphasis id="B919EACFEF5A3A3EFC9AF817FB8CF835" bold="true" box="[863,1097,2009,2036]" pageId="11" pageNumber="12">Descriptionofmale</emphasis>
. Body (
<figureCitation id="13562A58EF5A3A3EFB66F814FB3EF835" box="[1187,1275,2010,2036]" captionStart="FIGURE 3" captionStartId="11.[136,245,1567,1593]" captionTargetBox="[146,1444,210,1521]" captionTargetId="figure-211@11.[136,1451,200,1532]" captionTargetPageId="11" captionText="FIGURE 3. Bathynotodelphys defayeae gen. et sp. nov., male. A, habitus, dorsal; B, urosome, ventral; C, antennule. Scale bars: A, 0.5 mm; B, C, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4591144" httpUri="https://zenodo.org/record/4591144/files/figure.png" pageId="11" pageNumber="12">Fig. 3A</figureCitation>
) dorsoventrally depressed, tapering distally. Bodylength 2.20 mm. Prosome-urosome division indistinct. Urosome (
<figureCitation id="13562A58EF5D3A39FD6EFF63FCC4FF06" box="[683,769,173,199]" captionStart="FIGURE 3" captionStartId="11.[136,245,1567,1593]" captionTargetBox="[146,1444,210,1521]" captionTargetId="figure-211@11.[136,1451,200,1532]" captionTargetPageId="11" captionText="FIGURE 3. Bathynotodelphys defayeae gen. et sp. nov., male. A, habitus, dorsal; B, urosome, ventral; C, antennule. Scale bars: A, 0.5 mm; B, C, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4591144" httpUri="https://zenodo.org/record/4591144/files/figure.png" pageId="12" pageNumber="13">Fig. 3B</figureCitation>
) 6-segmented: fifth pedigerous somite (first urosomite) shorter than genital somite: genital somite 131×247 μm, with well-developed paired genital opercula ventrally: 4 abdominalsomites 167×196, 173×182, 135×160, and 135×145 μm, respectively. Caudal ramus about 5.6 times as long as wide (224×40 μm); outer lateral seta positioned at 40% of ramus length.
</paragraph>
<paragraph id="8BD236DDEF5D3A39FF78FE73FD0AFD9E" blockId="12.[136,778,173,1390]" pageId="12" pageNumber="13">Rostrum asin female. Antennule 10-segmented; non-geniculate, segmental fusion pattern as in generic diagnosis; armatureformula 3, 17, 2, 2, 2, 4, 2, 4, 3+aesthetasc, and 10+aesthetasc; one setaon first segment pinnate, all other setae naked. Antenna as in female.</paragraph>
<paragraph id="8BD236DDEF5D3A39FC75FD8BFA60FD04" pageId="12" pageNumber="13">
<table id="F96DC47DEF5DC5CAFCEFFD8BFA63FD07" box="[810,1446,581,710]" gridcols="5" gridrows="4" inLine="true" pageId="12" pageNumber="13">
<tr id="355D349FEF5DC5CAFCEFFD8BFA63FD9E" box="[810,1446,581,607]" gridrow="0" pageId="12" pageNumber="13" rowspan-0="1">
<th id="768C5DE3EF5DC5CAFC75FD8BFC2FFD9E" box="[944,1002,581,607]" gridcol="1" gridrow="0" pageId="12" pageNumber="13">Coxa</th>
<th id="768C5DE3EF5DC5CAFC32FD8BFBF7FD9E" box="[1015,1074,581,607]" gridcol="2" gridrow="0" pageId="12" pageNumber="13">Basis</th>
<th id="768C5DE3EF5DC5CAFBF8FD8BFB2DFD9E" box="[1085,1256,581,607]" gridcol="3" gridrow="0" pageId="12" pageNumber="13">Exopod</th>
<th id="768C5DE3EF5DC5CAFB3FFD8BFA63FD9E" box="[1274,1446,581,607]" gridcol="4" gridrow="0" pageId="12" pageNumber="13">Endopod</th>
</tr>
<tr id="355D349FEF5DC5CAFCEFFDA9FA63FD43" box="[810,1446,615,642]" gridrow="1" pageId="12" pageNumber="13">
<th id="768C5DE3EF5DC5CAFCEFFDA9FC61FD43" box="[810,932,615,642]" gridcol="0" gridrow="1" pageId="12" pageNumber="13">Leg 1</th>
<td id="768C5DE3EF5DC5CAFC75FDA9FC2FFD43" box="[944,1002,615,642]" gridcol="1" gridrow="1" pageId="12" pageNumber="13">0-1</td>
<td id="768C5DE3EF5DC5CAFC32FDA9FBF7FD43" box="[1015,1074,615,642]" gridcol="2" gridrow="1" pageId="12" pageNumber="13">1-I</td>
<td id="768C5DE3EF5DC5CAFBF8FDA9FB2DFD43" box="[1085,1256,615,642]" gridcol="3" gridrow="1" pageId="12" pageNumber="13">I-1; I-1; III, I, 4</td>
<td id="768C5DE3EF5DC5CAFB3FFDA9FA63FD43" box="[1274,1446,615,642]" gridcol="4" gridrow="1" pageId="12" pageNumber="13">0-1; 0-1; 1, 2, 3</td>
</tr>
<tr id="355D349FEF5DC5CAFCEFFD47FA63FD65" box="[810,1446,649,676]" gridrow="2" pageId="12" pageNumber="13">
<th id="768C5DE3EF5DC5CAFCEFFD47FC61FD65" box="[810,932,649,676]" gridcol="0" gridrow="2" pageId="12" pageNumber="13">
Legs 2 &amp;
<td id="768C5DE3EF5DC5CAFC75FD47FC2FFD65" box="[944,1002,649,676]" gridcol="1" gridrow="2" pageId="12" pageNumber="13">30-1</td>
</th>
<td id="768C5DE3EF5DC5CAFC32FD47FBF7FD65" box="[1015,1074,649,676]" gridcol="2" gridrow="2" pageId="12" pageNumber="13">1-0</td>
<td id="768C5DE3EF5DC5CAFBF8FD47FB2DFD65" box="[1085,1256,649,676]" gridcol="3" gridrow="2" pageId="12" pageNumber="13">1-1; 1-1; 3, 1, 5</td>
<td id="768C5DE3EF5DC5CAFB3FFD47FA63FD65" box="[1274,1446,649,676]" gridcol="4" gridrow="2" pageId="12" pageNumber="13">0-1; 0-2; 1, 2, 3</td>
</tr>
<tr id="355D349FEF5DC5CAFCEFFD65FA63FD07" box="[810,1446,683,710]" gridrow="3" pageId="12" pageNumber="13">
<th id="768C5DE3EF5DC5CAFCEFFD65FC61FD07" box="[810,932,683,710]" gridcol="0" gridrow="3" pageId="12" pageNumber="13">Leg 4</th>
<td id="768C5DE3EF5DC5CAFC75FD65FC2FFD07" box="[944,1002,683,710]" gridcol="1" gridrow="3" pageId="12" pageNumber="13">0-1</td>
<td id="768C5DE3EF5DC5CAFC32FD65FBF7FD07" box="[1015,1074,683,710]" gridcol="2" gridrow="3" pageId="12" pageNumber="13">1-0</td>
<td id="768C5DE3EF5DC5CAFBF8FD65FB2DFD07" box="[1085,1256,683,710]" gridcol="3" gridrow="3" pageId="12" pageNumber="13">1-1; 1-1; 2, 1, 5</td>
<td id="768C5DE3EF5DC5CAFB3FFD65FA63FD07" box="[1274,1446,683,710]" gridcol="4" gridrow="3" pageId="12" pageNumber="13">0-1; 0-2; 1, 2, 2</td>
</tr>
</table>
</paragraph>
<paragraph id="8BD236DDEF5D3A39FF78FDA9FD19FD04" blockId="12.[136,778,173,1390]" pageId="12" pageNumber="13">Labrum, maxilla, and legs 1–4 as in female. Mandible with 9 setae on second endopodal segment. Maxillule with 2 or 3 setae on first endopodal segment, as in female.</paragraph>
<paragraph id="8BD236DDEF5D3A39FF78FD00FDB7FC8C" blockId="12.[136,778,173,1390]" pageId="12" pageNumber="13">
Leg 5 (
<figureCitation id="13562A58EF5D3A39FEE5FD03FEBAFD26" box="[288,383,717,743]" captionStart="FIGURE 3" captionStartId="11.[136,245,1567,1593]" captionTargetBox="[146,1444,210,1521]" captionTargetId="figure-211@11.[136,1451,200,1532]" captionTargetPageId="11" captionText="FIGURE 3. Bathynotodelphys defayeae gen. et sp. nov., male. A, habitus, dorsal; B, urosome, ventral; C, antennule. Scale bars: A, 0.5 mm; B, C, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4591144" httpUri="https://zenodo.org/record/4591144/files/figure.png" pageId="12" pageNumber="13">Fig. 3B</figureCitation>
) protopod articulated from fifth pedigerous somite; free exopodal segment about 3.2 times as long as wide (80×25 μm). Leg 6 represented by 3 setae on genital operculum, innermost seta small.
