648 lines
81 KiB
XML
648 lines
81 KiB
XML
<document id="A3A7847C607CB045D3873633005E87A3" ID-CLB-Dataset="58039" ID-DOI="10.1111/syen.12590" ID-GBIF-Dataset="bfb878f3-8a74-46d3-a104-36485c32aaba" ID-Zenodo-Dep="7909395" IM.bibliography_approvedBy="admin" IM.illustrations_approvedBy="admin" IM.materialsCitations_approvedBy="tatiana" IM.metadata_approvedBy="tatiana" IM.tables_approvedBy="julia" IM.taxonomicNames_approvedBy="admin" IM.treatments_approvedBy="admin" checkinTime="1683588024562" checkinUser="julia" docAuthor="Espeland, Marianne, Nakahara, Shinichi, Zacca, Thamara, Barbosa, Eduardo P., Huertas, Blanca, Marín, Mario A., Lamas, Gerardo, Benmesbah, Mohamed, Brévignon, Christian, Casagrande, Mirna M., Fåhraeus, Christer, Grishin, Nick, Kawahara, Akito Y., Mielke, Olaf H. H., Miller, Jacqueline Y., Nakamura, Ichiro, Navas, Vanessa, Patrusky, Brooke, Pyrcz, Tomasz W., Richards, Lindsay, Tan, Denise, Tyler, Stephanie, Viloria, Angel, Warren, Andrew D., Xiao, Lei, Freitas, André V. L. & Willmott, Keith R." docDate="2023" docId="03EC879FFF9EFFE1A875A9DEFB681698" docLanguage="en" docName="SystEntomol.2023.1-73.pdf" docOrigin="Zoological Research 2023" docTitle="Pareuptychia Forster 1964" docType="treatment" docVersion="11" lastPageNumber="18" masterDocId="FFD5FFE7FF8FFFF0A80FAB7BFFAB1055" masterDocTitle="Combining target enrichment and Sanger sequencing data to clarify the systematics of the diverse Neotropical butterfly subtribe Euptychiina (Nymphalidae, Satyrinae)" masterLastPageNumber="73" masterPageNumber="1" pageNumber="18" updateTime="1684963631062" updateUser="ExternalLinkService">
|
||
<mods:mods id="69319E2D7E3C911AE6C95A79656AA3F4" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="7378ACB8950C67BAEB8BD8693F7FA698">
|
||
<mods:title id="D45ABBA941182F42484724A7B0CAB55C">Combining target enrichment and Sanger sequencing data to clarify the systematics of the diverse Neotropical butterfly subtribe Euptychiina (Nymphalidae, Satyrinae)</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="CE70134E0AEBB14140ED6FA50EC4ED54" type="personal">
|
||
<mods:role id="95913A750D9309C294F39B0CA4877E92">
|
||
<mods:roleTerm id="6366AACE8691609EA4D181C7655AA782">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="F81ED719362BCA6A5EA6D0FEF6B0067B">Espeland, Marianne</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="C8A83264E2236DB0AF0784655837462E" type="personal">
|
||
<mods:role id="2D00BFF3DFE53B441827348BF7A84010">
|
||
<mods:roleTerm id="DA70539E46A7C2E75EDACA2D2D7B4DF0">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="D6E612BC4D1FEF93F228D5B2F6B9AA3C">Nakahara, Shinichi</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="DBA6F5EC61B3C111CF70BE413F7C6CC3" type="personal">
|
||
<mods:role id="CF07044DBF586ABA2A51382B38E5E7E2">
|
||
<mods:roleTerm id="9602C0F29C86998F0CCAF6980491E878">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="FFC1CBB294E305FC796458A676DEA104">Zacca, Thamara</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="F674069299A0DFF05646E66CD081D6D0" type="personal">
|
||
<mods:role id="7215A973879C162E8D75CE99173A3E0E">
|
||
<mods:roleTerm id="E2EC8DBE7DCBF3ED5BC6399E24B7D03D">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="EAF2A43031821240BD2C338C99DD4E4A">Barbosa, Eduardo P.</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="466DF713C99024C94262222CA41E0E3F" type="personal">
|
||
<mods:role id="B557D3743C53BBB6612C38EBE98CAA6F">
|
||
<mods:roleTerm id="907AB42DC82DAF177DAE20A2FFBA2670">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="764751D2E10F89FA8080155A88C3A45D">Huertas, Blanca</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="24D5CACED8F2F5E5CA97109070EAD329" type="personal">
|
||
<mods:role id="27932F4B9C7BD99ABD1F3292CEB2D4F7">
|
||
<mods:roleTerm id="7891DCB7181F9844D8D4820896697AD2">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="9675FCEB3D8EA067AD8A44BAA9775924">Marín, Mario A.</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="D9772D562C2A44406FE7A91DC8FC66D4" type="personal">
|
||
<mods:role id="0302CAB7B2D9A775107A344C9A848EF1">
|
||
<mods:roleTerm id="1CC69E2F3C80682F1CBA6C663CD8DB32">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="BD6EB0D73BE08C07879D837BE241A3DD">Lamas, Gerardo</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="B23ADB0856BA236CB4C35B8584B97FBD" type="personal">
|
||
<mods:role id="DE273D08E3650855C66F18180FDF8542">
|
||
<mods:roleTerm id="D8782EDCC74017763BCA96B54BB0CBF7">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="75BF5D5067AB4540622693056A5803E5">Benmesbah, Mohamed</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="EFBDBA942A6BFA6FF93CF0565A9F6693" type="personal">
|
||
<mods:role id="D1AFA0F5F5AE35E1AFAA9E23E7B33BAD">
|
||
<mods:roleTerm id="27ED06813B0D0C00CA11D1843E73A99B">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="B2E9E8BC5C453E015CADD9CAA7699077">Brévignon, Christian</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="06F02799A37BCDA7FC48AAAC240BBC3A" type="personal">
|
||
<mods:role id="9345DF699C49A476B0CD52C1250DD114">
|
||
<mods:roleTerm id="8468F17A55FD9F5E40F7F92ABECFF32E">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="56B9D383FA9E9260A3CEA9BA9821516A">Casagrande, Mirna M.</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="B7B3EE3131C121AF81F22AE64F0768D2" type="personal">
|
||
<mods:role id="0570CB5CF93CA04D35EA555836200CB8">
|
||
<mods:roleTerm id="554814C4F9357E2AA7FCE8F81CDDFDDB">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="6BA6EC5EDC283240B731A7B04BBA20FC">Fåhraeus, Christer</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="78687783DEDCF73C9E9A9252FD1333B1" type="personal">
|
||
<mods:role id="BC6B8E9D9F62239CEAFD8057624C3480">
|
||
<mods:roleTerm id="761D63172D97606C226EEF6AB473916A">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="942795AA82C583F23482D3EBE682968B">Grishin, Nick</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="B693A75AD15C5FFF272D67E74EED846E" type="personal">
|
||
<mods:role id="F800E7AB90DC65B739667228052F2C35">
|
||
<mods:roleTerm id="D1C402F2C56A9AFBE227435ABF6BEDBE">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="2F0E1BBAC5ADFFD7A773B37AB3647A5D">Kawahara, Akito Y.</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="6C72EAAFACE92C8EC661BDF05D5DCCDB" type="personal">
|
||
<mods:role id="9262B883BA76A3330A62F71C16A85DC8">
|
||
<mods:roleTerm id="928125B01B0409E0FF45FA3214EFB959">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="B0E49C2AFB5B4500DEAD1AB0C277425F">Mielke, Olaf H. H.</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="8B96F7444F71333E36EA18C25EFF1C0C" type="personal">
|
||
<mods:role id="640466726C129F074E8554647FE6EA93">
|
||
<mods:roleTerm id="671ACF9B6C74D3B5B4E5FF1E55BE58E0">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="B240FE2D607526D245CF1896948417E8">Miller, Jacqueline Y.</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="BC80BC2F6B50B00822BB966B7B0C5A56" type="personal">
|
||
<mods:role id="9C798AE94E145CB0CA39F400BD6EA59C">
|
||
<mods:roleTerm id="5CF010FB542D860D82DC8B3D057D2F9C">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="33247BE63BB633A9C46783615390E67B">Nakamura, Ichiro</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="F338036E7E783D6D2739D768C924AE54" type="personal">
|
||
<mods:role id="9049754F53CB9350A72FF1B24E2856D3">
|
||
<mods:roleTerm id="CF19302ADEAE0ACDB46F9ED410FC48F9">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="93563CB43ED93478A1EBA5DB1869F38A">Navas, Vanessa</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="58E7447CA6A1E6547E7B2854FE198AE7" type="personal">
|
||
<mods:role id="7BCCFDD02AD959D24AE008ED6F67C75A">
|
||
<mods:roleTerm id="EA7944A5BD2EF698CC13FB814180EB0C">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="BFB88086A485256BEEB58DDDBD2A7F35">Patrusky, Brooke</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="0A7327DED26BD0F0CD806C9A45B9E638" type="personal">
|
||
<mods:role id="BB052932A7A18C0DF20463CBA5F716C2">
|
||
<mods:roleTerm id="46DFC3201545B02854201F6A1D129214">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="14A4DC2E8B32278AF1FB7BBFBCCEEE0E">Pyrcz, Tomasz W.</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="2C1A3BB9CF90029E976B4A29C99719BA" type="personal">
|
||
<mods:role id="F67D1DA6AEAD68E4910FCAFD9420D94F">
|
||
<mods:roleTerm id="151A7A966D96F0B89492C171B3C1D4F8">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="754506AF0D16EB5B1CCAF2C89FB2FBDC">Richards, Lindsay</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="D2065961B608BCCCD5BA162464D4EB2D" type="personal">
|
||
<mods:role id="C519E89B1B1AE96779A7B6BD9CB77F3B">
|
||
<mods:roleTerm id="621B825937791D60ADBC4E67D9F4B2D1">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="17F4B796951014840201EAA17521775C">Tan, Denise</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="3DEF3BEF3036F2D63586029504EC3EA0" type="personal">
|
||
<mods:role id="42F9958C220150F8451FC0C1D32284F6">
|
||
<mods:roleTerm id="EF04992A75A7F9799808E73C4FAFC965">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="878E1647F090166BD347A1C9D9E20703">Tyler, Stephanie</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="F1AFC7CE201158EB79B70D1E3780F170" type="personal">
|
||
<mods:role id="1D4E15D9B2E4766D83F3F72BB868EC68">
|
||
<mods:roleTerm id="1E4E7546FFDF31FF88AA603254DEF4AB">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="59CE3A4D9E8109DBE30B91020BBC76BD">Viloria, Angel</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="16DE8352214E8BED82412EAD651A5E50" type="personal">
|
||
<mods:role id="933679925514462EB242907D45367F37">
|
||
<mods:roleTerm id="75D9857AE39741F43B65C10E2AA2D74D">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="1AAC3CB3DB35FBA4AB79FFCA8A060EB7">Warren, Andrew D.</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="BB6DC044688AD256785D4C2C39000A44" type="personal">
|
||
<mods:role id="EB96F94D021DFD5BF144808AC0EC5A02">
|
||
<mods:roleTerm id="360BA333619E2FC2A137C64C0195CC2E">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="ECB05A50E94D343FDA76B9C7C32A7316">Xiao, Lei</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="633FF67C7F75F6179FC36834B5F7FAA8" type="personal">
|
||
<mods:role id="DFF350F87D2F9DED05CA6E6AE5A540A9">
|
||
