156 lines
20 KiB
XML
156 lines
20 KiB
XML
<document id="50D601AA5E7A5BDC6C357B8002E0AAF9" ID-CLB-Dataset="30626" ID-DOI="10.11646/zootaxa.4387.3.4" ID-GBIF-Dataset="43de8956-0706-4f8d-b5a9-f51c30c276d8" ID-ISSN="1175-5326" ID-Zenodo-Dep="1187697" ID-ZooBank="B53A6B20-5386-4E5A-A00F-A559EDF640DF" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1520059483769" checkinUser="plazi" docAuthor="Fernández, Mercedes, Fernicola, Juan Carlos, Cerdeño, Esperanza & Reguero, Marcelo A." docDate="2018" docId="03EC879BCF5E8317C1922AFF8F42FA10" docLanguage="en" docName="zootaxa.4387.3.4.pdf" docOrigin="Zootaxa 4387 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Protypotherium compressidens Ameghino 1891" docType="treatment" docVersion="7" lastPageNumber="486" masterDocId="FFD5FFE3CF53831AC10528498E40FFD5" masterDocTitle="Identification of type materials of the species of Protypotherium Ameghino, 1885 and Patriarchus Ameghino, 1889 (Notoungulata: Interatheriidae) erected by Florentino Ameghino" masterLastPageNumber="498" masterPageNumber="473" pageNumber="486" updateTime="1698526258101" updateUser="ExternalLinkService">
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<mods:title id="B3A44AD04260A75BC621B2F3BA50E346">Identification of type materials of the species of Protypotherium Ameghino, 1885 and Patriarchus Ameghino, 1889 (Notoungulata: Interatheriidae) erected by Florentino Ameghino</mods:title>
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<mods:namePart id="29D617D4984F86C484AC5658D6445D66">Fernández, Mercedes</mods:namePart>
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<mods:roleTerm id="1275C39330C8588B21F485B1840D743B">Author</mods:roleTerm>
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<mods:namePart id="4ED4712AC2D656E171405039E1683474">Fernicola, Juan Carlos</mods:namePart>
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<mods:namePart id="BACA5374694BDAD62CF73F9F7238110D">Cerdeño, Esperanza</mods:namePart>
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<mods:namePart id="EACA4DB2A3141B6C781A29C83EFC22CD">Reguero, Marcelo A.</mods:namePart>
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<mods:title id="72BAFDE594A3D97F58ABDEEC505D07E8">Zootaxa</mods:title>
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<mods:date id="CB1B6FFD3DBC15BDD18892914F975D1F">2018</mods:date>
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<mods:number id="6C3886B2116E21714DEEE7FC8906E5B8">2018-02-28</mods:number>
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<mods:number id="D77FBDD91D075F2F506E8544E761152D">4387</mods:number>
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<treatment id="03EC879BCF5E8317C1922AFF8F42FA10" ID-DOI="http://doi.org/10.5281/zenodo.5986695" ID-GBIF-Taxon="141264028" ID-Zenodo-Dep="5986695" LSID="urn:lsid:plazi:treatment:03EC879BCF5E8317C1922AFF8F42FA10" httpUri="http://treatment.plazi.org/id/03EC879BCF5E8317C1922AFF8F42FA10" lastPageNumber="486" pageId="13" pageNumber="486">
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<subSubSection id="C35F6506CF5E8317C1922AFF8C9EFD05" box="[151,734,694,720]" pageId="13" pageNumber="486" type="nomenclature">
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<paragraph id="8BFA368DCF5E8317C1922AFF8C9EFD05" blockId="13.[151,734,694,720]" box="[151,734,694,720]" pageId="13" pageNumber="486">
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<heading id="D0B281E1CF5E8317C1922AFF8C9EFD05" bold="true" box="[151,734,694,720]" fontSize="11" level="1" pageId="13" pageNumber="486" reason="1">
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<taxonomicName id="4C454D0ECF5E8317C1922AFF8C9EFD05" authority="Ameghino, 1891" authorityName="Ameghino" authorityYear="1891" box="[151,734,694,720]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="13" pageNumber="486" phylum="Chordata" rank="species" species="compressidens">
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<emphasis id="B931EA9FCF5E8317C1922AFF8C9EFD05" bold="true" box="[151,734,694,720]" pageId="13" pageNumber="486">