</paragraph>
</subSubSection>
<subSubSection id="C3776556EF5D3A39FF78FC9BFDE9FAAC" pageId="12" pageNumber="13" type="discussion">
<paragraph id="8BD236DDEF5D3A39FF78FC9BFF38FAE8" blockId="12.[136,778,173,1390]" pageId="12" pageNumber="13">
<emphasis id="B919EACFEF5D3A39FF78FC9BFEEFFCAE" bold="true" box="[189,298,853,879]" pageId="12" pageNumber="13">Remarks</emphasis>
. The sexual dimorphism in antennulary segmentation of the new genus is unique. In the female segments XII to XIV are fused whereas in the male they are free; segment XVII is incorporated into a compound segment XVII-XX in the female but segment XVII is separate from XVIII-XX in the male. More distally, segments XXIV and XXV are free in the female but incorporated into a compound apical segment XXIV-XXVIII in the male. So although the male of
<taxonomicName id="4C6D4D5EEF5D3A39FD42FBA8FCCFFBBE" authority="Kim &amp; Boxshall, 2020" authorityName="Kim &amp; Boxshall" authorityYear="2020" box="[647,778,1125,1152]" class="Maxillopoda" family="Ascidicolidae" genus="Bathynotodelphys" kingdom="Animalia" order="Cyclopoida" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="defayeae" status="gen. et sp. nov.">
<emphasis id="B919EACFEF5D3A39FD42FBA8FD5DFBBE" box="[647,664,1126,1151]" italics="true" pageId="12" pageNumber="13">B</emphasis>
.
<emphasis id="B919EACFEF5D3A39FD63FBABFCCFFBBE" box="[678,778,1125,1151]" italics="true" pageId="12" pageNumber="13">defayeae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22A57B4EF5D3A39FF4DFB49FEF8FB63" box="[136,317,1159,1186]" pageId="12" pageNumber="13" rank="species">
<emphasis id="B919EACFEF5D3A39FF4DFB49FEF2FB60" bold="true" box="[136,311,1159,1185]" pageId="12" pageNumber="13">gen. et sp. nov</emphasis>
.
</taxonomicNameLabel>
does not retain geniculate antennules in the male, the fusion pattern of its distal segments XXIV to XXVIII probably represents a vestige of the original pattern associated with the possession of a geniculate antennule.
</paragraph>
<paragraph id="8BD236DDEF5D3A39FF78FAFFFDE9FAAC" blockId="12.[136,778,173,1390]" pageId="12" pageNumber="13">
This species was found in deep water at a depth of
<quantity id="4C959B38EF5D3A39FF4DFA9DFEE6FAAF" box="[136,291,1363,1390]" metricMagnitude="3" metricUnit="m" metricValue="5.330499999999999" metricValueMax="5.332" metricValueMin="5.329" pageId="12" pageNumber="13" unit="m" value="5330.5" valueMax="5332.0" valueMin="5329.0">5329–5332 m</quantity>
in
<collectingCountry id="F37A764DEF5D3A39FE82FA9DFE18FAAC" box="[327,477,1363,1389]" name="Argentina" pageId="12" pageNumber="13">the Argentine</collectingCountry>
Basin.
</paragraph>
</subSubSection>
</treatment>
</document>
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