<mods:roleTerm id="85EA359308CFA3997421E34ED519C0E3">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="5E52A71B17A4F86DE29A653C4F378CC5">Freitas, André V. L.</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="6EC3CF6B5DFAB77B4CE695BF78103431" type="personal">
|
||
<mods:role id="7C4DEBA8A78A1BD71BB28B66D1423B8E">
|
||
<mods:roleTerm id="B43445826DB1D6D82FD54592B3A1793F">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="F452D27D043742DA912BFA793765FB2B">Willmott, Keith R.</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="D68C62B1606D590A802A2BD9A372E2EE">text</mods:typeOfResource>
|
||
<mods:relatedItem id="F3F8606F51E6C328CE66CAF44375FDD3" type="host">
|
||
<mods:titleInfo id="0D7C02BDFF2BDC68EF7958DFF3FABBCE">
|
||
<mods:title id="E90E7CA49C14F6C32913DC51CAA18EF8">Zoological Research</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="BBF123B09AAEC469A622BA540021AB48">
|
||
<mods:date id="EB78C02B0ADD5730F652FB9EA8CA94A1">2023</mods:date>
|
||
<mods:detail id="C73F39EE7826C224B05B4C7ED795E75A" type="pubDate">
|
||
<mods:number id="A0E59CD4583E0B9182E011887F164DDE">2023-02-15</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="4EA0001B085AC83EFCC83F77C2CEA40A" type="volume">
|
||
<mods:number id="B75834586A586A9DBB6140FC72AFDF67">2023</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="80043BB86E3B0B3479AF196319D2716C" unit="page">
|
||
<mods:start id="538678B7379B3181B1B81F4460BA7F59">1</mods:start>
|
||
<mods:end id="873C2F8C19F2EF101DB71EB3829C795D">73</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="10056EE559F470A16C756EA43C82511D">journal article</mods:classification>
|
||
<mods:identifier id="D472B4CC7D5D0378E9468967EC49CDD0" type="CLB-Dataset">58039</mods:identifier>
|
||
<mods:identifier id="6A53ACB4F5DC6F42AD5804848CD5426F" type="DOI">10.1111/syen.12590</mods:identifier>
|
||
<mods:identifier id="36F7340F1F056F2C2D665854DBEB38CE" type="GBIF-Dataset">bfb878f3-8a74-46d3-a104-36485c32aaba</mods:identifier>
|
||
<mods:identifier id="BAAB6B04DA7007129BE4C4183D987A88" type="Zenodo-Dep">7909395</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="03EC879FFF9EFFE1A875A9DEFB681698" ID-DOI="http://doi.org/10.5281/zenodo.7909457" ID-GBIF-Taxon="209589340" ID-Zenodo-Dep="7909457" LSID="urn:lsid:plazi:treatment:03EC879FFF9EFFE1A875A9DEFB681698" httpUri="http://treatment.plazi.org/id/03EC879FFF9EFFE1A875A9DEFB681698" lastPageNumber="18" pageId="17" pageNumber="18">
|
||
<subSubSection id="C35F6502FF9EFFE1A875A9DEFEE512EB" box="[122,334,677,702]" pageId="17" pageNumber="18" type="nomenclature">
|
||
<paragraph id="8BFA3689FF9EFFE1A875A9DEFEE512EB" blockId="17.[122,334,677,702]" box="[122,334,677,702]" pageId="17" pageNumber="18">
|
||
<heading id="D0B281E5FF9EFFE1A875A9DEFEE512EB" box="[122,334,677,702]" fontSize="10" level="3" pageId="17" pageNumber="18" reason="1">
|
||
<taxonomicName id="4C454D0AFF9EFFE1A875A9DEFEA012EB" ID-CoL="92J95" ID-ENA="270045" authority="Forster, 1964" authorityName="Forster" authorityYear="1964" box="[122,267,677,702]" class="Insecta" family="Nymphalidae" genus="Pareuptychia" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B931EA9BFF9EFFE1A875A9DEFEA012EB" box="[122,267,677,702]" italics="true" pageId="17" pageNumber="18">Pareuptychia</emphasis>
|
||
</taxonomicName>
|
||
clade
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35F6502FF9EFFE1A875A994FB681698" pageId="17" pageNumber="18" type="description">
|
||
<paragraph id="8BFA3689FF9EFFE1A875A994FF5B1526" blockId="17.[122,771,751,1949]" pageId="17" pageNumber="18">
|
||
The ‘
|
||
<taxonomicName id="4C454D0AFF9EFFE1A8A2A994FE8A1356" authority="Forster, 1964" box="[173,289,751,771]" class="Insecta" family="Nymphalidae" genus="Pareuptychia" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B931EA9BFF9EFFE1A8A2A994FE8A1356" box="[173,289,751,771]" italics="true" pageId="17" pageNumber="18">Pareuptychia</emphasis>
|
||
</taxonomicName>
|
||
clade’ was proposed first by
|
||
<bibRefCitation id="EFD44B78FF9EFFE1AA4BA994FF1D1370" author="Murray and Prowell" firstAuthor="Murray" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="17" pageNumber="18" pagination="67 - 80" part="34" refId="ref40211" refString="Murray, D. & Prowell, D. P. (2005) Molecular phylogenetics and evolutionary history of the neotropical Satyrine subtribe Euptychiina (Nymphalidae: Satyrinae). Molecular Phylogenetics and Evolution, 34, 67 - 80." title="Molecular phylogenetics and evolutionary history of the neotropical Satyrine subtribe Euptychiina (Nymphalidae: Satyrinae)" type="journal article" year="2005">Murray and Prowell (2005)</bibRefCitation>
|
||
and corroborated by increasingly comprehensive subsequent studies (
|
||
<bibRefCitation id="EFD44B78FF9EFFE1A8C8A84FFE33131D" author="Espeland" box="[199,408,820,840]" etAl="et al." firstAuthor="Espeland" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="17" pageNumber="18" pagination="116 - 124" part="131" refId="ref36456" refString="Espeland, M., Breinholt, J., Barbosa, E. P., Casagrande, M., Huertas, B., Lamas, G. et al. (2019 a) Four hundred shades of brown: higher level phylogeny of the problematic Euptychiina (Lepidoptera, Nymphalidae, Satyrinae) based on hybrid enrichment data. Molecular Phylogenetics and Evolution, 131, 116 - 124." title="Four hundred shades of brown: higher level phylogeny of the problematic Euptychiina (Lepidoptera, Nymphalidae, Satyrinae) based on hybrid enrichment data" type="journal article" year="2019" yearSuffix="a">Espeland et al., 2019a</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFD44B78FF9EFFE1A9AAA84FFDE4131D" author="Marin" box="[421,591,820,840]" etAl="et al." firstAuthor="Marin" journalOrPublisher="Systematic Entomology" pageId="17" pageNumber="18" pagination="768 - 785" part="42" refId="ref39677" refString="Marin, M. A., Pena, C., Uribe, S. I. & Freitas, A. V. L. (2017) Morphology agrees with molecular data: phylogenetic affinities of Euptychiina butterflies (Nymphalidae: Satyrinae). Systematic Entomology, 42, 768 - 785." title="Morphology agrees with molecular data: phylogenetic affinities of Euptychiina butterflies (Nymphalidae: Satyrinae)" type="journal article" year="2017">Marín et al., 2017</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFD44B78FF9EFFE1AA53A84FFD56131D" author="Pena" box="[604,765,820,840]" etAl="et al." firstAuthor="Pena" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="17" pageNumber="18" pagination="29 - 49" part="40" refId="ref42625" refString="Pena, C., Wahlberg, N., Weingartner, E., Kodandaramaiah, U., Nylin, S., Freitas, A. V. L. et al. (2006) Higher level phylogeny of Satyrinae butterflies (Lepidoptera: Nymphalidae) based on DNA sequence data. Molecular Phylogenetics and Evolution, 40, 29 - 49." title="Higher level phylogeny of Satyrinae butterflies (Lepidoptera: Nymphalidae) based on DNA sequence data" type="journal article" year="2006">Peña et al., 2006</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFD44B78FF9EFFE1A875A82CFF00133E" author="Pena" box="[122,171,855,875]" firstAuthor="Pena" journalOrPublisher="Zoologica Scripta" pageId="17" pageNumber="18" pagination="243 - 258" part="39" refId="ref42528" refString="Pena, C., Nylin, S., Freitas, A. V. L. & Wahlberg, N. (2010) Biogeographic history of the butterfly subtribe Euptychiina (Lepidoptera, Nymphalidae, Satyrinae). Zoologica Scripta, 39, 243 - 258." title="Biogeographic history of the butterfly subtribe Euptychiina (Lepidoptera, Nymphalidae, Satyrinae)" type="journal article" year="2010">2010</bibRefCitation>
|
||
). The clade contains 11 described genera,
|
||
<taxonomicName id="4C454D0AFF9EFFE1AA22A82CFD33133E" box="[557,664,855,875]" class="Insecta" family="Nymphalidae" genus="Neonympha" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B931EA9BFF9EFFE1AA22A82CFD33133E" box="[557,664,855,875]" italics="true" pageId="17" pageNumber="18">Neonympha</emphasis>
|
||
</taxonomicName>
|
||
(2 species),
|
||
<taxonomicName id="4C454D0AFF9EFFE1A875A802FF4513D8" authority="Forster, 1964" box="[122,238,889,909]" class="Insecta" family="Nymphalidae" genus="Pareuptychia" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B931EA9BFF9EFFE1A875A802FF4513D8" box="[122,238,889,909]" italics="true" pageId="17" pageNumber="18">Pareuptychia</emphasis>
|
||
</taxonomicName>
|
||
(9 species),
|
||
<taxonomicName id="4C454D0AFF9EFFE1A952A802FE6F13D8" authorityName="Forster" authorityYear="1964" box="[349,452,889,909]" class="Insecta" family="Nymphalidae" genus="Erichthodes" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B931EA9BFF9EFFE1A952A802FE6F13D8" box="[349,452,889,909]" italics="true" pageId="17" pageNumber="18">Erichthodes</emphasis>
|
||
</taxonomicName>
|
||
(monotypic),
|
||
<taxonomicName id="4C454D0AFF9EFFE1AA4FA802FD1613D8" baseAuthorityName="Janzen & Hallwachs" baseAuthorityYear="2022" box="[576,701,889,909]" class="Insecta" family="Nymphalidae" genus="Megeuptychia" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B931EA9BFF9EFFE1AA4FA802FD1613D8" box="[576,701,889,909]" italics="true" pageId="17" pageNumber="18">Megeuptychia</emphasis>
|
||
</taxonomicName>
|
||
(3 species),
|
||
<taxonomicName id="4C454D0AFF9EFFE1A8BBA8E7FE9813E5" authorityName="Forster" authorityYear="1964" box="[180,307,924,944]" class="Insecta" family="Nymphalidae" genus="Satyrotaygetis" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B931EA9BFF9EFFE1A8BBA8E7FE9813E5" box="[180,307,924,944]" italics="true" pageId="17" pageNumber="18">Satyrotaygetis</emphasis>
|
||
</taxonomicName>
|
||
(2 species),
|
||
<taxonomicName id="4C454D0AFF9EFFE1A9B7A8E7FD8613E5" authorityName="Forster" authorityYear="1964" box="[440,557,924,944]" class="Insecta" family="Nymphalidae" genus="Euptychoides" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B931EA9BFF9EFFE1A9B7A8E7FD8613E5" box="[440,557,924,944]" italics="true" pageId="17" pageNumber="18">Euptychoides</emphasis>
|
||
</taxonomicName>
|
||
(4 species),
|
||
<taxonomicName id="4C454D0AFF9EFFE1AABCA8E7FCAA13E5" baseAuthorityName="Freitas" baseAuthorityYear="2003" box="[691,769,924,944]" class="Insecta" family="Nymphalidae" genus="Taydebis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B931EA9BFF9EFFE1AABCA8E7FCAA13E5" box="[691,769,924,944]" italics="true" pageId="17" pageNumber="18">Taydebis</emphasis>
|
||
</taxonomicName>
|
||
(3 species),
|
||