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<emphasis id="B931EA9FCF5E8317C1922AFF8C4DFD05" bold="true" box="[151,525,694,720]" italics="true" pageId="13" pageNumber="486">Protypotherium compressidens</emphasis>
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<bibRefCitation id="EFD44B7CCF5E8317C3112AFF8C9EFD05" box="[532,734,694,720]" pageId="13" pageNumber="486" refId="ref17076" refString="Ameghino, F. (1891) Nuevos restos de mamiferos fosiles descubiertos por Carlos Ameghino en el Eoceno inferior de la Patagonia austral. Especies nuevas, adiciones y correcciones. Revista Argentina de Historia Natural, 1 (5), 289 - 328." type="journal article">Ameghino, 1891</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C35F6506CF5E8317C1922AB48DC7FCEF" pageId="13" pageNumber="486" type="materials_examined">
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<paragraph id="8BFA368DCF5E8317C1922AB48DC7FCEF" blockId="13.[151,1437,765,1477]" pageId="13" pageNumber="486">
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<emphasis id="B931EA9FCF5E8317C1922AB48F4BFCC3" bold="true" box="[151,267,765,790]" pageId="13" pageNumber="486">
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<typeStatus id="54FE882FCF5E8317C1922AB48F46FCC3" box="[151,262,765,790]" pageId="13" pageNumber="486" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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<collectionCode id="ED54AE48CF5E8317C0112AB78FC6FCC3" box="[276,390,766,790]" pageId="13" pageNumber="486">MACN-A</collectionCode>
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4029 (
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<figureCitation id="137E2A08CF5E8317C0DC2AB48C7EFCC3" box="[473,574,765,790]" captionStart="FIGURE 2" captionStartId="5.[151,249,1632,1654]" captionTargetBox="[308,1273,213,1595]" captionTargetId="figure@5.[305,1281,193,1611]" captionTargetPageId="5" captionText="FIGURE 2. Protypotherium attenuatum, syntype MACN-A 524, left maxilla with P2–M3 but lacking P1, in occlusal view (A); syntype MACN-A 628, right fragmented mandible with alveoli of i1–3 and c (broken)–m1 but lacking m2–3, in labial (B) and occlusal (C) views; reproductions of figures 21 (D), 20 (E) and 20a (F) from Ameghino (1889, plate 14). Protypotherium obstructum, holotype MACN-A 1677, lower premolar in labial, oclusal and lingual views (G); reproduction of figure 19 (H) from Ameghino (1889, plate 15). Protypotherium claudum, holotype MACN-A 551, right mandibular fragment with alveolus of p2 andp3–m2, in labial (I) and occlusal (J)views; reproductions of figures 22 (K) and 22a (L) from Ameghino (1889, plate 14). Protypotherium globosum, holotype MACN-A 4049, skull, in ventral (M) and dorsal (N) views. Protypotherium convexidens, holotype MACN-A 4054, right maxillary fragment with P3–M2 (broken), in occlusal view (O). Protypotherium diversidens, holotype MACN-A 4052, right maxillary fragment with alveolus of P1 and P2–M3, in occlusal view (P). Protypotheriumcompressidens, holotype MACN-A 4029, left maxillaryfragment withP3–M3, in occlusal view (Q); holotype MACN-A 4030, left mandibular fragment with p4 (talonid)–m3, in occlusal view (R). Protypotherium lineare, holotype MACN-A4039, right mandibular fragment with alveolus of p1 and p2–m1, in occlusal view (S); holotype MACN-A 4038, left mandibular fragment with p3–m2, in occlusal view (T). Scale bar = 10 mm." httpUri="https://zenodo.org/record/1187701/files/figure.png" pageId="13" pageNumber="486">Fig. 2 Q</figureCitation>
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), left maxillary fragment with P3–M3, and
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<collectionCode id="ED54AE48CF5E8317C5322AB78AE9FCC3" box="[1079,1193,766,790]" pageId="13" pageNumber="486">MACN-A</collectionCode>
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4030 (
