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<document ID-DOI="http://doi.org/10.5281/zenodo.3804006" ID-GBIF-Dataset="4d505549-2f45-49c2-b388-8fc8bfdbd5f3" ID-GBIF-Taxon="163737022" ID-Zenodo-Dep="3804006" checkinTime="1587658244385" checkinUser="plazi" docAuthor="Jałoszynski, Paweł, Luo, Xiao-Zhu, Hammel, Jorg U., Yamamoto, Sh ^ uhei &amp; Beutel, Rolf G." docDate="2020" docId="03E7BF21FF83C805FF50F903FEC2FA0F" docLanguage="en" docName="14772019.2020.1747561.pdf" docOrigin="Journal of Systematic Palaeontology 18" docStyle="DocumentStyle{}" docTitle="Lepiceratus Jaloszynski, Luo, Yamamoto &amp; 2020" docType="treatment" docVersion="6" lastPageId="7" lastPageNumber="8" masterDocId="FFDEC759FF87C802FFE7FFC1FF8DFFDA" masterDocTitle="The mid-Cretaceous † Lepiceratus Gen. nov. and the Evolution of the Relict Beetle Family Lepiceridae (Insecta: Coleoptera: Myxophaga)" masterLastPageNumber="14" masterPageNumber="1" pageId="4" pageNumber="5" updateTime="1643446050710" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>The mid-Cretaceous † Lepiceratus Gen. nov. and the Evolution of the Relict Beetle Family Lepiceridae (Insecta: Coleoptera: Myxophaga)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Jałoszynski, Paweł</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Luo, Xiao-Zhu</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hammel, Jorg U.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Yamamoto, Sh ^ uhei</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Beutel, Rolf G.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Journal of Systematic Palaeontology</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2020</mods:date>
<mods:detail type="pubDate">
<mods:number>2020-04-22</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>18</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>14</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.1080/14772019.2020.1747561</mods:identifier>
<mods:identifier type="GBIF-Dataset">4d505549-2f45-49c2-b388-8fc8bfdbd5f3</mods:identifier>
<mods:identifier type="Zenodo-Dep">3763694</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.3804006" ID-GBIF-Taxon="163737022" ID-Zenodo-Dep="3804006" LSID="urn:lsid:plazi:treatment:03E7BF21FF83C805FF50F903FEC2FA0F" httpUri="http://treatment.plazi.org/id/03E7BF21FF83C805FF50F903FEC2FA0F" lastPageId="7" lastPageNumber="8" pageId="4" pageNumber="5">
<subSubSection pageId="4" pageNumber="5" type="nomenclature">
<paragraph blockId="4.[183,757,1666,1786]" pageId="4" pageNumber="5">
Genus †
<taxonomicName authority="Jaloszynski, Luo, Yamamoto &amp;" authorityName="Jaloszynski, Luo, Yamamoto &amp;" authorityYear="2020" box="[273,757,1729,1753]" class="Insecta" family="Lepiceridae" genus="Lepiceratus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis bold="true" box="[273,402,1729,1753]" italics="true" pageId="4" pageNumber="5">Lepiceratus</emphasis>
Jałoszynski, Luo, Yamamoto &amp;
</taxonomicName>
Beutel, gen. nov.
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="5" type="type_taxon">
<paragraph blockId="4.[159,781,1844,1901]" pageId="4" pageNumber="5">
<emphasis bold="true" box="[159,319,1845,1868]" pageId="4" pageNumber="5">Type species.</emphasis>
†
<taxonomicName authority="Jaloszynski, Luo, Yamamoto &amp; Beutel" authorityName="Jaloszynski, Luo, Yamamoto &amp; Beutel" authorityYear="2020" class="Insecta" family="Lepiceridae" genus="Lepiceratus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="ankylosaurus" status="sp. nov.">
<emphasis box="[349,634,1845,1868]" italics="true" pageId="4" pageNumber="5">Lepiceratus ankylosaurus</emphasis>
Jałoszynski, Luo, Yamamoto &amp; Beutel
</taxonomicName>
,
<taxonomicNameLabel box="[459,546,1878,1901]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
(here designated).
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="5" type="diagnosis">
<paragraph blockId="4.[845,1467,1466,1622]" pageId="4" pageNumber="5">
<emphasis bold="true" box="[845,961,1466,1489]" pageId="4" pageNumber="5">Diagnosis.</emphasis>
Lepiceroid with heptamerous antennae; metathoracic discrimen not marked on ventral surface (but discernible on inner, dorsal surface of metaventrite); and prosternal process sub-triangular, elongate, gradually tapering toward pointed apex.
</paragraph>
</subSubSection>
<subSubSection lastPageId="6" lastPageNumber="7" pageId="4" pageNumber="5" type="description">
<paragraph blockId="4.[845,1467,1646,1901]" pageId="4" pageNumber="5">
<emphasis bold="true" box="[845,983,1647,1670]" pageId="4" pageNumber="5">Description.</emphasis>
Body (
<figureCitation box="[1070,1190,1646,1670]" captionStart="Figure 1" captionStartId="3.[159,226,1858,1879]" captionTargetBox="[175,1455,196,1830]" captionTargetId="figure@3.[159,1467,188,1840]" captionTargetPageId="3" captionText="Figure 1. †Lepiceratus ankylosaurus gen. and sp. nov., holotype, A, B, macrophotographs; C, artistic reconstruction; and D, schematic morphological details observable under light microscope. A, C, dorsal habitus; B, D, ventral habitus. Abbreviations: anf, antennal furrow; anp, antennal pocket; ce, compound eye; cx1–3, coxa; elex, elytral excavation; fm1–3, femur; hyp, hypomeron; kes, katepisternum; lb, labrum; mnt, mentum; ppr, prosternal process; r1–5, elytral rows of tubercles; st, abdominal sternite 2; tb1–3, tibia; tch1–3, trochanter; tr1–3, tarsus; v2, mesoventrite; v3, metaventrite." figureDoi="http://doi.org/10.5281/zenodo.3763696" httpUri="https://zenodo.org/record/3763696/files/figure.png" pageId="4" pageNumber="5">Figs 1A–C</figureCitation>
,
<figureCitation box="[1207,1274,1647,1670]" captionStart="Figure 2" captionStartId="4.[159,226,1321,1342]" captionTargetBox="[159,1454,189,1304]" captionTargetId="figure@4.[159,1467,188,1304]" captionTargetPageId="4" captionText="Figure 2. †Lepiceratus ankylosaurus gen. and sp. nov., holotype, synchrotron microtomography, surface reconstructions. A, dorsal habitus; B, lateral habitus; C, right antenna and eye in ventral view; D, ventral habitus. Abbreviations: anc, antennal club; anf, antennal furrow; an3–6, antennomere 3–6; ce, compound eye; elex, elytral excavation;?mxp, presumable maxillary palp; pd, pedicel; sc, scape; v2, mesoventrite." figureDoi="http://doi.org/10.5281/zenodo.3763698" httpUri="https://zenodo.org/record/3763698/files/figure.png" pageId="4" pageNumber="5">2A, B</figureCitation>
,
<figureCitation box="[1291,1324,1647,1670]" captionStart="Figure 3" captionStartId="5.[159,226,1539,1560]" captionTargetBox="[159,1467,188,1517]" captionTargetId="figure@5.[159,1467,188,1517]" captionTargetPageId="5" captionText="Figure 3. †Lepiceratus ankylosaurus gen. and sp. nov., holotype, synchrotron microtomography, volume renderings. A, dorsal habitus; B, submedian sagittal virtual section; C, right antenna, eye and anterolateral portion of prothorax in ventral view; D, distal surface of right antennal club; E, ventral habitus. Abbreviations: anc, antennal club; anf, antennal furrow; anp, antennal pocket; an3–6, antennomere 3–6; ce, compound eye; els, elytral suture; epi, elytral epipleura; pd, pedicel; ppr, prosternal process; sc, scape; scs, scutellar shield; wg, wings." figureDoi="http://doi.org/10.5281/zenodo.3763700" httpUri="https://zenodo.org/record/3763700/files/figure.png" pageId="4" pageNumber="5">3A</figureCitation>
) stout, compact (prothorax widely separated from posterior body as a post-mortem artefact) and moderately convex. Cuticle strongly sclerotized and conspicuously sculptured on dorsal side, sparsely setose.