<taxonomicName id="4C454D0AFF9EFFE1A8E5A8C4FECA1386" authorityName="Freitas, Barbosa & Zacca" authorityYear="2018" box="[234,353,959,979]" class="Insecta" family="Nymphalidae" genus="Nhambikuara" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B931EA9BFF9EFFE1A8E5A8C4FECA1386" box="[234,353,959,979]" italics="true" pageId="17" pageNumber="18">Nhambikuara</emphasis>
|
||
</taxonomicName>
|
||
(11 species),
|
||
<taxonomicName id="4C454D0AFF9EFFE1A9E9A8C4FDFA1386" box="[486,593,959,979]" class="Insecta" family="Nymphalidae" genus="Optimandes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B931EA9BFF9EFFE1A9E9A8C4FDFA1386" box="[486,593,959,979]" italics="true" pageId="17" pageNumber="18">Optimandes</emphasis>
|
||
</taxonomicName>
|
||
(2 species),
|
||
<taxonomicName id="4C454D0AFF9EFFE1AAC6A8C4FCAA1386" authorityName="Nakahara, Willmott & Espeland" authorityYear="2019" box="[713,769,959,979]" class="Insecta" family="Nymphalidae" genus="Amiga" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B931EA9BFF9EFFE1AAC6A8C4FCAA1386" box="[713,769,959,979]" italics="true" pageId="17" pageNumber="18">Amiga</emphasis>
|
||
</taxonomicName>
|
||
(2 species),
|
||
<taxonomicName id="4C454D0AFF9EFFE1A8E6A89AFEEB13A0" authority="Espeland1 & Nakahara & Zacca & Barbosa & Huertas & Marín & Lamas & Benmesbah & Brévignon & Casagrande & Fåhraeus & Grishin & Kawahara1 & Mielke & Miller & Nakamura & Navas & Patrusky & Pyrcz & Richards & Tan & Tyler & Viloria & Warren1 & Xiao & Freitas & Willmott, 2023" authorityName="Espeland1 & Nakahara & Zacca & Barbosa & Huertas & Marín & Lamas & Benmesbah & Brévignon & Casagrande & Fåhraeus & Grishin & Kawahara1 & Mielke & Miller & Nakamura & Navas & Patrusky & Pyrcz & Richards & Tan & Tyler & Viloria & Warren1 & Xiao & Freitas & Willmott" authorityYear="2023" box="[233,320,993,1013]" class="Insecta" family="Nymphalidae" genus="Xenovena" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus" status="gen. n.">
|
||
<emphasis id="B931EA9BFF9EFFE1A8E6A89AFEEB13A0" box="[233,320,993,1013]" italics="true" pageId="17" pageNumber="18">Xenovena</emphasis>
|
||
</taxonomicName>
|
||
gen. n. (monotypic) and an undescribed genus. The ‘
|
||
<taxonomicName id="4C454D0AFF9EFFE1A8A4AF7FFEB4144D" authority="Forster, 1964" box="[171,287,1028,1048]" class="Insecta" family="Nymphalidae" genus="Pareuptychia" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B931EA9BFF9EFFE1A8A4AF7FFEB4144D" box="[171,287,1028,1048]" italics="true" pageId="17" pageNumber="18">Pareuptychia</emphasis>
|
||
</taxonomicName>
|
||
clade’ is embedded within a larger clade also containing the ‘
|
||
<taxonomicName id="4C454D0AFF9EFFE1A8E6AF5CFE98146E" authorityName="Hubner" authorityYear="1819" box="[233,307,1063,1083]" class="Insecta" family="Nymphalidae" genus="Taygetis" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B931EA9BFF9EFFE1A8E6AF5CFE98146E" box="[233,307,1063,1083]" italics="true" pageId="17" pageNumber="18">Taygetis</emphasis>
|
||
</taxonomicName>
|
||
clade’, ‘
|
||
<taxonomicName id="4C454D0AFF9EFFE1A98FAF5CFDBA146E" authorityName="Forster" authorityYear="1964" box="[384,529,1063,1083]" class="Insecta" family="Nymphalidae" genus="Splendeuptychia" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B931EA9BFF9EFFE1A98FAF5CFDBA146E" box="[384,529,1063,1083]" italics="true" pageId="17" pageNumber="18">Splendeuptychia</emphasis>
|
||
</taxonomicName>
|
||
clade’ and ‘
|
||
<taxonomicName id="4C454D0AFF9EFFE1AA8DAF5CFCAA146E" authorityName="Forster" authorityYear="1964" box="[642,769,1063,1083]" class="Insecta" family="Nymphalidae" genus="Archeuptychia" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B931EA9BFF9EFFE1AA8DAF5CFCAA146E" box="[642,769,1063,1083]" italics="true" pageId="17" pageNumber="18">Archeuptychia</emphasis>
|
||
</taxonomicName>
|
||
clade’, and we also include the unplaced genus
|
||
<taxonomicName id="4C454D0AFF9EFFE1AA31AF32FD681408" authorityName="Forster" authorityYear="1964" box="[574,707,1097,1117]" class="Insecta" family="Nymphalidae" genus="Chloreuptychia" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B931EA9BFF9EFFE1AA31AF32FD681408" box="[574,707,1097,1117]" italics="true" pageId="17" pageNumber="18">Chloreuptychia</emphasis>
|
||
</taxonomicName>
|
||
in the discussions here (
|
||
<figureCitation id="137E2A0CFF9EFFE1A915AF17FE0714D5" box="[282,428,1132,1152]" captionStart="FIGURE 1" captionStartId="5.[122,218,1354,1373]" captionTargetBox="[155,1432,126,1307]" captionTargetId="graphics-326@5.[251,1339,165,1253]" captionText="FIGURE 1 Summary tree based on the FULL dataset showing the major clades supported in this study. Colours are the same as in Espeland et al. (2019a). Support for major clades is shown. Interpretation of support values can be found in the legend." figureDoi="http://doi.org/10.5281/zenodo.7968436" httpUri="https://zenodo.org/record/7968436/files/figure.png" pageId="17" pageNumber="18">
|
||
Figures 1,
|
||
<collectionCode id="ED54AE4CFF9EFFE1A975AF17FE2114D5" box="[378,394,1132,1152]" pageId="17" pageNumber="18">S</collectionCode>
|
||
1
|
||
<emphasis id="B931EA9BFF9EFFE1A99DAF16FE3514D5" box="[402,414,1133,1152]" italics="true" pageId="17" pageNumber="18">–</emphasis>
|
||
<collectionCode id="ED54AE4CFF9EFFE1A991AF17FE0714D5" box="[414,428,1132,1152]" pageId="17" pageNumber="18">S</collectionCode>
|
||
</figureCitation>
|
||
<figureCitation id="137E2A0CFF9EFFE1A9A2AF17FE1214D5" box="[429,441,1132,1152]" captionStart="FIGURE 3" captionStartId="7.[122,218,1308,1327]" captionTargetBox="[210,1466,123,1266]" captionTargetPageId="7" captionText="FIGURE 3 Phylogeny of the Cyllopsis clade based on the FULL dataset. Interpretation of support values can be found in the legend. The scale bars under the butterfly images equal 1 cm." figureDoi="http://doi.org/10.5281/zenodo.7909399" httpUri="https://zenodo.org/record/7909399/files/figure.png" pageId="17" pageNumber="18">3</figureCitation>
|
||
). In both FULL trees,
|
||
<taxonomicName id="4C454D0AFF9EFFE1AA73AF17FCAA14D5" authorityName="Forster" authorityYear="1964" box="[636,769,1132,1152]" class="Insecta" family="Nymphalidae" genus="Chloreuptychia" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B931EA9BFF9EFFE1AA73AF17FCAA14D5" box="[636,769,1132,1152]" italics="true" pageId="17" pageNumber="18">Chloreuptychia</emphasis>
|
||
</taxonomicName>
|
||
is sister to the ‘
|
||
<taxonomicName id="4C454D0AFF9EFFE1A91EAFF4FE2E14F6" authority="Forster, 1964" box="[273,389,1167,1187]" class="Insecta" family="Nymphalidae" genus="Pareuptychia" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B931EA9BFF9EFFE1A91EAFF4FE2E14F6" box="[273,389,1167,1187]" italics="true" pageId="17" pageNumber="18">Pareuptychia</emphasis>
|
||
</taxonomicName>
|
||
clade’, while in both 4GENES trees the genus is sister to the ‘
|
||
<taxonomicName id="4C454D0AFF9EFFE1A96FAFCAFE741490" authorityName="Forster" authorityYear="1964" box="[352,479,1201,1221]" class="Insecta" family="Nymphalidae" genus="Archeuptychia" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B931EA9BFF9EFFE1A96FAFCAFE741490" box="[352,479,1201,1221]" italics="true" pageId="17" pageNumber="18">Archeuptychia</emphasis>
|
||
</taxonomicName>
|
||
clade’. In the backbone tree (
|
||
<figureCitation id="137E2A0CFF9EFFE1A870AFAFFF7C14BD" box="[127,215,1236,1256]" captionStart="FIGURE 1" captionStartId="5.[122,218,1354,1373]" captionTargetBox="[155,1432,126,1307]" captionTargetId="graphics-326@5.[251,1339,165,1253]" captionText="FIGURE 1 Summary tree based on the FULL dataset showing the major clades supported in this study. Colours are the same as in Espeland et al. (2019a). Support for major clades is shown. Interpretation of support values can be found in the legend." figureDoi="http://doi.org/10.5281/zenodo.7968436" httpUri="https://zenodo.org/record/7968436/files/figure.png" pageId="17" pageNumber="18">
|
||
Figure
|
||
<collectionCode id="ED54AE4CFF9EFFE1A8CFAFAFFF7B14BD" box="[192,208,1236,1256]" pageId="17" pageNumber="18">S</collectionCode>
|
||
1
|
||
</figureCitation>
|
||
) the ‘
|
||
<taxonomicName id="4C454D0AFF9EFFE1A91FAFAFFE2F14BD" authority="Forster, 1964" box="[272,388,1236,1256]" class="Insecta" family="Nymphalidae" genus="Pareuptychia" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B931EA9BFF9EFFE1A91FAFAFFE2F14BD" box="[272,388,1236,1256]" italics="true" pageId="17" pageNumber="18">Pareuptychia</emphasis>
|
||
</taxonomicName>
|
||
clade’ is sister to the
|
||
<taxonomicName id="4C454D0AFF9EFFE1AA5BAFAFFD3514BD" authorityName="Hubner" authorityYear="1819" box="[596,670,1236,1256]" class="Insecta" family="Nymphalidae" genus="Taygetis" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B931EA9BFF9EFFE1AA5BAFAFFD3514BD" box="[596,670,1236,1256]" italics="true" pageId="17" pageNumber="18">Taygetis</emphasis>
|
||
</taxonomicName>
|
||
clade, and here
|
||
<taxonomicName id="4C454D0AFF9EFFE1A8A6AF8CFE85155E" authorityName="Forster" authorityYear="1964" box="[169,302,1271,1291]" class="Insecta" family="Nymphalidae" genus="Chloreuptychia" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B931EA9BFF9EFFE1A8A6AF8CFE85155E" box="[169,302,1271,1291]" italics="true" pageId="17" pageNumber="18">Chloreuptychia</emphasis>
|
||
</taxonomicName>
|
||
is found to be sister to a clade including the ‘
|
||
<taxonomicName id="4C454D0AFF9EFFE1AAD6AF8CFF081578" authorityName="Hubner" authorityYear="1819" class="Insecta" family="Nymphalidae" genus="Taygetis" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B931EA9BFF9EFFE1AAD6AF8CFF081578" italics="true" pageId="17" pageNumber="18">Taygetis</emphasis>
|
||
</taxonomicName>
|
||
clade’, ‘
|
||
<taxonomicName id="4C454D0AFF9EFFE1A8F9AE62FE2C1578" authorityName="Forster" authorityYear="1964" box="[246,391,1305,1325]" class="Insecta" family="Nymphalidae" genus="Splendeuptychia" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B931EA9BFF9EFFE1A8F9AE62FE2C1578" box="[246,391,1305,1325]" italics="true" pageId="17" pageNumber="18">Splendeuptychia</emphasis>
|
||
</taxonomicName>
|
||
clade’ and ‘
|
||
<taxonomicName id="4C454D0AFF9EFFE1A9F1AE62FDD61578" authorityName="Forster" authorityYear="1964" box="[510,637,1305,1325]" class="Insecta" family="Nymphalidae" genus="Archeuptychia" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B931EA9BFF9EFFE1A9F1AE62FDD61578" box="[510,637,1305,1325]" italics="true" pageId="17" pageNumber="18">Archeuptychia</emphasis>
|
||
</taxonomicName>
|
||
clade’. These relationships are not well supported and need to be further investigated.
|
||
</paragraph>
|
||
<paragraph id="8BFA3689FF9EFFE1A8ADAEFAFD5916A5" blockId="17.[122,771,751,1949]" pageId="17" pageNumber="18">
|
||
The small genus
|
||
<taxonomicName id="4C454D0AFF9EFFE1A94AAEFAFED615C0" authorityName="Nakahara, Willmott & Espeland" authorityYear="2019" box="[325,381,1409,1429]" class="Insecta" family="Nymphalidae" genus="Amiga" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B931EA9BFF9EFFE1A94AAEFAFED615C0" box="[325,381,1409,1429]" italics="true" pageId="17" pageNumber="18">Amiga</emphasis>
|
||
</taxonomicName>
|
||
is strongly supported as sister group to the remainder of the clade in all datasets. Most generic relationships are well-supported (
|
||
<figureCitation id="137E2A0CFF9EFFE1A937AEBCFE3A158E" box="[312,401,1479,1499]" captionStart="FIGURE 9" captionStartId="15.[122,218,1828,1847]" captionTargetBox="[256,1312,101,1786]" captionTargetPageId="15" captionText="FIGURE 9 Phylogeny of the ’Pareuptychia clade’ based on the FULL dataset. Interpretation of support values can be found in the legend. The scale bars under the butterfly images equal 1 cm." figureDoi="http://doi.org/10.5281/zenodo.7909413" httpUri="https://zenodo.org/record/7909413/files/figure.png" pageId="17" pageNumber="18">Figures 9</figureCitation>
|
||
,
|
||
<collectionCode id="ED54AE4CFF9EFFE1A990AEBCFE07158E" box="[415,428,1479,1499]" pageId="17" pageNumber="18">S</collectionCode>
|
||
<figureCitation id="137E2A0CFF9EFFE1A9A3AEBCFE1C158E" box="[428,439,1479,1499]" captionStart="FIGURE 2" captionStartId="6.[122,218,1092,1111]" captionTargetBox="[155,1113,122,1050]" captionTargetId="graphics-415@6.[156,985,123,1051]" captionTargetPageId="6" captionText="FIGURE 2 Phylogeny of Euptychia based on the FULL dataset. Interpretation of support values can be found in the legend. The scale bars under the butterfly images equal 1 cm." figureDoi="http://doi.org/10.5281/zenodo.7968438" httpUri="https://zenodo.org/record/7968438/files/figure.png" pageId="17" pageNumber="18">2</figureCitation>
|
||
and
|
||
<collectionCode id="ED54AE4CFF9EFFE1A9E5AEBCFE53158E" box="[490,504,1479,1499]" pageId="17" pageNumber="18">S</collectionCode>
|
||
<figureCitation id="137E2A0CFF9EFFE1A9F7AEBCFDAF158E" box="[504,516,1479,1499]" captionStart="FIGURE 3" captionStartId="7.[122,218,1308,1327]" captionTargetBox="[210,1466,123,1266]" captionTargetPageId="7" captionText="FIGURE 3 Phylogeny of the Cyllopsis clade based on the FULL dataset. Interpretation of support values can be found in the legend. The scale bars under the butterfly images equal 1 cm." figureDoi="http://doi.org/10.5281/zenodo.7909399" httpUri="https://zenodo.org/record/7909399/files/figure.png" pageId="17" pageNumber="18">3</figureCitation>
|
||
). The clade is divided into two well-supported subclades, the
|
||
<taxonomicName id="4C454D0AFF9EFFE1A9CDAE92FD8615A8" box="[450,557,1513,1533]" class="Insecta" family="Nymphalidae" genus="Optimandes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B931EA9BFF9EFFE1A9CDAE92FD8615A8" box="[450,557,1513,1533]" italics="true" pageId="17" pageNumber="18">Optimandes</emphasis>
|
||
</taxonomicName>
|
||
subclade (
|
||
<taxonomicName id="4C454D0AFF9EFFE1AA9EAE92FD5715A8" box="[657,764,1513,1533]" class="Insecta" family="Nymphalidae" genus="Optimandes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B931EA9BFF9EFFE1AA9EAE92FD5715A8" box="[657,764,1513,1533]" italics="true" pageId="17" pageNumber="18">Optimandes</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C454D0AFF9EFFE1A875AD77FF5A1675" authorityName="Freitas, Barbosa & Zacca" authorityYear="2018" box="[122,241,1548,1568]" class="Insecta" family="Nymphalidae" genus="Nhambikuara" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B931EA9BFF9EFFE1A875AD77FF5A1675" box="[122,241,1548,1568]" italics="true" pageId="17" pageNumber="18">Nhambikuara</emphasis>
|
||
</taxonomicName>
|
||
) and the
|
||
<taxonomicName id="4C454D0AFF9EFFE1A947AD77FE171675" authority="Forster, 1964" box="[328,444,1548,1568]" class="Insecta" family="Nymphalidae" genus="Pareuptychia" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B931EA9BFF9EFFE1A947AD77FE171675" box="[328,444,1548,1568]" italics="true" pageId="17" pageNumber="18">Pareuptychia</emphasis>
|
||
</taxonomicName>
|
||
subclade (
|
||
<taxonomicName id="4C454D0AFF9EFFE1AA12AD77FDDF1675" authority="Espeland1 & Nakahara & Zacca & Barbosa & Huertas & Marín & Lamas & Benmesbah & Brévignon & Casagrande & Fåhraeus & Grishin & Kawahara1 & Mielke & Miller & Nakamura & Navas & Patrusky & Pyrcz & Richards & Tan & Tyler & Viloria & Warren1 & Xiao & Freitas & Willmott, 2023" authorityName="Espeland1 & Nakahara & Zacca & Barbosa & Huertas & Marín & Lamas & Benmesbah & Brévignon & Casagrande & Fåhraeus & Grishin & Kawahara1 & Mielke & Miller & Nakamura & Navas & Patrusky & Pyrcz & Richards & Tan & Tyler & Viloria & Warren1 & Xiao & Freitas & Willmott" authorityYear="2023" box="[541,628,1548,1568]" class="Insecta" family="Nymphalidae" genus="Xenovena" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus" status="gen.n.">
|
||
<emphasis id="B931EA9BFF9EFFE1AA12AD77FDDF1675" box="[541,628,1548,1568]" italics="true" pageId="17" pageNumber="18">Xenovena</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B931EA9BFF9EFFE1AA75AD77FD191675" bold="true" box="[634,690,1548,1568]" pageId="17" pageNumber="18">gen.n.</emphasis>
|
||
,
|
||
<taxonomicName id="4C454D0AFF9EFFE1AAB3AD77FF1B1616" authority="Forster, 1964" class="Insecta" family="Nymphalidae" genus="Pareuptychia" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B931EA9BFF9EFFE1AAB3AD77FF1B1616" italics="true" pageId="17" pageNumber="18">Pareuptychia</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C454D0AFF9EFFE1A8B1AD54FE8E1616" authorityName="Forster" authorityYear="1964" box="[190,293,1583,1603]" class="Insecta" family="Nymphalidae" genus="Erichthodes" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B931EA9BFF9EFFE1A8B1AD54FE8E1616" box="[190,293,1583,1603]" italics="true" pageId="17" pageNumber="18">Erichthodes</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C454D0AFF9EFFE1A93BAD54FE341616" box="[308,415,1583,1603]" class="Insecta" family="Nymphalidae" genus="Neonympha" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B931EA9BFF9EFFE1A93BAD54FE341616" box="[308,415,1583,1603]" italics="true" pageId="17" pageNumber="18">Neonympha</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C454D0AFF9EFFE1A9A1AD54FD881616" authorityName="Forster" authorityYear="1964" box="[430,547,1583,1603]" class="Insecta" family="Nymphalidae" genus="Euptychoides" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B931EA9BFF9EFFE1A9A1AD54FD881616" box="[430,547,1583,1603]" italics="true" pageId="17" pageNumber="18">Euptychoides</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C454D0AFF9EFFE1AA3EAD54FD1B1616" authorityName="Forster" authorityYear="1964" box="[561,688,1583,1603]" class="Insecta" family="Nymphalidae" genus="Satyrotaygetis" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B931EA9BFF9EFFE1AA3EAD54FD1B1616" box="[561,688,1583,1603]" italics="true" pageId="17" pageNumber="18">Satyrotaygetis</emphasis>
|
||
</taxonomicName>
|
||
, and an undescribed generic-level clade containing some species currently in
|
||
<taxonomicName id="4C454D0AFF9EFFE1A875AD0FFF4A16DD" authorityName="Forster" authorityYear="1964" box="[122,225,1652,1672]" class="Insecta" family="Nymphalidae" genus="Erichthodes" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B931EA9BFF9EFFE1A875AD0FFF4A16DD" box="[122,225,1652,1672]" italics="true" pageId="17" pageNumber="18">Erichthodes</emphasis>
|
||
</taxonomicName>
|
||
), consistent with previous molecular studies (
|
||
<bibRefCitation id="EFD44B78FF9EFFE1AAACAD0FFF5016FE" author="Murray & Prowell" firstAuthor="Murray" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="17" pageNumber="18" pagination="67 - 80" part="34" refId="ref40211" refString="Murray, D. & Prowell, D. P. (2005) Molecular phylogenetics and evolutionary history of the neotropical Satyrine subtribe Euptychiina (Nymphalidae: Satyrinae). Molecular Phylogenetics and Evolution, 34, 67 - 80." title="Molecular phylogenetics and evolutionary history of the neotropical Satyrine subtribe Euptychiina (Nymphalidae: Satyrinae)" type="journal article" year="2005">Murray & Prowell, 2005</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFD44B78FF9EFFE1A908ADECFE0816FE" author="Pena" box="[263,419,1687,1707]" etAl="et al." firstAuthor="Pena" journalOrPublisher="Zoologica Scripta" pageId="17" pageNumber="18" pagination="243 - 258" part="39" refId="ref42528" refString="Pena, C., Nylin, S., Freitas, A. V. L. & Wahlberg, N. (2010) Biogeographic history of the butterfly subtribe Euptychiina (Lepidoptera, Nymphalidae, Satyrinae). Zoologica Scripta, 39, 243 - 258." title="Biogeographic history of the butterfly subtribe Euptychiina (Lepidoptera, Nymphalidae, Satyrinae)" type="journal article" year="2010">Peña et al., 2010</bibRefCitation>
|
||
). The placement of
|
||
<taxonomicName id="4C454D0AFF9EFFE1AA52ADECFD0016FE" baseAuthorityName="Freitas" baseAuthorityYear="2003" box="[605,683,1687,1707]" class="Insecta" family="Nymphalidae" genus="Taydebis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B931EA9BFF9EFFE1AA52ADECFD0016FE" box="[605,683,1687,1707]" italics="true" pageId="17" pageNumber="18">Taydebis</emphasis>
|
||
</taxonomicName>
|
||
needs to be further investigated; although it is found as sister to
|
||
<taxonomicName id="4C454D0AFF9EFFE1AA9FADC2FD501698" box="[656,763,1721,1741]" class="Insecta" family="Nymphalidae" genus="Optimandes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B931EA9BFF9EFFE1AA9FADC2FD501698" box="[656,763,1721,1741]" italics="true" pageId="17" pageNumber="18">Optimandes</emphasis>
|
||
</taxonomicName>
|
||
+
|
||
<taxonomicName id="4C454D0AFF9EFFE1A89FADA7FEAC16A5" authorityName="Freitas, Barbosa & Zacca" authorityYear="2018" box="[144,263,1756,1776]" class="Insecta" family="Nymphalidae" genus="Nhambikuara" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B931EA9BFF9EFFE1A89FADA7FEAC16A5" box="[144,263,1756,1776]" italics="true" pageId="17" pageNumber="18">Nhambikuara</emphasis>
|
||
</taxonomicName>
|
||
in all trees, the support for this branch is always low.
|
||
</paragraph>
|
||
<paragraph id="8BFA3689FF9EFFE1A8ADAD84FB9D1148" blockId="17.[122,771,751,1949]" lastBlockId="17.[818,1468,127,1741]" pageId="17" pageNumber="18">
|
||
Since the checklist
|
||
<bibRefCitation id="EFD44B78FF9EFFE1A95DAD84FE641746" author="Lamas" box="[338,463,1791,1811]" firstAuthor="Lamas" journalOrPublisher="Gainesville: Scientific Publishers / Association of Tropical Lepidoptera" pageId="17" pageNumber="18" pagination="439" refId="ref39197" refString="Lamas, G. (2004) Atlas of Neotropical Lepidoptera. Checklist: Part 4 A Hesperioidea - Papilionoidea. Gainesville: Scientific Publishers / Association of Tropical Lepidoptera, p. 439." title="Atlas of Neotropical Lepidoptera. Checklist: Part 4 A Hesperioidea - Papilionoidea" type="book chapter" year="2004">Lamas (2004)</bibRefCitation>
|
||
, a number of papers have helped revise the generic classification of the ‘
|
||
<taxonomicName id="4C454D0AFF9EFFE1A9FDAC5AFDCD1760" authority="Forster, 1964" box="[498,614,1825,1845]" class="Insecta" family="Nymphalidae" genus="Pareuptychia" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B931EA9BFF9EFFE1A9FDAC5AFDCD1760" box="[498,614,1825,1845]" italics="true" pageId="17" pageNumber="18">Pareuptychia</emphasis>
|
||
</taxonomicName>
|
||
clade’, with the description of new genera such as
|
||
<taxonomicName id="4C454D0AFF9EFFE1A9C6AC3FFD02170D" authority="(Freitas, 2003)" baseAuthorityName="Freitas" baseAuthorityYear="2003" box="[457,681,1860,1880]" class="Insecta" family="Nymphalidae" genus="Taydebis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B931EA9BFF9EFFE1A9C6AC3FFDBC170D" box="[457,535,1860,1880]" italics="true" pageId="17" pageNumber="18">Taydebis</emphasis>
|
||
(
|
||
<bibRefCitation id="EFD44B78FF9EFFE1AA2AAC3FFD09170D" author="Freitas" box="[549,674,1860,1880]" firstAuthor="Freitas" journalOrPublisher="Journal of the Lepidopterists' Society" pageId="17" pageNumber="18" pagination="100 - 106" part="57" refId="ref36881" refString="Freitas, A. V. L. (2003) Description of a new genus for " Euptychia peculiaris " (Nymphalidae: Satyrinae): immature stages and systematic position. Journal of the Lepidopterists' Society, 57, 100 - 106." title="Description of a new genus for " Euptychia peculiaris " (Nymphalidae: Satyrinae): immature stages and systematic position" type="journal article" year="2003">Freitas, 2003</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C454D0AFF9EFFE1AABBAC3FFF07172E" authorityName="Freitas, Barbosa & Zacca" authorityYear="2018" class="Insecta" family="Nymphalidae" genus="Nhambikuara" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B931EA9BFF9EFFE1AABBAC3FFF07172E" italics="true" pageId="17" pageNumber="18">Nhambikuara</emphasis>
|
||
</taxonomicName>
|
||
(Freitas, Barbosa, et al., 2018),
|
||
<taxonomicName id="4C454D0AFF9EFFE1AA06AC1CFDEA172E" authorityName="Nakahara, Willmott & Espeland" authorityYear="2019" box="[521,577,1895,1915]" class="Insecta" family="Nymphalidae" genus="Amiga" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B931EA9BFF9EFFE1AA06AC1CFDEA172E" box="[521,577,1895,1915]" italics="true" pageId="17" pageNumber="18">Amiga</emphasis>
|
||
</taxonomicName>
|
||
(Nakahara, Lamas, et al., 2019) and
|
||
<taxonomicName id="4C454D0AFF9EFFE1A92BACF2FDDB17C8" authority="(Willmott et al., 2019)" baseAuthorityName="Willmott" baseAuthorityYear="2019" box="[292,624,1929,1949]" class="Insecta" family="Nymphalidae" genus="Optimandes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B931EA9BFF9EFFE1A92BACF2FE2417C8" box="[292,399,1929,1949]" italics="true" pageId="17" pageNumber="18">Optimandes</emphasis>
|
||
(
|
||
<bibRefCitation id="EFD44B78FF9EFFE1A991ACF2FDC217C8" author="Willmott" box="[414,617,1929,1949]" etAl="et al." firstAuthor="Willmott" journalOrPublisher="Tropical Lepidoptera Research" pageId="17" pageNumber="18" pagination="29 - 44" part="29" refId="ref43893" refString="Willmott, K. R., Marin, M. A., Nakahara, S., Pomerantz, T., Lamas, G., Huertas, B. et al. (2019) A revision of the new Andean butterfly genus Optimandes Marin, Nakahara & Willmott, n. gen., with the description of a new species (Nymphalidae: Satyrinae: Euptychiina). Tropical Lepidoptera Research, 29, 29 - 44." title="A revision of the new Andean butterfly genus Optimandes Marin, Nakahara & Willmott, n. gen., with the description of a new species (Nymphalidae: Satyrinae: Euptychiina)" type="journal article" year="2019">Willmott et al., 2019</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, the sorting of species in previously described genera (
|
||
<bibRefCitation id="EFD44B78FF9EFFE1ACA5AB04FA1F10C6" author="Andrade-Correa" box="[1194,1460,127,147]" etAl="et al." firstAuthor="Andrade-Correa" journalOrPublisher="Anartia" pageId="17" pageNumber="18" pagination="26 - 46" part="28" refId="ref34200" refString="Andrade-Correa, G., Viloria, A´., Henao-Banol, E. R. & Le Crom, J. F. (2019) Generic reassignments of satyrine butterflies from Colombia and Venezuela (Lepidoptera: Nymphalidae). Anartia, 28, 26 - 46." title="Generic reassignments of satyrine butterflies from Colombia and Venezuela (Lepidoptera: Nymphalidae)" type="journal article" year="2019">Andrade-Correa et al., 2019</bibRefCitation>
|
||
) and descriptions of 11 new species (
|
||
<bibRefCitation id="EFD44B78FF9EFFE1ACF7ABDAFA1F10E0" author="Benmesbah" box="[1272,1460,161,181]" firstAuthor="Benmesbah" journalOrPublisher="Bulletin de la Societe Entomologique de France" pageId="17" pageNumber="18" pagination="147 - 156" part="120" refId="ref34889" refString="Benmesbah, M. (2015) Description de deux nouvelles especes de Satyrinae de Guyane franCaise (Lepidoptera, Nymphalidae). Bulletin de la Societe Entomologique de France, 120, 147 - 156." title="Description de deux nouvelles especes de Satyrinae de Guyane franCaise (Lepidoptera, Nymphalidae)" type="journal article" year="2015">Benmesbah, 2015</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFD44B78FF9EFFE1AB3DABBFFC64108D" author="Brevignon" box="[818,975,196,216]" firstAuthor="Brevignon" journalOrPublisher="Lambillionea" pageId="17" pageNumber="18" pagination="393 - 404" part="105" refId="ref35180" refString="Brevignon, C. (2005) Description de nouveaux Satyrinae provenant de Guyane franCaise (Lepidoptera, Nymphalidae). Lambillionea, 105, 393 - 404." title="Description de nouveaux Satyrinae provenant de Guyane franCaise (Lepidoptera, Nymphalidae)" type="journal article" year="2005">Brévignon, 2005</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFD44B78FF9EFFE1ABD1ABBFFB38108D" author="Freitas" box="[990,1171,196,216]" etAl="et al." firstAuthor="Freitas" journalOrPublisher="Neotropical Entomology" pageId="17" pageNumber="18" pagination="231 - 237" part="40" refId="ref37608" refString="Freitas, A., Mielke, O., Moser, A., Silva-Brandao t, K. & Iserhard, C. (2011) A new genus and species of Euptychiina (Lepidoptera: Nymphalidae: Satyrinae) from southern Brazil. Neotropical Entomology, 40, 231 - 237." title="A new genus and species of Euptychiina (Lepidoptera: Nymphalidae: Satyrinae) from southern Brazil" type="journal article" year="2011">Freitas et al., 2011</bibRefCitation>
|
||
; Freitas, Barbosa, et al., 2018;
|
||
<bibRefCitation id="EFD44B78FF9EFFE1AB3DAB9CFC1310AE" author="Huertas" box="[818,952,231,251]" firstAuthor="Huertas" journalOrPublisher="Zootaxa" pageId="17" pageNumber="18" pagination="63 - 68" part="2802" refId="ref38472" refString="Huertas, B. (2011) A new species of satyrid butterfly from Peru (Lepidoptera: Nymphalidae: Satyrinae: Euptychiini). Zootaxa, 2802, 63 - 68." title="A new species of satyrid butterfly from Peru (Lepidoptera: Nymphalidae: Satyrinae: Euptychiini)" type="journal article" year="2011">Huertas, 2011</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFD44B78FF9EFFE1ABCAAB9CFBD410AE" author="Huertas" box="[965,1151,231,251]" etAl="et al." firstAuthor="Huertas" journalOrPublisher="Zootaxa" pageId="17" pageNumber="18" pagination="51 - 58" part="2014" refId="ref38558" refString="Huertas, B., Rios-Malaver, C. & Le Crom, J. F. (2009) A new species of Splendeuptychia from the Magdalena Valley in Colombia (Lepidoptera: Nymphalidae: Satyrinae). Zootaxa, 2014, 51 - 58." title="A new species of Splendeuptychia from the Magdalena Valley in Colombia (Lepidoptera: Nymphalidae: Satyrinae)" type="journal article" year="2009">Huertas et al., 2009</bibRefCitation>
|
||
; Nakahara, Marin, & Neild, 2016;
|
||
