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<document id="85D6034A6BB1B38CA0BE0F97399E7756" ID-CLB-Dataset="45885" ID-DOI="10.5281/zenodo.205694" ID-GBIF-Dataset="fd0cd047-d39f-4ee1-9b28-d92386c1c437" ID-ISSN="1175-5326" ID-Zenodo-Dep="205694" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460414771815" checkinUser="plazi" docAuthor="Mikhaljova, Elena V., Golovatch, Sergei I. &amp; Chang, Hsueh-Wen" docDate="2011" docId="03A587F4D10FFFFFFF13FEE8B214F838" docLanguage="en" docName="zt03114p021.pdf" docOrigin="Zootaxa 3114" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Anaulaciulus multiarticulatus Mikhaljova, Golovatch &amp; Chang, 2011, sp. nov." docType="treatment" docVersion="11" lastPageNumber="11" masterDocId="FF9CFF8CD107FFF5FF84FF87B631FFAB" masterDocTitle="The millipede genus Anaulaciulus Pocock, 1895 in Taiwan, with descriptions of four new species (Diplopoda, Julida, Julidae)" masterLastPageNumber="21" masterPageNumber="1" pageNumber="9" updateTime="1698328502134" updateUser="ExternalLinkService">
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<mods:title id="8A386F5A3350CBC05950E86F45FFB0B0">The millipede genus Anaulaciulus Pocock, 1895 in Taiwan, with descriptions of four new species (Diplopoda, Julida, Julidae)</mods:title>
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<mods:namePart id="A3D55C875BB0C7E55699BA82E7535902">Mikhaljova, Elena V.</mods:namePart>
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<mods:namePart id="63DC28EA3DF770E4FC72B956F171C0D2">Golovatch, Sergei I.</mods:namePart>
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<mods:namePart id="D184993C72985D0924319B80D314E8EA">Chang, Hsueh-Wen</mods:namePart>
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<mods:date id="1DA449620D2105C4F3BEB763A89C5D4F">2011</mods:date>
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<mods:identifier id="78A8B2A6139802B2176895316E4D02C7" type="DOI">10.5281/zenodo.205694</mods:identifier>
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<treatment id="03A587F4D10FFFFFFF13FEE8B214F838" ID-DOI="http://doi.org/10.5281/zenodo.6183713" ID-GBIF-Taxon="119424640" ID-Zenodo-Dep="6183713" LSID="urn:lsid:plazi:treatment:03A587F4D10FFFFFFF13FEE8B214F838" httpUri="http://treatment.plazi.org/id/03A587F4D10FFFFFFF13FEE8B214F838" lastPageId="10" lastPageNumber="11" pageId="8" pageNumber="9">
<subSubSection id="C3166569D10FFFFDFF13FEE8B725FE01" pageId="8" pageNumber="9" type="nomenclature">
<paragraph id="8BB336E2D10FFFFDFF13FEE8B459FE22" blockId="8.[151,616,367,426]" box="[151,616,367,393]" pageId="8" pageNumber="9">
<heading id="D0FB818ED10FFFFDFF13FEE8B459FE22" bold="true" box="[151,616,367,393]" fontSize="11" level="1" pageId="8" pageNumber="9" reason="1">
<emphasis id="B978EAF0D10FFFFDFF13FEE8B459FE22" bold="true" box="[151,616,367,393]" pageId="8" pageNumber="9">
<taxonomicName id="4C0C4D61D10FFFFDFF13FEE8B432FE22" box="[151,515,367,393]" class="Diplopoda" family="Julidae" genus="Anaulaciulus" kingdom="Animalia" order="Julida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="multiarticulatus" status="sp. nov.">
<emphasis id="B978EAF0D10FFFFDFF13FEE8B432FE22" bold="true" box="[151,515,367,393]" italics="true" pageId="8" pageNumber="9">Anaulaciulus multiarticulatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24B578BD10FFFFDFD8EFEE8B459FE22" box="[522,616,367,393]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BB336E2D10FFFFDFF13FE15B725FE01" blockId="8.[151,616,367,426]" box="[151,276,402,426]" pageId="8" pageNumber="9">
<figureCitation id="13372A67D10FFFFDFF13FE15B725FE01" box="[151,276,402,426]" captionStart-0="FIGURES 27 34" captionStart-1="FIGURES 35 39" captionStartId-0="9.[151,264,1852,1875]" captionStartId-1="10.[151,264,989,1012]" captionTargetBox-0="[208,1393,320,1810]" captionTargetBox-1="[318,1266,216,966]" captionTargetId-0="figure@9.[181,1406,301,1832]" captionTargetId-1="figure@10.[309,1278,193,968]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionText-0="FIGURES 27 34. Anaulaciulus multiarticulatus sp. nov., male (27 32) and female (33 34) paratypes. 27, leg 1, front view; 28, leg 2 and penes; 29, promere, caudal view; 30, left opisthomere, caudal view; 31, left opisthomere, front view; 32, left opisthomere, lateral view; 33, right vulva of a smaller female from sample D 0293, caudal view; 34, distal part of vulva of a larger female from sample D 0293, caudal view; il, intermediate lamella of opisthomere; al, anterolateral lamella of opisthomere; ml, mesal lobe of opisthomere apex; fp, basal flap of opisthomere. Scale in mm." captionText-1="FIGURES 35 39. Anaulaciulus multiarticulatus sp. nov., male paratypes. Variation in shape of promere apex: 35 36, different individuals from sample D 0294, 37 39, different individuals from sample D 0293. Scale in mm." httpUri-0="https://zenodo.org/record/205701/files/figure.png" httpUri-1="https://zenodo.org/record/205702/files/figure.png" pageId="8" pageNumber="9">Figs 2739</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3166569D10FFFFDFF13FE5DB3ADFD29" pageId="8" pageNumber="9" type="materials_examined">
<paragraph id="8BB336E2D10FFFFDFF13FE5DB3ADFD29" blockId="8.[151,1437,474,2011]" pageId="8" pageNumber="9">
<emphasis id="B978EAF0D10FFFFDFF13FE5DB7B3FE58" bold="true" box="[151,386,474,499]" pageId="8" pageNumber="9">Material examined.</emphasis>
<typeStatus id="54B78840D10FFFFDFE08FE5CB7C3FE58" box="[396,498,475,499]" pageId="8" pageNumber="9" type="holotype">
<emphasis id="B978EAF0D10FFFFDFE08FE5CB7C3FE58" box="[396,498,475,499]" italics="true" pageId="8" pageNumber="9">Holotype</emphasis>
</typeStatus>
: male (
<collectionCode id="ED1DAE27D10FFFFDFDCFFE5DB491FE59" box="[587,672,474,498]" httpUri="http://grbio.org/cool/2nry-s1yg" name="National Museum of Natural Science" pageId="8" pageNumber="9">NMNS</collectionCode>
, 6702
<emphasis id="B978EAF0D10FFFFDFD62FE5CB4C5FE58" bold="true" box="[742,756,475,499]" pageId="8" pageNumber="9"></emphasis>
0 0 8, D
<emphasis id="B978EAF0D10FFFFDFCC7FE5CB560FE58" bold="true" box="[835,849,475,499]" pageId="8" pageNumber="9"></emphasis>
0293),
<collectingCountry id="F31B7672D10FFFFDFC26FE5DB5C6FE59" box="[930,1015,474,498]" name="Taiwan" pageId="8" pageNumber="9">Taiwan</collectingCountry>
, Nantou County, Huisun timberland,
<date id="FFB21022D10FFFFDFF13FE79B747FDBD" box="[151,374,510,534]" pageId="8" pageNumber="9" value="1997-12-27">27 December 1997</date>
, leg. S.H. Wu;
<typeStatus id="54B78840D10FFFFDFDA9FD87B4AFFDBC" box="[557,670,512,535]" pageId="8" pageNumber="9" type="paratype">
<emphasis id="B978EAF0D10FFFFDFDA9FD87B4AFFDBC" box="[557,670,512,535]" italics="true" pageId="8" pageNumber="9">Paratypes</emphasis>
</typeStatus>
:
<specimenCount id="9D0AFD6BD10FFFFDFD35FE78B53CFDBD" box="[689,781,510,534]" pageId="8" pageNumber="9" type="male">7 males</specimenCount>
,
<specimenCount id="9D0AFD6BD10FFFFDFC98FE78B5BFFDBD" box="[796,910,510,534]" pageId="8" pageNumber="9" type="female">7 females</specimenCount>
,
<specimenCount id="9D0AFD6BD10FFFFDFC19FE79B22CFDBD" box="[925,1053,510,534]" pageId="8" pageNumber="9" type="juvenile">5 juveniles</specimenCount>
, (
<collectionCode id="ED1DAE27D10FFFFDFBB0FE79B2B8FDBD" box="[1076,1161,510,534]" httpUri="http://grbio.org/cool/2nry-s1yg" name="National Museum of Natural Science" pageId="8" pageNumber="9">NMNS</collectionCode>
, 6702
<emphasis id="B978EAF0D10FFFFDFB4BFE78B2ECFDBC" bold="true" box="[1231,1245,511,535]" pageId="8" pageNumber="9"></emphasis>
0 0 9, D
<emphasis id="B978EAF0D10FFFFDFAA9FE78B30AFDBC" bold="true" box="[1325,1339,511,535]" pageId="8" pageNumber="9"></emphasis>
0293),
<specimenCount id="9D0AFD6BD10FFFFDFA0AFE79B6E1FD91" pageId="8" pageNumber="9" type="male">1 male</specimenCount>
,
<specimenCount id="9D0AFD6BD10FFFFDFF5FFDA5B777FD91" box="[219,326,546,570]" pageId="8" pageNumber="9" type="female">2 females</specimenCount>
(
<collectionCode id="ED1DAE27D10FFFFDFED1FDA5B7A4FD91" box="[341,405,546,570]" pageId="8" pageNumber="9">IBSS</collectionCode>
),
<specimenCount id="9D0AFD6BD10FFFFDFE2CFDA5B7C7FD91" box="[424,502,546,570]" pageId="8" pageNumber="9" type="male">1 male</specimenCount>
,
<specimenCount id="9D0AFD6BD10FFFFDFD84FDA5B450FD91" box="[512,609,546,570]" pageId="8" pageNumber="9" type="female">1 female</specimenCount>
(
<collectionCode id="ED1DAE27D10FFFFDFDF4FDA5B4FCFD91" box="[624,717,546,570]" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="8" pageNumber="9">ZMUM</collectionCode>
),
<specimenCount id="9D0AFD6BD10FFFFDFD5BFDA5B51CFD91" box="[735,813,546,570]" pageId="8" pageNumber="9" type="male">1 male</specimenCount>
,
<specimenCount id="9D0AFD6BD10FFFFDFCB3FDA5B5A9FD91" box="[823,920,546,570]" pageId="8" pageNumber="9" type="female">1 female</specimenCount>
(NSYSUB),
<specimenCount id="9D0AFD6BD10FFFFDFBA9FDA5B24AFD91" box="[1069,1147,546,570]" pageId="8" pageNumber="9" type="male">1 male</specimenCount>
,
<specimenCount id="9D0AFD6BD10FFFFDFB01FDA5B2D7FD91" box="[1157,1254,546,570]" pageId="8" pageNumber="9" type="female">1 female</specimenCount>
(
<collectionCode id="ED1DAE27D10FFFFDFB71FDA5B361FD91" LSID="urn:lsid:biocol.org:col:34991" box="[1269,1360,546,570]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="8" pageNumber="9">HNHM</collectionCode>
), same locality, together with
<typeStatus id="54B78840D10FFFFDFE19FDC1B433FDF5" box="[413,514,582,606]" pageId="8" pageNumber="9" type="holotype">holotype</typeStatus>
,
<date id="FFB21022D10FFFFDFD94FDC1B4DEFDF5" box="[528,751,582,606]" pageId="8" pageNumber="9" value="1997-12-27">27 December 1997</date>
;
<specimenCount id="9D0AFD6BD10FFFFDFD7BFDC1B569FDF5" box="[767,856,582,606]" pageId="8" pageNumber="9" type="male">4 males</specimenCount>
(
<specimenCount id="9D0AFD6BD10FFFFDFCEAFDC1B58AFDF5" box="[878,955,582,606]" pageId="8" pageNumber="9" type="male">1 male</specimenCount>
devoid of gonopods, presumably lost),
<specimenCount id="9D0AFD6BD10FFFFDFA0AFDC1B6D2FD29" pageId="8" pageNumber="9" type="female">1 female</specimenCount>
(
<collectionCode id="ED1DAE27D10FFFFDFF76FDEDB776FD29" box="[242,327,618,642]" httpUri="http://grbio.org/cool/2nry-s1yg" name="National Museum of Natural Science" pageId="8" pageNumber="9">NMNS</collectionCode>
, 6702
<emphasis id="B978EAF0D10FFFFDFE0EFDECB7A9FD28" bold="true" box="[394,408,619,643]" pageId="8" pageNumber="9"></emphasis>
0 10, D
<emphasis id="B978EAF0D10FFFFDFE60FDECB7C3FD28" bold="true" box="[484,498,619,643]" pageId="8" pageNumber="9"></emphasis>
0294),
<specimenCount id="9D0AFD6BD10FFFFDFDC5FDEDB4BEFD29" box="[577,655,618,642]" pageId="8" pageNumber="9" type="male">1 male</specimenCount>
,
<specimenCount id="9D0AFD6BD10FFFFDFD1FFDEDB4CDFD29" box="[667,764,618,642]" pageId="8" pageNumber="9" type="female">1 female</specimenCount>
(
<collectionCode id="ED1DAE27D10FFFFDFC8FFDEDB57AFD29" box="[779,843,618,642]" pageId="8" pageNumber="9">IBSS</collectionCode>
), same locality,
<date id="FFB21022D10FFFFDFC7BFDEDB2E6FD29" box="[1023,1239,618,642]" pageId="8" pageNumber="9" value="1997-12-27">27 December 1997</date>
, all leg. S.H. Wu.
</paragraph>
</subSubSection>
<subSubSection id="C3166569D10FFFFDFF41FD09B2F7FCB8" pageId="8" pageNumber="9" type="diagnosis">
<paragraph id="8BB336E2D10FFFFDFF41FD09B2F7FCB8" blockId="8.[151,1437,474,2011]" pageId="8" pageNumber="9">
<emphasis id="B978EAF0D10FFFFDFF41FD09B70EFD0C" bold="true" box="[197,319,654,679]" pageId="8" pageNumber="9">Diagnosis.</emphasis>
Differs from congeners mainly by the shape of the gonopod opisthomere showing an anterolateral lamella broadening at the lateral margin distally into a rounded lobe, as well as by the subrectangular promere with its apex ranging from rather clearly rounded to excavated. This new species differs from the obviously especially similar
<taxonomicName id="4C0C4D61D10FFFFDFF68FD7CB7FEFCB8" box="[236,463,763,787]" class="Diplopoda" family="Julidae" genus="Anaulaciulus" kingdom="Animalia" order="Julida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="oligosegmentatus" status="sp. nov.">
<emphasis id="B978EAF0D10FFFFDFF68FD7CB7FEFCB8" box="[236,463,763,787]" italics="true" pageId="8" pageNumber="9">A. oligosegmentatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24B578BD10FFFFDFE51FD7CB41EFCB8" box="[469,559,763,787]" pageId="8" pageNumber="9" rank="species">
<emphasis id="B978EAF0D10FFFFDFE51FD7CB41EFCB8" bold="true" box="[469,559,763,787]" pageId="8" pageNumber="9">sp. nov.</emphasis>
</taxonomicNameLabel>
by the lesser body diameter and more numerous segments.
