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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.949.51269" ID-GBIF-Dataset="e4d2c425-4596-487c-8c4f-b09330b48203" ID-PMC="PMC7381721" ID-Pensoft-Pub="1313-2970-949-1" ID-Pensoft-UUID="303D64593B3B550DBB7CE01AA547409D" ID-PubMed="32765176" ID-ZooBank="93BFA448BB7343CDBC4EA86090DA63D5" ModsDocID="1313-2970-949-1" checkinTime="1594832489974" checkinUser="pensoft" docAuthor="Salata, Sebastian &amp; Fisher, Brian L." docDate="2020" docId="A1F22F1F8FAA512A98455C9D9DD18978" docLanguage="en" docName="ZooKeys 949: 1-185" docOrigin="ZooKeys 949" docSource="http://dx.doi.org/10.3897/zookeys.949.51269" docTitle="Pheidole midongy Salata &amp; Fisher 2020, sp. nov." docType="treatment" docUuid="100E9394-CB7C-4A53-84F1-D208EC6EBFC1" docUuidSource="ZooBank" docVersion="4" id="303D64593B3B550DBB7CE01AA547409D" lastPageNumber="1" masterDocId="303D64593B3B550DBB7CE01AA547409D" masterDocTitle="Taxonomic revision of the Pheidole sikorae species group (Hymenoptera, Formicidae) from Madagascar" masterLastPageNumber="185" masterPageNumber="1" pageNumber="1" updateTime="1668169097828" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Taxonomic revision of the Pheidole sikorae species group (Hymenoptera, Formicidae) from Madagascar</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Salata, Sebastian</mods:namePart>
<mods:affiliation>Department of Entomology, California Academy of Sciences, San Francisco, CA 94118, USA</mods:affiliation>
<mods:nameIdentifier type="orcid">https://orcid.org/0000-0003-0811-2309</mods:nameIdentifier>
<mods:nameIdentifier type="email">sdsalata@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Fisher, Brian L.</mods:namePart>
<mods:affiliation>Department of Entomology, California Academy of Sciences, San Francisco, CA 94118, USA</mods:affiliation>
<mods:nameIdentifier type="orcid">https://orcid.org/0000-0002-4653-3270</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2020</mods:date>
<mods:detail type="volume">
<mods:number>949</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>185</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.949.51269</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.949.51269</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-949-1</mods:identifier>
<mods:identifier type="ZooBank">93BFA448BB7343CDBC4EA86090DA63D5</mods:identifier>
<mods:identifier type="Pensoft-UUID">303D64593B3B550DBB7CE01AA547409D</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="165090169" LSID="urn:lsid:zoobank.org:act:100E9394-CB7C-4A53-84F1-D208EC6EBFC1" httpUri="http://treatment.plazi.org/id/A1F22F1F8FAA512A98455C9D9DD18978" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName LSID="A1F22F1F-8FAA-512A-9845-5C9D9DD18978" authority="Salata &amp; Fisher, 2020" authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole midongy" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="midongy" status="sp. nov.">Pheidole midongy</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 45" captionStartId="F45" captionText="Figure 45. Pheidole midongy sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923275) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0119419)." figureDoi="10.3897/zookeys.949.51269.figure45" httpUri="https://binary.pensoft.net/fig/432214" pageId="0" pageNumber="1">Figs 45A-F</figureCitation>
<figureCitation captionStart="Figure 64" captionStartId="F64" captionText="Figure 64. Major worker, hypostomal teeth. Pheidole masoandro sp. nov. (A). Pheidole mavohavoana sp. nov. (B). Pheidole midongy sp. nov. (C). Pheidole mikros sp. nov. (D). Pheidole mivory sp. nov. (E). Pheidole nitidobruna sp. nov. (F). Pheidole parvula sp. nov. (G). Pheidole parvulogibba sp. nov. (H). Pheidole renirano sp. nov. (I). Pheidole sava sp. nov. (J). Pheidole sikorae Forel (K). Pheidole sofia sp. nov. (L). Pheidole sparsa sp. nov. (M). Pheidole tampony sp. nov. (N). Pheidole trichotos sp. nov. (O). Pheidole tsaravoniana sp. nov. (P). Pheidole vadum sp. nov. (Q). Pheidole veteratrix Forel (R). Pheidole volontany sp. nov. (S). Pheidole vony sp. nov. (T)." figureDoi="10.3897/zookeys.949.51269.figure64" httpUri="https://binary.pensoft.net/fig/432233" pageId="0" pageNumber="1">, 64C</figureCitation>
<figureCitation captionStart="Figure 66" captionStartId="F66" captionText="Figure 66. Distribution. Pheidole mahamavo sp. nov. (A). Pheidole mainty sp. nov. (B). Pheidole mamiratra sp. nov. (C). Pheidole manantenina sp. nov. (D). Pheidole masoandro sp. nov. (E). Pheidole mavohavoana sp. nov. (F). Pheidole midongy sp. nov. (G). Pheidole mikros sp. nov. (H). Pheidole mivory sp. nov. (I). Pheidole nitidobruna sp. nov. (J). Pheidole parvula sp. nov. (K). Pheidole parvulogibba sp. nov. (L)." figureDoi="10.3897/zookeys.949.51269.figure66" httpUri="https://binary.pensoft.net/fig/432235" pageId="0" pageNumber="1">, 66G</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type material">
<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Holotype.</emphasis>
Madagascar. • 1 major worker; Fianarantsoa; Parc National Befotaka-Midongy, Papango 27.7 km S Midongy-Sud, Mount Papango;
<geoCoordinate degrees="23.83517" direction="south" orientation="latitude" precision="1" value="-23.83517">-23.83517</geoCoordinate>
,
<geoCoordinate degrees="46.96367" direction="east" orientation="longitude" precision="1" value="46.96367">46.96367</geoCoordinate>
; alt. 940 m; 14 Nov 2006; B. L. Fisher et al. leg.; rainforest, ex rotten log; BLF14842; CASENT0119419 (CASC).
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Paratype.</emphasis>
• 1w.; same data as for holotype, CASENT0923275 (CASC).
</paragraph>
<caption doi="10.3897/zookeys.949.51269.figure45" httpUri="https://binary.pensoft.net/fig/432214" pageId="0" pageNumber="1" start="Figure 45" startId="F45">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 45.</emphasis>
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole midongy" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="midongy">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole midongy</emphasis>
</taxonomicName>
sp. nov., full-face view (
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
), profile (
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
), and dorsal view (
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
) of paratype minor worker (CASENT0923275) and full-face view (
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
), profile (
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
), and dorsal view (
<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
) of holotype major worker (CASENT0119419).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
Minute species.
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Major workers.</emphasis>
HL &lt;0.9 mm and WL &lt;0.7 mm; head in full-face view sub-oval, slightly widening posteriorly, with anterior and posterior sides convex, in lateral view sub-oval; ventral and dorsal faces convex; body blackish brown; sides of head with moderately dense, moderately short, suberect pilosity; entire head distinctly sculptured, medial part of frons with thick, longitudinal, and dense rugae and distinctly rugulate interspaces, rugae more irregular and directed slightly outward on posteromedial part; scape, when laid back, exceeding the midlength of head by one-fifth of its length; mesosoma distinctly foveolate, promesonotum with additional sparse, thin, and irregular rugae on dorsum; katepisternum and propodeum with sparser foveolae; inner hypostomal teeth distinct, large, closely spaced, triangular, with rounded apex directed upward; outer hypostomal teeth lobe-like, wider than inner teeth and approximately the same height; inner and outer hypostomal teeth closely spaced and not connected by concavity; base of first gastral tergite smooth.
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Minor workers.</emphasis>
HL &lt;0.5 mm and WL &lt;0.6 mm, scape, when laid back, exceeding the posterior head margin by less than one-fifth of its length; propodeal spines reduced to small tubercles; head relatively oval; frons with sparse and moderately thick, short, and predominantly longitudinal rugulae fading medially; vertex with transverse, sparse, moderately thick, and short rugulae; area posterolateral from eyes smooth; body dark yellow; mesosoma smooth.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Major workers.</emphasis>
Measurements (
<emphasis italics="true" pageId="0" pageNumber="1">N</emphasis>
= 1): HL: 0.89; HW: 0.91; SL: 0.5; EL: 0.12; WL: 0.74; PSL: 0.15; MTL: 0.46; PNW: 0.45; PTW: 0.12; PPW: 0.3; CI: 96.8; SI: 54.5; PSLI: 17.4; PPI: 40.5; PNI: 49.2; MTI: 50.0.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Head.</emphasis>
In full-face view sub-oval, slightly widening posteriorly, with anterior and posterior sides convex (Fig.
<figureCitation captionStart="Figure 45" captionStartId="F45" captionText="Figure 45. Pheidole midongy sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923275) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0119419)." figureDoi="10.3897/zookeys.949.51269.figure45" httpUri="https://binary.pensoft.net/fig/432214" pageId="0" pageNumber="1">45B</figureCitation>
). In lateral view sub-oval; ventral and dorsal faces convex; inner hypostomal teeth visible. Sides of the head with moderately dense, moderately short, suberect pilosity; whole head with dense, long, decumbent to erect pilosity. Medial part of frons with thick, longitudinal, and dense rugae and distinctly rugulate interspaces, rugae more irregular and directed slightly outward on posteromedial part; lateral sides with thick, sparse, and irregular rugae with distinctly rugofoveolate interspaces. Occipital lobes with dense, irregular rugae and distinctly rugoreticulate interspaces. Area posterolateral from eyes with dense rugofoveolae and additional sparse and irregular rugae. Gena with relatively sparse and thick, longitudinal rugae and smooth to indistinctly rugoreticulate interspaces. Centre of clypeus smooth and shiny, lateral sides with indistinct rugulae; median notch present, moderately wide, and shallow; median longitudinal carina absent; lateral longitudinal carinae absent. Scape, when laid back, exceeding the midlength of head by one-fifth of its length; pilosity subdecumbent to erect (Fig.
<figureCitation captionStart="Figure 45" captionStartId="F45" captionText="Figure 45. Pheidole midongy sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923275) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0119419)." figureDoi="10.3897/zookeys.949.51269.figure45" httpUri="https://binary.pensoft.net/fig/432214" pageId="0" pageNumber="1">45B, D</figureCitation>
). Inner hypostomal teeth distinct, large, closely spaced, triangular, with rounded apex directed upward; outer hypostomal teeth lobe-like, wider than inner teeth and approximately the same height; inner and outer hypostomal teeth closely spaced and not connected by concavity (Fig.
<figureCitation captionStart="Figure 64" captionStartId="F64" captionText="Figure 64. Major worker, hypostomal teeth. Pheidole masoandro sp. nov. (A). Pheidole mavohavoana sp. nov. (B). Pheidole midongy sp. nov. (C). Pheidole mikros sp. nov. (D). Pheidole mivory sp. nov. (E). Pheidole nitidobruna sp. nov. (F). Pheidole parvula sp. nov. (G). Pheidole parvulogibba sp. nov. (H). Pheidole renirano sp. nov. (I). Pheidole sava sp. nov. (J). Pheidole sikorae Forel (K). Pheidole sofia sp. nov. (L). Pheidole sparsa sp. nov. (M). Pheidole tampony sp. nov. (N). Pheidole trichotos sp. nov. (O). Pheidole tsaravoniana sp. nov. (P). Pheidole vadum sp. nov. (Q). Pheidole veteratrix Forel (R). Pheidole volontany sp. nov. (S). Pheidole vony sp. nov. (T)." figureDoi="10.3897/zookeys.949.51269.figure64" httpUri="https://binary.pensoft.net/fig/432233" pageId="0" pageNumber="1">64C</figureCitation>
).
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Mesosoma.</emphasis>
In lateral view, promesonotum short, angular, and moderately high, posterior mesonotum moderately steep, mesonotal process very indistinct, tubercle-like; promesonotal groove absent; metanotal groove absent; propodeal spines moderate, with wide base and acute apex; humeral area produced (Fig.
<figureCitation captionStart="Figure 45" captionStartId="F45" captionText="Figure 45. Pheidole midongy sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923275) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0119419)." figureDoi="10.3897/zookeys.949.51269.figure45" httpUri="https://binary.pensoft.net/fig/432214" pageId="0" pageNumber="1">45D</figureCitation>
). Surface shiny and distinctly foveolate; promesonotum with additional sparse, thin, and irregular rugae on dorsum; katepisternum and propodeum with sparser foveolae. Pilosity moderately sparse, moderately long, and erect (Fig.
<figureCitation captionStart="Figure 45" captionStartId="F45" captionText="Figure 45. Pheidole midongy sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923275) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0119419)." figureDoi="10.3897/zookeys.949.51269.figure45" httpUri="https://binary.pensoft.net/fig/432214" pageId="0" pageNumber="1">45D, F</figureCitation>
).
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Petiole.</emphasis>
Shiny with dense foveolae; node finely foveolate, triangular, with rounded and thin apex, in rear view node dorsoventrally convex; pilosity moderately sparse and erect (Fig.
<figureCitation captionStart="Figure 45" captionStartId="F45" captionText="Figure 45. Pheidole midongy sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923275) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0119419)." figureDoi="10.3897/zookeys.949.51269.figure45" httpUri="https://binary.pensoft.net/fig/432214" pageId="0" pageNumber="1">45D, F</figureCitation>
).
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Postpetiole.</emphasis>
Shiny and foveolate; dorsum with reduced sculpture; in dorsal view oval, lateral margins medially with two dentate projections; pilosity long, moderately sparse, and erect (Fig.
<figureCitation captionStart="Figure 45" captionStartId="F45" captionText="Figure 45. Pheidole midongy sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923275) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0119419)." figureDoi="10.3897/zookeys.949.51269.figure45" httpUri="https://binary.pensoft.net/fig/432214" pageId="0" pageNumber="1">45D, F</figureCitation>
).
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Gaster.</emphasis>
Shiny and smooth; pilosity moderately dense, moderately short, and erect (Fig.
<figureCitation captionStart="Figure 45" captionStartId="F45" captionText="Figure 45. Pheidole midongy sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923275) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0119419)." figureDoi="10.3897/zookeys.949.51269.figure45" httpUri="https://binary.pensoft.net/fig/432214" pageId="0" pageNumber="1">45D, F</figureCitation>
).
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Colour.</emphasis>
Blackish brown with yellow legs and antenna (Fig.
<figureCitation captionStart="Figure 45" captionStartId="F45" captionText="Figure 45. Pheidole midongy sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923275) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0119419)." figureDoi="10.3897/zookeys.949.51269.figure45" httpUri="https://binary.pensoft.net/fig/432214" pageId="0" pageNumber="1">45D, F</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Minor workers.</emphasis>
Measurements (
<emphasis italics="true" pageId="0" pageNumber="1">N</emphasis>
= 1): HL: 0.48; HW: 0.42; SL: 0.44; EL: 0.09; WL: 0.57; PSL: 0.05; MTL: 0.34; PNW: 0.29; PTW: 0.06; PPW: 0.12; CI: 114.8; SI: 105.7; PSLI: 11.0; PPI: 50.8; PNI: 70.0; MTI: 81.4.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Head.</emphasis>
Cephalic margin slightly concave (Fig.
<figureCitation captionStart="Figure 45" captionStartId="F45" captionText="Figure 45. Pheidole midongy sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923275) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0119419)." figureDoi="10.3897/zookeys.949.51269.figure45" httpUri="https://binary.pensoft.net/fig/432214" pageId="0" pageNumber="1">45A</figureCitation>
). Pilosity relatively sparse, moderately long, decumbent to subdecumbent. Sculpture shiny; frons with sparse and moderately thick, short, and predominantly longitudinal rugulae fading medially; vertex with transverse, sparse, moderately thick, and short rugulae; area posterolateral from eyes smooth; antennal sockets with few thick, curved outward rugae and smooth interspaces. Clypeus with median longitudinal carina absent; two lateral longitudinal carinae absent. Scape, when laid back, exceeding the posterior head margin by less than one-fifth of its length; pilosity dense, subdecumbent to erect (Fig.
<figureCitation captionStart="Figure 45" captionStartId="F45" captionText="Figure 45. Pheidole midongy sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923275) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0119419)." figureDoi="10.3897/zookeys.949.51269.figure45" httpUri="https://binary.pensoft.net/fig/432214" pageId="0" pageNumber="1">45A, C</figureCitation>
).
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Mesosoma.</emphasis>
In lateral view, promesonotum moderately high and short, arched; promesonotal groove absent; metanotal groove distinct; propodeal spines reduced to small tubercles (Fig.
<figureCitation captionStart="Figure 45" captionStartId="F45" captionText="Figure 45. Pheidole midongy sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923275) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0119419)." figureDoi="10.3897/zookeys.949.51269.figure45" httpUri="https://binary.pensoft.net/fig/432214" pageId="0" pageNumber="1">45C</figureCitation>
). Sculpture smooth; promesonotal dorsum with very sparse, short, and transverse rugulae; propodeum with indistinct and irregular, sparse rugae. Pilosity sparse, moderately long, and erect (Fig.
<figureCitation captionStart="Figure 45" captionStartId="F45" captionText="Figure 45. Pheidole midongy sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923275) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0119419)." figureDoi="10.3897/zookeys.949.51269.figure45" httpUri="https://binary.pensoft.net/fig/432214" pageId="0" pageNumber="1">45C, E</figureCitation>
).
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Gaster.</emphasis>
With sparse, erect pilosity (Fig.
<figureCitation captionStart="Figure 45" captionStartId="F45" captionText="Figure 45. Pheidole midongy sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923275) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0119419)." figureDoi="10.3897/zookeys.949.51269.figure45" httpUri="https://binary.pensoft.net/fig/432214" pageId="0" pageNumber="1">45C, E</figureCitation>
).
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Colour.</emphasis>
Dark yellow (Fig.
<figureCitation captionStart="Figure 45" captionStartId="F45" captionText="Figure 45. Pheidole midongy sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923275) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0119419)." figureDoi="10.3897/zookeys.949.51269.figure45" httpUri="https://binary.pensoft.net/fig/432214" pageId="0" pageNumber="1">45C, E</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="etymology">
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="1">From the type locality.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="biology_ecology">
<paragraph pageId="0" pageNumber="1">Biology.</paragraph>
<paragraph pageId="0" pageNumber="1">The species was collected at 940 m in elevation, in rainforest. Nest was located in a rotten log.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="comments">
<paragraph pageId="0" pageNumber="1">Comments.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole midongy" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="midongy">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole midongy</emphasis>
</taxonomicName>
sp. nov. belongs to the group of species characterised by small body size (major workers: HL &lt;1.05 mm, WL &lt;0.9 mm and minor workers HL &lt;0.5 mm, WL &lt;0.6 mm), sub-oval, slightly widening posteriorly head with anterior and posterior sides convex in major workers, and minor workers with yellow to brown body colouration and head foveolate or predominantly smooth and relatively oval. The group includes six species:
<taxonomicName lsidName="P. havoana" pageId="0" pageNumber="1" rank="species" species="havoana">
<emphasis italics="true" pageId="0" pageNumber="1">P. havoana</emphasis>
</taxonomicName>
sp. nov.,
<taxonomicName lsidName="P. kely" pageId="0" pageNumber="1" rank="species" species="kely">
<emphasis italics="true" pageId="0" pageNumber="1">P. kely</emphasis>
</taxonomicName>
sp. nov.,
<taxonomicName lsidName="P. parvula" pageId="0" pageNumber="1" rank="species" species="parvula">
<emphasis italics="true" pageId="0" pageNumber="1">P. parvula</emphasis>
</taxonomicName>
sp. nov.,
<taxonomicName lsidName="P. parvulogibba" pageId="0" pageNumber="1" rank="species" species="parvulogibba">
<emphasis italics="true" pageId="0" pageNumber="1">P. parvulogibba</emphasis>
</taxonomicName>
sp. nov.,
<taxonomicName lsidName="P. volontany" pageId="0" pageNumber="1" rank="species" species="volontany">
<emphasis italics="true" pageId="0" pageNumber="1">P. volontany</emphasis>
</taxonomicName>
sp. nov., and
<taxonomicName lsidName="P. midongy" pageId="0" pageNumber="1" rank="species" species="midongy">
<emphasis italics="true" pageId="0" pageNumber="1">P. midongy</emphasis>
</taxonomicName>
sp. nov. Because of dark body colouration
<taxonomicName lsidName="P. midongy" pageId="0" pageNumber="1" rank="species" species="midongy">
<emphasis italics="true" pageId="0" pageNumber="1">P. midongy</emphasis>
</taxonomicName>
sp. nov., described form Parc National Befotaka-Midongy in Fianarantsoa, is most similar to
<taxonomicName lsidName="P. volontany" pageId="0" pageNumber="1" rank="species" species="volontany">
<emphasis italics="true" pageId="0" pageNumber="1">P. volontany</emphasis>
</taxonomicName>
sp. nov., known from
<normalizedToken originalValue="Forêt">Foret</normalizedToken>
<normalizedToken originalValue="Classée">Classee</normalizedToken>
<normalizedToken originalValue="d’Analavelona">d'Analavelona</normalizedToken>
in Toliara. Major workers of
<taxonomicName lsidName="P. midongy" pageId="0" pageNumber="1" rank="species" species="midongy">
<emphasis italics="true" pageId="0" pageNumber="1">P. midongy</emphasis>
</taxonomicName>
sp. nov. differ from
<taxonomicName lsidName="P. volontany" pageId="0" pageNumber="1" rank="species" species="volontany">
<emphasis italics="true" pageId="0" pageNumber="1">P. volontany</emphasis>
</taxonomicName>
sp. nov. in frons with thick, longitudinal, and dense rugae and distinctly rugulate interspaces, and sides of the head with moderately dense, moderately short, and suberect pilosity; minor workers differ in frons with sparse and moderately thick, short, and predominantly longitudinal rugulae, vertex with transverse, sparse, and moderately thick short rugulae, oval head shape, and brighter body colouration.
</paragraph>
</subSubSection>
</treatment>
</document>
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