</paragraph>
<paragraph blockId="4.[845,1467,1646,1901]" lastBlockId="5.[845,1467,1718,1901]" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5">
Head large in relation to body, slightly inclined, only partly visible in dorsal view but largely exposed ventrally. Posterior-most part retracted into prothorax, constricted ‘neck’ region apparently absent. Dorsal side with distinct protuberances. Compound eyes (
<figureCitation box="[682,774,1750,1773]" captionStart="Figure 1" captionStartId="3.[159,226,1858,1879]" captionTargetBox="[175,1455,196,1830]" captionTargetId="figure@3.[159,1467,188,1840]" captionTargetPageId="3" captionText="Figure 1. †Lepiceratus ankylosaurus gen. and sp. nov., holotype, A, B, macrophotographs; C, artistic reconstruction; and D, schematic morphological details observable under light microscope. A, C, dorsal habitus; B, D, ventral habitus. Abbreviations: anf, antennal furrow; anp, antennal pocket; ce, compound eye; cx1–3, coxa; elex, elytral excavation; fm1–3, femur; hyp, hypomeron; kes, katepisternum; lb, labrum; mnt, mentum; ppr, prosternal process; r1–5, elytral rows of tubercles; st, abdominal sternite 2; tb1–3, tibia; tch1–3, trochanter; tr1–3, tarsus; v2, mesoventrite; v3, metaventrite." figureDoi="http://doi.org/10.5281/zenodo.3763696" httpUri="https://zenodo.org/record/3763696/files/figure.png" pageId="5" pageNumber="6">Figs 1D</figureCitation>
,
<figureCitation box="[159,190,1782,1805]" captionStart="Figure 2" captionStartId="4.[159,226,1321,1342]" captionTargetBox="[159,1454,189,1304]" captionTargetId="figure@4.[159,1467,188,1304]" captionTargetPageId="4" captionText="Figure 2. †Lepiceratus ankylosaurus gen. and sp. nov., holotype, synchrotron microtomography, surface reconstructions. A, dorsal habitus; B, lateral habitus; C, right antenna and eye in ventral view; D, ventral habitus. Abbreviations: anc, antennal club; anf, antennal furrow; an3–6, antennomere 3–6; ce, compound eye; elex, elytral excavation;?mxp, presumable maxillary palp; pd, pedicel; sc, scape; v2, mesoventrite." figureDoi="http://doi.org/10.5281/zenodo.3763698" httpUri="https://zenodo.org/record/3763698/files/figure.png" pageId="5" pageNumber="6">2C</figureCitation>
,
<figureCitation box="[209,240,1782,1805]" captionStart="Figure 3" captionStartId="5.[159,226,1539,1560]" captionTargetBox="[159,1467,188,1517]" captionTargetId="figure@5.[159,1467,188,1517]" captionTargetPageId="5" captionText="Figure 3. †Lepiceratus ankylosaurus gen. and sp. nov., holotype, synchrotron microtomography, volume renderings. A, dorsal habitus; B, submedian sagittal virtual section; C, right antenna, eye and anterolateral portion of prothorax in ventral view; D, distal surface of right antennal club; E, ventral habitus. Abbreviations: anc, antennal club; anf, antennal furrow; anp, antennal pocket; an3–6, antennomere 3–6; ce, compound eye; els, elytral suture; epi, elytral epipleura; pd, pedicel; ppr, prosternal process; sc, scape; scs, scutellar shield; wg, wings." figureDoi="http://doi.org/10.5281/zenodo.3763700" httpUri="https://zenodo.org/record/3763700/files/figure.png" pageId="5" pageNumber="6">3C</figureCitation>
) conspicuously large and protruding, composed of numerous small ommatidia with distinctly convex cornea lenses. Antennal furrow (
<figureCitation box="[528,618,1846,1869]" captionStart="Figure 1" captionStartId="3.[159,226,1858,1879]" captionTargetBox="[175,1455,196,1830]" captionTargetId="figure@3.[159,1467,188,1840]" captionTargetPageId="3" captionText="Figure 1. †Lepiceratus ankylosaurus gen. and sp. nov., holotype, A, B, macrophotographs; C, artistic reconstruction; and D, schematic morphological details observable under light microscope. A, C, dorsal habitus; B, D, ventral habitus. Abbreviations: anf, antennal furrow; anp, antennal pocket; ce, compound eye; cx1–3, coxa; elex, elytral excavation; fm1–3, femur; hyp, hypomeron; kes, katepisternum; lb, labrum; mnt, mentum; ppr, prosternal process; r1–5, elytral rows of tubercles; st, abdominal sternite 2; tb1–3, tibia; tch1–3, trochanter; tr1–3, tarsus; v2, mesoventrite; v3, metaventrite." figureDoi="http://doi.org/10.5281/zenodo.3763696" httpUri="https://zenodo.org/record/3763696/files/figure.png" pageId="5" pageNumber="6">Figs 1D</figureCitation>
,
<figureCitation box="[636,668,1846,1869]" captionStart="Figure 2" captionStartId="4.[159,226,1321,1342]" captionTargetBox="[159,1454,189,1304]" captionTargetId="figure@4.[159,1467,188,1304]" captionTargetPageId="4" captionText="Figure 2. †Lepiceratus ankylosaurus gen. and sp. nov., holotype, synchrotron microtomography, surface reconstructions. A, dorsal habitus; B, lateral habitus; C, right antenna and eye in ventral view; D, ventral habitus. Abbreviations: anc, antennal club; anf, antennal furrow; an3–6, antennomere 3–6; ce, compound eye; elex, elytral excavation;?mxp, presumable maxillary palp; pd, pedicel; sc, scape; v2, mesoventrite." figureDoi="http://doi.org/10.5281/zenodo.3763698" httpUri="https://zenodo.org/record/3763698/files/figure.png" pageId="5" pageNumber="6">2C</figureCitation>
,
<figureCitation box="[686,717,1846,1869]" captionStart="Figure 3" captionStartId="5.[159,226,1539,1560]" captionTargetBox="[159,1467,188,1517]" captionTargetId="figure@5.[159,1467,188,1517]" captionTargetPageId="5" captionText="Figure 3. †Lepiceratus ankylosaurus gen. and sp. nov., holotype, synchrotron microtomography, volume renderings. A, dorsal habitus; B, submedian sagittal virtual section; C, right antenna, eye and anterolateral portion of prothorax in ventral view; D, distal surface of right antennal club; E, ventral habitus. Abbreviations: anc, antennal club; anf, antennal furrow; anp, antennal pocket; an3–6, antennomere 3–6; ce, compound eye; els, elytral suture; epi, elytral epipleura; pd, pedicel; ppr, prosternal process; sc, scape; scs, scutellar shield; wg, wings." figureDoi="http://doi.org/10.5281/zenodo.3763700" httpUri="https://zenodo.org/record/3763700/files/figure.png" pageId="5" pageNumber="6">3C</figureCitation>
) present, sharply demarcated, as narrow as proximal flagellomeres and running along mesoventral margin of each eye. Clypeus, labrum and paired mouthparts poorly visible or not observable, except for elongated tetramerous structure that may be a posteriorly bent maxillary palp (
<figureCitation box="[911,998,1846,1869]" captionStart="Figure 2" captionStartId="4.[159,226,1321,1342]" captionTargetBox="[159,1454,189,1304]" captionTargetId="figure@4.[159,1467,188,1304]" captionTargetPageId="4" captionText="Figure 2. †Lepiceratus ankylosaurus gen. and sp. nov., holotype, synchrotron microtomography, surface reconstructions. A, dorsal habitus; B, lateral habitus; C, right antenna and eye in ventral view; D, ventral habitus. Abbreviations: anc, antennal club; anf, antennal furrow; an3–6, antennomere 3–6; ce, compound eye; elex, elytral excavation;?mxp, presumable maxillary palp; pd, pedicel; sc, scape; v2, mesoventrite." figureDoi="http://doi.org/10.5281/zenodo.3763698" httpUri="https://zenodo.org/record/3763698/files/figure.png" pageId="5" pageNumber="6">Fig. 2D</figureCitation>
). Mentum (
<figureCitation box="[1136,1223,1846,1869]" captionStart="Figure 1" captionStartId="3.[159,226,1858,1879]" captionTargetBox="[175,1455,196,1830]" captionTargetId="figure@3.[159,1467,188,1840]" captionTargetPageId="3" captionText="Figure 1. †Lepiceratus ankylosaurus gen. and sp. nov., holotype, A, B, macrophotographs; C, artistic reconstruction; and D, schematic morphological details observable under light microscope. A, C, dorsal habitus; B, D, ventral habitus. Abbreviations: anf, antennal furrow; anp, antennal pocket; ce, compound eye; cx1–3, coxa; elex, elytral excavation; fm1–3, femur; hyp, hypomeron; kes, katepisternum; lb, labrum; mnt, mentum; ppr, prosternal process; r1–5, elytral rows of tubercles; st, abdominal sternite 2; tb1–3, tibia; tch1–3, trochanter; tr1–3, tarsus; v2, mesoventrite; v3, metaventrite." figureDoi="http://doi.org/10.5281/zenodo.3763696" httpUri="https://zenodo.org/record/3763696/files/figure.png" pageId="5" pageNumber="6">Fig. 1D</figureCitation>
) large and plate-like, almost completely covering ventral side of head.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.3763700" ID-Zenodo-Dep="3763700" httpUri="https://zenodo.org/record/3763700/files/figure.png" pageId="5" pageNumber="6" startId="5.[159,226,1539,1560]" targetBox="[159,1467,188,1517]" targetPageId="5">
<paragraph blockId="5.[159,1467,1538,1666]" pageId="5" pageNumber="6">
<emphasis bold="true" box="[159,257,1539,1560]" pageId="5" pageNumber="6">Figure 3.</emphasis>
†
<taxonomicName authority="Jałoszynski &amp; Luo &amp; Hammel &amp; Yamamoto &amp; Beutel, 2020" authorityName="Jałoszynski &amp; Luo &amp; Hammel &amp; Yamamoto &amp; Beutel" authorityYear="2020" box="[281,540,1538,1560]" class="Insecta" family="Lepiceridae" genus="Lepiceratus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="ankylosaurus" status="gen. and sp. nov.">
<emphasis bold="true" box="[281,540,1538,1560]" italics="true" pageId="5" pageNumber="6">Lepiceratus ankylosaurus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[553,737,1539,1560]" pageId="5" pageNumber="6" rank="species">gen. and sp. nov.</taxonomicNameLabel>
, holotype, synchrotron microtomography, volume renderings.
<emphasis bold="true" box="[1372,1389,1539,1560]" pageId="5" pageNumber="6">A</emphasis>
, dorsal habitus;
<emphasis bold="true" box="[243,259,1565,1586]" pageId="5" pageNumber="6">B</emphasis>
, submedian sagittal virtual section;
<emphasis bold="true" box="[619,636,1565,1586]" pageId="5" pageNumber="6">C</emphasis>
, right antenna, eye and anterolateral portion of prothorax in ventral view;
<emphasis bold="true" box="[1383,1400,1565,1586]" pageId="5" pageNumber="6">D</emphasis>
, distal surface of right antennal club;
<emphasis bold="true" box="[473,489,1592,1613]" pageId="5" pageNumber="6">E</emphasis>
, ventral habitus.
<emphasis bold="true" box="[666,810,1592,1613]" pageId="5" pageNumber="6">Abbreviations</emphasis>
:
<emphasis bold="true" box="[827,863,1592,1613]" pageId="5" pageNumber="6">anc</emphasis>
, antennal club;
<emphasis bold="true" box="[1028,1061,1592,1613]" pageId="5" pageNumber="6">anf</emphasis>
, antennal furrow;
<emphasis bold="true" box="[1251,1289,1592,1613]" pageId="5" pageNumber="6">anp</emphasis>
, antennal pocket;
<emphasis bold="true" box="[159,220,1618,1640]" pageId="5" pageNumber="6">an3–6</emphasis>
, antennomere 3–6;
<emphasis bold="true" box="[416,437,1619,1640]" pageId="5" pageNumber="6">ce</emphasis>
, compound eye;
<emphasis bold="true" box="[608,634,1619,1640]" pageId="5" pageNumber="6">els</emphasis>
, elytral suture;
<emphasis bold="true" box="[791,821,1619,1640]" pageId="5" pageNumber="6">epi</emphasis>
, elytral epipleura;
<emphasis bold="true" box="[1008,1034,1619,1640]" pageId="5" pageNumber="6">pd</emphasis>
, pedicel;
<emphasis bold="true" box="[1132,1169,1619,1640]" pageId="5" pageNumber="6">ppr</emphasis>
, prosternal process;
<emphasis bold="true" box="[1373,1393,1619,1640]" pageId="5" pageNumber="6">sc</emphasis>
, scape;
<emphasis bold="true" box="[159,188,1645,1666]" pageId="5" pageNumber="6">scs</emphasis>
, scutellar shield;
<emphasis bold="true" box="[361,390,1645,1666]" pageId="5" pageNumber="6">wg</emphasis>
, wings.
</paragraph>
</caption>
<paragraph blockId="6.[159,782,189,1901]" pageId="6" pageNumber="7">
Antennae inserted in small lateroventrally open cavities; heptamerous, with broad basal part of scapus (
<figureCitation captionStart="Figure 2" captionStartId="4.[159,226,1321,1342]" captionTargetBox="[159,1454,189,1304]" captionTargetId="figure@4.[159,1467,188,1304]" captionTargetPageId="4" captionText="Figure 2. †Lepiceratus ankylosaurus gen. and sp. nov., holotype, synchrotron microtomography, surface reconstructions. A, dorsal habitus; B, lateral habitus; C, right antenna and eye in ventral view; D, ventral habitus. Abbreviations: anc, antennal club; anf, antennal furrow; an3–6, antennomere 3–6; ce, compound eye; elex, elytral excavation;?mxp, presumable maxillary palp; pd, pedicel; sc, scape; v2, mesoventrite." figureDoi="http://doi.org/10.5281/zenodo.3763698" httpUri="https://zenodo.org/record/3763698/files/figure.png" pageId="6" pageNumber="7">Figs 2C</figureCitation>
,
<figureCitation box="[211,242,254,277]" captionStart="Figure 3" captionStartId="5.[159,226,1539,1560]" captionTargetBox="[159,1467,188,1517]" captionTargetId="figure@5.[159,1467,188,1517]" captionTargetPageId="5" captionText="Figure 3. †Lepiceratus ankylosaurus gen. and sp. nov., holotype, synchrotron microtomography, volume renderings. A, dorsal habitus; B, submedian sagittal virtual section; C, right antenna, eye and anterolateral portion of prothorax in ventral view; D, distal surface of right antennal club; E, ventral habitus. Abbreviations: anc, antennal club; anf, antennal furrow; anp, antennal pocket; an3–6, antennomere 3–6; ce, compound eye; els, elytral suture; epi, elytral epipleura; pd, pedicel; ppr, prosternal process; sc, scape; scs, scutellar shield; wg, wings." figureDoi="http://doi.org/10.5281/zenodo.3763700" httpUri="https://zenodo.org/record/3763700/files/figure.png" pageId="6" pageNumber="7">3C</figureCitation>
) and narrower distal portion; pedicellus (
<figureCitation captionStart="Figure 2" captionStartId="4.[159,226,1321,1342]" captionTargetBox="[159,1454,189,1304]" captionTargetId="figure@4.[159,1467,188,1304]" captionTargetPageId="4" captionText="Figure 2. †Lepiceratus ankylosaurus gen. and sp. nov., holotype, synchrotron microtomography, surface reconstructions. A, dorsal habitus; B, lateral habitus; C, right antenna and eye in ventral view; D, ventral habitus. Abbreviations: anc, antennal club; anf, antennal furrow; an3–6, antennomere 3–6; ce, compound eye; elex, elytral excavation;?mxp, presumable maxillary palp; pd, pedicel; sc, scape; v2, mesoventrite." figureDoi="http://doi.org/10.5281/zenodo.3763698" httpUri="https://zenodo.org/record/3763698/files/figure.png" pageId="6" pageNumber="7">Figs 2C</figureCitation>
,
<figureCitation box="[210,241,286,309]" captionStart="Figure 3" captionStartId="5.[159,226,1539,1560]" captionTargetBox="[159,1467,188,1517]" captionTargetId="figure@5.[159,1467,188,1517]" captionTargetPageId="5" captionText="Figure 3. †Lepiceratus ankylosaurus gen. and sp. nov., holotype, synchrotron microtomography, volume renderings. A, dorsal habitus; B, submedian sagittal virtual section; C, right antenna, eye and anterolateral portion of prothorax in ventral view; D, distal surface of right antennal club; E, ventral habitus. Abbreviations: anc, antennal club; anf, antennal furrow; anp, antennal pocket; an3–6, antennomere 3–6; ce, compound eye; els, elytral suture; epi, elytral epipleura; pd, pedicel; ppr, prosternal process; sc, scape; scs, scutellar shield; wg, wings." figureDoi="http://doi.org/10.5281/zenodo.3763700" httpUri="https://zenodo.org/record/3763700/files/figure.png" pageId="6" pageNumber="7">3C</figureCitation>
) sub-cylindrical; antennomeres 3–6 indistinctly widening; apical antennomere 7 forming very large club (
<figureCitation box="[168,253,351,374]" captionStart="Figure 2" captionStartId="4.[159,226,1321,1342]" captionTargetBox="[159,1454,189,1304]" captionTargetId="figure@4.[159,1467,188,1304]" captionTargetPageId="4" captionText="Figure 2. †Lepiceratus ankylosaurus gen. and sp. nov., holotype, synchrotron microtomography, surface reconstructions. A, dorsal habitus; B, lateral habitus; C, right antenna and eye in ventral view; D, ventral habitus. Abbreviations: anc, antennal club; anf, antennal furrow; an3–6, antennomere 3–6; ce, compound eye; elex, elytral excavation;?mxp, presumable maxillary palp; pd, pedicel; sc, scape; v2, mesoventrite." figureDoi="http://doi.org/10.5281/zenodo.3763698" httpUri="https://zenodo.org/record/3763698/files/figure.png" pageId="6" pageNumber="7">Figs 2C</figureCitation>
,
<figureCitation box="[269,300,351,374]" captionStart="Figure 3" captionStartId="5.[159,226,1539,1560]" captionTargetBox="[159,1467,188,1517]" captionTargetId="figure@5.[159,1467,188,1517]" captionTargetPageId="5" captionText="Figure 3. †Lepiceratus ankylosaurus gen. and sp. nov., holotype, synchrotron microtomography, volume renderings. A, dorsal habitus; B, submedian sagittal virtual section; C, right antenna, eye and anterolateral portion of prothorax in ventral view; D, distal surface of right antennal club; E, ventral habitus. Abbreviations: anc, antennal club; anf, antennal furrow; anp, antennal pocket; an3–6, antennomere 3–6; ce, compound eye; els, elytral suture; epi, elytral epipleura; pd, pedicel; ppr, prosternal process; sc, scape; scs, scutellar shield; wg, wings." figureDoi="http://doi.org/10.5281/zenodo.3763700" httpUri="https://zenodo.org/record/3763700/files/figure.png" pageId="6" pageNumber="7">3C</figureCitation>
), broadening distad and obliquely truncated at apex; distal surface (
<figureCitation box="[459,534,384,407]" captionStart="Figure 3" captionStartId="5.[159,226,1539,1560]" captionTargetBox="[159,1467,188,1517]" captionTargetId="figure@5.[159,1467,188,1517]" captionTargetPageId="5" captionText="Figure 3. †Lepiceratus ankylosaurus gen. and sp. nov., holotype, synchrotron microtomography, volume renderings. A, dorsal habitus; B, submedian sagittal virtual section; C, right antenna, eye and anterolateral portion of prothorax in ventral view; D, distal surface of right antennal club; E, ventral habitus. Abbreviations: anc, antennal club; anf, antennal furrow; anp, antennal pocket; an3–6, antennomere 3–6; ce, compound eye; els, elytral suture; epi, elytral epipleura; pd, pedicel; ppr, prosternal process; sc, scape; scs, scutellar shield; wg, wings." figureDoi="http://doi.org/10.5281/zenodo.3763700" httpUri="https://zenodo.org/record/3763700/files/figure.png" pageId="6" pageNumber="7">Fig. 3</figureCitation>
) oval, with distinct circumferential rim.
