154 lines
18 KiB
XML
154 lines
18 KiB
XML
<document id="D0C584003487EF3DEC6EE730C7ED9B55" ID-CLB-Dataset="24540" ID-DOI="10.11646/zootaxa.4708.1.1" ID-GBIF-Dataset="653d580d-e1c7-499c-8604-4b5a7b1920ef" ID-ISSN="1175-5326" ID-Zenodo-Dep="3587181" ID-ZooBank="09F24F0B-5090-4089-A525-8E5150C91EF7" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1576825058981" checkinUser="plazi" docAuthor="Perkins, Philip D." docDate="2019" docId="03A587E7A849FFC80CBA1565FBA490A6" docLanguage="en" docName="zootaxa.4708.1.1.pdf" docOrigin="Zootaxa 4708 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Hydrochus pictus Perkins 2019, new species" docType="treatment" docVersion="8" lastPageNumber="45" masterDocId="FF9CFF9FA863FFE40C2D1244FFE19255" masterDocTitle="Taxonomy of Venezuelan water beetles in the genus Hydrochus Leach, 1817, and an analysis of male genitalia morphology (Coleoptera: Hydrochidae)" masterLastPageNumber="59" masterPageNumber="1" pageNumber="43" updateTime="1698775642300" updateUser="ExternalLinkService">
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<mods:mods id="254C33E2DF15FF674A3829E509EF7C54" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="F0351A56FAD9D5447307F28A613DA39C">
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<mods:title id="BA942AB088C888D0DCDF19C7A16B8837">Taxonomy of Venezuelan water beetles in the genus Hydrochus Leach, 1817, and an analysis of male genitalia morphology (Coleoptera: Hydrochidae)</mods:title>
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<mods:name id="6A288EB0363758A35ED0673F90DC37D3" type="personal">
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<mods:roleTerm id="6FB1C9664F1E929C055F9E3277941C97">Author</mods:roleTerm>
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<mods:namePart id="C5B4FB4E37627B8E261772B83292AC85">Perkins, Philip D.</mods:namePart>
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<mods:typeOfResource id="45439569272FC1693989AE3D4D872564">text</mods:typeOfResource>
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<mods:title id="F06E33AB2FED136361C3989FDEC8C11D">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="93FC91E1A4231A4A5F6916B1DBCF6084">
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<mods:date id="A54C83307F330912EAF9B183FFB15D10">2019</mods:date>
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<mods:number id="722CB54F52B7B58D281087BD92B47EBA">2019-12-12</mods:number>
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<mods:detail id="C69597A99B9D03845FA25E6798DC59EC" type="volume">
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<mods:number id="C1FEA01E664ED7FBB9828BACCF9B08C8">4708</mods:number>
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<mods:number id="E17C474178198D818ABC733F15B696A2">1</mods:number>
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<mods:start id="98EB1ABFF5944F3B133C4961E0040335">1</mods:start>
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<mods:classification id="FC39CF98390AA9A35F446F655D195B4A">journal article</mods:classification>
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<mods:identifier id="B9EADC70441C84DCAD4A289FA2E5FFA7" type="CLB-Dataset">24540</mods:identifier>
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<mods:identifier id="AE61CB1B5C8BCE52ED271980FFAC6C92" type="DOI">10.11646/zootaxa.4708.1.1</mods:identifier>
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<mods:identifier id="152D5A0CE74A6129AC1D0F389CE46252" type="GBIF-Dataset">653d580d-e1c7-499c-8604-4b5a7b1920ef</mods:identifier>
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<mods:identifier id="EF1D8AADCBE504ACDCBD751C1801885B" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="3A462C81DA7921E0A9BAB8E7ED5220AA" type="Zenodo-Dep">3587181</mods:identifier>
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</mods:mods>
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<treatment id="03A587E7A849FFC80CBA1565FBA490A6" ID-DOI="http://doi.org/10.5281/zenodo.5941805" ID-GBIF-Taxon="160798420" ID-Zenodo-Dep="5941805" LSID="urn:lsid:plazi:treatment:03A587E7A849FFC80CBA1565FBA490A6" httpUri="http://treatment.plazi.org/id/03A587E7A849FFC80CBA1565FBA490A6" lastPageId="44" lastPageNumber="45" pageId="42" pageNumber="43">
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<subSubSection id="C316657AA849FFCE0CBA1565FDE2956E" box="[151,515,1825,1852]" pageId="42" pageNumber="43" type="nomenclature">
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<paragraph id="8BB336F1A849FFCE0CBA1565FDE2956E" blockId="42.[151,515,1825,1887]" box="[151,515,1825,1852]" pageId="42" pageNumber="43">
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<heading id="D0FB819DA849FFCE0CBA1565FDE2956E" bold="true" box="[151,515,1825,1852]" fontSize="11" level="1" pageId="42" pageNumber="43" reason="1">
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<emphasis id="B978EAE3A849FFCE0CBA1565FDE2956E" bold="true" box="[151,515,1825,1852]" pageId="42" pageNumber="43">
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<taxonomicName id="4C0C4D72A849FFCE0CBA1565FE8D956E" ID-CoL="8Y4R4" authority="Perkins, 2019" authorityName="Perkins" authorityYear="2019" box="[151,364,1825,1852]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="pictus" status="sp. nov.">
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<emphasis id="B978EAE3A849FFCE0CBA1565FE8D956E" bold="true" box="[151,364,1825,1852]" italics="true" pageId="42" pageNumber="43">Hydrochus pictus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A24B5798A849FFCE0D5E1565FDE2956E" box="[371,515,1825,1851]" pageId="42" pageNumber="43" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C316657AA849FFCE0CBA1501FEEC950A" box="[151,269,1861,1887]" pageId="42" pageNumber="43" type="description">
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<paragraph id="8BB336F1A849FFCE0CBA1501FEEC950A" blockId="42.[151,515,1825,1887]" box="[151,269,1861,1887]" pageId="42" pageNumber="43">
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<figureCitation id="13372A74A849FFCE0CBA1501FF02950A" box="[151,227,1861,1887]" captionStart="FIGURE 1" captionStartId="11.[151,250,1894,1919]" captionTargetBox="[156,1422,330,1824]" captionTargetId="figure@11.[151,1435,306,1866]" captionTargetPageId="11" captionText="FIGURE 1. Dorsal habitus of Hydrochus species." figureDoi="http://doi.org/10.5281/zenodo.3587183" httpUri="https://zenodo.org/record/3587183/files/figure.png" pageId="42" pageNumber="43">Figs. 1</figureCitation>
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,
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<figureCitation id="13372A74A849FFCE0CDC1501FEEC950A" box="[241,269,1861,1887]" captionStart="FIGURE 24" captionStartId="43.[151,250,1841,1866]" captionTargetBox="[202,1397,292,1788]" captionTargetId="figure@43.[151,1435,273,1817]" captionTargetPageId="43" captionText="FIGURE 24. Hydrochus pictus n. sp., habitus and male genitalia of holotype." figureDoi="http://doi.org/10.5281/zenodo.3587231" httpUri="https://zenodo.org/record/3587231/files/figure.png" pageId="42" pageNumber="43">24</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C316657AA849FFCF0CBA15C9FBA792AE" lastPageId="43" lastPageNumber="44" pageId="42" pageNumber="43" type="materials_examined">
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<paragraph id="8BB336F1A849FFCE0CBA15C9FBA4959E" blockId="42.[151,1436,1932,2031]" pageId="42" pageNumber="43">
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<emphasis id="B978EAE3A849FFCE0CBA15C9FEA695F2" bold="true" box="[151,327,1932,1959]" pageId="42" pageNumber="43">Type Material.</emphasis>
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<materialsCitation id="3B643CACA849FFCE0D6315C9FBA4959E" ID-GBIF-Occurrence="2521263689" collectingDate="2010-01-29" collectionCode="MIZA" collectorName="Short" country="Venezuela" elevation="2" latitude="10.52365" location="El Pilar" longLatPrecision="1" longitude="-63.117832" pageId="42" pageNumber="43" specimenCount="1" stateProvince="Sucre" typeStatus="holotype">
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<emphasis id="B978EAE3A849FFCE0D6315C9FDF795F2" bold="true" box="[334,534,1932,1959]" pageId="42" pageNumber="43">
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<typeStatus id="54B78853A849FFCE0D6315C9FE5A95F2" box="[334,443,1933,1959]" pageId="42" pageNumber="43" type="holotype">Holotype</typeStatus>
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(male):
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</emphasis>
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“
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<collectingCountry id="F31B7661A849FFCE0E0B15C9FD4195F2" box="[550,672,1933,1959]" name="Venezuela" pageId="42" pageNumber="43">Venezuela</collectingCountry>
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:
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<collectingRegion id="49C8F813A849FFCE0E8715C9FCCD95F2" box="[682,812,1933,1959]" country="Venezuela" name="Sucre" pageId="42" pageNumber="43">Sucre State</collectingRegion>
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,
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<geoCoordinate id="EE385036A849FFCE0F1B15C9FC3495FD" box="[822,981,1933,1960]" degrees="10" direction="north" minutes="31.419" orientation="latitude" pageId="42" pageNumber="43" precision="1" value="10.52365">10° 31.419’ N</geoCoordinate>
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,
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<geoCoordinate id="EE385036A849FFCE0FCD15C9FB9495FD" box="[992,1141,1933,1960]" degrees="63" direction="west" minutes="7.070" orientation="longitude" pageId="42" pageNumber="43" precision="1" value="-63.117832">63° 7.070’ W</geoCoordinate>
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,
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<quantity id="4CF49B14A849FFCE085215C9FB4A95FD" box="[1151,1195,1933,1960]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="42" pageNumber="43" unit="m" value="2.0">
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<elevation id="0021D1C2A849FFCE085215C9FB4A95FD" box="[1151,1195,1933,1960]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="42" pageNumber="43" unit="m" value="2.0">2 m</elevation>
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</quantity>
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, ca.
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<quantity id="4CF49B14A849FFCE08F015C8FAF495F2" box="[1245,1301,1932,1959]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="42" pageNumber="43" unit="km" value="5.0">5 km</quantity>
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SE
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<location id="8ED3602AA849FFCE096F15C9FA7995F2" LSID="urn:lsid:plazi:treatment:03A587E7A849FFC80CBA1565FBA490A6:8ED3602AA849FFCE096F15C9FA7995F2" box="[1346,1432,1933,1959]" country="Venezuela" latitude="10.52365" longLatPrecision="1" longitude="-63.117832" name="El Pilar" pageId="42" pageNumber="43" stateProvince="Sucre">El Pilar</location>
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,
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<date id="FFB21031A849FFCE0CBA15F5FEE2959E" box="[151,259,1969,1995]" pageId="42" pageNumber="43" value="2010-01-29">
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<collectingDate id="EFF6E9D9A849FFCE0CBA15F5FEE2959E" box="[151,259,1969,1995]" pageId="42" pageNumber="43" value="2010-01-29">29.i.2010</collectingDate>
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</date>
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, leg.
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<collectorName id="26F95327A849FFCE0D1215F5FE9D959E" box="[319,380,1969,1995]" pageId="42" pageNumber="43">Short</collectorName>
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& Garcia; forested swamp by road; VZ10-0129-04A” (
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<collectionCode id="ED1DAE34A849FFCE0FC615F5FBD8959E" box="[1003,1081,1969,1995]" country="Venezuela" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33834" lsid="urn:lsid:biocol.org:col:33834" name="Museo del Instituto de Zoologia Agricola Francisco Fernandez Yepez" pageId="42" pageNumber="43" type="Museum">MIZA</collectionCode>
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).
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</materialsCitation>
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</paragraph>
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<paragraph id="8BB336F1A849FFCE0CEA1590FA7D95BA" blockId="42.[151,1436,1932,2031]" box="[199,1436,2004,2031]" pageId="42" pageNumber="43">
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<emphasis id="B978EAE3A849FFCE0CEA1590FE3795BA" bold="true" box="[199,470,2004,2031]" pageId="42" pageNumber="43">Differential Diagnosis.</emphasis>
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Among Venezuelan species, recognized by the combination of small size (ca. 2.31
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</paragraph>
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<paragraph id="8BB336F1A848FFCF0CBA12DDFBA792AE" blockId="43.[151,1437,153,251]" pageId="43" pageNumber="44">
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mm), the pronotum with very shallow impressions and rather uniform punctation, the pronotal shape, the very large black spots on the elytra, and the male genitalia (
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<figureCitation id="13372A74A848FFCF0EFD12F9FCC79282" box="[720,806,189,215]" captionStart="FIGURE 24" captionStartId="43.[151,250,1841,1866]" captionTargetBox="[202,1397,292,1788]" captionTargetId="figure@43.[151,1435,273,1817]" captionTargetPageId="43" captionText="FIGURE 24. Hydrochus pictus n. sp., habitus and male genitalia of holotype." figureDoi="http://doi.org/10.5281/zenodo.3587231" httpUri="https://zenodo.org/record/3587231/files/figure.png" pageId="43" pageNumber="44">Fig. 24</figureCitation>
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; described below). Differentiation from other similarly small sized species with similarly shaped pronotum will require dissection of males.
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</paragraph>
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</subSubSection>
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<caption id="DF736679A848FFCF0CBA1575FC4A951F" ID-DOI="http://doi.org/10.5281/zenodo.3587231" ID-Zenodo-Dep="3587231" box="[151,939,1841,1866]" httpUri="https://zenodo.org/record/3587231/files/figure.png" pageId="43" pageNumber="44" startId="43.[151,250,1841,1866]" targetBox="[202,1397,292,1788]" targetPageId="43">
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<paragraph id="8BB336F1A848FFCF0CBA1575FC4A951F" blockId="43.[151,939,1841,1866]" box="[151,939,1841,1866]" pageId="43" pageNumber="44">
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<emphasis id="B978EAE3A848FFCF0CBA1575FEC1951F" bold="true" box="[151,288,1841,1866]" pageId="43" pageNumber="44">FIGURE 24.</emphasis>
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<taxonomicName id="4C0C4D72A848FFCF0D0B1576FE39951F" authority="Perkins, 2019" authorityName="Perkins" authorityYear="2019" box="[294,472,1842,1866]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="pictus" status="sp. nov.">
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<emphasis id="B978EAE3A848FFCF0D0B1576FE39951F" box="[294,472,1842,1866]" italics="true" pageId="43" pageNumber="44">Hydrochus pictus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A24B5798A848FFCF0DF01577FDF7951F" box="[477,534,1843,1866]" pageId="43" pageNumber="44" rank="species">n. sp.</taxonomicNameLabel>
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, habitus and male genitalia of holotype.
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</paragraph>
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</caption>
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<subSubSection id="C316657AA848FFC80CEA153CFF36909A" lastPageId="44" lastPageNumber="45" pageId="43" pageNumber="44" type="description">
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<paragraph id="8BB336F1A848FFC80CEA153CFF0392AE" blockId="43.[151,1437,1912,2011]" lastBlockId="44.[151,1437,152,755]" lastPageId="44" lastPageNumber="45" pageId="43" pageNumber="44">
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<emphasis id="B978EAE3A848FFCF0CEA153CFEB095C6" bold="true" box="[199,337,1912,1939]" pageId="43" pageNumber="44">Description</emphasis>
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. Habitus as illustrated (
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<figureCitation id="13372A74A848FFCF0E49153CFD5B95C6" box="[612,698,1912,1939]" captionStart="FIGURE 24" captionStartId="43.[151,250,1841,1866]" captionTargetBox="[202,1397,292,1788]" captionTargetId="figure@43.[151,1435,273,1817]" captionTargetPageId="43" captionText="FIGURE 24. Hydrochus pictus n. sp., habitus and male genitalia of holotype." figureDoi="http://doi.org/10.5281/zenodo.3587231" httpUri="https://zenodo.org/record/3587231/files/figure.png" pageId="43" pageNumber="44">Fig. 24</figureCitation>
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). Size:
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<typeStatus id="54B78853A848FFCF0F3D153CFC9395C7" box="[784,882,1912,1938]" pageId="43" pageNumber="44" type="holotype">holotype</typeStatus>
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(length/width, mm): body (length to elytral apices) 2.31/0.86; head width 0.60; pronotum l/w 0.55/0.60; PA 0.60; PB 0.43; elytra 1.38/0.86. Body size small (ca.