</paragraph>
</subSubSection>
<subSubSection id="C3166569D10FFFFFFF41FC99B71FFBD7" lastPageId="10" lastPageNumber="11" pageId="8" pageNumber="9" type="description">
<paragraph id="8BB336E2D10FFFFDFF41FC99B50BFBFD" blockId="8.[151,1437,474,2011]" pageId="8" pageNumber="9">
<emphasis id="B978EAF0D10FFFFDFF41FC99B766FC9C" bold="true" box="[197,343,798,823]" pageId="8" pageNumber="9">Description.</emphasis>
Male. Length
<quantity id="4CF49B07D10FFFFDFD80FC99B4B3FC9D" box="[516,642,798,823]" metricMagnitude="-2" metricUnit="m" metricValue="2.7" metricValueMax="3.5" metricValueMin="1.9" pageId="8" pageNumber="9" unit="mm" value="27.0" valueMax="35.0" valueMin="19.0">
19
<emphasis id="B978EAF0D10FFFFDFDA4FC98B41FFC9C" bold="true" box="[544,558,799,823]" pageId="8" pageNumber="9"></emphasis>
35 mm
</quantity>
, vertical diameter
<quantity id="4CF49B07D10FFFFDFCDFFC99B5D4FC9D" box="[859,997,798,823]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" metricValueMax="1.1" metricValueMin="0.9" pageId="8" pageNumber="9" unit="mm" value="1.0" valueMax="1.1" valueMin="0.9">
0.9
<emphasis id="B978EAF0D10FFFFDFCFAFC98B5BDFC9C" bold="true" box="[894,908,799,823]" pageId="8" pageNumber="9"></emphasis>
1.1 mm
</quantity>
(the overwhelming majority of males show a vertical diameter of 1.0 mm), with 58(
<emphasis id="B978EAF0D10FFFFDFD26FCC4B481FCF0" bold="true" box="[674,688,835,859]" pageId="8" pageNumber="9"></emphasis>
8), 59(
<emphasis id="B978EAF0D10FFFFDFD78FCC4B53BFCF0" bold="true" box="[764,778,835,859]" pageId="8" pageNumber="9"></emphasis>
9), 60(
<emphasis id="B978EAF0D10FFFFDFCD1FCC4B552FCF0" bold="true" box="[853,867,835,859]" pageId="8" pageNumber="9"></emphasis>
9), 61(
<emphasis id="B978EAF0D10FFFFDFC2BFCC4B58CFCF0" bold="true" box="[943,957,835,859]" pageId="8" pageNumber="9"></emphasis>
5), 63(
<emphasis id="B978EAF0D10FFFFDFB8DFCC4B226FCF0" bold="true" box="[1033,1047,835,859]" pageId="8" pageNumber="9"></emphasis>
6), 64(
<emphasis id="B978EAF0D10FFFFDFBE7FCC4B240FCF0" bold="true" box="[1123,1137,835,859]" pageId="8" pageNumber="9"></emphasis>
4), 65(
<emphasis id="B978EAF0D10FFFFDFB38FCC4B2FBFCF0" bold="true" box="[1212,1226,835,859]" pageId="8" pageNumber="9"></emphasis>
6), 66(
<emphasis id="B978EAF0D10FFFFDFA92FCC4B315FCF0" bold="true" box="[1302,1316,835,859]" pageId="8" pageNumber="9"></emphasis>
5), 67(
<emphasis id="B978EAF0D10FFFFDFAEBFCC4B34CFCF0" bold="true" box="[1391,1405,835,859]" pageId="8" pageNumber="9"></emphasis>
7), 68(
<emphasis id="B978EAF0D10FFFFDFF38FCE0B6FBFCD4" bold="true" box="[188,202,871,895]" pageId="8" pageNumber="9"></emphasis>
6), 69(
<emphasis id="B978EAF0D10FFFFDFE90FCE0B713FCD4" bold="true" box="[276,290,871,895]" pageId="8" pageNumber="9"></emphasis>
5), 69(
<emphasis id="B978EAF0D10FFFFDFEE8FCE0B74BFCD4" bold="true" box="[364,378,871,895]" pageId="8" pageNumber="9"></emphasis>
7), 80(
<emphasis id="B978EAF0D10FFFFDFE40FCE0B7E3FCD4" bold="true" box="[452,466,871,895]" pageId="8" pageNumber="9"></emphasis>
3) segments, excluding telson. A male with 64(
<emphasis id="B978EAF0D10FFFFDFC64FCE0B5DFFCD4" bold="true" box="[992,1006,871,895]" pageId="8" pageNumber="9"></emphasis>
4) body segments showing last podous segment with legs developed only on the right side. Coloration brown or light brown with three narrow, longitudinal, dark brown stripes, one mid-dorsal and two lateral level to ozopores, so that dorsum showing two somewhat lighter, broad, ribbon-shaped, longitudinal stripes, these being less distinct as compared to those in
<taxonomicName id="4C0C4D61D10FFFFDFB62FC54B6FFFBA4" class="Diplopoda" family="Julidae" genus="Anaulaciulus" kingdom="Animalia" order="Julida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="oligosegmentatus" status="sp. nov.">
<emphasis id="B978EAF0D10FFFFDFB62FC54B6FFFBA4" italics="true" pageId="8" pageNumber="9">A. oligosegmentatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24B578BD10FFFFDFF52FC70B703FBA4" box="[214,306,1015,1039]" pageId="8" pageNumber="9" rank="species">
<emphasis id="B978EAF0D10FFFFDFF52FC70B703FBA4" bold="true" box="[214,306,1015,1039]" pageId="8" pageNumber="9">sp. nov.</emphasis>
</taxonomicNameLabel>
Collum with two transversely-oval, large, yellowish spots. Segments in front of telson can also be with two transversely-oval yellowish spots on dorsal side. Clypeolabral part of head lighter. Anal valves lighter, as usual. Antennae and legs brown or beige, eye patches black.
</paragraph>
<paragraph id="8BB336E2D10FFFFDFF41FBE4B405FB69" blockId="8.[151,1437,474,2011]" pageId="8" pageNumber="9">
Eye patches subtriangular, composed of at least 26 ocelli. Labral setae as in
<taxonomicName id="4C0C4D61D10FFFFDFBA1FBE4B282FBD0" box="[1061,1203,1123,1147]" class="Diplopoda" family="Julidae" genus="Anaulaciulus" kingdom="Animalia" order="Julida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="immensus" status="sp. nov.">
<emphasis id="B978EAF0D10FFFFDFBA1FBE4B282FBD0" box="[1061,1203,1123,1147]" italics="true" pageId="8" pageNumber="9">A. immensus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24B578BD10FFFFDFB38FBE4B327FBD0" box="[1212,1302,1123,1147]" pageId="8" pageNumber="9" rank="species">
<emphasis id="B978EAF0D10FFFFDFB38FBE4B327FBD0" bold="true" box="[1212,1302,1123,1147]" pageId="8" pageNumber="9">sp. nov.</emphasis>
</taxonomicNameLabel>
Gnathochilarium as in A.
<taxonomicName id="4C0C4D61D10FFFFDFEC3FB01B43AFB35" box="[327,523,1158,1182]" class="Diplopoda" family="Julidae" genus="Anaulaciulus" kingdom="Animalia" order="Julida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="oligosegmentatus" status="sp. nov.">
<emphasis id="B978EAF0D10FFFFDFEC3FB01B43AFB35" box="[327,523,1158,1182]" italics="true" pageId="8" pageNumber="9">oligosegmentatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24B578BD10FFFFDFD96FB00B45DFB34" box="[530,620,1159,1183]" pageId="8" pageNumber="9" rank="species">
<emphasis id="B978EAF0D10FFFFDFD96FB00B45DFB34" bold="true" box="[530,620,1159,1183]" pageId="8" pageNumber="9">sp. nov.</emphasis>
</taxonomicNameLabel>
Length ratios of antennomeres 27 as 2.7:2.4:2.4:2.3:1.5:1, width ratios as 1.1:1.1:1.2:1.5:1.4:1, respectively.
</paragraph>
<paragraph id="8BB336E2D10FFFFDFF41FB49B3ADFAF8" blockId="8.[151,1437,474,2011]" pageId="8" pageNumber="9">
Ozopores lying behind stricture without touching it. Striations on metazona not reaching caudal margin, thus making metazona look like being rimmed caudally; 15
<emphasis id="B978EAF0D10FFFFDFC88FB74B52BFAA0" bold="true" box="[780,794,1267,1291]" pageId="8" pageNumber="9"></emphasis>
16 striae on metazonital surface between dorsal midline and ozopore, or 7
<emphasis id="B978EAF0D10FFFFDFEE4FA90B75FFA84" bold="true" box="[352,366,1303,1327]" pageId="8" pageNumber="9"></emphasis>
8 striae per square lying between ozopore and axial line. Antennomeres 5 and 6, striations on dorsum and caudal projection of epiproct (including a tiny claw-shaped process) as in
<taxonomicName id="4C0C4D61D10FFFFDFBDDFABCB30DFAF8" box="[1113,1340,1339,1363]" class="Diplopoda" family="Julidae" genus="Anaulaciulus" kingdom="Animalia" order="Julida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="oligosegmentatus" status="sp. nov.">
<emphasis id="B978EAF0D10FFFFDFBDDFABCB30DFAF8" box="[1113,1340,1339,1363]" italics="true" pageId="8" pageNumber="9">A. oligosegmentatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24B578BD10FFFFDFAC7FABCB3ADFAF8" box="[1347,1436,1339,1363]" pageId="8" pageNumber="9" rank="species">
<emphasis id="B978EAF0D10FFFFDFAC7FABCB3ADFAF8" bold="true" box="[1347,1436,1339,1363]" pageId="8" pageNumber="9">sp. nov.</emphasis>
</taxonomicNameLabel>
</paragraph>
<paragraph id="8BB336E2D10FFFFDFF41FAD8B475F981" blockId="8.[151,1437,474,2011]" pageId="8" pageNumber="9">
Legs as in
<taxonomicName id="4C0C4D61D10FFFFDFEC6FAD9B417FADD" box="[322,550,1374,1398]" class="Diplopoda" family="Julidae" genus="Anaulaciulus" kingdom="Animalia" order="Julida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="oligosegmentatus" status="sp. nov.">
<emphasis id="B978EAF0D10FFFFDFEC6FAD9B417FADD" box="[322,550,1374,1398]" italics="true" pageId="8" pageNumber="9">A. oligosegmentatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24B578BD10FFFFDFDABFAD8B4BBFADC" box="[559,650,1375,1399]" pageId="8" pageNumber="9" rank="species">
<emphasis id="B978EAF0D10FFFFDFDABFAD8B4BBFADC" bold="true" box="[559,650,1375,1399]" pageId="8" pageNumber="9">sp. nov.</emphasis>
</taxonomicNameLabel>
, but pads on femora absent. Anterior legs with narrow pads on postfemora. Claw at base with a presumably long filament ventrally, but without accessory claw dorsally. All claws examined under a microscope (seven claws in 2
<emphasis id="B978EAF0D10FFFFDFD31FA20B4F2FA14" bold="true" box="[693,707,1447,1471]" pageId="8" pageNumber="9"></emphasis>
3 individuals) showed a ventrobasal stump which could presumably be considered as the rest of a long, broken-off filament, but hardly a short filament. Legs 1 with strong setae ventrally and a ventral papillate knob on telopoditomere 2, apex with several low papillae (
<figureCitation id="13372A67D10FFFFDFB4DFA69B310F9AD" box="[1225,1313,1518,1542]" captionStart="FIGURES 27 34" captionStartId="9.[151,264,1852,1875]" captionTargetBox="[208,1393,320,1810]" captionTargetId="figure@9.[181,1406,301,1832]" captionTargetPageId="9" captionText="FIGURES 27 34. Anaulaciulus multiarticulatus sp. nov., male (27 32) and female (33 34) paratypes. 27, leg 1, front view; 28, leg 2 and penes; 29, promere, caudal view; 30, left opisthomere, caudal view; 31, left opisthomere, front view; 32, left opisthomere, lateral view; 33, right vulva of a smaller female from sample D 0293, caudal view; 34, distal part of vulva of a larger female from sample D 0293, caudal view; il, intermediate lamella of opisthomere; al, anterolateral lamella of opisthomere; ml, mesal lobe of opisthomere apex; fp, basal flap of opisthomere. Scale in mm." httpUri="https://zenodo.org/record/205701/files/figure.png" pageId="8" pageNumber="9">Fig. 27</figureCitation>
). Penes of medium length, deeply bifid (
<figureCitation id="13372A67D10FFFFDFE67F995B409F981" box="[483,568,1554,1578]" captionStart="FIGURES 27 34" captionStartId="9.[151,264,1852,1875]" captionTargetBox="[208,1393,320,1810]" captionTargetId="figure@9.[181,1406,301,1832]" captionTargetPageId="9" captionText="FIGURES 27 34. Anaulaciulus multiarticulatus sp. nov., male (27 32) and female (33 34) paratypes. 27, leg 1, front view; 28, leg 2 and penes; 29, promere, caudal view; 30, left opisthomere, caudal view; 31, left opisthomere, front view; 32, left opisthomere, lateral view; 33, right vulva of a smaller female from sample D 0293, caudal view; 34, distal part of vulva of a larger female from sample D 0293, caudal view; il, intermediate lamella of opisthomere; al, anterolateral lamella of opisthomere; ml, mesal lobe of opisthomere apex; fp, basal flap of opisthomere. Scale in mm." httpUri="https://zenodo.org/record/205701/files/figure.png" pageId="8" pageNumber="9">Fig. 28</figureCitation>
).
</paragraph>
<paragraph id="8BB336E2D10FFFFDFF41F9B1B579F911" blockId="8.[151,1437,474,2011]" pageId="8" pageNumber="9">
Gonopods. Promere subquadrate, its mesal margin slightly curved rectangularly caudad; flagellum relatively long; rudimentary telopodites well-visible (
<figureCitation id="13372A67D10FFFFDFD08F9DDB4D4F9D9" box="[652,741,1626,1650]" captionStart="FIGURES 27 34" captionStartId="9.[151,264,1852,1875]" captionTargetBox="[208,1393,320,1810]" captionTargetId="figure@9.[181,1406,301,1832]" captionTargetPageId="9" captionText="FIGURES 27 34. Anaulaciulus multiarticulatus sp. nov., male (27 32) and female (33 34) paratypes. 27, leg 1, front view; 28, leg 2 and penes; 29, promere, caudal view; 30, left opisthomere, caudal view; 31, left opisthomere, front view; 32, left opisthomere, lateral view; 33, right vulva of a smaller female from sample D 0293, caudal view; 34, distal part of vulva of a larger female from sample D 0293, caudal view; il, intermediate lamella of opisthomere; al, anterolateral lamella of opisthomere; ml, mesal lobe of opisthomere apex; fp, basal flap of opisthomere. Scale in mm." httpUri="https://zenodo.org/record/205701/files/figure.png" pageId="8" pageNumber="9">Fig. 29</figureCitation>
). Apex of promere varying from relatively well rounded to excavated with a rounded mesal outgrowth (
<figureCitation id="13372A67D10FFFFDFD0BF9F9B522F93D" box="[655,787,1662,1687]" captionStart="FIGURES 35 39" captionStartId="10.[151,264,989,1012]" captionTargetBox="[318,1266,216,966]" captionTargetId="figure@10.[309,1278,193,968]" captionTargetPageId="10" captionText="FIGURES 35 39. Anaulaciulus multiarticulatus sp. nov., male paratypes. Variation in shape of promere apex: 35 36, different individuals from sample D 0294, 37 39, different individuals from sample D 0293. Scale in mm." httpUri="https://zenodo.org/record/205702/files/figure.png" pageId="8" pageNumber="9">
Figs 35
<emphasis id="B978EAF0D10FFFFDFD62F9F8B4C5F93C" bold="true" box="[742,756,1663,1687]" pageId="8" pageNumber="9"></emphasis>
39
</figureCitation>
), this variation being observed even within a single individual, possibly as a result of abrasion with age (
<figureCitation id="13372A67D10FFFFDFD3DF925B50AF911" box="[697,827,1698,1722]" captionStart="FIGURES 35 39" captionStartId="10.[151,264,989,1012]" captionTargetBox="[318,1266,216,966]" captionTargetId="figure@10.[309,1278,193,968]" captionTargetPageId="10" captionText="FIGURES 35 39. Anaulaciulus multiarticulatus sp. nov., male paratypes. Variation in shape of promere apex: 35 36, different individuals from sample D 0294, 37 39, different individuals from sample D 0293. Scale in mm." httpUri="https://zenodo.org/record/205702/files/figure.png" pageId="8" pageNumber="9">Figs 37, 39</figureCitation>
).
</paragraph>
<paragraph id="8BB336E2D10FFFFDFF41F941B2F6F975" blockId="8.[151,1437,474,2011]" box="[197,1223,1734,1759]" pageId="8" pageNumber="9">
Opisthomere (
<figureCitation id="13372A67D10FFFFDFEE0F941B7D5F975" box="[356,484,1734,1759]" captionStart="FIGURES 27 34" captionStartId="9.[151,264,1852,1875]" captionTargetBox="[208,1393,320,1810]" captionTargetId="figure@9.[181,1406,301,1832]" captionTargetPageId="9" captionText="FIGURES 27 34. Anaulaciulus multiarticulatus sp. nov., male (27 32) and female (33 34) paratypes. 27, leg 1, front view; 28, leg 2 and penes; 29, promere, caudal view; 30, left opisthomere, caudal view; 31, left opisthomere, front view; 32, left opisthomere, lateral view; 33, right vulva of a smaller female from sample D 0293, caudal view; 34, distal part of vulva of a larger female from sample D 0293, caudal view; il, intermediate lamella of opisthomere; al, anterolateral lamella of opisthomere; ml, mesal lobe of opisthomere apex; fp, basal flap of opisthomere. Scale in mm." httpUri="https://zenodo.org/record/205701/files/figure.png" pageId="8" pageNumber="9">
Figs 30
<emphasis id="B978EAF0D10FFFFDFE3DF940B7F6F974" bold="true" box="[441,455,1735,1759]" pageId="8" pageNumber="9"></emphasis>
32
</figureCitation>
) as in
<taxonomicName id="4C0C4D61D10FFFFDFDAAF941B520F975" box="[558,785,1734,1758]" class="Diplopoda" family="Julidae" genus="Anaulaciulus" kingdom="Animalia" order="Julida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="oligosegmentatus" status="sp. nov.">
<emphasis id="B978EAF0D10FFFFDFDAAF941B520F975" box="[558,785,1734,1758]" italics="true" pageId="8" pageNumber="9">A. oligosegmentatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24B578BD10FFFFDFC9CF940B543F974" box="[792,882,1735,1759]" pageId="8" pageNumber="9" rank="species">
<emphasis id="B978EAF0D10FFFFDFC9CF940B543F974" bold="true" box="[792,882,1735,1759]" pageId="8" pageNumber="9">sp. nov.</emphasis>
</taxonomicNameLabel>
, but distal part of
<emphasis id="B978EAF0D10FFFFDFBBBF940B264F974" bold="true" box="[1087,1109,1735,1759]" pageId="8" pageNumber="9">al</emphasis>
narrower.
</paragraph>
<paragraph id="8BB336E2D10FFFFCFF41F96DB76BFF7F" blockId="8.[151,1437,474,2011]" lastBlockId="9.[151,1436,151,248]" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9">
Female. Length
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17.5
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27 mm
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, vertical diameter 1.0
<emphasis id="B978EAF0D10FFFFDFC84F96CB53FF8A8" bold="true" box="[768,782,1771,1795]" pageId="8" pageNumber="9"></emphasis>
<quantity id="4CF49B07D10FFFFDFC8AF96DB556F8A9" box="[782,871,1770,1794]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="8" pageNumber="9" unit="mm" value="1.4">1.4 mm</quantity>
, with 53(
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6), 53(
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7), 54(
<emphasis id="B978EAF0D10FFFFDFB07F96CB2A0F8A8" bold="true" box="[1155,1169,1771,1795]" pageId="8" pageNumber="9"></emphasis>
7), 54(
<emphasis id="B978EAF0D10FFFFDFB58F96CB2DBF8A8" bold="true" box="[1244,1258,1771,1795]" pageId="8" pageNumber="9"></emphasis>
8), 57(
<emphasis id="B978EAF0D10FFFFDFAB1F96CB372F8A8" bold="true" box="[1333,1347,1771,1795]" pageId="8" pageNumber="9"></emphasis>
4), 58(
<emphasis id="B978EAF0D10FFFFDFA0AF96CB3ADF8A8" bold="true" box="[1422,1436,1771,1795]" pageId="8" pageNumber="9"></emphasis>
5), 59(
<emphasis id="B978EAF0D10FFFFDFF67F888B6C0F88C" bold="true" box="[227,241,1807,1831]" pageId="8" pageNumber="9"></emphasis>
5), 60(
<emphasis id="B978EAF0D10FFFFDFEB8F888B77BF88C" bold="true" box="[316,330,1807,1831]" pageId="8" pageNumber="9"></emphasis>
3), 63(
<emphasis id="B978EAF0D10FFFFDFE12F888B795F88C" bold="true" box="[406,420,1807,1831]" pageId="8" pageNumber="9"></emphasis>
4) segments, excluding telson. Claws with long ventral filaments only, these often broken off, leaving short stumps instead. Vulvae as in
<figureCitation id="13372A67D10FFFFDFD36F8B5B536F8E1" box="[690,775,1842,1866]" captionStart="FIGURES 27 34" captionStartId="9.[151,264,1852,1875]" captionTargetBox="[208,1393,320,1810]" captionTargetId="figure@9.[181,1406,301,1832]" captionTargetPageId="9" captionText="FIGURES 27 34. Anaulaciulus multiarticulatus sp. nov., male (27 32) and female (33 34) paratypes. 27, leg 1, front view; 28, leg 2 and penes; 29, promere, caudal view; 30, left opisthomere, caudal view; 31, left opisthomere, front view; 32, left opisthomere, lateral view; 33, right vulva of a smaller female from sample D 0293, caudal view; 34, distal part of vulva of a larger female from sample D 0293, caudal view; il, intermediate lamella of opisthomere; al, anterolateral lamella of opisthomere; ml, mesal lobe of opisthomere apex; fp, basal flap of opisthomere. Scale in mm." httpUri="https://zenodo.org/record/205701/files/figure.png" pageId="8" pageNumber="9">Fig. 33</figureCitation>
. Operculum longer than bursa, with two rows of setae on anterior face (6
<emphasis id="B978EAF0D10FFFFDFEC6F8D0B761F8C4" bold="true" box="[322,336,1879,1903]" pageId="8" pageNumber="9"></emphasis>
8 setae, i.e.
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3
<emphasis id="B978EAF0D10FFFFDFE60F8D0B7C3F8C4" bold="true" box="[484,498,1879,1903]" pageId="8" pageNumber="9"></emphasis>
4 in
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each row, in the vulvae examined). Apical margin of operculum with three low prominences, of which middle one either well-developed (
<figureCitation id="13372A67D10FFFFDFCA5F8FDB540F839" box="[801,881,1914,1938]" captionStart="FIGURES 27 34" captionStartId="9.[151,264,1852,1875]" captionTargetBox="[208,1393,320,1810]" captionTargetId="figure@9.[181,1406,301,1832]" captionTargetPageId="9" captionText="FIGURES 27 34. Anaulaciulus multiarticulatus sp. nov., male (27 32) and female (33 34) paratypes. 27, leg 1, front view; 28, leg 2 and penes; 29, promere, caudal view; 30, left opisthomere, caudal view; 31, left opisthomere, front view; 32, left opisthomere, lateral view; 33, right vulva of a smaller female from sample D 0293, caudal view; 34, distal part of vulva of a larger female from sample D 0293, caudal view; il, intermediate lamella of opisthomere; al, anterolateral lamella of opisthomere; ml, mesal lobe of opisthomere apex; fp, basal flap of opisthomere. Scale in mm." httpUri="https://zenodo.org/record/205701/files/figure.png" pageId="8" pageNumber="9">Fig. 34</figureCitation>
) or not (
<figureCitation id="13372A67D10FFFFDFC54F8FDB212F839" box="[976,1059,1914,1938]" captionStart="FIGURES 27 34" captionStartId="9.[151,264,1852,1875]" captionTargetBox="[208,1393,320,1810]" captionTargetId="figure@9.[181,1406,301,1832]" captionTargetPageId="9" captionText="FIGURES 27 34. Anaulaciulus multiarticulatus sp. nov., male (27 32) and female (33 34) paratypes. 27, leg 1, front view; 28, leg 2 and penes; 29, promere, caudal view; 30, left opisthomere, caudal view; 31, left opisthomere, front view; 32, left opisthomere, lateral view; 33, right vulva of a smaller female from sample D 0293, caudal view; 34, distal part of vulva of a larger female from sample D 0293, caudal view; il, intermediate lamella of opisthomere; al, anterolateral lamella of opisthomere; ml, mesal lobe of opisthomere apex; fp, basal flap of opisthomere. Scale in mm." httpUri="https://zenodo.org/record/205701/files/figure.png" pageId="8" pageNumber="9">Fig. 33</figureCitation>
). Lateral and mesal outgrowths at operculums apex varying from rounded (
<figureCitation id="13372A67D10FFFFDFDE9F819B48EF81D" box="[621,703,1950,1974]" captionStart="FIGURES 27 34" captionStartId="9.[151,264,1852,1875]" captionTargetBox="[208,1393,320,1810]" captionTargetId="figure@9.[181,1406,301,1832]" captionTargetPageId="9" captionText="FIGURES 27 34. Anaulaciulus multiarticulatus sp. nov., male (27 32) and female (33 34) paratypes. 27, leg 1, front view; 28, leg 2 and penes; 29, promere, caudal view; 30, left opisthomere, caudal view; 31, left opisthomere, front view; 32, left opisthomere, lateral view; 33, right vulva of a smaller female from sample D 0293, caudal view; 34, distal part of vulva of a larger female from sample D 0293, caudal view; il, intermediate lamella of opisthomere; al, anterolateral lamella of opisthomere; ml, mesal lobe of opisthomere apex; fp, basal flap of opisthomere. Scale in mm." httpUri="https://zenodo.org/record/205701/files/figure.png" pageId="8" pageNumber="9">Fig. 33</figureCitation>
) to acute (
<figureCitation id="13372A67D10FFFFDFCBCF819B5BCF81D" box="[824,909,1950,1974]" captionStart="FIGURES 27 34" captionStartId="9.[151,264,1852,1875]" captionTargetBox="[208,1393,320,1810]" captionTargetId="figure@9.[181,1406,301,1832]" captionTargetPageId="9" captionText="FIGURES 27 34. Anaulaciulus multiarticulatus sp. nov., male (27 32) and female (33 34) paratypes. 27, leg 1, front view; 28, leg 2 and penes; 29, promere, caudal view; 30, left opisthomere, caudal view; 31, left opisthomere, front view; 32, left opisthomere, lateral view; 33, right vulva of a smaller female from sample D 0293, caudal view; 34, distal part of vulva of a larger female from sample D 0293, caudal view; il, intermediate lamella of opisthomere; al, anterolateral lamella of opisthomere; ml, mesal lobe of opisthomere apex; fp, basal flap of opisthomere. Scale in mm." httpUri="https://zenodo.org/record/205701/files/figure.png" pageId="8" pageNumber="9">Fig. 34</figureCitation>
). Variation in structure of vulva in one and the same individual not noted,
<figureCitation id="13372A67D10FFFFDFE55F845B492F871" box="[465,675,1986,2010]" captionStart="FIGURES 27 34" captionStartId="9.[151,264,1852,1875]" captionTargetBox="[208,1393,320,1810]" captionTargetId="figure@9.[181,1406,301,1832]" captionTargetPageId="9" captionText="FIGURES 27 34. Anaulaciulus multiarticulatus sp. nov., male (27 32) and female (33 34) paratypes. 27, leg 1, front view; 28, leg 2 and penes; 29, promere, caudal view; 30, left opisthomere, caudal view; 31, left opisthomere, front view; 32, left opisthomere, lateral view; 33, right vulva of a smaller female from sample D 0293, caudal view; 34, distal part of vulva of a larger female from sample D 0293, caudal view; il, intermediate lamella of opisthomere; al, anterolateral lamella of opisthomere; ml, mesal lobe of opisthomere apex; fp, basal flap of opisthomere. Scale in mm." httpUri="https://zenodo.org/record/205701/files/figure.png" pageId="8" pageNumber="9">Figures 33 and 34</figureCitation>
showing vulvae of different specimens from the same sample D
<emphasis id="B978EAF0D10FFFFDFA0AF844B3ADF870" bold="true" box="[1422,1436,1987,2011]" pageId="8" pageNumber="9"></emphasis>
0 293. Bursa setose, with four setae both on lateral and mesal sclerites, six setae on each side of median plate of posterior sclerite.
</paragraph>
<paragraph id="8BB336E2D10EFFFCFF41FF58B2DEFF5C" blockId="9.[151,1436,151,248]" box="[197,1263,223,248]" pageId="9" pageNumber="10">
Juvenile. Length not exceeding
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9
<emphasis id="B978EAF0D10EFFFCFDBCFF67B477FF53" bold="true" box="[568,582,224,248]" pageId="9" pageNumber="10"></emphasis>
10 mm
</quantity>
, body with 9
<emphasis id="B978EAF0D10EFFFCFCACFF67B507FF53" bold="true" box="[808,822,224,248]" pageId="9" pageNumber="10"></emphasis>
10 apodous segments in front of telson.
</paragraph>
<caption id="DF73666AD10EFFFCFF13F8BBB5ACF865" httpUri="https://zenodo.org/record/205701/files/figure.png" pageId="9" pageNumber="10" targetBox="[208,1393,320,1810]" targetPageId="9">
<paragraph id="8BB336E2D10EFFFCFF13F8BBB5ACF865" blockId="9.[151,1436,1852,1998]" pageId="9" pageNumber="10">
<emphasis id="B978EAF0D10EFFFCFF13F8BBB77CF8F9" bold="true" box="[151,333,1852,1875]" pageId="9" pageNumber="10">FIGURES 2734</emphasis>
.
<emphasis id="B978EAF0D10EFFFCFEDFF8BAB4C5F8F9" bold="true" box="[347,756,1853,1875]" pageId="9" pageNumber="10">
<taxonomicName id="4C0C4D61D10EFFFCFEDFF8BAB4A4F8F8" box="[347,661,1853,1875]" class="Diplopoda" family="Julidae" genus="Anaulaciulus" kingdom="Animalia" order="Julida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="multiarticulatus" status="sp. nov.">
<emphasis id="B978EAF0D10EFFFCFEDFF8BAB4A4F8F8" bold="true" box="[347,661,1853,1875]" italics="true" pageId="9" pageNumber="10">Anaulaciulus multiarticulatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24B578BD10EFFFCFD18F8BAB4DEF8F9" box="[668,751,1853,1874]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
,
</emphasis>
male (27
<emphasis id="B978EAF0D10EFFFCFCD0F8BAB551F8F9" bold="true" box="[852,864,1853,1874]" pageId="9" pageNumber="10"></emphasis>
32
<emphasis id="B978EAF0D10EFFFCFCFEF8BBB5B2F8F8" bold="true" box="[890,899,1852,1875]" pageId="9" pageNumber="10">)</emphasis>
and female (33
<emphasis id="B978EAF0D10EFFFCFBA6F8BAB21EF8F9" bold="true" box="[1058,1071,1853,1874]" pageId="9" pageNumber="10"></emphasis>
34
<emphasis id="B978EAF0D10EFFFCFBCCF8BBB261F8F8" bold="true" box="[1096,1104,1852,1875]" pageId="9" pageNumber="10">)</emphasis>
paratypes. 27, leg 1, front view; 28, leg 2 and penes; 29, promere, caudal view; 30, left opisthomere, caudal view; 31, left opisthomere, front view; 32, left opisthomere, lateral view; 33, right vulva of a smaller female from sample D0293, caudal view; 34, distal part of vulva of a larger female from sample D0293, caudal view;
<emphasis id="B978EAF0D10EFFFCFDCFF81EB468F81B" bold="true" box="[587,601,1945,1968]" pageId="9" pageNumber="10">il</emphasis>
, intermediate lamella of opisthomere;
<emphasis id="B978EAF0D10EFFFCFC5BF81DB5C8F804" bold="true" box="[991,1017,1946,1967]" pageId="9" pageNumber="10">al,</emphasis>
anterolateral lamella of opisthomere;
<emphasis id="B978EAF0D10EFFFCFAFDF81DB3ADF804" bold="true" box="[1401,1436,1946,1967]" pageId="9" pageNumber="10">ml,</emphasis>
mesal lobe of opisthomere apex;
<emphasis id="B978EAF0D10EFFFCFE62F830B432F865" bold="true" box="[486,515,1975,1998]" pageId="9" pageNumber="10">fp,</emphasis>
basal flap of opisthomere. Scale in mm.
</paragraph>
</caption>
<caption id="DF73666AD10DFFFFFF13FC5AB291FBB9" httpUri="https://zenodo.org/record/205702/files/figure.png" pageId="10" pageNumber="11" targetBox="[318,1266,216,966]" targetPageId="10">
<paragraph id="8BB336E2D10DFFFFFF13FC5AB291FBB9" blockId="10.[151,1436,989,1042]" pageId="10" pageNumber="11">
<emphasis id="B978EAF0D10DFFFFFF13FC5AB77CFC58" bold="true" box="[151,333,989,1012]" pageId="10" pageNumber="11">FIGURES 3539</emphasis>
.
<emphasis id="B978EAF0D10DFFFFFEDEFC5AB4C2FC58" bold="true" box="[346,755,989,1011]" pageId="10" pageNumber="11">
<taxonomicName id="4C0C4D61D10DFFFFFEDEFC5AB4A5FC58" box="[346,660,989,1011]" class="Diplopoda" family="Julidae" genus="Anaulaciulus" kingdom="Animalia" order="Julida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="multiarticulatus" status="sp. nov.">
<emphasis id="B978EAF0D10DFFFFFEDEFC5AB4A5FC58" bold="true" box="[346,660,989,1011]" italics="true" pageId="10" pageNumber="11">Anaulaciulus multiarticulatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24B578BD10DFFFFFD1FFC59B4DFFC58" box="[667,750,990,1011]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
,
</emphasis>
male paratypes. Variation in shape of promere apex: 3536, different individuals from sample D0294, 3739, different individuals from sample D0293. Scale in mm.