</paragraph>
<paragraph blockId="6.[159,782,189,1901]" pageId="6" pageNumber="7">
Pronotum sub-trapezoidal, strongly transverse; anterior and posterior corners well-marked but blunt, obtuse-angled; lateral margins irregularly undulate. Anteroventral surface behind each eye with antennal pocket (
<figureCitation box="[254,347,579,602]" captionStart="Figure 1" captionStartId="3.[159,226,1858,1879]" captionTargetBox="[175,1455,196,1830]" captionTargetId="figure@3.[159,1467,188,1840]" captionTargetPageId="3" captionText="Figure 1. †Lepiceratus ankylosaurus gen. and sp. nov., holotype, A, B, macrophotographs; C, artistic reconstruction; and D, schematic morphological details observable under light microscope. A, C, dorsal habitus; B, D, ventral habitus. Abbreviations: anf, antennal furrow; anp, antennal pocket; ce, compound eye; cx1–3, coxa; elex, elytral excavation; fm1–3, femur; hyp, hypomeron; kes, katepisternum; lb, labrum; mnt, mentum; ppr, prosternal process; r1–5, elytral rows of tubercles; st, abdominal sternite 2; tb1–3, tibia; tch1–3, trochanter; tr1–3, tarsus; v2, mesoventrite; v3, metaventrite." figureDoi="http://doi.org/10.5281/zenodo.3763696" httpUri="https://zenodo.org/record/3763696/files/figure.png" pageId="6" pageNumber="7">Figs 1D</figureCitation>
,
<figureCitation box="[369,400,579,602]" captionStart="Figure 3" captionStartId="5.[159,226,1539,1560]" captionTargetBox="[159,1467,188,1517]" captionTargetId="figure@5.[159,1467,188,1517]" captionTargetPageId="5" captionText="Figure 3. †Lepiceratus ankylosaurus gen. and sp. nov., holotype, synchrotron microtomography, volume renderings. A, dorsal habitus; B, submedian sagittal virtual section; C, right antenna, eye and anterolateral portion of prothorax in ventral view; D, distal surface of right antennal club; E, ventral habitus. Abbreviations: anc, antennal club; anf, antennal furrow; anp, antennal pocket; an3–6, antennomere 3–6; ce, compound eye; els, elytral suture; epi, elytral epipleura; pd, pedicel; ppr, prosternal process; sc, scape; scs, scutellar shield; wg, wings." figureDoi="http://doi.org/10.5281/zenodo.3763700" httpUri="https://zenodo.org/record/3763700/files/figure.png" pageId="6" pageNumber="7">3C</figureCitation>
) to accommodate antennal club. Large median area of pronotal disc strongly convex and bearing longitudinal rows of large tubercles, except for longitudinal and impressed median region; lateral portions of pronotal disc abruptly demarcated, slightly concave dorsally, laterally forming complete lateral edges.
</paragraph>
<paragraph blockId="6.[159,782,189,1901]" pageId="6" pageNumber="7">
Prosternum short and laterally distinctly demarcated from broad pronotal hypomera (
<figureCitation box="[512,596,806,829]" captionStart="Figure 1" captionStartId="3.[159,226,1858,1879]" captionTargetBox="[175,1455,196,1830]" captionTargetId="figure@3.[159,1467,188,1840]" captionTargetPageId="3" captionText="Figure 1. †Lepiceratus ankylosaurus gen. and sp. nov., holotype, A, B, macrophotographs; C, artistic reconstruction; and D, schematic morphological details observable under light microscope. A, C, dorsal habitus; B, D, ventral habitus. Abbreviations: anf, antennal furrow; anp, antennal pocket; ce, compound eye; cx1–3, coxa; elex, elytral excavation; fm1–3, femur; hyp, hypomeron; kes, katepisternum; lb, labrum; mnt, mentum; ppr, prosternal process; r1–5, elytral rows of tubercles; st, abdominal sternite 2; tb1–3, tibia; tch1–3, trochanter; tr1–3, tarsus; v2, mesoventrite; v3, metaventrite." figureDoi="http://doi.org/10.5281/zenodo.3763696" httpUri="https://zenodo.org/record/3763696/files/figure.png" pageId="6" pageNumber="7">Fig. 1D</figureCitation>
); prosternal process (
<figureCitation box="[222,310,838,861]" captionStart="Figure 1" captionStartId="3.[159,226,1858,1879]" captionTargetBox="[175,1455,196,1830]" captionTargetId="figure@3.[159,1467,188,1840]" captionTargetPageId="3" captionText="Figure 1. †Lepiceratus ankylosaurus gen. and sp. nov., holotype, A, B, macrophotographs; C, artistic reconstruction; and D, schematic morphological details observable under light microscope. A, C, dorsal habitus; B, D, ventral habitus. Abbreviations: anf, antennal furrow; anp, antennal pocket; ce, compound eye; cx1–3, coxa; elex, elytral excavation; fm1–3, femur; hyp, hypomeron; kes, katepisternum; lb, labrum; mnt, mentum; ppr, prosternal process; r1–5, elytral rows of tubercles; st, abdominal sternite 2; tb1–3, tibia; tch1–3, trochanter; tr1–3, tarsus; v2, mesoventrite; v3, metaventrite." figureDoi="http://doi.org/10.5281/zenodo.3763696" httpUri="https://zenodo.org/record/3763696/files/figure.png" pageId="6" pageNumber="7">Figs 1D</figureCitation>
,
<figureCitation box="[326,356,838,861]" captionStart="Figure 3" captionStartId="5.[159,226,1539,1560]" captionTargetBox="[159,1467,188,1517]" captionTargetId="figure@5.[159,1467,188,1517]" captionTargetPageId="5" captionText="Figure 3. †Lepiceratus ankylosaurus gen. and sp. nov., holotype, synchrotron microtomography, volume renderings. A, dorsal habitus; B, submedian sagittal virtual section; C, right antenna, eye and anterolateral portion of prothorax in ventral view; D, distal surface of right antennal club; E, ventral habitus. Abbreviations: anc, antennal club; anf, antennal furrow; anp, antennal pocket; an3–6, antennomere 3–6; ce, compound eye; els, elytral suture; epi, elytral epipleura; pd, pedicel; ppr, prosternal process; sc, scape; scs, scutellar shield; wg, wings." figureDoi="http://doi.org/10.5281/zenodo.3763700" httpUri="https://zenodo.org/record/3763700/files/figure.png" pageId="6" pageNumber="7">3E</figureCitation>
) sub-triangular and elongate, with lateral margins convergent posterad toward pointed apex, posterior process nearly reaching hind margins of procoxae, its ventral surface with pair of elongate lateral bulges which anteriorly continue along posterior prosternal margin, posterior portion of process behind bulges distinctly concave. Pleura and trochantin not discernible.
