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<document id="7E25DEEB8D06D1C3ABD6C67AF9F89538" ID-DOI="10.11646/phytotaxa.311.3.9" ID-ISSN="1179-3163" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1725577108548" checkinUser="felipe" docAuthor="Ståhl, Bertil &amp; Persson, Claes" docDate="2017" docId="D217A851FFDEFF8BFF74FF4CD212FAED" docLanguage="en" docName="phytotaxa.311.3.9.pdf" docOrigin="Phytotaxa 311 (3)" docSource="http://dx.doi.org/10.11646/phytotaxa.311.3.9" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Simira colorata Stahl &amp; C. H. Persson 2017, sp. nov." docType="treatment" docVersion="1" lastPageNumber="285" masterDocId="2E2ED029FFDFFF89FFFCFFD2D077FFF8" masterDocTitle="Simira colorata (Rubiaceae), a new species from western Ecuador" masterLastPageNumber="287" masterPageNumber="283" pageNumber="284" updateTime="1725586765793" updateUser="GgImagineBatch" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="D3D7398560C3302F23D95789F780C76C" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="53ABF6C0419E3F7B5BA117D7DF69527C">
<mods:title id="3A2E9E85BB0A7071DCE7A4FBED1903BB">Simira colorata (Rubiaceae), a new species from western Ecuador</mods:title>
</mods:titleInfo>
<mods:name id="E1A61CC0CEAB0A47758E99BEE598FEEF" type="personal">
<mods:role id="FFA9A5B19A6398A38DEBAFB1A9502D76">
<mods:roleTerm id="93F50706BA8D8201850196567BDB60AE">Author</mods:roleTerm>
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<mods:namePart id="24779E8E793A9BFA2CCB2C480DD9A99B">Ståhl, Bertil</mods:namePart>
<mods:affiliation id="A0D0B09D638915FBE2F51C1620EEDF15">Department of Organismal Biology, Uppsala University Campus Gotland, SE- 62167 Visby, Sweden</mods:affiliation>
</mods:name>
<mods:name id="8DCD41862E54BE6C2CAE9BD9B2746B0E" type="personal">
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</mods:role>
<mods:namePart id="7A90D5C48B2CFDCAF5A6BF4D442B7E76">Persson, Claes</mods:namePart>
<mods:affiliation id="F3526B3BBB566D4F7427B376241E9E19">Department of Biological and Environmental Sciences, University of Gothenburg, P. O. Box 461, S- 40530 Göteborg, Sweden</mods:affiliation>
</mods:name>
<mods:typeOfResource id="C385333D1DDF883091836F4D68348AF9">text</mods:typeOfResource>
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<mods:title id="8A4C7F5E8A7DC8C66D5E4F60ADF4C129">Phytotaxa</mods:title>
</mods:titleInfo>
<mods:part id="BEC029FCE15D8A1B676207DB74FF58CD">
<mods:date id="32BF5A0F15ABCC1206DEF44AAB5A5095">2017</mods:date>
<mods:detail id="6139EC332BB96B298AE070A8A14D80E4" type="pubDate">
<mods:number id="9E9693725EC2D9328D38141294C9A13E">2017-06-30</mods:number>
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<mods:number id="E5454CCACFE41E8138E1D5F9D7ED2A7B">311</mods:number>
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<mods:url id="55DFC6366216CFB5C1A23BB2E9F20AE3">http://dx.doi.org/10.11646/phytotaxa.311.3.9</mods:url>
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<mods:classification id="617C2AF18B849378D2C9B928EEA1FF2E">journal article</mods:classification>
<mods:identifier id="4DA71F19813ADF9F1F0F9801B2D5B949" type="DOI">10.11646/phytotaxa.311.3.9</mods:identifier>
<mods:identifier id="F48517601F52C5D1147D94E43D621C16" type="ISSN">1179-3163</mods:identifier>
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<treatment id="D217A851FFDEFF8BFF74FF4CD212FAED" LSID="urn:lsid:plazi:treatment:D217A851FFDEFF8BFF74FF4CD212FAED" httpUri="http://treatment.plazi.org/id/D217A851FFDEFF8BFF74FF4CD212FAED" lastPageId="2" lastPageNumber="285" pageId="1" pageNumber="284">
<subSubSection id="12A44ACCFFDEFF88FF74FF4CD287FF41" box="[136,752,158,185]" pageId="1" pageNumber="284" type="nomenclature">
<paragraph id="5A011947FFDEFF88FF74FF4CD287FF41" blockId="1.[136,752,158,185]" box="[136,752,158,185]" pageId="1" pageNumber="284">
<heading id="0149AE2BFFDEFF88FF74FF4CD287FF41" box="[136,752,158,185]" fontSize="11" level="2" pageId="1" pageNumber="284" reason="3">
<taxonomicName id="9DBE62C4FFDEFF88FF74FF4CD241FF41" ID-CoL="8X66R" authority="Stahl &amp; C.H. Persson" authorityName="Stahl &amp; C. H. Persson" authorityYear="2017" box="[136,566,158,185]" class="Magnoliopsida" family="Rubiaceae" genus="Simira" kingdom="Plantae" order="Gentianales" pageId="1" pageNumber="284" phylum="Tracheophyta" rank="species" species="colorata" status="sp. nov.">
<emphasis id="68CAC555FFDEFF88FF74FF4CD14BFF41" bold="true" box="[136,316,158,185]" italics="true" pageId="1" pageNumber="284">Simira colorata</emphasis>
Ståhl &amp; C.H. Persson
</taxonomicName>
,
<emphasis id="68CAC555FFDEFF88FDBDFF72D2E1FF41" box="[577,662,160,185]" italics="true" pageId="1" pageNumber="284">
<taxonomicNameLabel id="73F9782EFFDEFF88FDBDFF72D2E1FF41" box="[577,662,160,185]" pageId="1" pageNumber="284" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="C28505C2FFDEFF88FD59FF72D292FF41" box="[677,741,160,185]" captionStart="FIGURE 1" captionStartId="1.[136,229,1595,1616]" captionTargetBox="[166,1420,300,1555]" captionTargetId="figure-49@1.[151,1436,281,1570]" captionTargetPageId="1" captionText="FIGURE 1. Simira colorata.A, Branch with inflorescences; B, Axillary shoots; C, Flower; D, Style; E, Stamen; F, Ovary in cross section; G, Seeds; H, Fruits. A from Cerón 16624 (holotype); B from Delinks &amp; Robles 356; C–F from Pons s.n.; G–H from Cornejo &amp; Bonifaz 886. Drawn by B. Ståhl." pageId="1" pageNumber="284">Fig. 1</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="12A44ACCFFDEFF8BFF74FF14D4C6FDD9" lastPageId="2" lastPageNumber="285" pageId="1" pageNumber="284" type="materials_examined">
<paragraph id="5A011947FFDEFF88FF74FF14D351FF07" blockId="1.[136,1452,198,255]" pageId="1" pageNumber="284">
<materialsCitation id="EAD6131AFFDEFF88FF74FF14D354FF07" collectingDate="1991-09-20" collectionCode="C, E, MO" country="Ecuador" county="Type" elevation="300" latitude="-1.5" location="San Sebastian" longLatPrecision="1308" longitude="-80.566666" municipality="Road" pageId="1" pageNumber="284" specimenCount="2" stateProvince="Manabi" typeStatus="holotype">
<collectingCounty id="B36061CBFFDEFF88FF74FF14D0CDFF23" box="[136,186,198,219]" pageId="1" pageNumber="284">Type</collectingCounty>
:—
<collectingCountry id="22A959D7FFDEFF88FF29FF14D13AFF23" box="[213,333,198,219]" name="Ecuador" pageId="1" pageNumber="284">ECUADOR</collectingCountry>
.