</paragraph>
</caption>
<paragraph id="8BB336E2D10DFFFFFF41FBB8B71FFBD7" blockId="10.[151,1436,1087,1940]" pageId="10" pageNumber="11">
<emphasis id="B978EAF0D10DFFFFFF41FBB8B723FBFC" bold="true" box="[197,274,1087,1111]" pageId="10" pageNumber="11">Name.</emphasis>
The specific epithet refers to the more numerous body segments as compared to that in
<taxonomicName id="4C0C4D61D10DFFFFFB62FBB8B6FCFBD0" class="Diplopoda" family="Julidae" genus="Anaulaciulus" kingdom="Animalia" order="Julida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="oligosegmentatus" status="sp. nov.">
<emphasis id="B978EAF0D10DFFFFFB62FBB8B6FCFBD0" italics="true" pageId="10" pageNumber="11">A. oligosegmentatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24B578BD10DFFFFFF50FBE3B71FFBD7" box="[212,302,1124,1148]" pageId="10" pageNumber="11" rank="species">
<emphasis id="B978EAF0D10DFFFFFF50FBE3B71FFBD7" bold="true" box="[212,302,1124,1148]" pageId="10" pageNumber="11">sp. nov.</emphasis>
</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="C3166569D10DFFFFFF41FB00B214F838" pageId="10" pageNumber="11" type="discussion">
<paragraph id="8BB336E2D10DFFFFFF41FB00B6FBFA48" blockId="10.[151,1436,1087,1940]" pageId="10" pageNumber="11">
<emphasis id="B978EAF0D10DFFFFFF41FB00B708FB34" bold="true" box="[197,313,1159,1183]" pageId="10" pageNumber="11">Remarks.</emphasis>
Superficially,
<taxonomicName id="4C0C4D61D10DFFFFFE5AFB00B480FB34" box="[478,689,1159,1183]" class="Diplopoda" family="Julidae" genus="Anaulaciulus" kingdom="Animalia" order="Julida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="multiarticulatus" status="sp. nov.">
<emphasis id="B978EAF0D10DFFFFFE5AFB00B480FB34" box="[478,689,1159,1183]" italics="true" pageId="10" pageNumber="11">A. multiarticulatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24B578BD10DFFFFFD3DFB00B525FB34" box="[697,788,1159,1183]" pageId="10" pageNumber="11" rank="species">
<emphasis id="B978EAF0D10DFFFFFD3DFB00B525FB34" bold="true" box="[697,788,1159,1183]" pageId="10" pageNumber="11">sp. nov.</emphasis>
</taxonomicNameLabel>
is rather similar to
<taxonomicName id="4C0C4D61D10DFFFFFC77FB00B2E7FB34" box="[1011,1238,1159,1183]" class="Diplopoda" family="Julidae" genus="Anaulaciulus" kingdom="Animalia" order="Julida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="oligosegmentatus" status="sp. nov.">
<emphasis id="B978EAF0D10DFFFFFC77FB00B2E7FB34" box="[1011,1238,1159,1183]" italics="true" pageId="10" pageNumber="11">A. oligosegmentatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24B578BD10DFFFFFB5AFB00B308FB34" box="[1246,1337,1159,1183]" pageId="10" pageNumber="11" rank="species">
<emphasis id="B978EAF0D10DFFFFFB5AFB00B308FB34" bold="true" box="[1246,1337,1159,1183]" pageId="10" pageNumber="11">sp. nov.</emphasis>
</taxonomicNameLabel>
, but differs mainly by the slenderer body and more numerous segments. In most cases, males of
<taxonomicName id="4C0C4D61D10DFFFFFB19FB2CB340FB68" box="[1181,1393,1195,1219]" class="Diplopoda" family="Julidae" genus="Anaulaciulus" kingdom="Animalia" order="Julida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="multiarticulatus" status="sp. nov.">
<emphasis id="B978EAF0D10DFFFFFB19FB2CB340FB68" box="[1181,1393,1195,1219]" italics="true" pageId="10" pageNumber="11">A. multiarticulatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24B578BD10DFFFFFAFEFB2BB6F9FB4C" pageId="10" pageNumber="11" rank="species">
<emphasis id="B978EAF0D10DFFFFFAFEFB2BB6F9FB4C" bold="true" pageId="10" pageNumber="11">sp. nov.</emphasis>
</taxonomicNameLabel>
are longer than those of
<taxonomicName id="4C0C4D61D10DFFFFFE5BFB48B4F2FB4C" box="[479,707,1231,1255]" class="Diplopoda" family="Julidae" genus="Anaulaciulus" kingdom="Animalia" order="Julida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="oligosegmentatus" status="sp. nov.">
<emphasis id="B978EAF0D10DFFFFFE5BFB48B4F2FB4C" box="[479,707,1231,1255]" italics="true" pageId="10" pageNumber="11">A. oligosegmentatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24B578BD10DFFFFFD4EFB48B515FB4C" box="[714,804,1231,1255]" pageId="10" pageNumber="11" rank="species">
<emphasis id="B978EAF0D10DFFFFFD4EFB48B515FB4C" bold="true" box="[714,804,1231,1255]" pageId="10" pageNumber="11">sp. nov.</emphasis>
</taxonomicNameLabel>
One could suggest that the slenderer body in the former species is attained due to its greater length. Yet this is not so. In these two species, males of the same or a similar length show different diameters:
<quantity id="4CF49B07D10DFFFFFD83FA90B4BCFA84" box="[519,653,1303,1327]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" metricValueMax="1.1" metricValueMin="0.9" pageId="10" pageNumber="11" unit="mm" value="1.0" valueMax="1.1" valueMin="0.9">
0.9
<emphasis id="B978EAF0D10DFFFFFDAEFA90B409FA84" bold="true" box="[554,568,1303,1327]" pageId="10" pageNumber="11"></emphasis>
1.1 mm
</quantity>
in
<taxonomicName id="4C0C4D61D10DFFFFFD34FA90B5B3FA84" box="[688,898,1303,1327]" class="Diplopoda" family="Julidae" genus="Anaulaciulus" kingdom="Animalia" order="Julida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="multiarticulatus" status="sp. nov.">
<emphasis id="B978EAF0D10DFFFFFD34FA90B5B3FA84" box="[688,898,1303,1327]" italics="true" pageId="10" pageNumber="11">A. multiarticulatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24B578BD10DFFFFFC0CFA90B5D0FA84" box="[904,993,1303,1327]" pageId="10" pageNumber="11" rank="species">
<emphasis id="B978EAF0D10DFFFFFC0CFA90B5D0FA84" bold="true" box="[904,993,1303,1327]" pageId="10" pageNumber="11">sp. nov.</emphasis>
</taxonomicNameLabel>
, versus 1.0
<quantity id="4CF49B07D10DFFFFFBF4FA90B2F5FA84" box="[1136,1220,1303,1327]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="10" pageNumber="11" unit="mm" value="1.3">1.3 mm</quantity>
in
<taxonomicName id="4C0C4D61D10DFFFFFB62FA90B6FCFAF8" class="Diplopoda" family="Julidae" genus="Anaulaciulus" kingdom="Animalia" order="Julida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="oligosegmentatus" status="sp. nov.">
<emphasis id="B978EAF0D10DFFFFFB62FA90B6FCFAF8" italics="true" pageId="10" pageNumber="11">A. oligosegmentatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24B578BD10DFFFFFF50FABBB71FFAFF" box="[212,302,1340,1364]" pageId="10" pageNumber="11" rank="species">
<emphasis id="B978EAF0D10DFFFFFF50FABBB71FFAFF" bold="true" box="[212,302,1340,1364]" pageId="10" pageNumber="11">sp. nov.</emphasis>
</taxonomicNameLabel>
In addition, longitudinal stripes are far less distinct, up to virtually missing, on the dorsum of
<taxonomicName id="4C0C4D61D10DFFFFFACEFABCB711FADC" class="Diplopoda" family="Julidae" genus="Anaulaciulus" kingdom="Animalia" order="Julida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="multiarticulatus" status="sp. nov.">
<emphasis id="B978EAF0D10DFFFFFACEFABCB711FADC" italics="true" pageId="10" pageNumber="11">A. multiarticulatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24B578BD10DFFFFFEA3FAD8B7B0FADC" box="[295,385,1375,1399]" pageId="10" pageNumber="11" rank="species">
<emphasis id="B978EAF0D10DFFFFFEA3FAD8B7B0FADC" bold="true" box="[295,385,1375,1399]" pageId="10" pageNumber="11">sp. nov.</emphasis>
</taxonomicNameLabel>
, with most of the specimens (except one) also showing more numerous apodous segments in front of the telson. This latter is evidence of further potential growth, meaning the animals would become still longer and attain more segments. In other words, this is a stable feature to securely distinguish both of these new species.
</paragraph>
<paragraph id="8BB336E2D10DFFFFFF41FA77B50FF974" blockId="10.[151,1436,1087,1940]" pageId="10" pageNumber="11">
As regards gonopod structure, the main differences are observed in the structure of the promere apex: however variable and partly overlapping (= regularly convex) in shape, it is sometimes excavate in
<taxonomicName id="4C0C4D61D10DFFFFFB1AF994B343F980" box="[1182,1394,1555,1579]" class="Diplopoda" family="Julidae" genus="Anaulaciulus" kingdom="Animalia" order="Julida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="multiarticulatus" status="sp. nov.">
<emphasis id="B978EAF0D10DFFFFFB1AF994B343F980" box="[1182,1394,1555,1579]" italics="true" pageId="10" pageNumber="11">A. multiarticulatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24B578BD10DFFFFFAFEF993B6F9F9E4" pageId="10" pageNumber="11" rank="species">
<emphasis id="B978EAF0D10DFFFFFAFEF993B6F9F9E4" bold="true" pageId="10" pageNumber="11">sp. nov.</emphasis>
</taxonomicNameLabel>
, but never so in
<taxonomicName id="4C0C4D61D10DFFFFFE00F9B0B459F9E4" box="[388,616,1591,1615]" class="Diplopoda" family="Julidae" genus="Anaulaciulus" kingdom="Animalia" order="Julida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="oligosegmentatus" status="sp. nov.">
<emphasis id="B978EAF0D10DFFFFFE00F9B0B459F9E4" box="[388,616,1591,1615]" italics="true" pageId="10" pageNumber="11">A. oligosegmentatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24B578BD10DFFFFFDEBF9B0B4F8F9E4" box="[623,713,1591,1615]" pageId="10" pageNumber="11" rank="species">
<emphasis id="B978EAF0D10DFFFFFDEBF9B0B4F8F9E4" bold="true" box="[623,713,1591,1615]" pageId="10" pageNumber="11">sp. nov.</emphasis>
</taxonomicNameLabel>
, ranging from regularly convex to excavate (
<figureCitation id="13372A67D10DFFFFFB41F9B0B374F9E4" box="[1221,1349,1591,1615]" captionStart="FIGURES 35 39" captionStartId="10.[151,264,989,1012]" captionTargetBox="[318,1266,216,966]" captionTargetId="figure@10.[309,1278,193,968]" captionTargetPageId="10" captionText="FIGURES 35 39. Anaulaciulus multiarticulatus sp. nov., male paratypes. Variation in shape of promere apex: 35 36, different individuals from sample D 0294, 37 39, different individuals from sample D 0293. Scale in mm." httpUri="https://zenodo.org/record/205702/files/figure.png" pageId="10" pageNumber="11">Figs 3539</figureCitation>
) versus oval to regularly convex (
<figureCitation id="13372A67D10DFFFFFE32F9DCB409F9D8" box="[438,568,1627,1651]" captionStart="FIGURES 21 23" captionStartId="7.[151,264,757,780]" captionTargetBox="[374,1202,218,665]" captionTargetId="figure@7.[355,1232,193,736]" captionTargetPageId="7" captionText="FIGURES 21 23. Anaulaciulus oligosegmentatus sp. nov., male paratypes. Variation in shape of promere apex: 21, individual from sample D 0620, 22, individual from sample D 0298, 23, individual from sample D 0011. Scale in mm." httpUri="https://zenodo.org/record/205699/files/figure.png" pageId="10" pageNumber="11">Figs 2123</figureCitation>
), respectively. In addition, the shape of the promere in
<taxonomicName id="4C0C4D61D10DFFFFFB25F9DCB345F9D8" box="[1185,1396,1627,1651]" class="Diplopoda" family="Julidae" genus="Anaulaciulus" kingdom="Animalia" order="Julida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="multiarticulatus" status="sp. nov.">
<emphasis id="B978EAF0D10DFFFFFB25F9DCB345F9D8" box="[1185,1396,1627,1651]" italics="true" pageId="10" pageNumber="11">A. multiarticulatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24B578BD10DFFFFFAFEF9DBB6F9F93C" pageId="10" pageNumber="11" rank="species">
<emphasis id="B978EAF0D10DFFFFFAFEF9DBB6F9F93C" bold="true" pageId="10" pageNumber="11">sp. nov.</emphasis>
</taxonomicNameLabel>
, is rather subquadrate, as opposed to subtriangular in
<taxonomicName id="4C0C4D61D10DFFFFFCAAF9F8B223F93C" box="[814,1042,1663,1687]" class="Diplopoda" family="Julidae" genus="Anaulaciulus" kingdom="Animalia" order="Julida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="oligosegmentatus" status="sp. nov.">
<emphasis id="B978EAF0D10DFFFFFCAAF9F8B223F93C" box="[814,1042,1663,1687]" italics="true" pageId="10" pageNumber="11">A. oligosegmentatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24B578BD10DFFFFFB9EF9F8B244F93C" box="[1050,1141,1663,1687]" pageId="10" pageNumber="11" rank="species">
<emphasis id="B978EAF0D10DFFFFFB9EF9F8B244F93C" bold="true" box="[1050,1141,1663,1687]" pageId="10" pageNumber="11">sp. nov.</emphasis>
</taxonomicNameLabel>
The breadth of lamella
<emphasis id="B978EAF0D10DFFFFFA02F9F8B3ADF93C" bold="true" box="[1414,1436,1663,1687]" pageId="10" pageNumber="11">al</emphasis>
on the opisthomere somewhat differs as well. The vulvae are morphologically very similar and highly variable, offering no reliable features to distinguish these two species.
</paragraph>
<paragraph id="8BB336E2D10DFFFFFF41F96BB214F838" blockId="10.[151,1436,1087,1940]" pageId="10" pageNumber="11">
It is also noteworthy that, based both on opisthomere structure (
<emphasis id="B978EAF0D10DFFFFFC14F96BB597F8AF" bold="true" box="[912,934,1772,1796]" pageId="10" pageNumber="11">al</emphasis>
and
<emphasis id="B978EAF0D10DFFFFFC58F96BB5CAF8AF" bold="true" box="[988,1019,1772,1796]" pageId="10" pageNumber="11">ml</emphasis>
breadths) and a vertical body diameter of 1.0 mm, males of
<taxonomicName id="4C0C4D61D10DFFFFFE26F888B4B4F88C" box="[418,645,1807,1831]" class="Diplopoda" family="Julidae" genus="Anaulaciulus" kingdom="Animalia" order="Julida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="oligosegmentatus" status="sp. nov.">
<emphasis id="B978EAF0D10DFFFFFE26F888B4B4F88C" box="[418,645,1807,1831]" italics="true" pageId="10" pageNumber="11">A. oligosegmentatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24B578BD10DFFFFFD08F888B4D4F88C" box="[652,741,1807,1831]" pageId="10" pageNumber="11" rank="species">
<emphasis id="B978EAF0D10DFFFFFD08F888B4D4F88C" bold="true" box="[652,741,1807,1831]" pageId="10" pageNumber="11">sp. nov.</emphasis>
</taxonomicNameLabel>
from Hualien County approach closely those of
<taxonomicName id="4C0C4D61D10DFFFFFA81F888B6EDF8E0" class="Diplopoda" family="Julidae" genus="Anaulaciulus" kingdom="Animalia" order="Julida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="multiarticulatus" status="sp. nov.">
<emphasis id="B978EAF0D10DFFFFFA81F888B6EDF8E0" italics="true" pageId="10" pageNumber="11">A. multiarticulatus</emphasis>
</taxonomicName>
<emphasis id="B978EAF0D10DFFFFFF60F8B3B70EF8E7" bold="true" box="[228,319,1844,1868]" pageId="10" pageNumber="11">
<taxonomicNameLabel id="A24B578BD10DFFFFFF60F8B3B70EF8E7" box="[228,319,1844,1868]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Yet the shapes of the promere, coupled with fewer body segments, i.e. 47(
<emphasis id="B978EAF0D10DFFFFFB1EF8B3B299F8E7" bold="true" box="[1178,1192,1844,1868]" pageId="10" pageNumber="11"></emphasis>
2), allow for a secure identification of these samples to be made as belonging to
<taxonomicName id="4C0C4D61D10DFFFFFCAEF8D0B23CF8C4" box="[810,1037,1879,1903]" class="Diplopoda" family="Julidae" genus="Anaulaciulus" kingdom="Animalia" order="Julida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="oligosegmentatus" status="sp. nov.">
<emphasis id="B978EAF0D10DFFFFFCAEF8D0B23CF8C4" box="[810,1037,1879,1903]" italics="true" pageId="10" pageNumber="11">A. oligosegmentatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24B578BD10DFFFFFB90F8D0B25FF8C4" box="[1044,1134,1879,1903]" pageId="10" pageNumber="11" rank="species">
<emphasis id="B978EAF0D10DFFFFFB90F8D0B25FF8C4" bold="true" box="[1044,1134,1879,1903]" pageId="10" pageNumber="11">sp. nov.</emphasis>
</taxonomicNameLabel>
Molecular analyses would be indispensable to further clarify the identities of these two very similar species.
</paragraph>
</subSubSection>
</treatment>
</document>