</paragraph>
<paragraph blockId="6.[159,782,189,1901]" pageId="6" pageNumber="7">
Mesonotum exposed between pronotum and elytra but details of scutum and main part of scutellum not recognizable, except for small triangular scutellar shield (
<figureCitation box="[168,267,1163,1186]" captionStart="Figure 3" captionStartId="5.[159,226,1539,1560]" captionTargetBox="[159,1467,188,1517]" captionTargetId="figure@5.[159,1467,188,1517]" captionTargetPageId="5" captionText="Figure 3. †Lepiceratus ankylosaurus gen. and sp. nov., holotype, synchrotron microtomography, volume renderings. A, dorsal habitus; B, submedian sagittal virtual section; C, right antenna, eye and anterolateral portion of prothorax in ventral view; D, distal surface of right antennal club; E, ventral habitus. Abbreviations: anc, antennal club; anf, antennal furrow; anp, antennal pocket; an3–6, antennomere 3–6; ce, compound eye; els, elytral suture; epi, elytral epipleura; pd, pedicel; ppr, prosternal process; sc, scape; scs, scutellar shield; wg, wings." figureDoi="http://doi.org/10.5281/zenodo.3763700" httpUri="https://zenodo.org/record/3763700/files/figure.png" pageId="6" pageNumber="7">Fig. 3A</figureCitation>
) between mesal edges of elytral base. Mesoventrite (
<figureCitation box="[317,401,1196,1219]" captionStart="Figure 1" captionStartId="3.[159,226,1858,1879]" captionTargetBox="[175,1455,196,1830]" captionTargetId="figure@3.[159,1467,188,1840]" captionTargetPageId="3" captionText="Figure 1. †Lepiceratus ankylosaurus gen. and sp. nov., holotype, A, B, macrophotographs; C, artistic reconstruction; and D, schematic morphological details observable under light microscope. A, C, dorsal habitus; B, D, ventral habitus. Abbreviations: anf, antennal furrow; anp, antennal pocket; ce, compound eye; cx1–3, coxa; elex, elytral excavation; fm1–3, femur; hyp, hypomeron; kes, katepisternum; lb, labrum; mnt, mentum; ppr, prosternal process; r1–5, elytral rows of tubercles; st, abdominal sternite 2; tb1–3, tibia; tch1–3, trochanter; tr1–3, tarsus; v2, mesoventrite; v3, metaventrite." figureDoi="http://doi.org/10.5281/zenodo.3763696" httpUri="https://zenodo.org/record/3763696/files/figure.png" pageId="6" pageNumber="7">Fig. 1D</figureCitation>
) sub-rhomboidal in shape, lacking visible discrimen and transverse suture; anterior margin sub-triangular with slightly sinuate sides, forming anteromedian process strongly projecting anterad (
<figureCitation box="[680,766,1293,1316]" captionStart="Figure 2" captionStartId="4.[159,226,1321,1342]" captionTargetBox="[159,1454,189,1304]" captionTargetId="figure@4.[159,1467,188,1304]" captionTargetPageId="4" captionText="Figure 2. †Lepiceratus ankylosaurus gen. and sp. nov., holotype, synchrotron microtomography, surface reconstructions. A, dorsal habitus; B, lateral habitus; C, right antenna and eye in ventral view; D, ventral habitus. Abbreviations: anc, antennal club; anf, antennal furrow; an3–6, antennomere 3–6; ce, compound eye; elex, elytral excavation;?mxp, presumable maxillary palp; pd, pedicel; sc, scape; v2, mesoventrite." figureDoi="http://doi.org/10.5281/zenodo.3763698" httpUri="https://zenodo.org/record/3763698/files/figure.png" pageId="6" pageNumber="7">Fig. 2B</figureCitation>
), which likely articulates with caudal concave portion of prosternal process when prothorax is in normal position (i.e. as in living beetles). Median area of mesoventrite between mesocoxae distinctly impressed. Mesotrochantin not recognizable.
</paragraph>
<paragraph blockId="6.[159,782,189,1901]" pageId="6" pageNumber="7">
Metanotum not visible. Metaventrite (
<figureCitation box="[615,703,1488,1511]" captionStart="Figure 1" captionStartId="3.[159,226,1858,1879]" captionTargetBox="[175,1455,196,1830]" captionTargetId="figure@3.[159,1467,188,1840]" captionTargetPageId="3" captionText="Figure 1. †Lepiceratus ankylosaurus gen. and sp. nov., holotype, A, B, macrophotographs; C, artistic reconstruction; and D, schematic morphological details observable under light microscope. A, C, dorsal habitus; B, D, ventral habitus. Abbreviations: anf, antennal furrow; anp, antennal pocket; ce, compound eye; cx1–3, coxa; elex, elytral excavation; fm1–3, femur; hyp, hypomeron; kes, katepisternum; lb, labrum; mnt, mentum; ppr, prosternal process; r1–5, elytral rows of tubercles; st, abdominal sternite 2; tb1–3, tibia; tch1–3, trochanter; tr1–3, tarsus; v2, mesoventrite; v3, metaventrite." figureDoi="http://doi.org/10.5281/zenodo.3763696" httpUri="https://zenodo.org/record/3763696/files/figure.png" pageId="6" pageNumber="7">Fig. 1D</figureCitation>
) lacks externally visible discrimen (but visible on inner, dorsal surface of metaventrite in tomographic sections), with complete transverse suture, broad and short anteromedian intercoxal process overlapping with posterior mesoventrite. Median and lateral regions of metaventrite distinctly impressed, with anterior and lateral margins elevated. Katepisternum (
<figureCitation box="[334,421,1716,1739]" captionStart="Figure 1" captionStartId="3.[159,226,1858,1879]" captionTargetBox="[175,1455,196,1830]" captionTargetId="figure@3.[159,1467,188,1840]" captionTargetPageId="3" captionText="Figure 1. †Lepiceratus ankylosaurus gen. and sp. nov., holotype, A, B, macrophotographs; C, artistic reconstruction; and D, schematic morphological details observable under light microscope. A, C, dorsal habitus; B, D, ventral habitus. Abbreviations: anf, antennal furrow; anp, antennal pocket; ce, compound eye; cx1–3, coxa; elex, elytral excavation; fm1–3, femur; hyp, hypomeron; kes, katepisternum; lb, labrum; mnt, mentum; ppr, prosternal process; r1–5, elytral rows of tubercles; st, abdominal sternite 2; tb1–3, tibia; tch1–3, trochanter; tr1–3, tarsus; v2, mesoventrite; v3, metaventrite." figureDoi="http://doi.org/10.5281/zenodo.3763696" httpUri="https://zenodo.org/record/3763696/files/figure.png" pageId="6" pageNumber="7">Fig. 1D</figureCitation>
) large, widening laterally, with two paramedian posterior (intercoxal) triangular processes separated by rounded median excavation.
</paragraph>
<paragraph blockId="6.[159,782,189,1901]" lastBlockId="6.[844,1467,189,1351]" pageId="6" pageNumber="7">
Elytra (
<figureCitation box="[269,390,1812,1836]" captionStart="Figure 1" captionStartId="3.[159,226,1858,1879]" captionTargetBox="[175,1455,196,1830]" captionTargetId="figure@3.[159,1467,188,1840]" captionTargetPageId="3" captionText="Figure 1. †Lepiceratus ankylosaurus gen. and sp. nov., holotype, A, B, macrophotographs; C, artistic reconstruction; and D, schematic morphological details observable under light microscope. A, C, dorsal habitus; B, D, ventral habitus. Abbreviations: anf, antennal furrow; anp, antennal pocket; ce, compound eye; cx1–3, coxa; elex, elytral excavation; fm1–3, femur; hyp, hypomeron; kes, katepisternum; lb, labrum; mnt, mentum; ppr, prosternal process; r1–5, elytral rows of tubercles; st, abdominal sternite 2; tb1–3, tibia; tch1–3, trochanter; tr1–3, tarsus; v2, mesoventrite; v3, metaventrite." figureDoi="http://doi.org/10.5281/zenodo.3763696" httpUri="https://zenodo.org/record/3763696/files/figure.png" pageId="6" pageNumber="7">Figs 1A–D</figureCitation>
,
<figureCitation box="[407,509,1813,1836]" captionStart="Figure 2" captionStartId="4.[159,226,1321,1342]" captionTargetBox="[159,1454,189,1304]" captionTargetId="figure@4.[159,1467,188,1304]" captionTargetPageId="4" captionText="Figure 2. †Lepiceratus ankylosaurus gen. and sp. nov., holotype, synchrotron microtomography, surface reconstructions. A, dorsal habitus; B, lateral habitus; C, right antenna and eye in ventral view; D, ventral habitus. Abbreviations: anc, antennal club; anf, antennal furrow; an3–6, antennomere 3–6; ce, compound eye; elex, elytral excavation;?mxp, presumable maxillary palp; pd, pedicel; sc, scape; v2, mesoventrite." figureDoi="http://doi.org/10.5281/zenodo.3763698" httpUri="https://zenodo.org/record/3763698/files/figure.png" pageId="6" pageNumber="7">2A, B, D</figureCitation>
,
<figureCitation box="[526,590,1813,1836]" captionStart="Figure 3" captionStartId="5.[159,226,1539,1560]" captionTargetBox="[159,1467,188,1517]" captionTargetId="figure@5.[159,1467,188,1517]" captionTargetPageId="5" captionText="Figure 3. †Lepiceratus ankylosaurus gen. and sp. nov., holotype, synchrotron microtomography, volume renderings. A, dorsal habitus; B, submedian sagittal virtual section; C, right antenna, eye and anterolateral portion of prothorax in ventral view; D, distal surface of right antennal club; E, ventral habitus. Abbreviations: anc, antennal club; anf, antennal furrow; anp, antennal pocket; an3–6, antennomere 3–6; ce, compound eye; els, elytral suture; epi, elytral epipleura; pd, pedicel; ppr, prosternal process; sc, scape; scs, scutellar shield; wg, wings." figureDoi="http://doi.org/10.5281/zenodo.3763700" httpUri="https://zenodo.org/record/3763700/files/figure.png" pageId="6" pageNumber="7">3A, E</figureCitation>
) slightly broader than pronotum. Humeral angles strongly pronounced; elytral suture (
<figureCitation box="[321,406,1878,1901]" captionStart="Figure 3" captionStartId="5.[159,226,1539,1560]" captionTargetBox="[159,1467,188,1517]" captionTargetId="figure@5.[159,1467,188,1517]" captionTargetPageId="5" captionText="Figure 3. †Lepiceratus ankylosaurus gen. and sp. nov., holotype, synchrotron microtomography, volume renderings. A, dorsal habitus; B, submedian sagittal virtual section; C, right antenna, eye and anterolateral portion of prothorax in ventral view; D, distal surface of right antennal club; E, ventral habitus. Abbreviations: anc, antennal club; anf, antennal furrow; anp, antennal pocket; an3–6, antennomere 3–6; ce, compound eye; els, elytral suture; epi, elytral epipleura; pd, pedicel; ppr, prosternal process; sc, scape; scs, scutellar shield; wg, wings." figureDoi="http://doi.org/10.5281/zenodo.3763700" httpUri="https://zenodo.org/record/3763700/files/figure.png" pageId="6" pageNumber="7">Fig. 3A</figureCitation>
) well-marked. Elytral disc with a pattern of strongly developed tubercles arranged in four complete rows (
<figureCitation box="[1033,1123,221,244]" captionStart="Figure 1" captionStartId="3.[159,226,1858,1879]" captionTargetBox="[175,1455,196,1830]" captionTargetId="figure@3.[159,1467,188,1840]" captionTargetPageId="3" captionText="Figure 1. †Lepiceratus ankylosaurus gen. and sp. nov., holotype, A, B, macrophotographs; C, artistic reconstruction; and D, schematic morphological details observable under light microscope. A, C, dorsal habitus; B, D, ventral habitus. Abbreviations: anf, antennal furrow; anp, antennal pocket; ce, compound eye; cx1–3, coxa; elex, elytral excavation; fm1–3, femur; hyp, hypomeron; kes, katepisternum; lb, labrum; mnt, mentum; ppr, prosternal process; r1–5, elytral rows of tubercles; st, abdominal sternite 2; tb1–3, tibia; tch1–3, trochanter; tr1–3, tarsus; v2, mesoventrite; v3, metaventrite." figureDoi="http://doi.org/10.5281/zenodo.3763696" httpUri="https://zenodo.org/record/3763696/files/figure.png" pageId="6" pageNumber="7">Fig. 1A</figureCitation>
), and additionally with short row just laterad proximal portion of r4 (
<figureCitation box="[1305,1391,254,277]" captionStart="Figure 1" captionStartId="3.[159,226,1858,1879]" captionTargetBox="[175,1455,196,1830]" captionTargetId="figure@3.[159,1467,188,1840]" captionTargetPageId="3" captionText="Figure 1. †Lepiceratus ankylosaurus gen. and sp. nov., holotype, A, B, macrophotographs; C, artistic reconstruction; and D, schematic morphological details observable under light microscope. A, C, dorsal habitus; B, D, ventral habitus. Abbreviations: anf, antennal furrow; anp, antennal pocket; ce, compound eye; cx1–3, coxa; elex, elytral excavation; fm1–3, femur; hyp, hypomeron; kes, katepisternum; lb, labrum; mnt, mentum; ppr, prosternal process; r1–5, elytral rows of tubercles; st, abdominal sternite 2; tb1–3, tibia; tch1–3, trochanter; tr1–3, tarsus; v2, mesoventrite; v3, metaventrite." figureDoi="http://doi.org/10.5281/zenodo.3763696" httpUri="https://zenodo.org/record/3763696/files/figure.png" pageId="6" pageNumber="7">Fig. 1A</figureCitation>
); continuous longitudinal costae absent. Elytra with complete lateral carina and broad epipleura (
<figureCitation box="[1241,1325,319,342]" captionStart="Figure 3" captionStartId="5.[159,226,1539,1560]" captionTargetBox="[159,1467,188,1517]" captionTargetId="figure@5.[159,1467,188,1517]" captionTargetPageId="5" captionText="Figure 3. †Lepiceratus ankylosaurus gen. and sp. nov., holotype, synchrotron microtomography, volume renderings. A, dorsal habitus; B, submedian sagittal virtual section; C, right antenna, eye and anterolateral portion of prothorax in ventral view; D, distal surface of right antennal club; E, ventral habitus. Abbreviations: anc, antennal club; anf, antennal furrow; anp, antennal pocket; an3–6, antennomere 3–6; ce, compound eye; els, elytral suture; epi, elytral epipleura; pd, pedicel; ppr, prosternal process; sc, scape; scs, scutellar shield; wg, wings." figureDoi="http://doi.org/10.5281/zenodo.3763700" httpUri="https://zenodo.org/record/3763700/files/figure.png" pageId="6" pageNumber="7">Fig. 3E</figureCitation>
), each bearing two excavations (
<figureCitation box="[1081,1169,352,375]" captionStart="Figure 1" captionStartId="3.[159,226,1858,1879]" captionTargetBox="[175,1455,196,1830]" captionTargetId="figure@3.[159,1467,188,1840]" captionTargetPageId="3" captionText="Figure 1. †Lepiceratus ankylosaurus gen. and sp. nov., holotype, A, B, macrophotographs; C, artistic reconstruction; and D, schematic morphological details observable under light microscope. A, C, dorsal habitus; B, D, ventral habitus. Abbreviations: anf, antennal furrow; anp, antennal pocket; ce, compound eye; cx1–3, coxa; elex, elytral excavation; fm1–3, femur; hyp, hypomeron; kes, katepisternum; lb, labrum; mnt, mentum; ppr, prosternal process; r1–5, elytral rows of tubercles; st, abdominal sternite 2; tb1–3, tibia; tch1–3, trochanter; tr1–3, tarsus; v2, mesoventrite; v3, metaventrite." figureDoi="http://doi.org/10.5281/zenodo.3763696" httpUri="https://zenodo.org/record/3763696/files/figure.png" pageId="6" pageNumber="7">Figs 1D</figureCitation>
,
<figureCitation box="[1184,1217,352,375]" captionStart="Figure 2" captionStartId="4.[159,226,1321,1342]" captionTargetBox="[159,1454,189,1304]" captionTargetId="figure@4.[159,1467,188,1304]" captionTargetPageId="4" captionText="Figure 2. †Lepiceratus ankylosaurus gen. and sp. nov., holotype, synchrotron microtomography, surface reconstructions. A, dorsal habitus; B, lateral habitus; C, right antenna and eye in ventral view; D, ventral habitus. Abbreviations: anc, antennal club; anf, antennal furrow; an3–6, antennomere 3–6; ce, compound eye; elex, elytral excavation;?mxp, presumable maxillary palp; pd, pedicel; sc, scape; v2, mesoventrite." figureDoi="http://doi.org/10.5281/zenodo.3763698" httpUri="https://zenodo.org/record/3763698/files/figure.png" pageId="6" pageNumber="7">2D</figureCitation>
) for mesofemur (anterolaterally) and for metafemur (in median region). Elytra interlocked with metapleuron anterad metacoxae by conspicuous mesally directed process. Lateral outline of elytral edge (
<figureCitation box="[1034,1121,482,505]" captionStart="Figure 2" captionStartId="4.[159,226,1321,1342]" captionTargetBox="[159,1454,189,1304]" captionTargetId="figure@4.[159,1467,188,1304]" captionTargetPageId="4" captionText="Figure 2. †Lepiceratus ankylosaurus gen. and sp. nov., holotype, synchrotron microtomography, surface reconstructions. A, dorsal habitus; B, lateral habitus; C, right antenna and eye in ventral view; D, ventral habitus. Abbreviations: anc, antennal club; anf, antennal furrow; an3–6, antennomere 3–6; ce, compound eye; elex, elytral excavation;?mxp, presumable maxillary palp; pd, pedicel; sc, scape; v2, mesoventrite." figureDoi="http://doi.org/10.5281/zenodo.3763698" httpUri="https://zenodo.org/record/3763698/files/figure.png" pageId="6" pageNumber="7">Fig. 2B</figureCitation>
) strongly bisinuate, with submedian region forming broad sub-triangular ‘tooth’.
</paragraph>
<paragraph blockId="6.[844,1467,189,1351]" pageId="6" pageNumber="7">
Hind wings not exposed, but well-visible in microtomographic sections, folded in sub-elytral space (
<figureCitation box="[853,936,612,635]" captionStart="Figure 3" captionStartId="5.[159,226,1539,1560]" captionTargetBox="[159,1467,188,1517]" captionTargetId="figure@5.[159,1467,188,1517]" captionTargetPageId="5" captionText="Figure 3. †Lepiceratus ankylosaurus gen. and sp. nov., holotype, synchrotron microtomography, volume renderings. A, dorsal habitus; B, submedian sagittal virtual section; C, right antenna, eye and anterolateral portion of prothorax in ventral view; D, distal surface of right antennal club; E, ventral habitus. Abbreviations: anc, antennal club; anf, antennal furrow; anp, antennal pocket; an3–6, antennomere 3–6; ce, compound eye; els, elytral suture; epi, elytral epipleura; pd, pedicel; ppr, prosternal process; sc, scape; scs, scutellar shield; wg, wings." figureDoi="http://doi.org/10.5281/zenodo.3763700" httpUri="https://zenodo.org/record/3763700/files/figure.png" pageId="6" pageNumber="7">Fig. 3B</figureCitation>
).
</paragraph>
<paragraph blockId="6.[844,1467,189,1351]" pageId="6" pageNumber="7">
Legs moderately long and robust; coxae (
<figureCitation box="[1324,1406,644,667]" captionStart="Figure 1" captionStartId="3.[159,226,1858,1879]" captionTargetBox="[175,1455,196,1830]" captionTargetId="figure@3.[159,1467,188,1840]" captionTargetPageId="3" captionText="Figure 1. †Lepiceratus ankylosaurus gen. and sp. nov., holotype, A, B, macrophotographs; C, artistic reconstruction; and D, schematic morphological details observable under light microscope. A, C, dorsal habitus; B, D, ventral habitus. Abbreviations: anf, antennal furrow; anp, antennal pocket; ce, compound eye; cx1–3, coxa; elex, elytral excavation; fm1–3, femur; hyp, hypomeron; kes, katepisternum; lb, labrum; mnt, mentum; ppr, prosternal process; r1–5, elytral rows of tubercles; st, abdominal sternite 2; tb1–3, tibia; tch1–3, trochanter; tr1–3, tarsus; v2, mesoventrite; v3, metaventrite." figureDoi="http://doi.org/10.5281/zenodo.3763696" httpUri="https://zenodo.org/record/3763696/files/figure.png" pageId="6" pageNumber="7">Fig. 1D</figureCitation>
) flat, not elevated ventrad beyond ventral surface of body; trochanters (
<figureCitation box="[954,1042,709,732]" captionStart="Figure 1" captionStartId="3.[159,226,1858,1879]" captionTargetBox="[175,1455,196,1830]" captionTargetId="figure@3.[159,1467,188,1840]" captionTargetPageId="3" captionText="Figure 1. †Lepiceratus ankylosaurus gen. and sp. nov., holotype, A, B, macrophotographs; C, artistic reconstruction; and D, schematic morphological details observable under light microscope. A, C, dorsal habitus; B, D, ventral habitus. Abbreviations: anf, antennal furrow; anp, antennal pocket; ce, compound eye; cx1–3, coxa; elex, elytral excavation; fm1–3, femur; hyp, hypomeron; kes, katepisternum; lb, labrum; mnt, mentum; ppr, prosternal process; r1–5, elytral rows of tubercles; st, abdominal sternite 2; tb1–3, tibia; tch1–3, trochanter; tr1–3, tarsus; v2, mesoventrite; v3, metaventrite." figureDoi="http://doi.org/10.5281/zenodo.3763696" httpUri="https://zenodo.org/record/3763696/files/figure.png" pageId="6" pageNumber="7">Fig. 1D</figureCitation>
) sub-triangular, similar in size in all three pairs of legs; trochantero-femoral articulation oblique; femora (
<figureCitation box="[1037,1122,774,797]" captionStart="Figure 1" captionStartId="3.[159,226,1858,1879]" captionTargetBox="[175,1455,196,1830]" captionTargetId="figure@3.[159,1467,188,1840]" captionTargetPageId="3" captionText="Figure 1. †Lepiceratus ankylosaurus gen. and sp. nov., holotype, A, B, macrophotographs; C, artistic reconstruction; and D, schematic morphological details observable under light microscope. A, C, dorsal habitus; B, D, ventral habitus. Abbreviations: anf, antennal furrow; anp, antennal pocket; ce, compound eye; cx1–3, coxa; elex, elytral excavation; fm1–3, femur; hyp, hypomeron; kes, katepisternum; lb, labrum; mnt, mentum; ppr, prosternal process; r1–5, elytral rows of tubercles; st, abdominal sternite 2; tb1–3, tibia; tch1–3, trochanter; tr1–3, tarsus; v2, mesoventrite; v3, metaventrite." figureDoi="http://doi.org/10.5281/zenodo.3763696" httpUri="https://zenodo.org/record/3763696/files/figure.png" pageId="6" pageNumber="7">Fig. 1D</figureCitation>
) flattened, only profemora laterally reaching beyond lateral body margins; tibiae (
<figureCitation captionStart="Figure 1" captionStartId="3.[159,226,1858,1879]" captionTargetBox="[175,1455,196,1830]" captionTargetId="figure@3.[159,1467,188,1840]" captionTargetPageId="3" captionText="Figure 1. †Lepiceratus ankylosaurus gen. and sp. nov., holotype, A, B, macrophotographs; C, artistic reconstruction; and D, schematic morphological details observable under light microscope. A, C, dorsal habitus; B, D, ventral habitus. Abbreviations: anf, antennal furrow; anp, antennal pocket; ce, compound eye; cx1–3, coxa; elex, elytral excavation; fm1–3, femur; hyp, hypomeron; kes, katepisternum; lb, labrum; mnt, mentum; ppr, prosternal process; r1–5, elytral rows of tubercles; st, abdominal sternite 2; tb1–3, tibia; tch1–3, trochanter; tr1–3, tarsus; v2, mesoventrite; v3, metaventrite." figureDoi="http://doi.org/10.5281/zenodo.3763696" httpUri="https://zenodo.org/record/3763696/files/figure.png" pageId="6" pageNumber="7">Fig. 1D</figureCitation>
) sub-cylindrical, narrower than femora; tarsi (
<figureCitation box="[1375,1458,839,862]" captionStart="Figure 1" captionStartId="3.[159,226,1858,1879]" captionTargetBox="[175,1455,196,1830]" captionTargetId="figure@3.[159,1467,188,1840]" captionTargetPageId="3" captionText="Figure 1. †Lepiceratus ankylosaurus gen. and sp. nov., holotype, A, B, macrophotographs; C, artistic reconstruction; and D, schematic morphological details observable under light microscope. A, C, dorsal habitus; B, D, ventral habitus. Abbreviations: anf, antennal furrow; anp, antennal pocket; ce, compound eye; cx1–3, coxa; elex, elytral excavation; fm1–3, femur; hyp, hypomeron; kes, katepisternum; lb, labrum; mnt, mentum; ppr, prosternal process; r1–5, elytral rows of tubercles; st, abdominal sternite 2; tb1–3, tibia; tch1–3, trochanter; tr1–3, tarsus; v2, mesoventrite; v3, metaventrite." figureDoi="http://doi.org/10.5281/zenodo.3763696" httpUri="https://zenodo.org/record/3763696/files/figure.png" pageId="6" pageNumber="7">Fig. 1D</figureCitation>
) only slightly shorter than tibiae, slender, sub-cylindrical, either indistinctly divided into four units (protarsus) or completely undivided (meso- and metatarsus), with rows of short sub-erect spines on anterior and posterior margins and each with one long and weakly curved claw.
</paragraph>
<paragraph blockId="6.[844,1467,189,1351]" pageId="6" pageNumber="7">
Abdomen (
<figureCitation box="[995,1118,1034,1057]" captionStart="Figure 1" captionStartId="3.[159,226,1858,1879]" captionTargetBox="[175,1455,196,1830]" captionTargetId="figure@3.[159,1467,188,1840]" captionTargetPageId="3" captionText="Figure 1. †Lepiceratus ankylosaurus gen. and sp. nov., holotype, A, B, macrophotographs; C, artistic reconstruction; and D, schematic morphological details observable under light microscope. A, C, dorsal habitus; B, D, ventral habitus. Abbreviations: anf, antennal furrow; anp, antennal pocket; ce, compound eye; cx1–3, coxa; elex, elytral excavation; fm1–3, femur; hyp, hypomeron; kes, katepisternum; lb, labrum; mnt, mentum; ppr, prosternal process; r1–5, elytral rows of tubercles; st, abdominal sternite 2; tb1–3, tibia; tch1–3, trochanter; tr1–3, tarsus; v2, mesoventrite; v3, metaventrite." figureDoi="http://doi.org/10.5281/zenodo.3763696" httpUri="https://zenodo.org/record/3763696/files/figure.png" pageId="6" pageNumber="7">Figs 1B, D</figureCitation>
,
<figureCitation box="[1135,1168,1034,1057]" captionStart="Figure 2" captionStartId="4.[159,226,1321,1342]" captionTargetBox="[159,1454,189,1304]" captionTargetId="figure@4.[159,1467,188,1304]" captionTargetPageId="4" captionText="Figure 2. †Lepiceratus ankylosaurus gen. and sp. nov., holotype, synchrotron microtomography, surface reconstructions. A, dorsal habitus; B, lateral habitus; C, right antenna and eye in ventral view; D, ventral habitus. Abbreviations: anc, antennal club; anf, antennal furrow; an3–6, antennomere 3–6; ce, compound eye; elex, elytral excavation;?mxp, presumable maxillary palp; pd, pedicel; sc, scape; v2, mesoventrite." figureDoi="http://doi.org/10.5281/zenodo.3763698" httpUri="https://zenodo.org/record/3763698/files/figure.png" pageId="6" pageNumber="7">2D</figureCitation>
,
<figureCitation box="[1184,1248,1034,1057]" captionStart="Figure 3" captionStartId="5.[159,226,1539,1560]" captionTargetBox="[159,1467,188,1517]" captionTargetId="figure@5.[159,1467,188,1517]" captionTargetPageId="5" captionText="Figure 3. †Lepiceratus ankylosaurus gen. and sp. nov., holotype, synchrotron microtomography, volume renderings. A, dorsal habitus; B, submedian sagittal virtual section; C, right antenna, eye and anterolateral portion of prothorax in ventral view; D, distal surface of right antennal club; E, ventral habitus. Abbreviations: anc, antennal club; anf, antennal furrow; anp, antennal pocket; an3–6, antennomere 3–6; ce, compound eye; els, elytral suture; epi, elytral epipleura; pd, pedicel; ppr, prosternal process; sc, scape; scs, scutellar shield; wg, wings." figureDoi="http://doi.org/10.5281/zenodo.3763700" httpUri="https://zenodo.org/record/3763700/files/figure.png" pageId="6" pageNumber="7">3B, E</figureCitation>
) with seven visible sternites, lacking plastron. Sternite II forms broad median process between metacoxae, anteriorly fitting with katepisternal concavity; narrowed behind posterior metacoxal margin but widening laterad. Border between sternites II and III poorly visible medially but distinct at lateral edges. Posterior sternites form distinct bulges along their posterior edges and are gradually tapering towards rounded abdominal apex; median protuberance of terminal sternite not discernible.
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="7" type="etymology">
<paragraph blockId="6.[844,1467,1376,1431]" pageId="6" pageNumber="7">
<heading bold="true" box="[844,1127,1376,1399]" centered="true" fontSize="10" level="4" pageId="6" pageNumber="7" reason="4">
<emphasis bold="true" box="[844,1127,1376,1399]" pageId="6" pageNumber="7">Derivation of name.</emphasis>
</heading>
<heading box="[1162,1263,1376,1399]" centered="true" fontSize="10" level="5" pageId="6" pageNumber="7" reason="4">Variation</heading>
of
<heading box="[1355,1467,1376,1399]" centered="true" fontSize="10" level="5" pageId="6" pageNumber="7" reason="4">
<taxonomicName authorityName="Motschulsky" authorityYear="1855" box="[1355,1460,1376,1399]" class="Insecta" family="Curculionidae" genus="Lepicerus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis box="[1355,1460,1376,1399]" italics="true" pageId="6" pageNumber="7">Lepicerus</emphasis>
</taxonomicName>
.
</heading>
Gender masculine.
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="7" type="distribution">
<paragraph blockId="6.[844,1433,1456,1480]" box="[844,1433,1456,1480]" pageId="6" pageNumber="7">
<emphasis bold="true" box="[844,1078,1456,1479]" pageId="6" pageNumber="7">Stratigraphic range.</emphasis>
Late Albian–early Cenomanian.
</paragraph>
<paragraph blockId="6.[844,1466,1504,1560]" pageId="6" pageNumber="7">
<heading bold="true" box="[844,1097,1504,1527]" centered="true" fontSize="10" level="4" pageId="6" pageNumber="7" reason="4">
<emphasis bold="true" box="[844,1097,1504,1527]" pageId="6" pageNumber="7">Geographical range.</emphasis>
</heading>
<heading box="[1123,1219,1504,1527]" centered="true" fontSize="10" level="5" pageId="6" pageNumber="7" reason="4">Northern</heading>
<heading box="[1246,1356,1504,1527]" centered="true" fontSize="10" level="5" pageId="6" pageNumber="7" reason="4">Myanmar,</heading>
<heading box="[1383,1466,1504,1527]" centered="true" fontSize="10" level="5" pageId="6" pageNumber="7" reason="4">Kachin,</heading>
Hukawng Valley.
</paragraph>
</subSubSection>
<subSubSection lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7" type="discussion">
<paragraph blockId="6.[844,1467,1585,1901]" lastBlockId="7.[159,781,1044,1493]" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
<emphasis bold="true" box="[844,955,1585,1608]" pageId="6" pageNumber="7">Remarks.</emphasis>
The new genus is unambiguously placed in
<taxonomicName authorityName="Hinton" authorityYear="1936" box="[844,970,1617,1640]" class="Insecta" family="Lepiceridae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="family">Lepiceridae</taxonomicName>
on the basis of the following synapomorphies: body strongly sculptured; head large, macrophthalmous, with mouthparts partly internalized; antennae with reduced number of antennomeres, with the terminal one forming a very large undivided club; pronotum with distinctly marked anterior and posterior corners; head with antennal furrows and prothorax with antennal pockets; tarsomeres largely fused, with only indistinct septa visible on surface of protarsi; elytral epipleura with sinuate concavities to accommodate femora; elytra with longitudinal rows of convexities; prosternal process prominent and ‘U’- or ‘V’-shaped; mesoventral process anteriorly broadly sub-triangular and projecting; abdomen with seven visible sternites.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.3763702" ID-Zenodo-Dep="3763702" httpUri="https://zenodo.org/record/3763702/files/figure.png" pageId="7" pageNumber="8" startId="7.[159,226,813,834]" targetBox="[159,1467,189,795]" targetPageId="7">
<paragraph blockId="7.[159,1468,813,994]" pageId="7" pageNumber="8">
<emphasis bold="true" box="[159,252,813,834]" pageId="7" pageNumber="8">Figure 4.</emphasis>
Strict consensus tree of four most parsimonious trees obtained by analysis of morphological characters of
<taxonomicName authorityName="Crowson" authorityYear="1955" box="[1302,1416,813,834]" class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="subOrder" subOrder="Myxophaga">Myxophaga</taxonomicName>
with NONA (tree length 177 steps; consistency index
<emphasis bold="true" box="[657,675,839,861]" pageId="7" pageNumber="8">¼</emphasis>
0.76; retention index
<emphasis bold="true" box="[903,921,839,861]" pageId="7" pageNumber="8">¼</emphasis>
0.89); problematic †‘
<taxonomicName authorityName="Kirejtshuk &amp; Poinar" authorityYear="2006" box="[1142,1262,840,861]" class="Insecta" family="Lepiceridae" genus="Haplochelus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis box="[1142,1262,840,861]" italics="true" pageId="7" pageNumber="8">Haplochelus</emphasis>
</taxonomicName>
’ and † ‘
<taxonomicName authorityName="Kirejtshuk &amp; Poinar" authorityYear="2017" box="[1356,1459,840,861]" class="Insecta" family="Curculionidae" genus="Lepichelus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis box="[1356,1459,840,861]" italics="true" pageId="7" pageNumber="8">Lepichelus</emphasis>
</taxonomicName>
’ excluded. Bremer support value indicated beneath branches; unambiguously optimized character changes plotted along the internodes (black circles indicate unique character changes; white circles indicate parallelisms or reversals; character numbers are above circles; character states are below circles). Examples of extant and extinct myxophagan beetles are illustrated on the tree, from upper left: †
<taxonomicName authority="Jałoszynski &amp; Luo &amp; Hammel &amp; Yamamoto &amp; Beutel, 2020" authorityName="Jałoszynski &amp; Luo &amp; Hammel &amp; Yamamoto &amp; Beutel" authorityYear="2020" box="[171,429,946,967]" class="Insecta" family="Lepiceridae" genus="Lepiceratus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="ankylosaurus" status="sp. nov.">
<emphasis box="[171,429,946,967]" italics="true" pageId="7" pageNumber="8">Lepiceratus ankylosaurus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[449,540,946,967]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
;
<taxonomicName authorityName="Jaloszynski &amp; Yamamoto" authorityYear="2017" box="[563,741,946,967]" class="Insecta" family="Curculionidae" genus="Lepicerus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="mumia">
<emphasis box="[563,741,946,967]" italics="true" pageId="7" pageNumber="8">Lepicerus mumia</emphasis>
</taxonomicName>
;
<taxonomicName authorityName="Hinton" authorityYear="1934" box="[767,922,946,967]" class="Insecta" family="Curculionidae" genus="Lepicerus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="bufo">
<emphasis box="[767,922,946,967]" italics="true" pageId="7" pageNumber="8">Lepicerus bufo</emphasis>
</taxonomicName>
;
<taxonomicName authorityName="Waltl" authorityYear="1838" box="[949,1157,946,967]" class="Insecta" family="Sphaeriusidae" genus="Sphaerius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="acaroides">
<emphasis box="[949,1157,946,967]" italics="true" pageId="7" pageNumber="8">Sphaerius acaroides</emphasis>
</taxonomicName>
;
<taxonomicName box="[1183,1405,946,967]" class="Insecta" family="Lepiceridae" genus="Hydroscapaha" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="natans">
<emphasis box="[1183,1405,946,967]" italics="true" pageId="7" pageNumber="8">Hydroscapaha natans</emphasis>
</taxonomicName>
; and
<taxonomicName box="[159,290,973,994]" class="Insecta" family="Torridincolidae" genus="Satonius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="stysi">
<emphasis box="[159,290,973,994]" italics="true" pageId="7" pageNumber="8">Satonius stysi</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph blockId="7.[159,781,1044,1493]" pageId="7" pageNumber="8">
Apart from the unique characters listed in the diagnosis, the sole known species of †
<taxonomicName authorityName="Jaloszynski, Luo, Yamamoto &amp;" authorityYear="2020" box="[514,640,1240,1263]" class="Insecta" family="Lepiceridae" genus="Lepiceratus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis box="[514,640,1240,1263]" italics="true" pageId="7" pageNumber="8">Lepiceratus</emphasis>
</taxonomicName>
differs from all extant and extinct
<taxonomicName authorityName="Hinton" authorityYear="1936" box="[408,534,1273,1296]" class="Insecta" family="Lepiceridae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="family">Lepiceridae</taxonomicName>
by rows of distinctly separated and very long tubercles, well-defined, especially on the posterior half of the elytra; this differs from other described species of the family, where the elytra have either continuous costae or rows of weakly elevated tubercles fused one to another and forming elongate bulges.
</paragraph>
</subSubSection>
</treatment>
</document>
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