<collectingRegion id="987AD7A5FFDEFF88FEA4FF14D1D2FF23" box="[344,421,198,219]" country="Ecuador" name="Manabi" pageId="1" pageNumber="284">Manabí</collectingRegion>
:
<collectingMunicipality id="BA65833DFFDEFF88FE53FF14D195FF23" box="[431,482,198,219]" pageId="1" pageNumber="284">Road</collectingMunicipality>
from
<location id="5F614F9CFFDEFF88FDDCFF14D2D0FF23" LSID="urn:lsid:plazi:treatment:D217A851FFDEFF8BFF74FF4CD212FAED:5F614F9CFFDEFF88FDDCFF14D2D0FF23" box="[544,679,198,219]" country="Ecuador" county="Type" latitude="-1.5" longLatPrecision="1308" longitude="-80.566666" municipality="Road" name="San Sebastian" pageId="1" pageNumber="284" stateProvince="Manabi">San Sebastián</location>
to
<location id="5F614F9CFFDEFF88FD34FF14D33FFF23" LSID="urn:lsid:plazi:treatment:D217A851FFDEFF8BFF74FF4CD212FAED:5F614F9CFFDEFF88FD34FF14D33FFF23" box="[712,840,198,219]" country="Ecuador" county="Type" latitude="-1.5" longLatPrecision="1308" longitude="-80.566666" municipality="Road" name="Agua Blanca" pageId="1" pageNumber="284" stateProvince="Manabi">Agua Blanca</location>
,
<geoCoordinate id="3F8A7F80FFDEFF88FCAFFF14D3D2FF23" box="[851,933,198,219]" degrees="01" direction="south" minutes="30" orientation="latitude" pageId="1" pageNumber="284" precision="925" value="-1.5">01°30’S</geoCoordinate>
,
<geoCoordinate id="3F8A7F80FFDEFF88FC4CFF14D47EFF23" box="[944,1033,198,219]" degrees="80" direction="west" minutes="34" orientation="longitude" pageId="1" pageNumber="284" precision="925" value="-80.566666">80°34’W</geoCoordinate>
,
<quantity id="9D46B4A2FFDEFF88FBEFFF14D4F3FF23" box="[1043,1156,198,219]" metricMagnitude="2" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="1" pageNumber="284" unit="m" value="300.0" valueMax="400.0" valueMin="200.0">
<elevation id="D193FE74FFDEFF88FBEFFF14D4F3FF23" box="[1043,1156,198,219]" metricMagnitude="2" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="1" pageNumber="284" unit="m" value="300.0" valueMax="400.0" valueMin="200.0">200–400 m</elevation>
</quantity>
,
<date id="2E003F87FFDEFF88FB73FF14D53AFF23" box="[1167,1357,198,219]" pageId="1" pageNumber="284" value="1991-09-20">
<collectingDate id="3E44C66FFFDEFF88FB73FF14D53AFF23" box="[1167,1357,198,219]" pageId="1" pageNumber="284" value="1991-09-20">20 September 1991</collectingDate>
</date>
(fl),
<emphasis id="68CAC555FFDEFF88FA7DFF14D14CFF07" italics="true" pageId="1" pageNumber="284">
<collectionCode id="3CAF8182FFDEFF88FA7DFF14D5E4FF23" box="[1409,1427,198,219]" country="Denmark" name="University of Copenhagen" pageId="1" pageNumber="284" type="Herbarium">C</collectionCode>
.
<collectionCode id="3CAF8182FFDEFF88FA6AFF14D5DFFF23" box="[1430,1448,198,219]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="1" pageNumber="284" type="Herbarium">E</collectionCode>
. Cerón 16624
</emphasis>
(
<typeStatus id="8505A7E5FFDEFF88FEB4FF38D1E9FF07" box="[328,414,234,255]" pageId="1" pageNumber="284" type="holotype">holotype</typeStatus>
GB-01150505;
<typeStatus id="8505A7E5FFDEFF88FDC4FF38D20AFF07" box="[568,637,234,255]" pageId="1" pageNumber="284" type="isotype">isotype</typeStatus>
<collectionCode id="3CAF8182FFDEFF88FD78FF38D2DCFF07" box="[644,683,234,255]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="1" pageNumber="284" type="Herbarium">MO</collectionCode>
web-photo)
</materialsCitation>
.
</paragraph>
<caption id="0EC149CFFFDEFF88FF74F9E9D118F960" pageId="1" pageNumber="284" startId="1.[136,229,1595,1616]" targetBox="[166,1420,300,1555]" targetPageId="1" targetType="figure">
<paragraph id="5A011947FFDEFF88FF74F9E9D118F960" blockId="1.[136,1452,1595,1688]" pageId="1" pageNumber="284">
<emphasis id="68CAC555FFDEFF88FF74F9E9D080F9A8" bold="true" box="[136,247,1595,1616]" pageId="1" pageNumber="284">FIGURE 1</emphasis>
.
<taxonomicName id="9DBE62C4FFDEFF88FEFEF9E9D1EFF9A8" authorityName="Stahl &amp; C. H. Persson" authorityYear="2017" box="[258,408,1595,1616]" class="Magnoliopsida" family="Rubiaceae" genus="Simira" kingdom="Plantae" order="Gentianales" pageId="1" pageNumber="284" phylum="Tracheophyta" rank="species" species="colorata">
<emphasis id="68CAC555FFDEFF88FEFEF9E9D1EFF9A8" box="[258,408,1595,1616]" italics="true" pageId="1" pageNumber="284">Simira colorata</emphasis>
</taxonomicName>
. A, Branch with inflorescences; B, Axillary shoots; C, Flower; D, Style; E, Stamen; F, Ovary in cross section; G, Seeds; H, Fruits. A from
<emphasis id="68CAC555FFDEFF88FE65F98DD260F98C" box="[409,535,1631,1652]" italics="true" pageId="1" pageNumber="284">Cerón 16624</emphasis>
(holotype); B from
<emphasis id="68CAC555FFDEFF88FD27F98DD3C6F98C" box="[731,945,1631,1652]" italics="true" pageId="1" pageNumber="284">Delinks &amp; Robles 356</emphasis>
; C–F from
<emphasis id="68CAC555FFDEFF88FBDFF98DD40BF98C" box="[1059,1148,1631,1652]" italics="true" pageId="1" pageNumber="284">
<taxonomicName id="9DBE62C4FFDEFF88FBDFF98DD424F98C" authorityName="Johnson" authorityYear="1992" box="[1059,1107,1631,1652]" class="Insecta" family="Lycaenidae" genus="Pons" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="284" phylum="Arthropoda" rank="genus">Pons</taxonomicName>
s.n.
</emphasis>
; G–H from
<emphasis id="68CAC555FFDEFF88FB0FF98DD0DBF960" italics="true" pageId="1" pageNumber="284">Cornejo &amp; Bonifaz 886</emphasis>
. Drawn by
<emphasis id="68CAC555FFDEFF88FEE1F951D11EF960" box="[285,361,1667,1688]" italics="true" pageId="1" pageNumber="284">B. Ståhl</emphasis>
.
</paragraph>
</caption>
<paragraph id="5A011947FFDEFF8BFF74F91BD4C6FDD9" blockId="1.[136,1452,1737,2086]" lastBlockId="2.[136,1452,160,1301]" lastPageId="2" lastPageNumber="285" pageId="1" pageNumber="284">
Tree
<quantity id="9D46B4A2FFDEFF88FF3DF91BD161F91A" box="[193,278,1737,1762]" metricMagnitude="1" metricUnit="m" metricValue="1.1" metricValueMax="1.7" metricValueMin="0.5" pageId="1" pageNumber="284" unit="m" value="11.0" valueMax="17.0" valueMin="5.0">5–17 m</quantity>
tall; dbh to
<quantity id="9D46B4A2FFDEFF88FE61F91BD191F91A" box="[413,486,1737,1762]" metricMagnitude="-1" metricUnit="m" metricValue="2.5" pageId="1" pageNumber="284" unit="cm" value="25.0">25 cm</quantity>
; wood turning red when exposed to air. Branchlets
<quantity id="9D46B4A2FFDEFF88FBD8F91BD4E1F91A" box="[1060,1174,1737,1762]" metricMagnitude="-3" metricUnit="m" metricValue="3.25" metricValueMax="4.0" metricValueMin="2.5" pageId="1" pageNumber="284" unit="mm" value="3.25" valueMax="4.0" valueMin="2.5">2.5–4 mm</quantity>
thick, glabrous. Stipules caducous, narrowly triangular, 13–20 ×
<quantity id="9D46B4A2FFDEFF88FDA9F93FD2BCF8FE" box="[597,715,1773,1798]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="1" pageNumber="284" unit="mm" value="1.75" valueMax="2.0" valueMin="1.5">1.5–2 mm</quantity>
, slightly twisted, glabrous outside, apex narrowly acute. Leaves opposite; petioles 6–11 ×
<quantity id="9D46B4A2FFDEFF88FE56F8C3D19AF8D2" box="[426,493,1809,1834]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="284" unit="mm" value="1.0">1 mm</quantity>
, glabrous; blades elliptic, (7–)9–18.5 × (2.5–)
<quantity id="9D46B4A2FFDEFF88FC0DF8C3D42AF8D2" box="[1009,1117,1809,1834]" metricMagnitude="-2" metricUnit="m" metricValue="5.25" metricValueMax="7.0" metricValueMin="3.5" pageId="1" pageNumber="284" unit="cm" value="5.25" valueMax="7.0" valueMin="3.5">3.5–7 cm</quantity>
, papyraceous, acute to obtuse at base, acuminate at apex with acumen 1–1.5(–2.5) cm long, glabrous above and below, the midvein prominent, secondary veins 10–14 on each side of the midrib, prominulous, tertiary and quaternary venation forming a distinct network below; tuft domatia present. Inflorescences thyrsoid, pyramidal, 50–200-flowered, with 2–3 pairs of secondary branches, these
<quantity id="9D46B4A2FFDEFF88FEC2F873D1CFF842" box="[318,440,1953,1978]" metricMagnitude="-2" metricUnit="m" metricValue="1.65" metricValueMax="1.8" metricValueMin="1.5" pageId="1" pageNumber="284" unit="mm" value="16.5" valueMax="18.0" valueMin="15.0">15–18 mm</quantity>
long; pedicels to
<quantity id="9D46B4A2FFDEFF88FD7AF873D2BEF842" box="[646,713,1953,1978]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="284" unit="mm" value="1.0">1 mm</quantity>
long, bracts and bracteoles triangular, ca. 1 ×
<quantity id="9D46B4A2FFDEFF88FB22F873D555F842" box="[1246,1314,1953,1978]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="284" unit="mm" value="1.0">1 mm</quantity>
, with round apex, inflorescence branches and bracts minutely puberulent. Flowers sessile or short-pedicellate; pedicels to
<quantity id="9D46B4A2FFDEFF88FA95F817D5DBF826" box="[1385,1452,1989,2014]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="284" unit="mm" value="1.0">1 mm</quantity>
long. Hypanthium laterally compressed, ca. 1.5 ×
<quantity id="9D46B4A2FFDEFF88FD32F83BD363F7FA" box="[718,788,2025,2050]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="284" unit="mm" value="1.0">1 mm</quantity>
, minutely puberulent. Calyx tube absent, lobes 4, widely triangular, 0.5 ×
<quantity id="9D46B4A2FFDEFF88FEB8F7DFD1EBF7DE" box="[324,412,2061,2086]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="1" pageNumber="284" unit="mm" value="0.8">0.8 mm</quantity>
, without calycophylls, outside minutely puberulent, inside with 4–5 colleters at base of each calyx lobe. Corolla funnelform, 4-lobed with imbricate aestivation,
<quantity id="9D46B4A2FFDDFF8BFC88FF72D3B8FF41" box="[884,975,160,185]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="2" pageNumber="285" unit="mm" value="3.5" valueMax="4.0" valueMin="3.0">3–4 mm</quantity>
long, lilac, drying reddish brown; tube
<quantity id="9D46B4A2FFDDFF8BFA7EFF72D0C3FF25" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="2" pageNumber="285" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">2–3 mm</quantity>
long,
<quantity id="9D46B4A2FFDDFF8BFF07FF16D125FF25" box="[251,338,196,221]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="2" pageNumber="285" unit="mm" value="1.5">1.5 mm</quantity>
wide at base, very finely puberulent outside, with a ring of dense hairs at
<quantity id="9D46B4A2FFDDFF8BFB69FF16D4A0FF25" box="[1173,1239,196,221]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="285" unit="mm" value="1.0">1 mm</quantity>
above base inside, below glabrous, without a membranaceous fold inserted between corolla tube and stamens insertion, above finely pubescent and gradually more sparsely towards apex; lobes widely triangular, 1.5 ×
<quantity id="9D46B4A2FFDDFF8BFBB0FEDED4E6FEDD" box="[1100,1169,268,293]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="285" unit="mm" value="1.0">1 mm</quantity>
, acute to obtuse at apex. Stamens 4, exserted, filaments flat,
<quantity id="9D46B4A2FFDDFF8BFDE1FEE2D228FEB1" box="[541,607,304,329]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="2" pageNumber="285" unit="mm" value="4.0">4 mm</quantity>
long, inserted ca.
<quantity id="9D46B4A2FFDDFF8BFCCDFEE2D304FEB1" box="[817,883,304,329]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="285" unit="mm" value="1.0">1 mm</quantity>
above base of the corolla tube, densely pubescent at base, finely pubescent above; anthers oblong, 1.5 ×
<quantity id="9D46B4A2FFDDFF8BFD17FE86D334FE95" box="[747,835,340,365]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="285" unit="mm" value="0.5">0.5 mm</quantity>
, subdorsifixed, apex rounded, base truncate, pale pink, drying orange. Style ca.
<quantity id="9D46B4A2FFDDFF8BFE66FEAAD186FE69" box="[410,497,376,401]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="2" pageNumber="285" unit="mm" value="4.5">4.5 mm</quantity>
long, sparsely pilose, glabrous towards tip; style branches 2, ca.
<quantity id="9D46B4A2FFDDFF8BFB35FEAAD568FE69" box="[1225,1311,376,401]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="2" pageNumber="285" unit="mm" value="1.5">1.5 mm</quantity>
long. Ovary bilocular, with a straight placenta in each locule; ovules 10–12 per locule. Capsules loculicidally dehiscent, depressedglobose, 1–1.5 × 1.1–2 ×
<quantity id="9D46B4A2FFDDFF8BFE54FE12D25FFE21" box="[424,552,448,473]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" metricValueMax="1.5" metricValueMin="1.3" pageId="2" pageNumber="285" unit="cm" value="1.4" valueMax="1.5" valueMin="1.3">1.3–1.5 cm</quantity>
, each valve with secondary splitting (resulting in four lobes at capsule complete dehiscence), woody, glabrous, greyish brown. Seeds 10–12 on each placenta, attached horizontally in 2 somewhat overlapping rows, flat with a slightly lunate outline, winged, 9–10 ×
<quantity id="9D46B4A2FFDDFF8BFC76FDDAD390FDD9" box="[906,999,520,545]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="2" pageNumber="285" unit="mm" value="3.5" valueMax="4.0" valueMin="3.0">3–4 mm</quantity>
(including wing).
</paragraph>
</subSubSection>
<subSubSection id="12A44ACCFFDDFF8BFF41FDFED212FAED" pageId="2" pageNumber="285" type="distribution">
<paragraph id="5A011947FFDDFF8BFF41FDFED5ECFD91" blockId="2.[136,1452,160,1301]" pageId="2" pageNumber="285">
<emphasis id="68CAC555FFDDFF8BFF41FDFED19DFDBD" bold="true" box="[189,490,556,581]" pageId="2" pageNumber="285">Distribution and habitat:</emphasis>
—
<taxonomicName id="9DBE62C4FFDDFF8BFDFAFDFED2C1FDBD" authorityName="Stahl &amp; C. H. Persson" authorityYear="2017" box="[518,694,556,581]" class="Magnoliopsida" family="Rubiaceae" genus="Simira" kingdom="Plantae" order="Gentianales" pageId="2" pageNumber="285" phylum="Tracheophyta" rank="species" species="colorata">
<emphasis id="68CAC555FFDDFF8BFDFAFDFED2C1FDBD" box="[518,694,556,581]" italics="true" pageId="2" pageNumber="285">Simira colorata</emphasis>
</taxonomicName>
is endemic to the adjacent lowlands and lower slopes of the coastal mountains in western
<collectingCountry id="22A959D7FFDDFF8BFE83FD82D1A8FD91" box="[383,479,592,617]" name="Ecuador" pageId="2" pageNumber="285">Ecuador</collectingCountry>
. It occurs in remnants of seasonally dry, semideciduous forests, at
<quantity id="9D46B4A2FFDDFF8BFB34FD82D54EFD91" box="[1224,1337,592,617]" metricMagnitude="2" metricUnit="m" metricValue="2.5" metricValueMax="4.5" metricValueMin="0.5" pageId="2" pageNumber="285" unit="m" value="250.0" valueMax="450.0" valueMin="50.0">50–450 m</quantity>
altitude.
</paragraph>
<paragraph id="5A011947FFDDFF8BFF41FDA6D290FCB9" blockId="2.[136,1452,160,1301]" pageId="2" pageNumber="285">
<emphasis id="68CAC555FFDDFF8BFF41FDA6D1B8FD75" bold="true" box="[189,463,628,653]" pageId="2" pageNumber="285">Conservation status:—</emphasis>
According to people living in the vicinity of the town of Balzar, in the northern province of
<collectingRegion id="987AD7A5FFDDFF8BFF59FD4AD08BFD49" box="[165,252,664,689]" country="Ecuador" name="Guayas" pageId="2" pageNumber="285">Guayas</collectingRegion>
,
<taxonomicName id="9DBE62C4FFDDFF8BFEFAFD4AD1F7FD49" authorityName="Stahl &amp; C. H. Persson" authorityYear="2017" box="[262,384,664,689]" class="Magnoliopsida" family="Rubiaceae" genus="Simira" kingdom="Plantae" order="Gentianales" pageId="2" pageNumber="285" phylum="Tracheophyta" rank="species" species="colorata">
<emphasis id="68CAC555FFDDFF8BFEFAFD4AD1F7FD49" box="[262,384,664,689]" italics="true" pageId="2" pageNumber="285">S. colorata</emphasis>
</taxonomicName>
was still plentiful in the surrounding forests in the 1950s and 1960s, and its wood was often used for house constructions. Since most natural forests in the area have now been converted into pastures or plantations, especially of teak,
<taxonomicName id="9DBE62C4FFDDFF8BFEA7FD32D1A1FD01" authorityName="Stahl &amp; C. H. Persson" authorityYear="2017" box="[347,470,736,761]" class="Magnoliopsida" family="Rubiaceae" genus="Simira" kingdom="Plantae" order="Gentianales" pageId="2" pageNumber="285" phylum="Tracheophyta" rank="species" species="colorata">
<emphasis id="68CAC555FFDDFF8BFEA7FD32D1A1FD01" box="[347,470,736,761]" italics="true" pageId="2" pageNumber="285">S. colorata</emphasis>
</taxonomicName>
has become very rare or disappeared in much of its natural range. Today individuals or small stands of
<taxonomicName id="9DBE62C4FFDDFF8BFECAFCD6D1C6FCE5" authorityName="Stahl &amp; C. H. Persson" authorityYear="2017" box="[310,433,772,797]" class="Magnoliopsida" family="Rubiaceae" genus="Simira" kingdom="Plantae" order="Gentianales" pageId="2" pageNumber="285" phylum="Tracheophyta" rank="species" species="colorata">
<emphasis id="68CAC555FFDDFF8BFECAFCD6D1C6FCE5" box="[310,433,772,797]" italics="true" pageId="2" pageNumber="285">S. colorata</emphasis>
</taxonomicName>
are restricted to forest fragments, often along streams. Following the IUCN criteria (
<bibRefCitation id="3E2F64B6FFDDFF8BFA99FCD6D0B3FCB9" author="IUCN" pageId="2" pageNumber="285" refId="ref3246" refString="IUCN. (2012) IUCN Red List Categories and Criteria: Version 3.1. Second edition. IUCN, Gland, Switzerland and Cambridge, UK." type="book" year="2012">IUCN 2012</bibRefCitation>
),
<taxonomicName id="9DBE62C4FFDDFF8BFF2BFCFAD125FCB9" authorityName="Stahl &amp; C. H. Persson" authorityYear="2017" box="[215,338,808,833]" class="Magnoliopsida" family="Rubiaceae" genus="Simira" kingdom="Plantae" order="Gentianales" pageId="2" pageNumber="285" phylum="Tracheophyta" rank="species" species="colorata">
<emphasis id="68CAC555FFDDFF8BFF2BFCFAD125FCB9" box="[215,338,808,833]" italics="true" pageId="2" pageNumber="285">S. colorata</emphasis>
</taxonomicName>
is best considered vulnerable (VU).
</paragraph>
<paragraph id="5A011947FFDDFF8BFF41FC9ED1DDFC71" blockId="2.[136,1452,160,1301]" pageId="2" pageNumber="285">
<emphasis id="68CAC555FFDDFF8BFF41FC9ED1F1FC9D" bold="true" box="[189,390,844,869]" pageId="2" pageNumber="285">Common names:</emphasis>
―Colorado (Cerón 16624; Cornejo &amp; Bonifaz 807; Persson &amp; Peña 1357); colorado de montaña (Cornejo &amp; Bonifaz 886).
</paragraph>
<paragraph id="5A011947FFDDFF8BFF41FC46D305FC55" blockId="2.[136,1452,160,1301]" box="[189,882,916,941]" pageId="2" pageNumber="285">
<emphasis id="68CAC555FFDDFF8BFF41FC46D08AFC55" bold="true" box="[189,253,916,941]" pageId="2" pageNumber="285">Uses:</emphasis>
―The wood is used for construction and for grill sticks.
</paragraph>
<paragraph id="5A011947FFDDFF8BFF41FC6AD212FAED" blockId="2.[136,1452,160,1301]" pageId="2" pageNumber="285">
<materialsCitation id="EAD6131AFFDDFF8BFF41FC6AD5DFFC0D" collectingDate="2007-02-03" collectionCode="N, B, GB, GUAY" collectorName="Hacienda Monocongo &amp; Empalme &amp; Stahl" country="Ecuador" elevation="55" latitude="-1.25" location="22 km N of Balzar" longLatPrecision="1308" longitude="-79.816666" pageId="2" pageNumber="285" specimenCount="2" stateProvince="Guayas">
<emphasis id="68CAC555FFDDFF8BFF41FC6AD281FC29" bold="true" box="[189,758,952,977]" pageId="2" pageNumber="285">
<typeStatus id="8505A7E5FFDDFF8BFF41FC6AD238FC29" box="[189,591,952,977]" isOtherTypeStatus="true" pageId="2" pageNumber="285">Additional specimens examined</typeStatus>
(
<typeStatus id="8505A7E5FFDDFF8BFD96FC6AD29EFC29" box="[618,745,952,977]" pageId="2" pageNumber="285" type="paratype">Paratypes</typeStatus>
):
</emphasis>
―
<collectingCountry id="22A959D7FFDDFF8BFCEEFC6AD3EAFC29" box="[786,925,952,977]" name="Ecuador" pageId="2" pageNumber="285">ECUADOR</collectingCountry>
.
<emphasis id="68CAC555FFDDFF8BFC48FC6AD460FC29" bold="true" box="[948,1047,952,977]" pageId="2" pageNumber="285">
<collectingRegion id="987AD7A5FFDDFF8BFC48FC6AD464FC29" box="[948,1043,952,977]" country="Ecuador" name="Guayas" pageId="2" pageNumber="285">Guayas</collectingRegion>
:
</emphasis>
<collectorName id="F74B7C91FFDDFF8BFBD6FC6AD546FC29" box="[1066,1329,952,977]" pageId="2" pageNumber="285">Hacienda Monocongo</collectorName>
,
<geoCoordinate id="3F8A7F80FFDDFF8BFAB4FC6AD5D1FC29" box="[1352,1446,952,977]" degrees="01" direction="south" minutes="15" orientation="latitude" pageId="2" pageNumber="285" precision="925" value="-1.25">01°15’S</geoCoordinate>
,
<geoCoordinate id="3F8A7F80FFDDFF8BFF74FC0ED098FC0D" box="[136,239,988,1013]" degrees="79" direction="west" minutes="49" orientation="longitude" pageId="2" pageNumber="285" precision="925" value="-79.816666">79°49’W</geoCoordinate>
, ca.
<location id="5F614F9CFFDDFF8BFEDEFC0ED184FC0D" LSID="urn:lsid:plazi:treatment:D217A851FFDEFF8BFF74FF4CD212FAED:5F614F9CFFDDFF8BFEDEFC0ED184FC0D" box="[290,499,988,1013]" country="Ecuador" latitude="-1.25" longLatPrecision="1308" longitude="-79.816666" name="22 km N of Balzar" pageId="2" pageNumber="285" stateProvince="Guayas">
<quantity id="9D46B4A2FFDDFF8BFEDEFC0ED11EFC0D" box="[290,361,988,1013]" metricMagnitude="4" metricUnit="m" metricValue="2.2" pageId="2" pageNumber="285" unit="km" value="22.0">22 km</quantity>
<collectionCode id="3CAF8182FFDDFF8BFE8CFC0ED1F3FC0D" box="[368,388,988,1013]" country="China" lsid="urn:lsid:biocol.org:col:13092" name="Nanjing University" pageId="2" pageNumber="285" type="Herbarium">N</collectionCode>
of Balzar
</location>
on road to
<collectorName id="F74B7C91FFDDFF8BFD88FC0ED295FC0D" box="[628,738,988,1013]" pageId="2" pageNumber="285">Empalme</collectorName>
,
<quantity id="9D46B4A2FFDDFF8BFD11FC0ED325FC0D" box="[749,850,988,1013]" metricMagnitude="1" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="2" pageNumber="285" unit="m" value="55.0" valueMax="60.0" valueMin="50.0">
<elevation id="D193FE74FFDDFF8BFD11FC0ED325FC0D" box="[749,850,988,1013]" metricMagnitude="1" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="2" pageNumber="285" unit="m" value="55.0" valueMax="60.0" valueMin="50.0">50–60 m</elevation>
</quantity>
,
<date id="2E003F87FFDDFF8BFCA3FC0ED46FFC0D" box="[863,1048,988,1013]" pageId="2" pageNumber="285" value="2007-02-03">
<collectingDate id="3E44C66FFFDDFF8BFCA3FC0ED46FFC0D" box="[863,1048,988,1013]" pageId="2" pageNumber="285" value="2007-02-03">3 February 2007</collectingDate>
</date>
(ster.),
<emphasis id="68CAC555FFDDFF8BFB92FC0ED571FC0D" box="[1134,1286,988,1013]" italics="true" pageId="2" pageNumber="285">
<collectionCode id="3CAF8182FFDDFF8BFB92FC0ED4F5FC0D" box="[1134,1154,988,1013]" country="Germany" lsid="urn:lsid:biocol.org:col:15534" name="Botanischer Garten und Botanisches Museum Berlin-Dahlem, Zentraleinrichtung der Freien Universitaet" pageId="2" pageNumber="285" type="Herbarium">B</collectionCode>
.
<collectorName id="F74B7C91FFDDFF8BFB71FC0ED4B0FC0D" box="[1165,1223,988,1013]" pageId="2" pageNumber="285">Ståhl</collectorName>
6802
</emphasis>
(
<collectionCode id="3CAF8182FFDDFF8BFAEAFC0ED537FC0D" box="[1302,1344,988,1013]" country="Sweden" name="University of Gothenburg" pageId="2" pageNumber="285" type="Herbarium">GB</collectionCode>
,
<collectionCode id="3CAF8182FFDDFF8BFAB0FC0ED5D7FC0D" box="[1356,1440,988,1013]" country="Ecuador" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15759" lsid="urn:lsid:biocol.org:col:15759" name="Universidad de Guayaquil" pageId="2" pageNumber="285" type="Herbarium">GUAY</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="EAD6131AFFDDFF8BFF74FBD2D44BFBE1" box="[136,1084,1024,1049]" collectingDate="2008-05-10" collectionCode="C, F, AAU, GB, GUAY" collectorName="Persson &amp; Pena" country="Ecuador" elevation="55" latitude="-1.25" location="22 km N of Balzar" longLatPrecision="1308" longitude="-79.816666" pageId="2" pageNumber="285" specimenCount="1" stateProvince="Guayas">
same locality,
<date id="2E003F87FFDDFF8BFED6FBD2D1B6FBE1" box="[298,449,1024,1049]" pageId="2" pageNumber="285" value="2008-05-10">
<collectingDate id="3E44C66FFFDDFF8BFED6FBD2D1B6FBE1" box="[298,449,1024,1049]" pageId="2" pageNumber="285" value="2008-05-10">10 May 2008</collectingDate>
</date>
(ster.),
<emphasis id="68CAC555FFDDFF8BFDE4FBD2D339FBE1" box="[536,846,1024,1049]" italics="true" pageId="2" pageNumber="285">
<collectionCode id="3CAF8182FFDDFF8BFDE4FBD2D259FBE1" box="[536,558,1024,1049]" country="Denmark" name="University of Copenhagen" pageId="2" pageNumber="285" type="Herbarium">C</collectionCode>
.
<collectorName id="F74B7C91FFDDFF8BFDC5FBD2D2E4FBE1" box="[569,659,1024,1049]" pageId="2" pageNumber="285">Persson</collectorName>
&amp;
<collectionCode id="3CAF8182FFDDFF8BFD45FBD2D2BDFBE1" box="[697,714,1024,1049]" country="USA" lsid="urn:lsid:biocol.org:col:15707" name="Field Museum of Natural History, Botany Department" pageId="2" pageNumber="285" type="Herbarium">F</collectionCode>
.
<collectorName id="F74B7C91FFDDFF8BFD29FBD2D378FBE1" box="[725,783,1024,1049]" pageId="2" pageNumber="285">Peña</collectorName>
1357
</emphasis>
(
<collectionCode id="3CAF8182FFDDFF8BFCA2FBD2D3E8FBE1" box="[862,927,1024,1049]" country="Ethiopia" httpUri="http://grbio.org/cool/5dxx-xet2" name="Addis Ababa University, Department of Biology" pageId="2" pageNumber="285">AAU</collectionCode>
,
<collectionCode id="3CAF8182FFDDFF8BFC57FBD2D3A3FBE1" box="[939,980,1024,1049]" country="Sweden" name="University of Gothenburg" pageId="2" pageNumber="285" type="Herbarium">GB</collectionCode>
,
<collectionCode id="3CAF8182FFDDFF8BFC1CFBD2D443FBE1" box="[992,1076,1024,1049]" country="Ecuador" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15759" lsid="urn:lsid:biocol.org:col:15759" name="Universidad de Guayaquil" pageId="2" pageNumber="285" type="Herbarium">GUAY</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="EAD6131AFFDDFF8BFBBBFBD2D240FBC5" collectingDate="2007-04-16" collectionCode="B, J, GB" country="Ecuador" location="Lara" municipality="Stahl" pageId="2" pageNumber="285" specimenCount="1" stateProvince="Guayas">
same plant as
<emphasis id="68CAC555FFDDFF8BFB1BFBD2D5F6FBE1" box="[1255,1409,1024,1049]" italics="true" pageId="2" pageNumber="285">
<collectionCode id="3CAF8182FFDDFF8BFB1BFBD2D48CFBE1" box="[1255,1275,1024,1049]" country="Germany" lsid="urn:lsid:biocol.org:col:15534" name="Botanischer Garten und Botanisches Museum Berlin-Dahlem, Zentraleinrichtung der Freien Universitaet" pageId="2" pageNumber="285" type="Herbarium">B</collectionCode>
.
<collectingMunicipality id="BA65833DFFDDFF8BFAFBFBD2D536FBE1" box="[1287,1345,1024,1049]" pageId="2" pageNumber="285">Ståhl</collectingMunicipality>
6802
</emphasis>
,
<date id="2E003F87FFDDFF8BFA73FBD2D176FBC5" pageId="2" pageNumber="285" value="2007-04-16">
<collectingDate id="3E44C66FFFDDFF8BFA73FBD2D176FBC5" pageId="2" pageNumber="285" value="2007-04-16">16 April 2007</collectingDate>
</date>
(fl),
<emphasis id="68CAC555FFDDFF8BFEC4FBF6D183FBC5" box="[312,500,1060,1085]" italics="true" pageId="2" pageNumber="285">
<collectionCode id="3CAF8182FFDDFF8BFEC4FBF6D13FFBC5" box="[312,328,1060,1085]" country="South Africa" lsid="urn:lsid:biocol.org:col:14647" name="University of the Witwatersrand" pageId="2" pageNumber="285" type="Herbarium">J</collectionCode>
.
<taxonomicName id="9DBE62C4FFDDFF8BFEAEFBF6D1FDFBC5" authorityName="Johnson" authorityYear="1992" box="[338,394,1060,1085]" class="Insecta" family="Lycaenidae" genus="Pons" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="285" phylum="Arthropoda" rank="genus">Pons</taxonomicName>
<location id="5F614F9CFFDDFF8BFE6CFBF6D1B1FBC5" LSID="urn:lsid:plazi:treatment:D217A851FFDEFF8BFF74FF4CD212FAED:5F614F9CFFDDFF8BFE6CFBF6D1B1FBC5" box="[400,454,1060,1085]" country="Ecuador" municipality="Stahl" name="Lara" pageId="2" pageNumber="285" stateProvince="Guayas">Lara</location>
s.n.
</emphasis>
(
<collectionCode id="3CAF8182FFDDFF8BFDFFFBF6D259FBC5" box="[515,558,1060,1085]" country="Sweden" name="University of Gothenburg" pageId="2" pageNumber="285" type="Herbarium">GB</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="EAD6131AFFDDFF8BFDBDFBF6D51AFB99" collectingDate="1999-10-06" collectionCode="S, T, C, GB, MO" collectorName="Jama &amp; Camarones &amp; Delinks &amp; Robles" country="Ecuador" elevation="100" latitude="-0.09388889" location="24 km S of Pedernales" longLatPrecision="21" longitude="-80.14555" pageId="2" pageNumber="285" specimenCount="1" stateProvince="Manabi">
<emphasis id="68CAC555FFDDFF8BFDBDFBF6D2D0FBC5" bold="true" box="[577,679,1060,1085]" pageId="2" pageNumber="285">
<collectingRegion id="987AD7A5FFDDFF8BFDBDFBF6D2D4FBC5" box="[577,675,1060,1085]" country="Ecuador" name="Manabi" pageId="2" pageNumber="285">Manabí</collectingRegion>
:
</emphasis>
<collectorName id="F74B7C91FFDDFF8BFD51FBF6D29EFBC5" box="[685,745,1060,1085]" pageId="2" pageNumber="285">Jama</collectorName>
,
<collectorName id="F74B7C91FFDDFF8BFD08FBF6D302FBC5" box="[756,885,1060,1085]" pageId="2" pageNumber="285">Camarones</collectorName>
,
<location id="5F614F9CFFDDFF8BFC83FBF6D40CFBC5" LSID="urn:lsid:plazi:treatment:D217A851FFDEFF8BFF74FF4CD212FAED:5F614F9CFFDDFF8BFC83FBF6D40CFBC5" box="[895,1147,1060,1085]" country="Ecuador" latitude="-0.09388889" longLatPrecision="21" longitude="-80.14555" name="24 km S of Pedernales" pageId="2" pageNumber="285" stateProvince="Manabi">
<quantity id="9D46B4A2FFDDFF8BFC83FBF6D3B2FBC5" box="[895,965,1060,1085]" metricMagnitude="4" metricUnit="m" metricValue="2.4" pageId="2" pageNumber="285" unit="km" value="24.0">24 km</quantity>
<collectionCode id="3CAF8182FFDDFF8BFC37FBF6D3ACFBC5" box="[971,987,1060,1085]" country="Sweden" lsid="urn:lsid:biocol.org:col:15668" name="Department of Botany, Swedish Museum of Natural History" pageId="2" pageNumber="285" type="Herbarium">S</collectionCode>
of Pedernales
</location>
, SE of the coastal highway,
<geoCoordinate id="3F8A7F80FFDDFF8BFF74FB9AD178FB99" box="[136,271,1096,1121]" degrees="00" direction="south" minutes="05" orientation="latitude" pageId="2" pageNumber="285" precision="15" seconds="38" value="-0.09388889">00°05’38”S</geoCoordinate>
,
<geoCoordinate id="3F8A7F80FFDDFF8BFEE0FB9AD1DBFB99" box="[284,428,1096,1121]" degrees="80" direction="west" minutes="08" orientation="longitude" pageId="2" pageNumber="285" precision="15" seconds="44" value="-80.14555">80°08’44”W</geoCoordinate>
,
<quantity id="9D46B4A2FFDDFF8BFE45FB9AD25AFB99" box="[441,557,1096,1121]" metricMagnitude="2" metricUnit="m" metricValue="1.0" metricValueMax="1.25" metricValueMin="0.75" pageId="2" pageNumber="285" unit="m" value="100.0" valueMax="125.0" valueMin="75.0">
<elevation id="D193FE74FFDDFF8BFE45FB9AD25AFB99" box="[441,557,1096,1121]" metricMagnitude="2" metricUnit="m" metricValue="1.0" metricValueMax="1.25" metricValueMin="0.75" pageId="2" pageNumber="285" unit="m" value="100.0" valueMax="125.0" valueMin="75.0">75–125 m</elevation>
</quantity>
,
<date id="2E003F87FFDDFF8BFDC7FB9AD29BFB99" box="[571,748,1096,1121]" pageId="2" pageNumber="285" value="1999-10-06">
<collectingDate id="3E44C66FFFDDFF8BFDC7FB9AD29BFB99" box="[571,748,1096,1121]" pageId="2" pageNumber="285" value="1999-10-06">6 October 1999</collectingDate>
</date>
(fr.),
<emphasis id="68CAC555FFDDFF8BFCD3FB9AD41DFB99" box="[815,1130,1096,1121]" italics="true" pageId="2" pageNumber="285">
<collectionCode id="3CAF8182FFDDFF8BFCD3FB9AD337FB99" box="[815,832,1096,1121]" name="Tavera, Department of Geology and Geophysics" pageId="2" pageNumber="285">T</collectionCode>
.
<collectorName id="F74B7C91FFDDFF8BFCB0FB9AD3D5FB99" box="[844,930,1096,1121]" pageId="2" pageNumber="285">Delinks</collectorName>
&amp;
<collectionCode id="3CAF8182FFDDFF8BFC35FB9AD3A8FB99" box="[969,991,1096,1121]" country="Denmark" name="University of Copenhagen" pageId="2" pageNumber="285" type="Herbarium">C</collectionCode>
.
<collectorName id="F74B7C91FFDDFF8BFC17FB9AD440FB99" box="[1003,1079,1096,1121]" pageId="2" pageNumber="285">Robles</collectorName>
356
</emphasis>
(
<collectionCode id="3CAF8182FFDDFF8BFB87FB9AD4D2FB99" box="[1147,1189,1096,1121]" country="Sweden" name="University of Gothenburg" pageId="2" pageNumber="285" type="Herbarium">GB</collectionCode>
,
<collectionCode id="3CAF8182FFDDFF8BFB4EFB9AD4A8FB99" box="[1202,1247,1096,1121]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="2" pageNumber="285" type="Herbarium">MO</collectionCode>
web-photo)
</materialsCitation>
;
<materialsCitation id="EAD6131AFFDDFF8BFA86FB9AD1F6FB51" collectingDate="1991-09-20" collectionCode="C, E, GB, MO" collectorName="San Sebastian &amp; Agua Blanca &amp; Ceron" country="Ecuador" elevation="300" latitude="-1.5" longLatPrecision="1308" longitude="-80.566666" pageId="2" pageNumber="285" specimenCount="1" stateProvince="Manabi">
road from
<collectorName id="F74B7C91FFDDFF8BFF35FBBED11EFB7D" box="[201,361,1132,1157]" pageId="2" pageNumber="285">San Sebastián</collectorName>
to
<collectorName id="F74B7C91FFDDFF8BFE6EFBBED25CFB7D" box="[402,555,1132,1157]" pageId="2" pageNumber="285">Agua Blanca</collectorName>
,
<geoCoordinate id="3F8A7F80FFDDFF8BFDC5FBBED2E0FB7D" box="[569,663,1132,1157]" degrees="01" direction="south" minutes="30" orientation="latitude" pageId="2" pageNumber="285" precision="925" value="-1.5">01°30’S</geoCoordinate>
,
<geoCoordinate id="3F8A7F80FFDDFF8BFD5BFBBED379FB7D" box="[679,782,1132,1157]" degrees="80" direction="west" minutes="34" orientation="longitude" pageId="2" pageNumber="285" precision="925" value="-80.566666">80°34’W</geoCoordinate>
,
<quantity id="9D46B4A2FFDDFF8BFCE1FBBED3D6FB7D" box="[797,929,1132,1157]" metricMagnitude="2" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="2" pageNumber="285" unit="m" value="300.0" valueMax="400.0" valueMin="200.0">
<elevation id="D193FE74FFDDFF8BFCE1FBBED3D6FB7D" box="[797,929,1132,1157]" metricMagnitude="2" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="2" pageNumber="285" unit="m" value="300.0" valueMax="400.0" valueMin="200.0">200–400 m</elevation>
</quantity>
,
<date id="2E003F87FFDDFF8BFC4DFBBED4E6FB7D" box="[945,1169,1132,1157]" pageId="2" pageNumber="285" value="1991-09-20">
<collectingDate id="3E44C66FFFDDFF8BFC4DFBBED4E6FB7D" box="[945,1169,1132,1157]" pageId="2" pageNumber="285" value="1991-09-20">20 September 1991</collectingDate>
</date>
(fl.),
<emphasis id="68CAC555FFDDFF8BFB24FBBED5DCFB7D" box="[1240,1451,1132,1157]" italics="true" pageId="2" pageNumber="285">
<collectionCode id="3CAF8182FFDDFF8BFB24FBBED49AFB7D" box="[1240,1261,1132,1157]" country="Denmark" name="University of Copenhagen" pageId="2" pageNumber="285" type="Herbarium">C</collectionCode>
.
<collectionCode id="3CAF8182FFDDFF8BFB0DFBBED571FB7D" box="[1265,1286,1132,1157]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="2" pageNumber="285" type="Herbarium">E</collectionCode>
.
<collectorName id="F74B7C91FFDDFF8BFAE9FBBED52CFB7D" box="[1301,1371,1132,1157]" pageId="2" pageNumber="285">Cerón</collectorName>
16624
</emphasis>
(
<collectionCode id="3CAF8182FFDDFF8BFF6CFB42D0CDFB51" box="[144,186,1168,1193]" country="Sweden" name="University of Gothenburg" pageId="2" pageNumber="285" type="Herbarium">GB</collectionCode>
,
<collectionCode id="3CAF8182FFDDFF8BFF3BFB42D083FB51" box="[199,244,1168,1193]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="2" pageNumber="285" type="Herbarium">MO</collectionCode>
web-photo)
</materialsCitation>
;
<materialsCitation id="EAD6131AFFDDFF8BFE70FB42D2F6FB35" collectingDate="1993-11-26" collectionCode="C, GB, GUAY" collectorName="X. Cornejo &amp; Bonifaz" country="Ecuador" elevation="350" latitude="-1.6333333" location="Cerro Cabeza de Vaca" longLatPrecision="1308" longitude="-80.63333" municipality="Rio Ayampe" pageId="2" pageNumber="285" specimenCount="1" stateProvince="Manabi">
<collectingMunicipality id="BA65833DFFDDFF8BFE70FB42D26DFB51" box="[396,538,1168,1193]" pageId="2" pageNumber="285">Río Ayampe</collectingMunicipality>
,
<location id="5F614F9CFFDDFF8BFDDAFB42D355FB51" LSID="urn:lsid:plazi:treatment:D217A851FFDEFF8BFF74FF4CD212FAED:5F614F9CFFDDFF8BFDDAFB42D355FB51" box="[550,802,1168,1193]" country="Ecuador" latitude="-1.6333333" longLatPrecision="1308" longitude="-80.63333" municipality="Rio Ayampe" name="Cerro Cabeza de Vaca" pageId="2" pageNumber="285" stateProvince="Manabi">Cerro Cabeza de Vaca</location>
,
<geoCoordinate id="3F8A7F80FFDDFF8BFCD2FB42D3FAFB51" box="[814,909,1168,1193]" degrees="01" direction="south" minutes="38" orientation="latitude" pageId="2" pageNumber="285" precision="925" value="-1.6333333">01°38’S</geoCoordinate>
,
<geoCoordinate id="3F8A7F80FFDDFF8BFC65FB42D477FB51" box="[921,1024,1168,1193]" degrees="80" direction="west" minutes="38" orientation="longitude" pageId="2" pageNumber="285" precision="925" value="-80.63333">80°38’W</geoCoordinate>
,
<quantity id="9D46B4A2FFDDFF8BFBF0FB42D421FB51" box="[1036,1110,1168,1193]" metricMagnitude="2" metricUnit="m" metricValue="3.5" pageId="2" pageNumber="285" unit="m" value="350.0">
<elevation id="D193FE74FFDDFF8BFBF0FB42D421FB51" box="[1036,1110,1168,1193]" metricMagnitude="2" metricUnit="m" metricValue="3.5" pageId="2" pageNumber="285" unit="m" value="350.0">350 m</elevation>
</quantity>
,
<date id="2E003F87FFDDFF8BFB9FFB42D54BFB51" box="[1123,1340,1168,1193]" pageId="2" pageNumber="285" value="1993-11-26">
<collectingDate id="3E44C66FFFDDFF8BFB9FFB42D54BFB51" box="[1123,1340,1168,1193]" pageId="2" pageNumber="285" value="1993-11-26">26 November 1993</collectingDate>
</date>
(ster.),
<emphasis id="68CAC555FFDDFF8BFA6FFB42D1C6FB35" italics="true" pageId="2" pageNumber="285">
<collectorName id="F74B7C91FFDDFF8BFA6FFB42D093FB35" pageId="2" pageNumber="285">X. Cornejo</collectorName>
&amp;
<collectionCode id="3CAF8182FFDDFF8BFEF5FB66D168FB35" box="[265,287,1204,1229]" country="Denmark" name="University of Copenhagen" pageId="2" pageNumber="285" type="Herbarium">C</collectionCode>
.
<collectorName id="F74B7C91FFDDFF8BFED6FB66D1F7FB35" box="[298,384,1204,1229]" pageId="2" pageNumber="285">Bonifaz</collectorName>
807
</emphasis>
(
<collectionCode id="3CAF8182FFDDFF8BFE3DFB66D19CFB35" box="[449,491,1204,1229]" country="Sweden" name="University of Gothenburg" pageId="2" pageNumber="285" type="Herbarium">GB</collectionCode>
,
<collectionCode id="3CAF8182FFDDFF8BFE04FB66D232FB35" box="[504,581,1204,1229]" country="Ecuador" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15759" lsid="urn:lsid:biocol.org:col:15759" name="Universidad de Guayaquil" pageId="2" pageNumber="285" type="Herbarium">GUAY</collectionCode>
<taxonomicNameLabel id="73F9782EFFDDFF8BFDB1FB66D200FB35" box="[589,631,1204,1229]" pageId="2" pageNumber="285" rank="variety">n.v.</taxonomicNameLabel>
)
</materialsCitation>
.
<materialsCitation id="EAD6131AFFDDFF8BFD71FB66D216FAED" collectingDate="1993-11-28" collectionCode="C, GB, GUAY" collectorName="X. Cornejo &amp; Bonifaz" country="Ecuador" county="Rio Ayampe" elevation="450" latitude="-1.6833334" location="Cerro El Burro" longLatPrecision="1308" longitude="-80.666664" municipality="Cerro El Burro" pageId="2" pageNumber="285" specimenCode="MA 816024" specimenCount="1" stateProvince="Santa Elena">
<emphasis id="68CAC555FFDDFF8BFD71FB66D351FB35" bold="true" box="[653,806,1204,1229]" pageId="2" pageNumber="285">
<collectingRegion id="987AD7A5FFDDFF8BFD71FB66D355FB35" box="[653,802,1204,1229]" country="Ecuador" name="Santa Elena" pageId="2" pageNumber="285">Santa Elena</collectingRegion>
:
</emphasis>
<collectingCounty id="B36061CBFFDDFF8BFCD2FB66D3CBFB35" box="[814,956,1204,1229]" pageId="2" pageNumber="285">Río Ayampe</collectingCounty>
,
<location id="5F614F9CFFDDFF8BFC3BFB66D404FB35" LSID="urn:lsid:plazi:treatment:D217A851FFDEFF8BFF74FF4CD212FAED:5F614F9CFFDDFF8BFC3BFB66D404FB35" box="[967,1139,1204,1229]" country="Ecuador" county="Rio Ayampe" latitude="-1.6833334" longLatPrecision="1308" longitude="-80.666664" municipality="Cerro El Burro" name="Cerro El Burro" pageId="2" pageNumber="285" stateProvince="Santa Elena">
<collectingMunicipality id="BA65833DFFDDFF8BFC3BFB66D404FB35" box="[967,1139,1204,1229]" pageId="2" pageNumber="285">Cerro El Burro</collectingMunicipality>
</location>
,
<geoCoordinate id="3F8A7F80FFDDFF8BFB83FB66D4AAFB35" box="[1151,1245,1204,1229]" degrees="01" direction="south" minutes="41" orientation="latitude" pageId="2" pageNumber="285" precision="925" value="-1.6833334">01°41’S</geoCoordinate>
,
<geoCoordinate id="3F8A7F80FFDDFF8BFB16FB66D526FB35" box="[1258,1361,1204,1229]" degrees="80" direction="west" minutes="40" orientation="longitude" pageId="2" pageNumber="285" precision="925" value="-80.666664">80°40’W</geoCoordinate>
,
<quantity id="9D46B4A2FFDDFF8BFAA1FB66D5D1FB35" box="[1373,1446,1204,1229]" metricMagnitude="2" metricUnit="m" metricValue="4.5" pageId="2" pageNumber="285" unit="m" value="450.0">
<elevation id="D193FE74FFDDFF8BFAA1FB66D5D1FB35" box="[1373,1446,1204,1229]" metricMagnitude="2" metricUnit="m" metricValue="4.5" pageId="2" pageNumber="285" unit="m" value="450.0">450 m</elevation>
</quantity>
,
<date id="2E003F87FFDDFF8BFF74FB0AD115FB09" box="[136,354,1240,1265]" pageId="2" pageNumber="285" value="1993-11-28">
<collectingDate id="3E44C66FFFDDFF8BFF74FB0AD115FB09" box="[136,354,1240,1265]" pageId="2" pageNumber="285" value="1993-11-28">28 November 1993</collectingDate>
</date>
(ster.),
<emphasis id="68CAC555FFDDFF8BFE45FB0AD375FB09" box="[441,770,1240,1265]" italics="true" pageId="2" pageNumber="285">
<collectorName id="F74B7C91FFDDFF8BFE45FB0AD242FB09" box="[441,565,1240,1265]" pageId="2" pageNumber="285">X. Cornejo</collectorName>
&amp;
<collectionCode id="3CAF8182FFDDFF8BFDA6FB0AD207FB09" box="[602,624,1240,1265]" country="Denmark" name="University of Copenhagen" pageId="2" pageNumber="285" type="Herbarium">C</collectionCode>
.
<collectorName id="F74B7C91FFDDFF8BFD87FB0AD2A6FB09" box="[635,721,1240,1265]" pageId="2" pageNumber="285">Bonifaz</collectorName>
886
</emphasis>
(
<collectionCode id="3CAF8182FFDDFF8BFCEFFB0AD34AFB09" box="[787,829,1240,1265]" country="Sweden" name="University of Gothenburg" pageId="2" pageNumber="285" type="Herbarium">GB</collectionCode>
,
<collectionCode id="3CAF8182FFDDFF8BFCB5FB0AD3E1FB09" box="[841,918,1240,1265]" country="Ecuador" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15759" lsid="urn:lsid:biocol.org:col:15759" name="Universidad de Guayaquil" pageId="2" pageNumber="285" type="Herbarium">GUAY</collectionCode>
<taxonomicNameLabel id="73F9782EFFDDFF8BFC62FB0AD3BEFB09" box="[926,969,1240,1265]" pageId="2" pageNumber="285" rank="variety">n.v.</taxonomicNameLabel>
).
<emphasis id="68CAC555FFDDFF8BFC23FB0AD4CDFB09" bold="true" box="[991,1210,1240,1265]" pageId="2" pageNumber="285">
<location id="5F614F9CFFDDFF8BFC23FB0AD4C2FB09" LSID="urn:lsid:plazi:treatment:D217A851FFDEFF8BFF74FF4CD212FAED:5F614F9CFFDDFF8BFC23FB0AD4C2FB09" box="[991,1205,1240,1265]" country="Ecuador" county="Rio Ayampe" latitude="-1.6833334" longLatPrecision="1308" longitude="-80.666664" municipality="Cerro El Burro" name="Without province" pageId="2" pageNumber="285" stateProvince="Santa Elena">Without province</location>
:
</emphasis>
1802 (fr.), “Herbario de
<emphasis id="68CAC555FFDDFF8BFF55FB2ED130FAED" box="[169,327,1276,1301]" italics="true" pageId="2" pageNumber="285">Ruiz &amp; Pavón</emphasis>
”,
<emphasis id="68CAC555FFDDFF8BFE9DFB2ED1CAFAED" box="[353,445,1276,1301]" italics="true" pageId="2" pageNumber="285">s.d., s.n.</emphasis>
(
<specimenCode id="0A18B13CFFDDFF8BFE37FB2ED22EFAED" box="[459,601,1276,1301]" collectionCode="MA" country="Spain" lsid="urn:lsid:biocol.org:col:15682" name="Real Jardín Botánico" pageId="2" pageNumber="285" type="Herbarium">MA 816024</specimenCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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