<bibRefCitation id="EFD44B78FF9EFFE1AB3DAA72FC5B1148" author="Willmott" box="[818,1008,265,285]" etAl="et al." firstAuthor="Willmott" journalOrPublisher="Tropical Lepidoptera Research" pageId="17" pageNumber="18" pagination="39 - 45" part="28" refId="ref43822" refString="Willmott, K. R., Lamas, G., Radford, J. T., Marin, M. A., Nakahara, S., Espeland, M. et al. (2018) A distinctive new species of cloud forest Euptychiina (Lepidoptera: Nymphalidae: Satyrinae) from Ecuador and Peru. Tropical Lepidoptera Research, 28, 39 - 45." title="A distinctive new species of cloud forest Euptychiina (Lepidoptera: Nymphalidae: Satyrinae) from Ecuador and Peru" type="journal article" year="2018">Willmott et al., 2018</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFD44B78FF9EFFE1ABF5AA72FB801148" author="Willmott" box="[1018,1067,265,285]" firstAuthor="Willmott" journalOrPublisher="Global Ecology and Biogeography" pageId="17" pageNumber="18" pagination="1118 - 1132" part="28" refId="ref35539" refString="Chazot, N., Willmott, K. R., Lamas, G., Freitas, A. V. L., Piron-Prunier, F., Arias, C. F. et al. (2019) Renewed diversification following Miocene landscape turnover in a Neotropical butterfly radiation. Global Ecology and Biogeography, 28, 1118 - 1132." title="Renewed diversification following Miocene landscape turnover in a Neotropical butterfly radiation" type="journal article" year="2019">2019</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BFA3689FF9EFFE1AB55AA57FB2D13D8" blockId="17.[818,1468,127,1741]" pageId="17" pageNumber="18">
|
||
We found strong support for a clade containing the
|
||
<typeStatus id="54FE882BFF9EFFE1AD3EAA57FAF11115" box="[1329,1370,300,320]" pageId="17" pageNumber="18">type</typeStatus>
|
||
species of
|
||
<taxonomicName id="4C454D0AFF9EFFE1AB3DAA34FC0C1136" authorityName="Freitas, Barbosa & Zacca" authorityYear="2018" box="[818,935,335,355]" class="Insecta" family="Nymphalidae" genus="Nhambikuara" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B931EA9BFF9EFFE1AB3DAA34FC0C1136" box="[818,935,335,355]" italics="true" pageId="17" pageNumber="18">Nhambikuara</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C454D0AFF9EFFE1ABB8AA34FB9E1136" authorityName="Freitas, Barbosa & Zacca" authorityYear="2018" box="[951,1077,335,355]" class="Insecta" family="Nymphalidae" genus="Nhambikuara" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="cerradensis">
|
||
<emphasis id="B931EA9BFF9EFFE1ABB8AA34FC6D1136" box="[951,966,335,355]" italics="true" pageId="17" pageNumber="18">N</emphasis>
|
||
.
|
||
<emphasis id="B931EA9BFF9EFFE1ABDAAA34FB9E1136" box="[981,1077,335,355]" italics="true" pageId="17" pageNumber="18">cerradensis</emphasis>
|
||
</taxonomicName>
|
||
, along with nine other species formerly placed in
|
||
<taxonomicName id="4C454D0AFF9EFFE1AB84AA0AFBB211D0" authorityName="Forster" authorityYear="1964" box="[907,1049,369,389]" class="Insecta" family="Nymphalidae" genus="Splendeuptychia" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B931EA9BFF9EFFE1AB84AA0AFBB211D0" box="[907,1049,369,389]" italics="true" pageId="17" pageNumber="18">Splendeuptychia</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EFD44B78FF9EFFE1AC2AAA0AFB3111D0" author="Lamas" box="[1061,1178,369,389]" firstAuthor="Lamas" journalOrPublisher="Gainesville: Scientific Publishers / Association of Tropical Lepidoptera" pageId="17" pageNumber="18" pagination="439" refId="ref39197" refString="Lamas, G. (2004) Atlas of Neotropical Lepidoptera. Checklist: Part 4 A Hesperioidea - Papilionoidea. Gainesville: Scientific Publishers / Association of Tropical Lepidoptera, p. 439." title="Atlas of Neotropical Lepidoptera. Checklist: Part 4 A Hesperioidea - Papilionoidea" type="book chapter" year="2004">Lamas, 2004</bibRefCitation>
|
||
), a relationship partly indicated in Freitas, Barbosa, et al. (2018). Morphological characters also support this as a relatively compact group of species (
|
||
<bibRefCitation id="EFD44B78FF9EFFE1ACEEAACCFACC119E" author="Huertas" box="[1249,1383,439,459]" firstAuthor="Huertas" journalOrPublisher="University College London" pageId="17" pageNumber="18" refId="ref38505" refString="Huertas, B. (2014) Systematics of the genus Splendeuptychia (Forster, 1964) and implications of taxonomy in conservation assessments. Chapter 4. In Evaluating the conservation status of Neotropical butterflies and the impact of systematics on threat assessments (PhD thesis). University College London." title="Systematics of the genus Splendeuptychia (Forster, 1964) and implications of taxonomy in conservation assessments. Chapter 4" type="book" volumeTitle="Evaluating the conservation status of Neotropical butterflies and the impact of systematics on threat assessments (PhD thesis)" year="2014">Huertas, 2014</bibRefCitation>
|
||
) and we thus transfer the nine former
|
||
<taxonomicName id="4C454D0AFF9EFFE1AC4FAAA2FABE11B8" box="[1088,1301,473,493]" pageId="17" pageNumber="18">
|
||
<emphasis id="B931EA9BFF9EFFE1AC4FAAA2FB6511B8" box="[1088,1230,473,493]" italics="true" pageId="17" pageNumber="18">Splendeuptychia</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
into
|
||
<taxonomicName id="4C454D0AFF9EFFE1AD4BAAA2FA1211B8" authorityName="Freitas, Barbosa & Zacca" authorityYear="2018" box="[1348,1465,473,493]" class="Insecta" family="Nymphalidae" genus="Nhambikuara" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B931EA9BFF9EFFE1AD4BAAA2FA1211B8" box="[1348,1465,473,493]" italics="true" pageId="17" pageNumber="18">Nhambikuara</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<emphasis id="B931EA9BFF9EFFE1AB37AA87FC2A1245" bold="true" box="[824,897,508,528]" pageId="17" pageNumber="18">comb.n.</emphasis>
|
||
). We found strong support for the recently described genus
|
||
<taxonomicName id="4C454D0AFF9EFFE1AB3DA964FB071266" authority="(Andrade-Correa et al., 2019)" baseAuthorityName="Andrade-Correa" baseAuthorityYear="2019" box="[818,1196,543,563]" genus="Macrocissia" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="17" pageNumber="18" rank="genus">
|
||
<emphasis id="B931EA9BFF9EFFE1AB3DA964FC3C1266" box="[818,919,543,563]" italics="true" pageId="17" pageNumber="18">Macrocissia</emphasis>
|
||
(
|
||
<bibRefCitation id="EFD44B78FF9EFFE1ABACA964FB0D1266" author="Andrade-Correa" box="[931,1190,543,563]" etAl="et al." firstAuthor="Andrade-Correa" journalOrPublisher="Anartia" pageId="17" pageNumber="18" pagination="26 - 46" part="28" refId="ref34200" refString="Andrade-Correa, G., Viloria, A´., Henao-Banol, E. R. & Le Crom, J. F. (2019) Generic reassignments of satyrine butterflies from Colombia and Venezuela (Lepidoptera: Nymphalidae). Anartia, 28, 26 - 46." title="Generic reassignments of satyrine butterflies from Colombia and Venezuela (Lepidoptera: Nymphalidae)" type="journal article" year="2019">Andrade-Correa et al., 2019</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
not being monophyletic, and the two species previously included in this genus are transferred to
|
||
<taxonomicName id="4C454D0AFF9EFFE1AB3DA91FFC05122D" authorityName="Forster" authorityYear="1964" box="[818,942,612,632]" class="Insecta" family="Nymphalidae" genus="Satyrotaygetis" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B931EA9BFF9EFFE1AB3DA91FFC05122D" box="[818,942,612,632]" italics="true" pageId="17" pageNumber="18">Satyrotaygetis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName id="4C454D0AFF9EFFE1ABB0A91FFBF4122D" authorityName="Espeland1 & Nakahara & Zacca & Barbosa & Huertas & Marín & Lamas & Benmesbah & Brévignon & Casagrande & Fåhraeus & Grishin & Kawahara1 & Mielke & Miller & Nakamura & Navas & Patrusky & Pyrcz & Richards & Tan & Tyler & Viloria & Warren1 & Xiao & Freitas & Willmott" authorityYear="2023" baseAuthorityName="C. Felder & R. Felder" baseAuthorityYear="1867" box="[959,1119,612,632]" class="Insecta" family="Nymphalidae" genus="Satyrotaygetis" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="iris">
|
||
<emphasis id="B931EA9BFF9EFFE1ABB0A91FFBF4122D" box="[959,1119,612,632]" italics="true" pageId="17" pageNumber="18">Satyrotaygetis iris</emphasis>
|
||
</taxonomicName>
|
||
[C. Felder & R. Felder, 1867]) and
|
||
<taxonomicName id="4C454D0AFF9EFFE1AB3DA9FCFC0E12CE" authorityName="Forster" authorityYear="1964" box="[818,933,647,667]" class="Insecta" family="Nymphalidae" genus="Euptychoides" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B931EA9BFF9EFFE1AB3DA9FCFC0E12CE" box="[818,933,647,667]" italics="true" pageId="17" pageNumber="18">Euptychoides</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName id="4C454D0AFF9EFFE1ABB9A9FCFBF512CE" authorityName="Espeland1 & Nakahara & Zacca & Barbosa & Huertas & Marín & Lamas & Benmesbah & Brévignon & Casagrande & Fåhraeus & Grishin & Kawahara1 & Mielke & Miller & Nakamura & Navas & Patrusky & Pyrcz & Richards & Tan & Tyler & Viloria & Warren1 & Xiao & Freitas & Willmott" authorityYear="2023" baseAuthorityName="Staudinger" baseAuthorityYear="1886" box="[950,1118,647,667]" class="Insecta" family="Nymphalidae" genus="Euptychoides" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="inani">
|
||
<emphasis id="B931EA9BFF9EFFE1ABB9A9FCFBF512CE" box="[950,1118,647,667]" italics="true" pageId="17" pageNumber="18">Euptychoides inani</emphasis>
|
||
</taxonomicName>
|
||
[Staudinger, 1886]) in concordance with the phylogeny. The combination of morphological characters used by
|
||
<bibRefCitation id="EFD44B78FF9EFFE1AB41A9B7FBFF12B5" author="Andrade-Correa" box="[846,1108,716,736]" etAl="et al." firstAuthor="Andrade-Correa" journalOrPublisher="Anartia" pageId="17" pageNumber="18" pagination="26 - 46" part="28" refId="ref34200" refString="Andrade-Correa, G., Viloria, A´., Henao-Banol, E. R. & Le Crom, J. F. (2019) Generic reassignments of satyrine butterflies from Colombia and Venezuela (Lepidoptera: Nymphalidae). Anartia, 28, 26 - 46." title="Generic reassignments of satyrine butterflies from Colombia and Venezuela (Lepidoptera: Nymphalidae)" type="journal article" year="2019">Andrade-Correa et al. (2019)</bibRefCitation>
|
||
to support
|
||
<taxonomicName id="4C454D0AFF9EFFE1ACB1A9B7FA8812B5" box="[1214,1315,716,736]" genus="Macrocissia" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="17" pageNumber="18" rank="genus">
|
||
<emphasis id="B931EA9BFF9EFFE1ACB1A9B7FA8812B5" box="[1214,1315,716,736]" italics="true" pageId="17" pageNumber="18">Macrocissia</emphasis>
|
||
</taxonomicName>
|
||
, ‘vestigial recurrent vein into the discal cell’ in the forewing, and ‘valvae with a hook-like distal process’ and a ‘finely serrate cucullus in the male genitalia’, is a combination of characters present in previously described genera such as
|
||
<taxonomicName id="4C454D0AFF9EFFE1AB86A82CFBAE133E" authorityName="Forster" authorityYear="1964" box="[905,1029,855,875]" class="Insecta" family="Nymphalidae" genus="Satyrotaygetis" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B931EA9BFF9EFFE1AB86A82CFBAE133E" box="[905,1029,855,875]" italics="true" pageId="17" pageNumber="18">Satyrotaygetis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C454D0AFF9EFFE1AC4FA82CFB19133E" authorityName="Forster" authorityYear="1964" box="[1088,1202,855,875]" class="Insecta" family="Nymphalidae" genus="Euptychoides" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B931EA9BFF9EFFE1AC4FA82CFB19133E" box="[1088,1202,855,875]" italics="true" pageId="17" pageNumber="18">Euptychoides</emphasis>
|
||
</taxonomicName>
|
||
, and we thus synonymize
|
||
<taxonomicName id="4C454D0AFF9EFFE1AB3DA802FC3C13D8" box="[818,919,889,909]" genus="Macrocissia" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="17" pageNumber="18" rank="genus" status="syn.n.">
|
||
<emphasis id="B931EA9BFF9EFFE1AB3DA802FC3C13D8" box="[818,919,889,909]" italics="true" pageId="17" pageNumber="18">Macrocissia</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B931EA9BFF9EFFE1AB92A802FC7813D8" bold="true" box="[925,979,889,909]" pageId="17" pageNumber="18">
|
||
<taxonomicNameLabel id="A20257E0FF9EFFE1AB92A802FC7813D8" box="[925,979,889,909]" pageId="17" pageNumber="18">syn.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
with
|
||
<taxonomicName id="4C454D0AFF9EFFE1AC0AA802FB2A13D8" authorityName="Forster" authorityYear="1964" box="[1029,1153,889,909]" class="Insecta" family="Nymphalidae" genus="Satyrotaygetis" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B931EA9BFF9EFFE1AC0AA802FB2A13D8" box="[1029,1153,889,909]" italics="true" pageId="17" pageNumber="18">Satyrotaygetis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BFA3689FF9EFFE1AB55A8E7FB681698" blockId="17.[818,1468,127,1741]" pageId="17" pageNumber="18">
|
||
The ‘
|
||
<taxonomicName id="4C454D0AFF9EFFE1AB82A8E7FC5413E5" authority="Forster, 1964" box="[909,1023,924,944]" class="Insecta" family="Nymphalidae" genus="Pareuptychia" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B931EA9BFF9EFFE1AB82A8E7FC5413E5" box="[909,1023,924,944]" italics="true" pageId="17" pageNumber="18">Pareuptychia</emphasis>
|
||
</taxonomicName>
|
||
clade’ contains 44 described species, with the genera
|
||
<taxonomicName id="4C454D0AFF9EFFE1AB7AA8C4FC4C1386" authority="Forster, 1964" box="[885,999,959,979]" class="Insecta" family="Nymphalidae" genus="Pareuptychia" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B931EA9BFF9EFFE1AB7AA8C4FC4C1386" box="[885,999,959,979]" italics="true" pageId="17" pageNumber="18">Pareuptychia</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C454D0AFF9EFFE1AC1CA8C4FB231386" authorityName="Freitas, Barbosa & Zacca" authorityYear="2018" box="[1043,1160,959,979]" class="Insecta" family="Nymphalidae" genus="Nhambikuara" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B931EA9BFF9EFFE1AC1CA8C4FB231386" box="[1043,1160,959,979]" italics="true" pageId="17" pageNumber="18">Nhambikuara</emphasis>
|
||
</taxonomicName>
|
||
, with nine and 11 species respectively, being the most diverse. Taxonomic revisions for these two genera are in progress, in addition to work on
|
||
<taxonomicName id="4C454D0AFF9EFFE1ACC9AF7FFAB9144D" baseAuthorityName="Freitas" baseAuthorityYear="2003" box="[1222,1298,1028,1048]" class="Insecta" family="Nymphalidae" genus="Taydebis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B931EA9BFF9EFFE1ACC9AF7FFAB9144D" box="[1222,1298,1028,1048]" italics="true" pageId="17" pageNumber="18">Taydebis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C454D0AFF9EFFE1AD11AF7FFA3A144D" authorityName="Forster" authorityYear="1964" box="[1310,1425,1028,1048]" class="Insecta" family="Nymphalidae" genus="Euptychoides" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B931EA9BFF9EFFE1AD11AF7FFA3A144D" box="[1310,1425,1028,1048]" italics="true" pageId="17" pageNumber="18">Euptychoides</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C454D0AFF9EFFE1AB3DAF5CFC3D146E" authorityName="Forster" authorityYear="1964" box="[818,918,1063,1083]" class="Insecta" family="Nymphalidae" genus="Erichthodes" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B931EA9BFF9EFFE1AB3DAF5CFC3D146E" box="[818,918,1063,1083]" italics="true" pageId="17" pageNumber="18">Erichthodes</emphasis>
|
||
</taxonomicName>
|
||
. Members of the ‘
|
||
<taxonomicName id="4C454D0AFF9EFFE1AC4BAF5CFB1D146E" authority="Forster, 1964" box="[1092,1206,1063,1083]" class="Insecta" family="Nymphalidae" genus="Pareuptychia" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B931EA9BFF9EFFE1AC4BAF5CFB1D146E" box="[1092,1206,1063,1083]" italics="true" pageId="17" pageNumber="18">Pareuptychia</emphasis>
|
||
</taxonomicName>
|
||
clade’ have a wide distribution, ranging from southern
|
||
<collectingCountry id="F3527619FF9EFFE1AC32AF32FB2A1408" box="[1085,1153,1097,1117]" name="Canada" pageId="17" pageNumber="18" value="Canada">Canada</collectingCountry>
|
||
to northern
|
||
<collectingCountry id="F3527619FF9EFFE1ACF1AF32FAF21408" box="[1278,1369,1097,1117]" name="Argentina" pageId="17" pageNumber="18" value="Argentina">Argentina</collectingCountry>
|
||
, and from sea level to almost
|
||
<quantity id="4CBD9B6CFF9EFFE1ABE6AF17FB9914D5" box="[1001,1074,1132,1152]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="17" pageNumber="18" unit="m" value="2600.0">2600 m</quantity>
|
||
, with the highest diversity in the Amazon basin and the Andes (Marín et al. in preparation). According to
|
||
<bibRefCitation id="EFD44B78FF9EFFE1AD8BAFF4FC0E1490" author="Marin" etAl="et al." firstAuthor="Marin" journalOrPublisher="Systematic Entomology" pageId="17" pageNumber="18" pagination="768 - 785" part="42" refId="ref39677" refString="Marin, M. A., Pena, C., Uribe, S. I. & Freitas, A. V. L. (2017) Morphology agrees with molecular data: phylogenetic affinities of Euptychiina butterflies (Nymphalidae: Satyrinae). Systematic Entomology, 42, 768 - 785." title="Morphology agrees with molecular data: phylogenetic affinities of Euptychiina butterflies (Nymphalidae: Satyrinae)" type="journal article" year="2017">Marín et al. (2017)</bibRefCitation>
|
||
, possible synapomorphies (not unique) and/or distinctive characters for the clade include the following: (a) obvious sclerotized cornuti in aedeagus; (b) in the valva, posterior ending of the cucullus acute. Hostplants for members of this clade are mostly bamboos or grasses in the family
|
||
<taxonomicName id="4C454D0AFF9EFFE1ABFCAE47FBE91505" box="[1011,1090,1340,1360]" class="Liliopsida" family="Poaceae" kingdom="Plantae" order="Poales" pageId="17" pageNumber="18" phylum="Tracheophyta" rank="family">Poaceae</taxonomicName>
|
||
, with records of
|
||
<taxonomicName id="4C454D0AFF9EFFE1ACD2AE47FAEC1505" box="[1245,1351,1340,1360]" class="Liliopsida" family="Cyperaceae" kingdom="Plantae" order="Poales" pageId="17" pageNumber="18" phylum="Tracheophyta" rank="family">Cyperaceae</taxonomicName>
|
||
in
|
||
<taxonomicName id="4C454D0AFF9EFFE1AD6AAE47FBF61526" authority="Forster, 1964" class="Insecta" family="Nymphalidae" genus="Pareuptychia" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B931EA9BFF9EFFE1AD6AAE47FCFD1526" italics="true" pageId="17" pageNumber="18">Pareuptychia</emphasis>
|
||
(
|
||
<bibRefCitation id="EFD44B78FF9EFFE1AB6DAE24FBFC1526" accessDate="2022-01-01" author="Janzen & Hallwachs" box="[866,1111,1375,1395]" firstAuthor="Janzen" journalOrPublisher="Guanacaste (ACG), northwestern Costa Rica - caterpillars, pupae, butterflies & moths [WWW document]" pageId="17" pageNumber="18" publicationUrl="http://janzen.sas.upenn.edu/caterpillars/database.lasso" refId="ref38713" refString="Janzen, D. H. & Hallwachs, W. (2022) Area de Conservacion´Guanacaste (ACG), northwestern Costa Rica - caterpillars, pupae, butterflies & moths [WWW document]. Available from: http: // janzen. sas. upenn. edu / caterpillars / database. lasso [Accessed on 1 st January 2022]" title="Area de Conservacion" type="url" year="2022">Janzen & Hallwachs, 2022</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C454D0AFF9EFFE1AC82AE24FA1C1526" authority="(Hamm et al., 2013)" baseAuthorityName="Hamm" baseAuthorityYear="2013" box="[1165,1463,1375,1395]" class="Insecta" family="Nymphalidae" genus="Neonympha" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B931EA9BFF9EFFE1AC82AE24FB5D1526" box="[1165,1270,1375,1395]" italics="true" pageId="17" pageNumber="18">Neonympha</emphasis>
|
||
(
|
||
<bibRefCitation id="EFD44B78FF9EFFE1AD0DAE24FA1B1526" author="Hamm" box="[1282,1456,1375,1395]" etAl="et al." firstAuthor="Hamm" journalOrPublisher="Journal of the Lepidopterists' Society" pageId="17" pageNumber="18" pagination="15 - 28" part="67" refId="ref38211" refString="Hamm, C. A., Williams, B. L. & Landis, D. A. (2013) Natural history and conservation status of the endangered Mitchell ' s satyr butterfly: synthesis and expansion of our knowledge regarding Neonympha mitchellii mitchellii French 1889. Journal of the Lepidopterists' Society, 67, 15 - 28." title="Natural history and conservation status of the endangered Mitchell ' s satyr butterfly: synthesis and expansion of our knowledge regarding Neonympha mitchellii mitchellii French 1889" type="journal article" year="2013">Hamm et al., 2013</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C454D0AFF9EFFE1AB54AEFAFC7A15C0" box="[859,977,1409,1429]" class="Liliopsida" family="Marantaceae" kingdom="Plantae" order="Zingiberales" pageId="17" pageNumber="18" phylum="Tracheophyta" rank="family">Marantaceae</taxonomicName>
|
||
in
|
||
<taxonomicName id="4C454D0AFF9EFFE1ABFDAEFAFADE15C0" authority="(Janzen & Hallwachs, 2022)" baseAuthorityName="Janzen & Hallwachs" baseAuthorityYear="2022" box="[1010,1397,1409,1429]" class="Insecta" family="Nymphalidae" genus="Megeuptychia" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B931EA9BFF9EFFE1ABFDAEFAFBC715C0" box="[1010,1132,1409,1429]" italics="true" pageId="17" pageNumber="18">Megeuptychia</emphasis>
|
||
(
|
||
<bibRefCitation id="EFD44B78FF9EFFE1AC77AEFAFAC515C0" accessDate="2022-01-01" author="Janzen & Hallwachs" box="[1144,1390,1409,1429]" firstAuthor="Janzen" journalOrPublisher="Guanacaste (ACG), northwestern Costa Rica - caterpillars, pupae, butterflies & moths [WWW document]" pageId="17" pageNumber="18" publicationUrl="http://janzen.sas.upenn.edu/caterpillars/database.lasso" refId="ref38713" refString="Janzen, D. H. & Hallwachs, W. (2022) Area de Conservacion´Guanacaste (ACG), northwestern Costa Rica - caterpillars, pupae, butterflies & moths [WWW document]. Available from: http: // janzen. sas. upenn. edu / caterpillars / database. lasso [Accessed on 1 st January 2022]" title="Area de Conservacion" type="url" year="2022">Janzen & Hallwachs, 2022</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. Immature stages have been described or illustrated for
|
||
<taxonomicName id="4C454D0AFF9EFFE1ACF3AEDFFA9815ED" authorityName="Nakahara, Willmott & Espeland" authorityYear="2019" box="[1276,1331,1444,1464]" class="Insecta" family="Nymphalidae" genus="Amiga" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B931EA9BFF9EFFE1ACF3AEDFFA9815ED" box="[1276,1331,1444,1464]" italics="true" pageId="17" pageNumber="18">Amiga</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C454D0AFF9EFFE1AD31AEDFFA2115ED" baseAuthorityName="Freitas" baseAuthorityYear="2003" box="[1342,1418,1444,1464]" class="Insecta" family="Nymphalidae" genus="Taydebis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B931EA9BFF9EFFE1AD31AEDFFA2115ED" box="[1342,1418,1444,1464]" italics="true" pageId="17" pageNumber="18">Taydebis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C454D0AFF9EFFE1AD9AAEDFFC2D158E" authority="Forster, 1964" class="Insecta" family="Nymphalidae" genus="Pareuptychia" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B931EA9BFF9EFFE1AD9AAEDFFC2D158E" italics="true" pageId="17" pageNumber="18">Pareuptychia</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C454D0AFF9EFFE1AB9AAEBCFC55158E" box="[917,1022,1479,1499]" class="Insecta" family="Nymphalidae" genus="Optimandes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B931EA9BFF9EFFE1AB9AAEBCFC55158E" box="[917,1022,1479,1499]" italics="true" pageId="17" pageNumber="18">Optimandes</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C454D0AFF9EFFE1AC03AEBCFB23158E" authorityName="Forster" authorityYear="1964" box="[1036,1160,1479,1499]" class="Insecta" family="Nymphalidae" genus="Satyrotaygetis" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B931EA9BFF9EFFE1AC03AEBCFB23158E" box="[1036,1160,1479,1499]" italics="true" pageId="17" pageNumber="18">Satyrotaygetis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C454D0AFF9EFFE1AC99AEBCFABB158E" baseAuthorityName="Janzen & Hallwachs" baseAuthorityYear="2022" box="[1174,1296,1479,1499]" class="Insecta" family="Nymphalidae" genus="Megeuptychia" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B931EA9BFF9EFFE1AC99AEBCFABB158E" box="[1174,1296,1479,1499]" italics="true" pageId="17" pageNumber="18">Megeuptychia</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C454D0AFF9EFFE1AD4BAEBCFA12158E" authorityName="Freitas, Barbosa & Zacca" authorityYear="2018" box="[1348,1465,1479,1499]" class="Insecta" family="Nymphalidae" genus="Nhambikuara" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B931EA9BFF9EFFE1AD4BAEBCFA12158E" box="[1348,1465,1479,1499]" italics="true" pageId="17" pageNumber="18">Nhambikuara</emphasis>
|
||
</taxonomicName>
|
||
(Corahua-Espinoza, Nakahara, Kabir, et al., 2022; Corahua-Espinoza, Nakahara, Shellman, et al., 2022;
|
||
<bibRefCitation id="EFD44B78FF9EFFE1AC76AD77FB5E1675" author="Freitas" box="[1145,1269,1548,1568]" firstAuthor="Freitas" journalOrPublisher="Journal of the Lepidopterists' Society" pageId="17" pageNumber="18" pagination="100 - 106" part="57" refId="ref36881" refString="Freitas, A. V. L. (2003) Description of a new genus for " Euptychia peculiaris " (Nymphalidae: Satyrinae): immature stages and systematic position. Journal of the Lepidopterists' Society, 57, 100 - 106." title="Description of a new genus for " Euptychia peculiaris " (Nymphalidae: Satyrinae): immature stages and systematic position" type="journal article" year="2003">Freitas, 2003</bibRefCitation>
|
||
; Freitas, Barbosa, & Marín, 2016;
|
||
<bibRefCitation id="EFD44B78FF9EFFE1ABB6AD54FB111616" accessDate="2022-01-01" author="Janzen & Hallwachs" box="[953,1210,1583,1603]" firstAuthor="Janzen" journalOrPublisher="Guanacaste (ACG), northwestern Costa Rica - caterpillars, pupae, butterflies & moths [WWW document]" pageId="17" pageNumber="18" publicationUrl="http://janzen.sas.upenn.edu/caterpillars/database.lasso" refId="ref38713" refString="Janzen, D. H. & Hallwachs, W. (2022) Area de Conservacion´Guanacaste (ACG), northwestern Costa Rica - caterpillars, pupae, butterflies & moths [WWW document]. Available from: http: // janzen. sas. upenn. edu / caterpillars / database. lasso [Accessed on 1 st January 2022]" title="Area de Conservacion" type="url" year="2022">Janzen & Hallwachs, 2022</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFD44B78FF9EFFE1ACC4AD54FAF21616" author="Murray" bookContentInfo="367 pp." box="[1227,1369,1583,1603]" firstAuthor="Murray" journalOrPublisher="Louisiana State University" pageId="17" pageNumber="18" refId="ref40253" refString="Murray, D. L. (2001 b) Systematics of euptychiine butterflies (Nymphalidae: Satyrinae: Euptychiina) based on larval morphology and DNA sequence data and the evolution of life history traits (PhD thesis). Louisiana State University. 367 pp." title="Systematics of euptychiine butterflies (Nymphalidae: Satyrinae: Euptychiina) based on larval morphology and DNA sequence data and the evolution of life history traits" type="book" year="2001" yearSuffix="b">Murray, 2001b</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFD44B78FF9EFFE1AD65AD54FC371630" author="Willmott" etAl="et al." firstAuthor="Willmott" journalOrPublisher="Tropical Lepidoptera Research" pageId="17" pageNumber="18" pagination="29 - 44" part="29" refId="ref43893" refString="Willmott, K. R., Marin, M. A., Nakahara, S., Pomerantz, T., Lamas, G., Huertas, B. et al. (2019) A revision of the new Andean butterfly genus Optimandes Marin, Nakahara & Willmott, n. gen., with the description of a new species (Nymphalidae: Satyrinae: Euptychiina). Tropical Lepidoptera Research, 29, 29 - 44." title="A revision of the new Andean butterfly genus Optimandes Marin, Nakahara & Willmott, n. gen., with the description of a new species (Nymphalidae: Satyrinae: Euptychiina)" type="journal article" year="2019">Willmott et al., 2019</bibRefCitation>
|
||
). These are morphologically simple and similar to those of several other species of Neotropical
|
||
<taxonomicName id="4C454D0AFF9EFFE1AC80AD0FFB5216DD" box="[1167,1273,1652,1672]" class="Insecta" family="Nymphalidae" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="53" phylum="Arthropoda" rank="subTribe" subTribe="Euptychiina">Euptychiina</taxonomicName>
|
||
larvae, lacking body scoli, and having short- to medium-sized head horns and caudal filaments, and the pupae are short and smooth.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |