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<document id="4B45DE36C2A82096E00EFF8508DB7476" ID-DOI="10.11646/zootaxa.4000.1.1" ID-GBIF-Dataset="507a3784-35f5-46f5-9492-e0884a502f40" ID-ISSN="1175-5326" ID-Zenodo-Dep="289219" ID-ZooBank="00769C76-A2D4-4792-A5DD-5F2B7EA352CB" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461184674989" checkinUser="plazi" docAuthor="Allegro, Gianni &amp; Giachino, Pier Mauro" docDate="2015" docId="039087C7FFA4FF8DE28C4E0F343A689B" docLanguage="en" docName="zt04000p048.pdf" docOrigin="Zootaxa 4000 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Blennidus (Agraphoderus) negrei Straneo 1993" docType="treatment" docVersion="6" lastPageNumber="12" masterDocId="FFA9FFBFFFAFFF86E21B4B7535296F7A" masterDocTitle="Annotated checklist of the Blennidus subgenus Agraphoderus species from Peru with description of B. bombonensis n. sp. and synonymic notes (Coleoptera: Carabidae: Pterostichinae)" masterLastPageNumber="48" masterPageNumber="1" pageNumber="12" updateTime="1698599231025" updateUser="plazi">
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<mods:title id="ABBEF0B5E26A953D27D8CFAB64E90222">Annotated checklist of the Blennidus subgenus Agraphoderus species from Peru with description of B. bombonensis n. sp. and synonymic notes (Coleoptera: Carabidae: Pterostichinae)</mods:title>
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<mods:namePart id="FAD0C9591601038DFBE6B93FB215982D">Allegro, Gianni</mods:namePart>
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<mods:namePart id="2F4451872030E590AF8FC251298EAB07">Giachino, Pier Mauro</mods:namePart>
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<treatment id="039087C7FFA4FF8DE28C4E0F343A689B" ID-DOI="http://doi.org/10.5281/zenodo.6096051" ID-GBIF-Taxon="127692759" ID-Zenodo-Dep="6096051" LSID="urn:lsid:plazi:treatment:039087C7FFA4FF8DE28C4E0F343A689B" httpUri="http://treatment.plazi.org/id/039087C7FFA4FF8DE28C4E0F343A689B" lastPageNumber="12" pageId="11" pageNumber="12">
<subSubSection id="C323655AFFA4FF8DE28C4E0F344C6ACC" pageId="11" pageNumber="12" type="nomenclature">
<paragraph id="8B8636D1FFA4FF8DE28C4E0F37DB6AEF" blockId="11.[151,754,1402,1462]" box="[151,754,1402,1429]" pageId="11" pageNumber="12">
<heading id="D0CE81BDFFA4FF8DE28C4E0F37DB6AEF" bold="true" box="[151,754,1402,1429]" fontSize="11" level="1" pageId="11" pageNumber="12" reason="1">
<taxonomicName id="4C394D52FFA4FF8DE28C4E0F37DB6AEF" ID-CoL="M55Q" authority="Straneo, 1993" authorityName="Straneo" authorityYear="1993" box="[151,754,1402,1429]" class="Insecta" family="Carabidae" genus="Blennidus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="negrei" subGenus="Agraphoderus">
<emphasis id="B94DEAC3FFA4FF8DE28C4E0F37DB6AEF" bold="true" box="[151,754,1402,1429]" pageId="11" pageNumber="12">
<emphasis id="B94DEAC3FFA4FF8DE28C4E0F343A6AEE" bold="true" box="[151,275,1402,1428]" italics="true" pageId="11" pageNumber="12">Blennidus</emphasis>
(
<emphasis id="B94DEAC3FFA4FF8DE33F4E0F34F96AEE" bold="true" box="[292,464,1402,1428]" italics="true" pageId="11" pageNumber="12">Agraphoderus</emphasis>
)
<emphasis id="B94DEAC3FFA4FF8DE3FA4E0F37076AEE" bold="true" box="[481,558,1402,1428]" italics="true" pageId="11" pageNumber="12">negrei</emphasis>
(
<bibRefCitation id="EFA84B20FFA4FF8DE0264E0E37C36AEF" author="Straneo" box="[573,746,1403,1429]" pageId="11" pageNumber="12" refString="Straneo, S. L. (1993) Nuove specie del genere Ogmopleura Tschitscherine (Coleoptera, Carabidae, Pterostichinae) del Peru e dell'Ecuador e chiave per la loro determinazione. Annali del Museo civico di Storia naturale &quot; G. Doria &quot;, 89, 351 - 399." type="journal article" year="1993">Straneo, 1993</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8B8636D1FFA4FF8DE28C4EE8344C6ACC" blockId="11.[151,754,1402,1462]" box="[151,357,1437,1462]" pageId="11" pageNumber="12">
(
<figureCitation id="13022A54FFA4FF8DE2844EE835D56ACC" box="[159,252,1437,1462]" captionStart="FIGURES 28 31" captionStartId="31.[151,264,1953,1976]" captionTargetBox="[223,1375,207,1911]" captionTargetId="figure@31.[193,1393,193,1932]" captionTargetPageId="31" captionText="FIGURES 28 31. Habitus of Blennidus: longiloba male HT (28); mateui male HT (29); mesotibialis male HT (30); negrei male HT (31)." httpUri="https://zenodo.org/record/289227/files/figure.png" pageId="11" pageNumber="12">Figs. 31</figureCitation>
,
<figureCitation id="13022A54FFA4FF8DE3134EE8340F6ACC" box="[264,294,1437,1462]" captionStart="FIGURES 66 73" captionStartId="39.[151,264,1916,1939]" captionTargetBox="[161,1422,202,1884]" captionTargetId="figure@39.[151,1436,193,1895]" captionTargetPageId="39" captionText="FIGURES 66 73. Pronotum of Blennidus: longiloba male HT (66); mateui male HT (67); mesotibialis male HT (68); negrei male HT (69); nigritulus male HT (70); olivaceus male HT (71); orbicollis male PT (72); pachycerus male HT (73)." httpUri="https://zenodo.org/record/289234/files/figure.png" pageId="11" pageNumber="12">69</figureCitation>
,
<figureCitation id="13022A54FFA4FF8DE3294EE834746ACC" box="[306,349,1437,1462]" captionStart="FIGURES 100 105" captionStartId="44.[151,264,1953,1976]" captionTargetBox="[236,1352,219,1908]" captionTargetId="figure@44.[217,1370,193,1932]" captionTargetPageId="44" captionText="FIGURES 100 105. Aedeagus (left lateral and dorsal aspects) of Blennidus: jelskii LT (100); languens LT (101); longiloba HT (102); mateui HT (103); mesotibialis PT (104); negrei HT (105)." httpUri="https://zenodo.org/record/289239/files/figure.png" pageId="11" pageNumber="12">105</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C323655AFFA4FF8DE28C4E903612692D" pageId="11" pageNumber="12" type="reference_group">
<paragraph id="8B8636D1FFA4FF8DE28C4E90370B6A81" blockId="11.[151,827,1509,1623]" box="[151,546,1509,1531]" pageId="11" pageNumber="12">
<treatmentCitationGroup id="AB2911FFFFA4FF8DE28C4E90370B6A81" box="[151,546,1509,1531]" pageId="11" pageNumber="12">
<taxonomicName id="4C394D52FFA4FF8DE28C4E90370B6A81" ID-CoL="48Z6Z" authority="Straneo, 1993: 397" authorityName="Straneo" authorityPageNumber="397" authorityYear="1993" box="[151,546,1509,1531]" class="Insecta" family="Carabidae" genus="Ogmopleura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="negrei">
<emphasis id="B94DEAC3FFA4FF8DE28C4E9034746A81" box="[151,349,1509,1531]" italics="true" pageId="11" pageNumber="12">Ogmopleura negrei</emphasis>
<treatmentCitation id="0A9810C0FFA4FF8DE3784E90370B6A81" author="Straneo" box="[355,546,1509,1531]" page="397" pageId="11" pageNumber="12" year="1993">
<bibRefCitation id="EFA84B20FFA4FF8DE3784E9034DA6A81" author="Straneo" box="[355,499,1509,1531]" pageId="11" pageNumber="12" refString="Straneo, S. L. (1993) Nuove specie del genere Ogmopleura Tschitscherine (Coleoptera, Carabidae, Pterostichinae) del Peru e dell'Ecuador e chiave per la loro determinazione. Annali del Museo civico di Storia naturale &quot; G. Doria &quot;, 89, 351 - 399." type="journal article" year="1993">Straneo, 1993</bibRefCitation>
: 397
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</paragraph>
<paragraph id="8B8636D1FFA4FF8DE28C4D713612692D" blockId="11.[151,827,1509,1623]" pageId="11" pageNumber="12">
<treatmentCitationGroup id="AB2911FFFFA4FF8DE28C4D713612692D" pageId="11" pageNumber="12">
<taxonomicName id="4C394D52FFA4FF8DE28C4D7134C06960" ID-CoL="M55Q" authority="Straneo, 1993" authorityName="Straneo" authorityYear="1993" box="[151,489,1540,1562]" class="Insecta" family="Carabidae" genus="Blennidus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="negrei">
<emphasis id="B94DEAC3FFA4FF8DE28C4D71346B6960" box="[151,322,1540,1562]" italics="true" pageId="11" pageNumber="12">Blennidus negrei</emphasis>
(
<bibRefCitation id="EFA84B20FFA4FF8DE3544D7134C86960" author="Straneo" box="[335,481,1540,1562]" pageId="11" pageNumber="12" refString="Straneo, S. L. (1993) Nuove specie del genere Ogmopleura Tschitscherine (Coleoptera, Carabidae, Pterostichinae) del Peru e dell'Ecuador e chiave per la loro determinazione. Annali del Museo civico di Storia naturale &quot; G. Doria &quot;, 89, 351 - 399." type="journal article" year="1993">Straneo, 1993</bibRefCitation>
)
</taxonomicName>
:
<treatmentCitation id="0A9810C0FFA4FF8DE3E84D7037916960" author="Lorenz" box="[499,696,1540,1562]" page="263" pageId="11" pageNumber="12" year="2005">
<bibRefCitation id="EFA84B20FFA4FF8DE3E84D7037A06960" author="Lorenz" box="[499,649,1540,1562]" pageId="11" pageNumber="12" refString="Lorenz, W. (2005 a) Systematic list of extant ground beetles of the world (Insecta Coleoptera ' Geadephaga': Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). Second Edition. Published by the author, Tutzing, 530 pp." type="book" year="2005" yearSuffix="a">Lorenz, 2005a</bibRefCitation>
: 263
</treatmentCitation>
<taxonomicName id="4C394D52FFA4FF8DE28C4D5734C06942" ID-CoL="M55Q" authority="Straneo, 1993" authorityName="Straneo" authorityYear="1993" box="[151,489,1570,1592]" class="Insecta" family="Carabidae" genus="Blennidus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="negrei">
<emphasis id="B94DEAC3FFA4FF8DE28C4D57346B6942" box="[151,322,1570,1592]" italics="true" pageId="11" pageNumber="12">Blennidus negrei</emphasis>
(
<bibRefCitation id="EFA84B20FFA4FF8DE3544D5734C86942" author="Straneo" box="[335,481,1570,1592]" pageId="11" pageNumber="12" refString="Straneo, S. L. (1993) Nuove specie del genere Ogmopleura Tschitscherine (Coleoptera, Carabidae, Pterostichinae) del Peru e dell'Ecuador e chiave per la loro determinazione. Annali del Museo civico di Storia naturale &quot; G. Doria &quot;, 89, 351 - 399." type="journal article" year="1993">Straneo, 1993</bibRefCitation>
)
</taxonomicName>
:
<treatmentCitation id="0A9810C0FFA4FF8DE3E84D5637936942" author="Lorenz" box="[499,698,1570,1592]" page="672" pageId="11" pageNumber="12" year="2005">
<bibRefCitation id="EFA84B20FFA4FF8DE3E84D5637A36942" author="Lorenz" box="[499,650,1570,1592]" pageId="11" pageNumber="12" refString="Lorenz, W. (2005 b) Nomina Carabidarum. A directory of the scientific names of ground beetles (Insecta Coleoptera ' Geadephaga': Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). 2 nd Edition. Published by the author, Tutzing, 993 pp." type="book" year="2005" yearSuffix="b">Lorenz, 2005b</bibRefCitation>
: 672
</treatmentCitation>
<taxonomicName id="4C394D52FFA4FF8DE28C4D3434C0692D" ID-CoL="M55Q" authority="Straneo, 1993" authorityName="Straneo" authorityYear="1993" box="[151,489,1601,1623]" class="Insecta" family="Carabidae" genus="Blennidus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="negrei">
<emphasis id="B94DEAC3FFA4FF8DE28C4D34346B692D" box="[151,322,1601,1623]" italics="true" pageId="11" pageNumber="12">Blennidus negrei</emphasis>
(
<bibRefCitation id="EFA84B20FFA4FF8DE3544D3434C8692D" author="Straneo" box="[335,481,1601,1623]" pageId="11" pageNumber="12" refString="Straneo, S. L. (1993) Nuove specie del genere Ogmopleura Tschitscherine (Coleoptera, Carabidae, Pterostichinae) del Peru e dell'Ecuador e chiave per la loro determinazione. Annali del Museo civico di Storia naturale &quot; G. Doria &quot;, 89, 351 - 399." type="journal article" year="1993">Straneo, 1993</bibRefCitation>
)
</taxonomicName>
:
<treatmentCitation id="0A9810C0FFA4FF8DE3E84D343612692D" author="Allegro" box="[499,827,1601,1623]" page="173" pageId="11" pageNumber="12" year="2011">
<bibRefCitation id="EFA84B20FFA4FF8DE3E84D343625692D" author="Allegro" box="[499,780,1601,1623]" pageId="11" pageNumber="12" refString="Allegro, G. &amp; Giachino, P. M. (2011 b) Studies on the Blennidus Motschulsky subgen. Agraphoderus Bates from Peru: the orbicollis species group (Coleoptera: Carabidae: Pterostichini). Veroeffentlichungen des Naturkunde Museums Erfurt, 30, 167 - 173." type="journal article" year="2011" yearSuffix="b">Allegro &amp; Giachino, 2011b</bibRefCitation>
: 173
</treatmentCitation>
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</paragraph>
</subSubSection>
<subSubSection id="C323655AFFA4FF8DE28C4DF131FE687D" pageId="11" pageNumber="12" type="materials_examined">
<paragraph id="8B8636D1FFA4FF8DE28C4DF137EC69E6" blockId="11.[151,1436,1668,2017]" box="[151,709,1668,1693]" pageId="11" pageNumber="12">
<emphasis id="B94DEAC3FFA4FF8DE28C4DF1341C69E7" bold="true" box="[151,309,1668,1693]" pageId="11" pageNumber="12">
<typeStatus id="54828873FFA4FF8DE28C4DF135FB69E7" box="[151,210,1668,1693]" pageId="11" pageNumber="12">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F32E7641FFA4FF8DE3274DF0345B69E6" box="[316,370,1669,1692]" name="Peru" pageId="11" pageNumber="12">Peru</collectingCountry>
, Cordillera Cumullca,
<quantity id="4CC19B34FFA4FF8DE06A4DF137EC69E6" box="[625,709,1668,1693]" metricMagnitude="3" metricUnit="m" metricValue="3.6" pageId="11" pageNumber="12" unit="m" value="3600.0">3600 m</quantity>
</paragraph>
<paragraph id="8B8636D1FFA4FF8DE2DC4DD2361E69BA" blockId="11.[151,1436,1668,2017]" box="[199,823,1703,1728]" pageId="11" pageNumber="12">
<emphasis id="B94DEAC3FFA4FF8DE2DC4DD2349A69BA" bold="true" box="[199,435,1703,1728]" pageId="11" pageNumber="12">Examined material.</emphasis>
HT ♂ (CSt at
<collectionCode id="ED28AE14FFA4FF8DE0434DDD379B69BA" box="[600,690,1704,1728]" pageId="11" pageNumber="12">MSNM</collectionCode>
);
<specimenCount id="9D3FFD58FFA4FF8DE0DE4DDD37C669C5" box="[709,751,1704,1728]" pageId="11" pageNumber="12" type="generic">1 ♂</specimenCount>
(CAl)
</paragraph>
<paragraph id="8B8636D1FFA4FF8DE2DC4DB931FE687D" blockId="11.[151,1436,1668,2017]" pageId="11" pageNumber="12">
<emphasis id="B94DEAC3FFA4FF8DE2DC4DB9342F699F" bold="true" box="[199,262,1740,1765]" pageId="11" pageNumber="12">Note.</emphasis>
Species distinguished by a robust subapical inner spine on ♂ mesotibiae and by the morphology of aedeagus, which appears to relate it to
<taxonomicName id="4C394D52FFA4FF8DE0524D8437986872" box="[585,689,1776,1800]" class="Insecta" family="Carabidae" genus="Blennidus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="bellesi">
<emphasis id="B94DEAC3FFA4FF8DE0524D8437986872" box="[585,689,1776,1800]" italics="true" pageId="11" pageNumber="12">B. bellesi</emphasis>
</taxonomicName>
, a species that lives in the same geographic area.
</paragraph>
</subSubSection>
<subSubSection id="C323655AFFA4FF8DE2DC4C61343A689B" pageId="11" pageNumber="12" type="distribution">
<paragraph id="8B8636D1FFA4FF8DE2DC4C6134B4680E" blockId="11.[151,1436,1668,2017]" pageId="11" pageNumber="12">
<emphasis id="B94DEAC3FFA4FF8DE2DC4C6134496857" bold="true" box="[199,352,1812,1837]" pageId="11" pageNumber="12">Distribution.</emphasis>
The two recorded specimens are from the Andes of northern
<collectingCountry id="F32E7641FFA4FF8DE60B4C60316F6856" box="[1040,1094,1813,1836]" name="Peru" pageId="11" pageNumber="12">Peru</collectingCountry>
, Dept. Cajamarca. Besides the HT (legit Weyroud 1947), this species is known from one specimen recently collected in Cajamarca, Cumbe Mayo (legit G. Allegro 2008).
</paragraph>
<paragraph id="8B8636D1FFA4FF8DE2DC4C0A343A689B" blockId="11.[151,1436,1668,2017]" pageId="11" pageNumber="12">
<emphasis id="B94DEAC3FFA4FF8DE2DC4C0A340368E2" bold="true" box="[199,298,1919,1944]" pageId="11" pageNumber="12">Habitat.</emphasis>
The recent capture of a specimen under a stone in Andean grassland at
<quantity id="4CC19B34FFA4FF8DE6574C0A318168ED" box="[1100,1192,1919,1944]" metricMagnitude="3" metricUnit="m" metricValue="3.75" pageId="11" pageNumber="12" unit="m" value="3750.0">3,750 m</quantity>
a.s.l. (Cumbe Mayo), as well as the data reported on the HT label, indicate a likely preference of this species for the high altitude grasslands.
</paragraph>
</subSubSection>
</treatment>
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