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<quantity id="4CF49B14A848FFCF0CBA1585FF1E958E" box="[151,255,1985,2011]" metricMagnitude="-3" metricUnit="m" metricValue="2.31" pageId="43" pageNumber="44" unit="mm" value="2.31">2.31 mm</quantity>
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). Dorsum grey, with moderately strong iridescence, elytra with several large black spots. Legs testaceous, with tibiofemoral joints darker. Punctation on elytra ca. 2–
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<date id="FFB21031A84FFFC80F1C12DDFCAA92E6" box="[817,843,153,179]" pageId="44" pageNumber="45">3x</date>
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that of pronotum. Elytra interstriae ca. 1–1.5x strial puncture diameter. Interstria 9
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<superScript id="7C799BB9A84FFFC80DC512F8FE14929F" attach="left" box="[488,501,188,202]" fontSize="6" pageId="44" pageNumber="45">th</superScript>
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more convex than others, overhanging 10
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<superScript id="7C799BB9A84FFFC80FE112F8FC38929F" attach="left" box="[972,985,188,202]" fontSize="6" pageId="44" pageNumber="45">th</superScript>
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interstria. Usual area of callus on 5
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<superScript id="7C799BB9A84FFFC8094812F8FA93929F" attach="left" box="[1381,1394,188,202]" fontSize="6" pageId="44" pageNumber="45">th</superScript>
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not raised.
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</paragraph>
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<paragraph id="8BB336F1A84FFFC80CEA1341FF3E9332" blockId="44.[151,1437,152,755]" pageId="44" pageNumber="45">Pronotum slightly wider than long (as ca. 55/60), widest at anterior margin, markedly narrowed at base, sides straight, smooth; anterior margin bisinuate; depressions very shallow, or absent; punctation quite uniform over pronotum</paragraph>
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<paragraph id="8BB336F1A84FFFC80CEA1335FBCF93FA" blockId="44.[151,1437,152,755]" pageId="44" pageNumber="45">Elytra with spaces between strial punctures ca. 0.5x their diameter, to narrow ridges; apices rather sharply conjointly rounded in dorsal view, in lateral view outer margin with slight angulation.</paragraph>
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<paragraph id="8BB336F1A84FFFC80CEA13FDFA799386" blockId="44.[151,1437,152,755]" box="[199,1432,441,467]" pageId="44" pageNumber="45">Ventral characters: Mentum moderately coarsely punctate, median fovea shallow; submentum without foveae.</paragraph>
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<paragraph id="8BB336F1A84FFFC80CEA1399FEA79036" blockId="44.[151,1437,152,755]" pageId="44" pageNumber="45">
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Male genitalia (
|
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<figureCitation id="13372A74A84FFFC80D541399FE2D93A2" box="[377,460,477,503]" captionStart="FIGURE 24" captionStartId="43.[151,250,1841,1866]" captionTargetBox="[202,1397,292,1788]" captionTargetId="figure@43.[151,1435,273,1817]" captionTargetPageId="43" captionText="FIGURE 24. Hydrochus pictus n. sp., habitus and male genitalia of holotype." figureDoi="http://doi.org/10.5281/zenodo.3587231" httpUri="https://zenodo.org/record/3587231/files/figure.png" pageId="44" pageNumber="45">Fig. 24</figureCitation>
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) general characters: parameres long, tapering to very fine apices, very narrow and extending well beyond distal end of aedeagus, tips in dorsal/ventral views very thin and sharply pointed, in lateral view tips very thin and arcuate ventrad; basal piece shorter than parameres, in lateral view markedly arcuate, orifice with rim sclerotized.
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</paragraph>
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<paragraph id="8BB336F1A84FFFC80CEA1029FC1490D2" blockId="44.[151,1437,152,755]" box="[199,1013,621,647]" pageId="44" pageNumber="45">Dorsal surface: adtl lacking; pdmm slightly overlapping alm; adbl small.</paragraph>
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<paragraph id="8BB336F1A84FFFC80CEA10D5FF36909A" blockId="44.[151,1437,152,755]" pageId="44" pageNumber="45">Ventral surface: avtl near basal 1/3 of aedeagus; pvmm overlapping alm, medial margins very narrowly separated.</paragraph>
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</subSubSection>
|
||
<subSubSection id="C316657AA84FFFC80CEA109DFBA490A6" box="[199,1093,729,755]" pageId="44" pageNumber="45" type="etymology">
|
||
<paragraph id="8BB336F1A84FFFC80CEA109DFBA490A6" blockId="44.[151,1437,152,755]" box="[199,1093,729,755]" pageId="44" pageNumber="45">
|
||
<emphasis id="B978EAE3A84FFFC80CEA109DFEAF90A6" bold="true" box="[199,334,729,755]" pageId="44" pageNumber="45">Etymology.</emphasis>
|
||
Named in reference to the numerous large black spots of the elytra.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |