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<document id="7F0D931F138AD66DC9698425B475C937" ID-CLB-Dataset="4270" ID-DOI="10.18590/euscorpius.2016.vol2016.iss215.1" ID-GBIF-Dataset="1b8f4b62-3bc6-4c83-912c-dc6ba2343866" ID-ISSN="1536-9307" ID-Zenodo-Dep="5510002" ID-ZooBank="85BF1181-214F-4471-B504-9A37A075145C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1617134602639" checkinUser="carolina" docAuthor="Mirza, Zeeshan A., Sanap, Rajesh V. &amp; Kunte, Krushnamegh" docDate="2016" docId="03E7B92AF409FFAAFED86503F7EAFA01" docLanguage="en" docName="Euscorpius.215.3-11.pdf" docOrigin="Euscorpius 215" docStyle="DocumentStyle:05AD4CDCECA6C90C60787CEF3EC4672A.3:Euscorpius.2002-2019.journal_article" docStyleId="05AD4CDCECA6C90C60787CEF3EC4672A" docStyleName="Euscorpius.2002-2019.journal_article" docStyleVersion="3" docTitle="Thaicharmus guptai Mirza &amp; Sanap &amp; Kunte 2016, sp. nov." docType="treatment" docUuid="683BBC4A-4711-48EB-998F-3E51C589D6BF" docUuidSource="ZooBank" docVersion="8" lastPageNumber="7" masterDocId="FFDEC152F40DFFACFFB26368F335FFDA" masterDocTitle="A new species of the genus Thaicharmus Kovařík, 1995 (Scorpiones: Buthidae) from northeast India" masterLastPageNumber="11" masterPageNumber="3" pageNumber="5" updateTime="1698935400404" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
<mods:mods id="5D73ADAEE4A8D49F5B74E9BAEA2073F6" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="268DBDAADBE6039FD200A1550F7342F3">
<mods:title id="1777CAD019D9C089BAA10DF88B5A9299">A new species of the genus Thaicharmus Kovařík, 1995 (Scorpiones: Buthidae) from northeast India</mods:title>
</mods:titleInfo>
<mods:name id="32C096E838E2E9EC3A0724113B8E5E66" type="personal">
<mods:role id="FC83951F7D1640A03FC67A8A353600E4">
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<mods:namePart id="5A0F14130755591F8B31803FCDDC6F45">Mirza, Zeeshan A.</mods:namePart>
</mods:name>
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<mods:role id="C1F1997617EF2AF79DE260C70A6B3236">
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<mods:namePart id="E51AD3768700E83101BB799CD84F8E5C">Sanap, Rajesh V.</mods:namePart>
</mods:name>
<mods:name id="82ABDCB3712AA58EC32362C68A12F380" type="personal">
<mods:role id="D603BE564893EA9BA0A824BC10B3872A">
<mods:roleTerm id="5D710006F5D6CA0552A66871ABDB3DE3">Author</mods:roleTerm>
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<mods:namePart id="07D3A6C07DAD05B0EFB490582300D905">Kunte, Krushnamegh</mods:namePart>
</mods:name>
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<mods:title id="E9F77F77363AC695E0439CD3EACDABB1">Euscorpius</mods:title>
</mods:titleInfo>
<mods:part id="A25E23A05B7F077CF2EA173B8CB5353C">
<mods:date id="12202D2E7B4746D0E2C2A85634026293">2016</mods:date>
<mods:detail id="3B3846985CE8FD7675DAF722B15167DD" type="pubDate">
<mods:number id="620F5C6A29E6E73CC9334925E51B138A">2016-12-31</mods:number>
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<mods:detail id="C58938324D7D35BA59F21F85AE4588C7" type="volume">
<mods:number id="AE103B62352E18E75F217C51F01368F9">215</mods:number>
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<mods:start id="B8BAEC2DFDB086AD78D419BD78D3ACE8">3</mods:start>
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<mods:classification id="A701579A8F43C1FF761687F62D3F6FC9">journal article</mods:classification>
<mods:identifier id="656EE67650FF9222F2AA2951C4D54C3B" type="CLB-Dataset">4270</mods:identifier>
<mods:identifier id="D8F9989F34ADF2F440EE6F08FC5BA2BF" type="DOI">10.18590/euscorpius.2016.vol2016.iss215.1</mods:identifier>
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<mods:identifier id="BC50717A7E18E493AD49EB1C19790E63" type="ISSN">1536-9307</mods:identifier>
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<treatment id="03E7B92AF409FFAAFED86503F7EAFA01" ID-DOI="http://doi.org/10.5281/zenodo.5510010" ID-GBIF-Taxon="187974113" ID-Zenodo-Dep="5510010" LSID="urn:lsid:zoobank.org:act:683BBC4A-4711-48EB-998F-3E51C589D6BF" httpUri="http://treatment.plazi.org/id/03E7B92AF409FFAAFED86503F7EAFA01" lastPageId="6" lastPageNumber="7" pageId="4" pageNumber="5">
<subSubSection id="C3545BB7F409FFA8FED86503F19DF959" box="[362,680,1643,1667]" pageId="4" pageNumber="5" type="nomenclature">
<paragraph id="8BF1083CF409FFA8FED86503F19DF959" blockId="4.[362,680,1643,1667]" box="[362,680,1643,1667]" pageId="4" pageNumber="5">
<heading id="D0B9BF50F409FFA8FED86503F19DF959" box="[362,680,1643,1667]" centered="true" fontSize="11" level="2" pageId="4" pageNumber="5" reason="3">
<emphasis id="B93AD42EF409FFA8FED86503F19DF959" bold="true" box="[362,680,1643,1667]" pageId="4" pageNumber="5">
<taxonomicName id="4C4E73BFF409FFA8FED86503F17EF959" ID-CoL="55YSW" authority="Mirza &amp; Sanap &amp; Kunte, 2016" authorityName="Mirza &amp; Sanap &amp; Kunte" authorityYear="2016" box="[362,587,1643,1667]" class="Arachnida" family="Buthidae" genus="Thaicharmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="guptai" status="sp. nov.">
<emphasis id="B93AD42EF409FFA8FED86503F17EF959" bold="true" box="[362,587,1643,1667]" italics="true" pageId="4" pageNumber="5">Thaicharmus guptai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2096955F409FFA8FDE36503F19DF959" box="[593,680,1643,1667]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3545BB7F409FFA8FE2D65E1F1F0F8DB" pageId="4" pageNumber="5" type="description">
<paragraph id="8BF1083CF409FFA8FE2D65E1F146F97B" blockId="4.[415,627,1673,1697]" box="[415,627,1673,1697]" pageId="4" pageNumber="5">
(
<figureCitation id="137514B9F409FFA8FE1565E1F138F97B" box="[423,525,1673,1697]" captionStart-0="Figure 2" captionStart-1="Figure 3" captionStart-2="Figure 4" captionStart-3="Figure 5" captionStart-4="Figure 6" captionStartId-0="3.[192,267,1756,1780]" captionStartId-1="5.[192,267,1428,1452]" captionStartId-2="6.[192,267,1342,1366]" captionStartId-3="7.[192,267,1823,1847]" captionStartId-4="8.[192,267,843,867]" captionTargetBox-0="[191,1440,935,1722]" captionTargetBox-1="[199,1428,234,1390]" captionTargetBox-2="[199,1436,199,1301]" captionTargetBox-3="[208,1413,156,1776]" captionTargetBox-4="[199,1432,197,807]" captionTargetId-0="figure-87@3.[192,1439,936,1721]" captionTargetId-1="figure-265@5.[192,1439,229,1399]" captionTargetId-2="figure-284@6.[192,1439,192,1309]" captionTargetId-3="figure-55@7.[192,1438,144,1785]" captionTargetId-4="figure-64@8.[192,1439,191,816]" captionTargetPageId-0="3" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="7" captionTargetPageId-4="8" captionText-0="Figure 2: Thaicharmus guptai sp. nov. holotype male NCBS-AG800, live coloration." captionText-1="Figure 3: Thaicharmus guptai sp. nov. holotype male NCBS-AG800,, habitus (a) dorsal, (b) ventral." captionText-2="Figure 4: Thaicharmus guptai sp. nov. holotype male NCBS-AG800, (a) carapace, (b) fixed finger of chelicerae, ventral view, (c) movable finger of chelicerae, ventral view, (d) right pectine, (e) sternum, genital operculum and pectines." captionText-3="Figure 5: Thaicharmus guptai sp. nov. holotype male NCBS-AG800, right pedipalp. (a) femur, dorsal view, (b) patella dorsal view, (c) patella external view, (d) movable finger, (e) manus dorsal view, (f) manus external view, (g) manus ventral view. Visible trichobothria indicated with green circles." captionText-4="Figure 6: Thaicharmus guptai sp. nov. holotype male NCBS-AG800, metasoma (a) ventral view, (b) lateral view, (c), dorsal view. Not to scale." figureDoi-0="http://doi.org/10.5281/zenodo.5510008" figureDoi-1="http://doi.org/10.5281/zenodo.5510012" figureDoi-2="http://doi.org/10.5281/zenodo.5510014" figureDoi-3="http://doi.org/10.5281/zenodo.5510016" figureDoi-4="http://doi.org/10.5281/zenodo.5510018" httpUri-0="https://zenodo.org/record/5510008/files/figure.png" httpUri-1="https://zenodo.org/record/5510012/files/figure.png" httpUri-2="https://zenodo.org/record/5510014/files/figure.png" httpUri-3="https://zenodo.org/record/5510016/files/figure.png" httpUri-4="https://zenodo.org/record/5510018/files/figure.png" pageId="4" pageNumber="5">Figs. 2–6</figureCitation>
,
<tableCitation id="C6CC3D87F409FFA8FDAB65E1F15DF97B" box="[537,616,1673,1697]" captionStart="Table 1" captionStartId="4.[192,257,1334,1358]" captionTargetBox="[258,732,216,1275]" captionTargetId="graphics-16@4.[243,751,207,1277]" captionText="Table 1: Morphometric values (in mm) of Thaicharmus guptai sp. nov." httpUri="http://table.plazi.org/id/DF3158B4F409FFA8FF72665EF26AFAB3" pageId="4" pageNumber="5" tableUuid="DF3158B4F409FFA8FF72665EF26AFAB3">Table 1</tableCitation>
)
</paragraph>
<paragraph id="8BF1083CF409FFA8FF7365A2F015F938" blockId="4.[193,800,1738,1793]" box="[193,800,1738,1762]" pageId="4" pageNumber="5">
<emphasis id="B93AD42EF409FFA8FF7365A2F015F938" bold="true" box="[193,800,1738,1762]" pageId="4" pageNumber="5">http://zoobank.org/urn:lsid:zoobank.org:act:683BBC</emphasis>
</paragraph>
<paragraph id="8BF1083CF409FFA8FEAF6581F1F0F8DB" blockId="4.[193,800,1738,1793]" box="[285,709,1769,1793]" pageId="4" pageNumber="5">
<emphasis id="B93AD42EF409FFA8FEAF6581F1F0F8DB" bold="true" box="[285,709,1769,1793]" pageId="4" pageNumber="5">4A-4711-48EB-998F-3E51C589D6BF</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C3545BB7F409FFA8FF426444F7D5FF26" pageId="4" pageNumber="5" type="materials_examined">
<paragraph id="8BF1083CF409FFA8FF426444F7D5FF26" blockId="4.[192,802,1836,1890]" lastBlockId="4.[831,1440,198,252]" pageId="4" pageNumber="5">
<materialsCitation id="3B260261F409FFA8FF426444F017F8B8" ID-GBIF-Occurrence="3352115302" country="India" county="North" location="Phuldungsei" municipality="Jampui Hills" pageId="4" pageNumber="5" specimenCount="1" specimenCount-male="1" stateProvince="Tripura" typeStatus="holotype">
<emphasis id="B93AD42EF409FFA8FF426444F26DF89E" bold="true" box="[240,344,1836,1860]" pageId="4" pageNumber="5">
<typeStatus id="54F5B69EF409FFA8FF426444F26DF89E" box="[240,344,1836,1860]" pageId="4" pageNumber="5" type="holotype">Holotype</typeStatus>
</emphasis>
<specimenCount id="9D48C3B5F409FFA8FED26444F241F899" box="[352,372,1836,1859]" pageId="4" pageNumber="5" type="male">♂</specimenCount>
NCBS- AG800, near
<location id="8E915EE7F409FFA8FDEE6444F1D7F89E" LSID="urn:lsid:plazi:treatment:03E7B92AF409FFAAFED86503F7EAFA01:8E915EE7F409FFA8FDEE6444F1D7F89E" box="[604,738,1836,1860]" country="India" county="North" municipality="Jampui Hills" name="Phuldungsei" pageId="4" pageNumber="5" stateProvince="Tripura">Phuldungsei</location>
,
<collectingMunicipality id="6B959246F409FFA8FD5C6444F210F8B8" pageId="4" pageNumber="5">Jampui Hills</collectingMunicipality>
,
<collectingCounty id="629070B0F409FFA8FE856422F240F8B8" box="[311,373,1866,1890]" pageId="4" pageNumber="5">North</collectingCounty>
<collectingRegion id="498AC6DEF409FFA8FE306422F107F8B8" box="[386,562,1866,1890]" country="India" name="Tripura" pageId="4" pageNumber="5">Tripura District</collectingRegion>
,
<collectingRegion id="498AC6DEF409FFA8FDF66422F1ECF8B8" box="[580,729,1866,1890]" country="India" name="Tripura" pageId="4" pageNumber="5">Tripura State</collectingRegion>
,
<collectingCountry id="F35948ACF409FFA8FD596422F017F8B8" box="[747,802,1866,1890]" name="India" pageId="4" pageNumber="5">India</collectingCountry>
</materialsCitation>
(
<geoCoordinate id="EE7A6EFBF409FFA8FCF463AEF0E4FF04" box="[838,977,198,222]" degrees="23" direction="north" minutes="48.704" orientation="latitude" pageId="4" pageNumber="5" precision="1" value="23.811733">23°48.704'N</geoCoordinate>
,
<geoCoordinate id="EE7A6EFBF409FFA8FC6D63AEF747FF04" box="[991,1138,198,222]" degrees="092" direction="east" minutes="15.667" orientation="longitude" pageId="4" pageNumber="5" precision="1" value="92.261116">092°15.667'E</geoCoordinate>
, elevation
<quantity id="4CB6A5D9F409FFA8FB5F63AEF60DFF04" box="[1261,1336,198,222]" metricMagnitude="2" metricUnit="m" metricValue="9.2" pageId="4" pageNumber="5" unit="m" value="920.0">920 m</quantity>
),
<materialsCitation id="3B260261F409FFA8FAFC63AEF7E9FF26" ID-GBIF-Occurrence="3352115301" collectingDate="2014-11-01" collectionCode="R" collectorName="Sanap &amp; Z. Mirza" pageId="4" pageNumber="5" specimenCount="1">
coll.
<collectionCode id="ED5F90F9F409FFA8FA3563AEF6A9FF07" box="[1415,1436,198,221]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="4" pageNumber="5">R</collectionCode>
.
<collectorName id="26BB6DEAF409FFA8FC8D638CF0B5FF26" box="[831,896,228,252]" pageId="4" pageNumber="5">Sanap</collectorName>
&amp;
<collectorName id="26BB6DEAF409FFA8FC10638DF734FF26" box="[930,1025,228,252]" pageId="4" pageNumber="5">Z. Mirza</collectorName>
,
<date id="FFF02EFCF409FFA8FBBE638CF7E9FF26" box="[1036,1244,227,252]" pageId="4" pageNumber="5" value="2014-11-01">
<collectingDate id="EFB4D714F409FFA8FBBE638CF7E9FF26" box="[1036,1244,227,252]" pageId="4" pageNumber="5" value="2014-11-01">
1
<superScript id="7C3BA574F409FFA8FBA8638BF713FF2B" attach="left" box="[1050,1062,227,241]" fontSize="6" pageId="4" pageNumber="5">st</superScript>
November 2014
</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3545BB7F409FFA8FCDD624DF08FFE0C" pageId="4" pageNumber="5" type="etymology">
<paragraph id="8BF1083CF409FFA8FCDD624DF08FFE0C" blockId="4.[831,1440,293,470]" pageId="4" pageNumber="5">
<emphasis id="B93AD42EF409FFA8FCDD624DF0C7FEE7" bold="true" box="[879,1010,293,317]" pageId="4" pageNumber="5">Etymology:</emphasis>
The specific epithet is a patronym honoring Atul Gupta, IFS, Principal Chief Conservator of Forest and the Director of
<collectingRegion id="498AC6DEF409FFA8FBC5620AF7F2FEA0" box="[1143,1223,354,378]" country="India" name="Tripura" pageId="4" pageNumber="5">Tripura</collectingRegion>
Biodiversity Board, for initiating and supporting the
<collectingRegion id="498AC6DEF409FFA8FB7362E9F624FE43" box="[1217,1297,385,409]" country="India" name="Tripura" pageId="4" pageNumber="5">Tripura</collectingRegion>
biodiversity documentation project during which the new species was discovered.
</paragraph>
</subSubSection>
<subSubSection id="C3545BB7F409FFA8FCDD6297F6AAFC1E" pageId="4" pageNumber="5" type="diagnosis">
<paragraph id="8BF1083CF409FFA8FCDD6297F6AAFC1E" blockId="4.[831,1441,510,964]" pageId="4" pageNumber="5">
<emphasis id="B93AD42EF409FFA8FCDD6297F0D0FDCD" bold="true" box="[879,997,511,535]" pageId="4" pageNumber="5">Diagnosis:</emphasis>
<taxonomicName id="4C4E73BFF409FFA8FC446296F7EDFDCC" authority="Mirza &amp; Sanap &amp; Kunte, 2016" authorityName="Mirza &amp; Sanap &amp; Kunte" authorityYear="2016" box="[1014,1240,510,534]" class="Arachnida" family="Buthidae" genus="Thaicharmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="guptai" status="sp. nov.">
<emphasis id="B93AD42EF409FFA8FC446296F7EDFDCC" box="[1014,1240,510,534]" italics="true" pageId="4" pageNumber="5">Thaicharmus guptai</emphasis>
</taxonomicName>
<emphasis id="B93AD42EF409FFA8FB5B6297F67FFDCD" bold="true" box="[1257,1354,511,535]" pageId="4" pageNumber="5">
<taxonomicNameLabel id="A2096955F409FFA8FB5B6297F67FFDCD" box="[1257,1354,511,535]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from all known members of the genus
<taxonomicName id="4C4E73BFF409FFA8FB466175F6B4FDEF" authorityName="Kovarik" authorityYear="1995" box="[1268,1409,541,565]" class="Arachnida" family="Buthidae" genus="Thaicharmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B93AD42EF409FFA8FB466175F6B4FDEF" box="[1268,1409,541,565]" italics="true" pageId="4" pageNumber="5">Thaicharmus</emphasis>
</taxonomicName>
in bearing the following set of morphological characters: trichobothrium
<emphasis id="B93AD42EF409FFA8FC556132F0C8FDA9" box="[999,1021,602,627]" italics="true" pageId="4" pageNumber="5">
d
<subScript id="17CA0A79F409FFA8FC46610DF0C8FDA9" attach="left" box="[1012,1021,613,627]" fontSize="6" pageId="4" pageNumber="5">2</subScript>
</emphasis>
distal to
<emphasis id="B93AD42EF409FFA8FBD06133F747FDA9" box="[1122,1138,603,627]" italics="true" pageId="4" pageNumber="5">
i
<subScript id="17CA0A79F409FFA8FBDB610DF747FDA9" attach="left" box="[1129,1138,613,627]" fontSize="6" pageId="4" pageNumber="5">1</subScript>
</emphasis>
on femur; trichobothria
<emphasis id="B93AD42EF409FFA8FACE6133F06BFD4B" italics="true" pageId="4" pageNumber="5">est, em</emphasis>
and
<emphasis id="B93AD42EF409FFA8FC2B6112F099FD4B" box="[921,940,634,657]" italics="true" pageId="4" pageNumber="5">et</emphasis>
on external surface of patella do not form a straight line; subaculear tubercle absent; movable finger of pedipalp chela with 12 cutting rows of denticles; movable finger much longer than pedipalp patella; pectinal teeth number 20/19; pectines with distinct lamellae and fulcra; two horn-like projections on ventrolateral aspect of metasomal segment V. The female is unknown.
<taxonomicName id="4C4E73BFF409FFA8FC256038F747FCB2" authority="Mirza &amp; Sanap &amp; Kunte, 2016" authorityName="Mirza &amp; Sanap &amp; Kunte" authorityYear="2016" box="[919,1138,848,872]" class="Arachnida" family="Buthidae" genus="Thaicharmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="guptai" status="sp. nov.">
<emphasis id="B93AD42EF409FFA8FC256038F747FCB2" box="[919,1138,848,872]" italics="true" pageId="4" pageNumber="5">Thaicharmus guptai</emphasis>
</taxonomicName>
<emphasis id="B93AD42EF409FFA8FBCE6038F7E0FCB2" bold="true" box="[1148,1237,848,872]" pageId="4" pageNumber="5">
<taxonomicNameLabel id="A2096955F409FFA8FBCE6038F7E0FCB2" box="[1148,1237,848,872]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is most similar to
<taxonomicName id="4C4E73BFF409FFA8FC8D6007F0F2FC5C" authorityName="Kovarik" authorityYear="1995" box="[831,967,878,902]" class="Arachnida" family="Buthidae" genus="Thaicharmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="mahunkai">
<emphasis id="B93AD42EF409FFA8FC8D6007F0F2FC5C" box="[831,967,878,902]" italics="true" pageId="4" pageNumber="5">T. mahunkai</emphasis>
</taxonomicName>
in bearing 12 cutting rows of denticles on movable finger of pedipalp chela; however, it differs from the latter species in lacking the subaculear tubercle.
</paragraph>
</subSubSection>
<subSubSection id="C3545BB7F409FFAAFC8D6085F7EAFA01" lastPageId="6" lastPageNumber="7" pageId="4" pageNumber="5" type="description">
<paragraph id="8BF1083CF409FFA8FC8D6085F602FBDD" blockId="4.[831,1335,1005,1032]" box="[831,1335,1005,1032]" pageId="4" pageNumber="5">
<heading id="D0B9BF50F409FFA8FC8D6085F602FBDD" box="[831,1335,1005,1032]" pageId="4" pageNumber="5">
<emphasis id="B93AD42EF409FFA8FC8D6085F756FBDD" bold="true" box="[831,1123,1005,1031]" pageId="4" pageNumber="5">
Description of
<typeStatus id="54F5B69EF409FFA8FC456085F756FBDD" box="[1015,1123,1005,1031]" pageId="4" pageNumber="5" type="holotype">holotype</typeStatus>
</emphasis>
<specimenCount id="9D48C3B5F409FFA8FBD86086F7B5FBD2" box="[1130,1152,1006,1032]" pageId="4" pageNumber="5" type="male">♂</specimenCount>
<emphasis id="B93AD42EF409FFA8FB356085F602FBDD" bold="true" box="[1159,1335,1005,1031]" pageId="4" pageNumber="5">NCBS-AG800</emphasis>
</heading>
</paragraph>
<paragraph id="8BF1083CF409FFA8FCDD6745F0BFFA09" blockId="4.[831,1441,1068,1890]" pageId="4" pageNumber="5">
<emphasis id="B93AD42EF409FFA8FCDD6745F61DFB9F" bold="true" box="[879,1320,1069,1093]" pageId="4" pageNumber="5">
Coloration (live) (
<figureCitation id="137514B9F409FFA8FBF56745F7AFFB9F" box="[1095,1178,1069,1093]" captionStart="Figure 2" captionStartId="3.[192,267,1756,1780]" captionTargetBox="[191,1440,935,1722]" captionTargetId="figure-87@3.[192,1439,936,1721]" captionTargetPageId="3" captionText="Figure 2: Thaicharmus guptai sp. nov. holotype male NCBS-AG800, live coloration." figureDoi="http://doi.org/10.5281/zenodo.5510008" httpUri="https://zenodo.org/record/5510008/files/figure.png" pageId="4" pageNumber="5">Figs. 2</figureCitation>
,
<figureCitation id="137514B9F409FFA8FB1C6745F62FFB9F" box="[1198,1306,1069,1093]" captionStart="Figure 3" captionStartId="5.[192,267,1428,1452]" captionTargetBox="[199,1428,234,1390]" captionTargetId="figure-265@5.[192,1439,229,1399]" captionTargetPageId="5" captionText="Figure 3: Thaicharmus guptai sp. nov. holotype male NCBS-AG800,, habitus (a) dorsal, (b) ventral." figureDoi="http://doi.org/10.5281/zenodo.5510012" httpUri="https://zenodo.org/record/5510012/files/figure.png" pageId="4" pageNumber="5">3a &amp; 3b</figureCitation>
):
</emphasis>
Carapace, mesosoma and metasoma in shade of black; pedipalp in shade of brown, chelicerae light brown with dark reticulation, cheliceral teeth on movable and immovable finger dark brown to black, femur and patella of legs almost black fading to dark brown distally, the brown color from patella further fades to pale yellow towards the tarsus. Pectines, genital operculum and all sternites (except the last one, which is of a darker shade) pale yellowish. Sternum distally dark brown, fading to yellow towards the center and base. Coloration in preserved condition much faded and paler. Metasomal segment fades from black to a dark brown color. Telson dark brown.
</paragraph>
<paragraph id="8BF1083CF409FFA9FCDD66B2F676F8BD" blockId="4.[831,1441,1068,1890]" lastBlockId="5.[831,1441,1503,1895]" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5">
<emphasis id="B93AD42EF409FFA8FCDD66B2F730FA28" bold="true" box="[879,1029,1498,1522]" pageId="4" pageNumber="5">Morphology:</emphasis>
Carapace trapezoidal with sinuous anterior, bearing a beady gloss and sparse granulation, more concentrated on lateral aspect (
<figureCitation id="137514B9F409FFA8FB65657FF61DF9F5" box="[1239,1320,1559,1583]" captionStart="Figure 4" captionStartId="6.[192,267,1342,1366]" captionTargetBox="[199,1436,199,1301]" captionTargetId="figure-284@6.[192,1439,192,1309]" captionTargetPageId="6" captionText="Figure 4: Thaicharmus guptai sp. nov. holotype male NCBS-AG800, (a) carapace, (b) fixed finger of chelicerae, ventral view, (c) movable finger of chelicerae, ventral view, (d) right pectine, (e) sternum, genital operculum and pectines." figureDoi="http://doi.org/10.5281/zenodo.5510014" httpUri="https://zenodo.org/record/5510014/files/figure.png" pageId="4" pageNumber="5">Fig. 4a</figureCitation>
). The eyetubercle slightly elevated, a deep longitudinal furrow from anterior border to the posterior border is present. Median eyes are situated anteriorly in a ratio of 1:3.51. Five lateral eyes are present comprising three large and two small ones. A moderately developed median epistome on its anterior border arises in a shallow concavity (
<figureCitation id="137514B9F409FFA8FCF46586F0ABF8DC" box="[838,926,1774,1798]" captionStart="Figure 4" captionStartId="6.[192,267,1342,1366]" captionTargetBox="[199,1436,199,1301]" captionTargetId="figure-284@6.[192,1439,192,1309]" captionTargetPageId="6" captionText="Figure 4: Thaicharmus guptai sp. nov. holotype male NCBS-AG800, (a) carapace, (b) fixed finger of chelicerae, ventral view, (c) movable finger of chelicerae, ventral view, (d) right pectine, (e) sternum, genital operculum and pectines." figureDoi="http://doi.org/10.5281/zenodo.5510014" httpUri="https://zenodo.org/record/5510014/files/figure.png" pageId="4" pageNumber="5">Fig. 4a</figureCitation>
). Carapace with only median ocular carina slightly elevated whereas others are absent. Chelicerae typical of family
<taxonomicName id="4C4E73BFF409FFA8FC486443F633F899" authority="(Sissom, 1990)" baseAuthorityName="Sissom" baseAuthorityYear="1990" box="[1018,1286,1835,1859]" class="Arachnida" family="Buthidae" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="5" phylum="Arthropoda" rank="family">
Buthidae (
<bibRefCitation id="EFDF75CDF409FFA8FBD86443F7CBF899" author="SISSOM" box="[1130,1278,1835,1859]" pageId="4" pageNumber="5" pagination="64 - 160" refId="ref3168" refString="SISSOM, W. D. 1990. Systematics, biogeography and paleontology. Pp. 64 - 160 in: Polis, G. A. (ed.), The Biology of Scorpions. Stanford: Stanford University Press." type="book chapter" year="1990">Sissom, 1990</bibRefCitation>
)
</taxonomicName>
(
<figureCitation id="137514B9F409FFA8FAA46443F654F899" box="[1302,1377,1835,1859]" captionStart="Figure 4" captionStartId="6.[192,267,1342,1366]" captionTargetBox="[199,1436,199,1301]" captionTargetId="figure-284@6.[192,1439,192,1309]" captionTargetPageId="6" captionText="Figure 4: Thaicharmus guptai sp. nov. holotype male NCBS-AG800, (a) carapace, (b) fixed finger of chelicerae, ventral view, (c) movable finger of chelicerae, ventral view, (d) right pectine, (e) sternum, genital operculum and pectines." figureDoi="http://doi.org/10.5281/zenodo.5510014" httpUri="https://zenodo.org/record/5510014/files/figure.png" pageId="4" pageNumber="5">Figs. 4</figureCitation>
b-4c). Mesosoma: Tergites I–VI smooth, lustrous throughout, lacking granulation with a single slightly elevated median smooth carinae (
<figureCitation id="137514B9F408FFA9FE676696F11DF9CC" box="[469,552,1534,1558]" captionStart="Figure 3" captionStartId="5.[192,267,1428,1452]" captionTargetBox="[199,1428,234,1390]" captionTargetId="figure-265@5.[192,1439,229,1399]" captionTargetPageId="5" captionText="Figure 3: Thaicharmus guptai sp. nov. holotype male NCBS-AG800,, habitus (a) dorsal, (b) ventral." figureDoi="http://doi.org/10.5281/zenodo.5510012" httpUri="https://zenodo.org/record/5510012/files/figure.png" pageId="5" pageNumber="6">Fig. 3a</figureCitation>
). Tergite VII has two pairs of granular carinae (
<figureCitation id="137514B9F408FFA9FE6A6575F11FF9EF" box="[472,554,1565,1589]" captionStart="Figure 3" captionStartId="5.[192,267,1428,1452]" captionTargetBox="[199,1428,234,1390]" captionTargetId="figure-265@5.[192,1439,229,1399]" captionTargetPageId="5" captionText="Figure 3: Thaicharmus guptai sp. nov. holotype male NCBS-AG800,, habitus (a) dorsal, (b) ventral." figureDoi="http://doi.org/10.5281/zenodo.5510012" httpUri="https://zenodo.org/record/5510012/files/figure.png" pageId="5" pageNumber="6">Fig. 3b</figureCitation>
). Granular throughout. Sternites smooth throughout with a few sparse punctuations, densely concentrated in the last segment. Stigmata are slit-like. The sternum is pentagonal with a distinct longitudinal median furrow. Pectinal teeth 20/
<quantity id="4CB6A5D9F408FFA9FCBA65FFF3E1F914" metricMagnitude="-1" metricUnit="m" metricValue="4.826" pageId="5" pageNumber="6" unit="in" value="19.0">19 in</quantity>
number. Pectines are well-developed, with nine lamellae and with fulcra at base of each tooth excluding the first and last pectinal teeth (
<figureCitation id="137514B9F408FFA9FDAC659BF15EF8D1" box="[542,619,1779,1803]" captionStart="Figure 4" captionStartId="6.[192,267,1342,1366]" captionTargetBox="[199,1436,199,1301]" captionTargetId="figure-284@6.[192,1439,192,1309]" captionTargetPageId="6" captionText="Figure 4: Thaicharmus guptai sp. nov. holotype male NCBS-AG800, (a) carapace, (b) fixed finger of chelicerae, ventral view, (c) movable finger of chelicerae, ventral view, (d) right pectine, (e) sternum, genital operculum and pectines." figureDoi="http://doi.org/10.5281/zenodo.5510014" httpUri="https://zenodo.org/record/5510014/files/figure.png" pageId="5" pageNumber="6">Figs. 4</figureCitation>
d-4e). Metasoma (
<figureCitation id="137514B9F408FFA9FF7A647AF224F8F0" box="[200,273,1810,1834]" captionStart="Figure 6" captionStartId="8.[192,267,843,867]" captionTargetBox="[199,1432,197,807]" captionTargetId="figure-64@8.[192,1439,191,816]" captionTargetPageId="8" captionText="Figure 6: Thaicharmus guptai sp. nov. holotype male NCBS-AG800, metasoma (a) ventral view, (b) lateral view, (c), dorsal view. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5510018" httpUri="https://zenodo.org/record/5510018/files/figure.png" pageId="5" pageNumber="6">Fig. 6</figureCitation>
): All segments robust, densely punctuated on their ventral aspect and also hirsute. Segment V has two distinct horn-like projections on its ventro-lateral aspect. Vesicle bulbous, lacking a distinct aculear tubercle. Pedipalp: Smooth and lustrous throughout, but densely granular between crenulated dorsal and internal carinae of femur. Patella lacks distinct carinae, with only a few granules on its internal aspect. Movable finger of pedipalp chela with 12 cutting rows of denticles (
<figureCitation id="137514B9F408FFA9FAF26511F6A6F94B" box="[1344,1427,1657,1681]" captionStart="Figure 5" captionStartId="7.[192,267,1823,1847]" captionTargetBox="[208,1413,156,1776]" captionTargetId="figure-55@7.[192,1438,144,1785]" captionTargetPageId="7" captionText="Figure 5: Thaicharmus guptai sp. nov. holotype male NCBS-AG800, right pedipalp. (a) femur, dorsal view, (b) patella dorsal view, (c) patella external view, (d) movable finger, (e) manus dorsal view, (f) manus external view, (g) manus ventral view. Visible trichobothria indicated with green circles." figureDoi="http://doi.org/10.5281/zenodo.5510016" httpUri="https://zenodo.org/record/5510016/files/figure.png" pageId="5" pageNumber="6">Fig. 5d</figureCitation>
). The movable finger is much longer than pedipalp patella. Chela lacking carinae. All segments lacking setae. Dorsal trichobothria on femur in
<emphasis id="B93AD42EF408FFA9FA8C65BDF652F937" box="[1342,1383,1749,1773]" italics="true" pageId="5" pageNumber="6">alfa</emphasis>
-configuration (
<figureCitation id="137514B9F408FFA9FC0A659BF73DF8D1" box="[952,1032,1779,1803]" captionStart="Figure 5" captionStartId="7.[192,267,1823,1847]" captionTargetBox="[208,1413,156,1776]" captionTargetId="figure-55@7.[192,1438,144,1785]" captionTargetPageId="7" captionText="Figure 5: Thaicharmus guptai sp. nov. holotype male NCBS-AG800, right pedipalp. (a) femur, dorsal view, (b) patella dorsal view, (c) patella external view, (d) movable finger, (e) manus dorsal view, (f) manus external view, (g) manus ventral view. Visible trichobothria indicated with green circles." figureDoi="http://doi.org/10.5281/zenodo.5510016" httpUri="https://zenodo.org/record/5510016/files/figure.png" pageId="5" pageNumber="6">Fig. 5a</figureCitation>
). Trichobothrium
<emphasis id="B93AD42EF408FFA9FB7D659CF7D8F8D1" box="[1231,1261,1780,1803]" italics="true" pageId="5" pageNumber="6">est</emphasis>
basal to
<emphasis id="B93AD42EF408FFA9FAE0659CF644F8D1" box="[1362,1393,1780,1803]" italics="true" pageId="5" pageNumber="6">em</emphasis>
and
<emphasis id="B93AD42EF408FFA9FC8D647BF067F8F0" box="[831,850,1811,1834]" italics="true" pageId="5" pageNumber="6">et</emphasis>
on the external surface of patella.
<emphasis id="B93AD42EF408FFA9FB47647BF622F8F0" box="[1269,1303,1811,1834]" italics="true" pageId="5" pageNumber="6">Est</emphasis>
,
<emphasis id="B93AD42EF408FFA9FA9E647BF67EF8F0" box="[1324,1355,1811,1834]" italics="true" pageId="5" pageNumber="6">em</emphasis>
and
<emphasis id="B93AD42EF408FFA9FA3F647BF695F8F0" box="[1421,1440,1811,1834]" italics="true" pageId="5" pageNumber="6">et</emphasis>
aligned angularly, not forming a straight line (
<figureCitation id="137514B9F408FFA9FAF36458F6A6F892" box="[1345,1427,1840,1864]" captionStart="Figure 5" captionStartId="7.[192,267,1823,1847]" captionTargetBox="[208,1413,156,1776]" captionTargetId="figure-55@7.[192,1438,144,1785]" captionTargetPageId="7" captionText="Figure 5: Thaicharmus guptai sp. nov. holotype male NCBS-AG800, right pedipalp. (a) femur, dorsal view, (b) patella dorsal view, (c) patella external view, (d) movable finger, (e) manus dorsal view, (f) manus external view, (g) manus ventral view. Visible trichobothria indicated with green circles." figureDoi="http://doi.org/10.5281/zenodo.5510016" httpUri="https://zenodo.org/record/5510016/files/figure.png" pageId="5" pageNumber="6">Fig. 5c</figureCitation>
). Trichobothrium
<emphasis id="B93AD42EF408FFA9FC5C6427F731F8BD" box="[1006,1028,1871,1895]" italics="true" pageId="5" pageNumber="6">
d
<subScript id="17CA0A79F408FFA9FC496431F731F8BD" attach="left" box="[1019,1028,1881,1895]" fontSize="6" pageId="5" pageNumber="6">2</subScript>
</emphasis>
distal to
<emphasis id="B93AD42EF408FFA9FBD66438F740F8BD" box="[1124,1141,1872,1895]" italics="true" pageId="5" pageNumber="6">
i
<subScript id="17CA0A79F408FFA9FBDE6431F740F8BD" attach="left" box="[1132,1141,1881,1895]" fontSize="6" pageId="5" pageNumber="6">1</subScript>
</emphasis>
on femur (
<figureCitation id="137514B9F408FFA9FB5A6427F603F8BD" box="[1256,1334,1871,1895]" captionStart="Figure 5" captionStartId="7.[192,267,1823,1847]" captionTargetBox="[208,1413,156,1776]" captionTargetId="figure-55@7.[192,1438,144,1785]" captionTargetPageId="7" captionText="Figure 5: Thaicharmus guptai sp. nov. holotype male NCBS-AG800, right pedipalp. (a) femur, dorsal view, (b) patella dorsal view, (c) patella external view, (d) movable finger, (e) manus dorsal view, (f) manus external view, (g) manus ventral view. Visible trichobothria indicated with green circles." figureDoi="http://doi.org/10.5281/zenodo.5510016" httpUri="https://zenodo.org/record/5510016/files/figure.png" pageId="5" pageNumber="6">Fig. 5a</figureCitation>
).
</paragraph>
<caption id="DF3158B4F408FFA9FF7266FCF7A2FA71" ID-DOI="http://doi.org/10.5281/zenodo.5510012" ID-Zenodo-Dep="5510012" box="[192,1175,1428,1452]" httpUri="https://zenodo.org/record/5510012/files/figure.png" pageId="5" pageNumber="6" startId="5.[192,267,1428,1452]" targetBox="[199,1428,234,1390]" targetPageId="5">
<paragraph id="8BF1083CF408FFA9FF7266FCF7A2FA71" blockId="5.[192,1175,1428,1452]" box="[192,1175,1428,1452]" pageId="5" pageNumber="6">
<emphasis id="B93AD42EF408FFA9FF7266FCF21DFA76" bold="true" box="[192,296,1428,1452]" pageId="5" pageNumber="6">Figure 3:</emphasis>
<taxonomicName id="4C4E73BFF408FFA9FE9D66FDF2C5FA71" authority="Mirza &amp; Sanap &amp; Kunte, 2016" authorityName="Mirza &amp; Sanap &amp; Kunte" authorityYear="2016" box="[303,496,1429,1451]" class="Arachnida" family="Buthidae" genus="Thaicharmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="guptai" status="sp. nov.">
<emphasis id="B93AD42EF408FFA9FE9D66FDF2C5FA71" box="[303,496,1429,1451]" italics="true" pageId="5" pageNumber="6">Thaicharmus guptai</emphasis>
</taxonomicName>
<emphasis id="B93AD42EF408FFA9FE4566FDF171FA70" bold="true" box="[503,580,1429,1450]" pageId="5" pageNumber="6">
<taxonomicNameLabel id="A2096955F408FFA9FE4566FDF171FA70" box="[503,580,1429,1450]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
holotype male NCBS-AG800,, habitus
<emphasis id="B93AD42EF408FFA9FC7666FDF0F9FA70" bold="true" box="[964,972,1429,1450]" pageId="5" pageNumber="6">(</emphasis>
a) dorsal, (b) ventral.
</paragraph>
</caption>
<caption id="DF3158B4F40BFFAAFF726656F7E0FAAB" ID-DOI="http://doi.org/10.5281/zenodo.5510014" ID-Zenodo-Dep="5510014" httpUri="https://zenodo.org/record/5510014/files/figure.png" pageId="6" pageNumber="7" startId="6.[192,267,1342,1366]" targetBox="[199,1436,199,1301]" targetPageId="6">
<paragraph id="8BF1083CF40BFFAAFF726656F7E0FAAB" blockId="6.[192,1440,1342,1393]" pageId="6" pageNumber="7">
<emphasis id="B93AD42EF40BFFAAFF726656F21CFA8C" bold="true" box="[192,297,1342,1366]" pageId="6" pageNumber="7">Figure 4:</emphasis>
<taxonomicName id="4C4E73BFF40BFFAAFE836657F2C6FA8F" authority="Mirza &amp; Sanap &amp; Kunte, 2016" authorityName="Mirza &amp; Sanap &amp; Kunte" authorityYear="2016" box="[305,499,1343,1365]" class="Arachnida" family="Buthidae" genus="Thaicharmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="guptai" status="sp. nov.">
<emphasis id="B93AD42EF40BFFAAFE836657F2C6FA8F" box="[305,499,1343,1365]" italics="true" pageId="6" pageNumber="7">Thaicharmus guptai</emphasis>
</taxonomicName>
<emphasis id="B93AD42EF40BFFAAFE496657F17CFA8E" bold="true" box="[507,585,1343,1364]" pageId="6" pageNumber="7">
<taxonomicNameLabel id="A2096955F40BFFAAFE496657F17CFA8E" box="[507,585,1343,1364]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
holotype male NCBS-AG800, (a) carapace, (b) fixed finger of chelicerae, ventral view, (c) movable finger of chelicerae, ventral view, (d) right pectine, (e) sternum, genital operculum and pectines.
</paragraph>
</caption>
<paragraph id="8BF1083CF40BFFAAFF4266CDF7EAFA01" blockId="6.[192,802,1445,1868]" lastBlockId="6.[831,1439,1445,1500]" pageId="6" pageNumber="7">
<emphasis id="B93AD42EF40BFFAAFF4266CDF14BFA67" bold="true" box="[240,638,1445,1469]" pageId="6" pageNumber="7">Natural history and distribution:</emphasis>
The
<typeStatus id="54F5B69EF40BFFAAFD7766CDF017FA67" box="[709,802,1445,1469]" pageId="6" pageNumber="7" type="holotype">holotype</typeStatus>
male was found actively moving on a rocky escarpment in degraded evergreen forest along a stream. The type locality is Jampui Hills, on the border of Tripura and Mizoram States (
<figureCitation id="137514B9F40BFFAAFECE6577F2F5F9ED" box="[380,448,1567,1591]" captionStart="Figure 7" captionStartId="8.[192,267,1816,1840]" captionTargetBox="[191,1440,943,1776]" captionTargetId="figure-65@8.[192,1439,945,1776]" captionTargetPageId="8" captionText="Figure 7: Landscape view of the type locality showing forest of Tripura in the foreground and forests of Mizoram in the background." figureDoi="http://doi.org/10.5281/zenodo.5510020" httpUri="https://zenodo.org/record/5510020/files/figure.png" pageId="6" pageNumber="7">Fig. 7</figureCitation>
). Hence, the species is likely to be present in Mizoram and also perhaps in
<collectingCountry id="F35948ACF40BFFAAFD166556F014F98C" box="[676,801,1598,1622]" name="Bangladesh" pageId="6" pageNumber="7">Bangladesh</collectingCountry>
and
<collectingCountry id="F35948ACF40BFFAAFF436536F26DF9AC" box="[241,344,1630,1654]" name="Myanmar" pageId="6" pageNumber="7">Myanmar</collectingCountry>
as the hill range extends into the neighboring states and countries sharing a similar habitat. Other scorpions observed at the type locality were
<taxonomicName id="4C4E73BFF40BFFAAFF7265D1F245F90B" baseAuthorityName="Pocock" baseAuthorityYear="1890" box="[192,368,1721,1745]" class="Arachnida" family="Chaerilidae" genus="Chaerilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="pictus">
<emphasis id="B93AD42EF40BFFAAFF7265D1F245F90B" box="[192,368,1721,1745]" italics="true" pageId="6" pageNumber="7">Chaerilus pictus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C4E73BFF40BFFAAFECC65D2F1B3F90B" baseAuthorityName="Pocock" baseAuthorityYear="1893" box="[382,646,1721,1745]" class="Arachnida" family="Euscorpiidae" genus="Euscorpiops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="longimanus">
<emphasis id="B93AD42EF40BFFAAFECC65D2F1B3F90B" box="[382,646,1721,1745]" italics="true" pageId="6" pageNumber="7">Euscorpiops longimanus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C4E73BFF40BFFAAFD0E65D1F017F90B" authorityName="Sundevall" authorityYear="1833" box="[700,802,1721,1745]" class="Arachnida" family="Hemiscorpiidae" genus="Liocheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B93AD42EF40BFFAAFD0E65D1F017F90B" box="[700,802,1721,1745]" italics="true" pageId="6" pageNumber="7">Liocheles</emphasis>
</taxonomicName>
sp. The new species was found during a biodiversity survey of Tripura. The fieldwork was conducted across the entire southern part of the state for approximately eight weeks, and only on one occasion the new species was observed. The
<typeStatus id="54F5B69EF40BFFAAFBA766CDF747FA67" box="[1045,1138,1445,1469]" pageId="6" pageNumber="7" type="holotype">holotype</typeStatus>
was found in a ravine that can only be accessed in the dry season.
</paragraph>
</subSubSection>
</treatment>
</document>
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