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<figureCitation id="137E2A08CF5E8317C5F92AB48B21FCC3" box="[1276,1377,765,790]" captionStart="FIGURE 2" captionStartId="5.[151,249,1632,1654]" captionTargetBox="[308,1273,213,1595]" captionTargetId="figure@5.[305,1281,193,1611]" captionTargetPageId="5" captionText="FIGURE 2. Protypotherium attenuatum, syntype MACN-A 524, left maxilla with P2–M3 but lacking P1, in occlusal view (A); syntype MACN-A 628, right fragmented mandible with alveoli of i1–3 and c (broken)–m1 but lacking m2–3, in labial (B) and occlusal (C) views; reproductions of figures 21 (D), 20 (E) and 20a (F) from Ameghino (1889, plate 14). Protypotherium obstructum, holotype MACN-A 1677, lower premolar in labial, oclusal and lingual views (G); reproduction of figure 19 (H) from Ameghino (1889, plate 15). Protypotherium claudum, holotype MACN-A 551, right mandibular fragment with alveolus of p2 andp3–m2, in labial (I) and occlusal (J)views; reproductions of figures 22 (K) and 22a (L) from Ameghino (1889, plate 14). Protypotherium globosum, holotype MACN-A 4049, skull, in ventral (M) and dorsal (N) views. Protypotherium convexidens, holotype MACN-A 4054, right maxillary fragment with P3–M2 (broken), in occlusal view (O). Protypotherium diversidens, holotype MACN-A 4052, right maxillary fragment with alveolus of P1 and P2–M3, in occlusal view (P). Protypotheriumcompressidens, holotype MACN-A 4029, left maxillaryfragment withP3–M3, in occlusal view (Q); holotype MACN-A 4030, left mandibular fragment with p4 (talonid)–m3, in occlusal view (R). Protypotherium lineare, holotype MACN-A4039, right mandibular fragment with alveolus of p1 and p2–m1, in occlusal view (S); holotype MACN-A 4038, left mandibular fragment with p3–m2, in occlusal view (T). Scale bar = 10 mm." httpUri="https://zenodo.org/record/1187701/files/figure.png" pageId="13" pageNumber="486">Fig. 2 R</figureCitation>
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), left mandibular fragment with p4 (talonid)–m3, of the same individual.
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</paragraph>
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</subSubSection>
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<subSubSection id="C35F6506CF5E8317C1C22B0C8CA8FBEC" pageId="13" pageNumber="486" type="diagnosis">
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<paragraph id="8BFA368DCF5E8317C1C22B0C8F61FC18" blockId="13.[151,1437,765,1477]" pageId="13" pageNumber="486">
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<emphasis id="B931EA9FCF5E8317C1C22B0C8F7AFC8B" bold="true" box="[199,314,837,862]" pageId="13" pageNumber="486">Diagnosis</emphasis>
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(
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<bibRefCitation id="EFD44B7CCF5E8317C04A2B0C8C11FC8B" box="[335,593,837,862]" pageId="13" pageNumber="486" refId="ref17076" refString="Ameghino, F. (1891) Nuevos restos de mamiferos fosiles descubiertos por Carlos Ameghino en el Eoceno inferior de la Patagonia austral. Especies nuevas, adiciones y correcciones. Revista Argentina de Historia Natural, 1 (5), 289 - 328." type="journal article">Ameghino 1891: 292</bibRefCitation>
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). “
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<emphasis id="B931EA9FCF5E8317C37D2B0F8D30FC88" box="[632,880,837,862]" italics="true" pageId="13" pageNumber="486">Tamaño casi igual a</emphasis>
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<taxonomicName id="4C454D0ECF5E8317C2782B0C8AD2FC8B" authorityName="Ameghino. Se" baseAuthorityName="Moreno" baseAuthorityYear="1882" box="[893,1170,837,862]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="13" pageNumber="486" phylum="Chordata" rank="species" species="australe">Protypotherium australe</taxonomicName>
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,
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<emphasis id="B931EA9FCF5E8317C5A22B0F8F4EFC18" italics="true" pageId="13" pageNumber="486">pero con las muelas comprimidas lateralmente y mucho más angostas. Verdaderos molares superiores de cara externa suavemente ondulada. Longitud del p.3 al m.3, 28 milímetros. Ancho máximo de los verdaderos molares superiores, 3 milímetros</emphasis>
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”.
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</paragraph>
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<paragraph id="8BFA368DCF5E8317C1C22B918CA8FBEC" blockId="13.[151,1437,765,1477]" pageId="13" pageNumber="486">
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English translation. Almost equal size to
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<taxonomicName id="4C454D0ECF5E8317C3A32B918DFDFC24" authorityName="Ameghino. Se" baseAuthorityName="Moreno" baseAuthorityYear="1882" box="[678,957,984,1009]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="13" pageNumber="486" phylum="Chordata" rank="species" species="australe">
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<emphasis id="B931EA9FCF5E8317C3A32B918DFDFC24" box="[678,957,984,1009]" italics="true" pageId="13" pageNumber="486">Protypotherium australe</emphasis>
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</taxonomicName>
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, but with laterally compressed and much narrower teeth. Upper molars with slightly undulated external face. Length from P3 to M3, 28 millimetres. Maximum width of true upper molars, 3 millimetres.
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</paragraph>
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</subSubSection>
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<subSubSection id="C35F6506CF5E8317C1C22C0D8F42FA10" pageId="13" pageNumber="486" type="discussion">
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<paragraph id="8BFA368DCF5E8317C1C22C0D8C80FAA9" blockId="13.[151,1437,765,1477]" pageId="13" pageNumber="486">
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<emphasis id="B931EA9FCF5E8317C1C22C0D8F0FFB88" bold="true" box="[199,335,1092,1117]" pageId="13" pageNumber="486">Comments.</emphasis>
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<bibRefCitation id="EFD44B7CCF5E8317C0592C0D8C6AFB88" box="[348,554,1092,1117]" pageId="13" pageNumber="486" refId="ref17076" refString="Ameghino, F. (1891) Nuevos restos de mamiferos fosiles descubiertos por Carlos Ameghino en el Eoceno inferior de la Patagonia austral. Especies nuevas, adiciones y correcciones. Revista Argentina de Historia Natural, 1 (5), 289 - 328." type="journal article">Ameghino (1891)</bibRefCitation>
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did not add any illustration to his description. According to Ameghino’s catalogue and to
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<bibRefCitation id="EFD44B7CCF5E8317C0752C208C59FB54" box="[368,537,1128,1153]" pageId="13" pageNumber="486" refId="ref18248" refString="Mones A. (1986) Palaeovertebrata Sudamericana. Catalogo Sistematico de los Vertebrados Fosiles de America del Sur, Parte I. Preliminary List and Bibliography. Courier Forschungsinstitut Senckenberg, 82, 1 - 625." type="journal article">Mones (1986)</bibRefCitation>
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, MACN-A 4029 and MACN-A 4030, fragmented maxilla and mandible, respectively, are the
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<typeStatus id="54FE882FCF5E8317C08C2CC48FF9FB70" box="[393,441,1165,1189]" pageId="13" pageNumber="486">type</typeStatus>
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specimens of the species; following Ameghino’s catalogue, both belong to the same individual, a fact that was corroborated by us since they articulate. These specimens are the only available materials assigned to
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<taxonomicName id="4C454D0ECF5E8317C08C2C9C8C09FB38" authorityName="Ameghino" authorityYear="1891" box="[393,585,1236,1261]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="13" pageNumber="486" phylum="Chordata" rank="species" species="compressidens">
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<emphasis id="B931EA9FCF5E8317C08C2C9C8C09FB38" box="[393,585,1236,1261]" italics="true" pageId="13" pageNumber="486">P. compressidens</emphasis>
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</taxonomicName>
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, and both match the original diagnosis (
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<bibRefCitation id="EFD44B7CCF5E8317C50E2C9D8A84FB38" box="[1035,1220,1236,1261]" pageId="13" pageNumber="486" refId="ref17076" refString="Ameghino, F. (1891) Nuevos restos de mamiferos fosiles descubiertos por Carlos Ameghino en el Eoceno inferior de la Patagonia austral. Especies nuevas, adiciones y correcciones. Revista Argentina de Historia Natural, 1 (5), 289 - 328." type="journal article">Ameghino 1891</bibRefCitation>
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) because the upper and lower teeth are strongly labio-lingually compressed. Besides, the length of P3–M3 of MACN-A 4029 is
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<quantity id="4CBD9B68CF5E8317C46E2CB18E85FAE1" metricMagnitude="-2" metricUnit="m" metricValue="2.86" pageId="13" pageNumber="486" unit="mm" value="28.6">28.6 mm</quantity>
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, and the maximum width of the upper molars (M1 being the widest) is
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<quantity id="4CBD9B68CF5E8317C2FA2D548A19FAE1" box="[1023,1113,1309,1333]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="13" pageNumber="486" unit="mm" value="3.4">3.4 mm</quantity>
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, which are similar values to those provided by
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<bibRefCitation id="EFD44B7CCF5E8317C0752D098C7EFA8D" box="[368,574,1344,1369]" pageId="13" pageNumber="486" refId="ref17076" refString="Ameghino, F. (1891) Nuevos restos de mamiferos fosiles descubiertos por Carlos Ameghino en el Eoceno inferior de la Patagonia austral. Especies nuevas, adiciones y correcciones. Revista Argentina de Historia Natural, 1 (5), 289 - 328." type="journal article">Ameghino (1891)</bibRefCitation>
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. Therefore, we confirm that the individual MACN-A 4029 plus MACN-A 4030 constitute the
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<typeStatus id="54FE882FCF5E8317C0762D2D8F95FAA8" box="[371,469,1380,1405]" pageId="13" pageNumber="486" type="holotype">holotype</typeStatus>
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of
|
||
<taxonomicName id="4C454D0ECF5E8317C0FF2D2C8CFAFAA8" authorityName="Ameghino" authorityYear="1891" box="[506,698,1380,1405]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="13" pageNumber="486" phylum="Chordata" rank="species" species="compressidens">
|
||
<emphasis id="B931EA9FCF5E8317C0FF2D2C8CFAFAA8" box="[506,698,1380,1405]" italics="true" pageId="13" pageNumber="486">P. compressidens</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BFA368DCF5E8317C1C22DC18F42FA10" blockId="13.[151,1437,765,1477]" pageId="13" pageNumber="486">
|
||
<materialsCitation id="3B2D3CD0CF5E8317C1C22DC18F42FA10" ID-GBIF-Occurrence="1828873148" collectionCode="SALMA" location="Santa Cruz Province" pageId="13" pageNumber="486" specimenCount="1" stateProvince="Santa Cruz Province" typeStatus="holotype">
|
||
<emphasis id="B931EA9FCF5E8317C1C22DC18DBEFA74" bold="true" box="[199,1022,1416,1441]" pageId="13" pageNumber="486">
|
||
Chronological and geographical distribution of the
|
||
<typeStatus id="54FE882FCF5E8317C24C2DC18D3DFA74" box="[841,893,1416,1441]" pageId="13" pageNumber="486">type</typeStatus>
|
||
specimen.
|
||
</emphasis>
|
||
Santacrucian
|
||
<collectionCode id="ED54AE48CF5E8317C5A32DC18B4BFA75" box="[1190,1291,1416,1440]" pageId="13" pageNumber="486">SALMA</collectionCode>
|
||
,
|
||
<collectingRegion id="4981F86FCF5E8317C41E2DC18EBFFA10" country="Argentina" name="Santa Cruz" pageId="13" pageNumber="486">
|
||
Santa
|
||
<collectingRegion id="4981F86FCF5E8317C4632DC18EBFFA10" inferredBySuffix="true" pageId="13" pageNumber="486">Cruz Province</collectingRegion>
|
||
</collectingRegion>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |