209 lines
25 KiB
XML
209 lines
25 KiB
XML
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<mods:title id="897B91B9511F7FEADCBF9B0CADD5BF19">Review of the subgenus Lyrothorax of the genus Pterostichus (Coleoptera: Carabidae) in the Western Palaearctic, with descriptions of two new species</mods:title>
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<mods:namePart id="4B7C4FC21E5592C0C5E36E7E6ACD33DF">Muilwijk, Jan</mods:namePart>
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<mods:nameIdentifier id="EF0394A8EA3C7A87E8143B384A21621A" type="ORCID">0000-0002-0230-226X</mods:nameIdentifier>
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<mods:nameIdentifier id="C85CA8DBAE288ADEA03957A140C160CC" type="email">uilwijk@gmail.com</mods:nameIdentifier>
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<mods:namePart id="BF599B1A7BFFD9F799AD6E1ED02365E1">Kataev, Boris M.</mods:namePart>
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<mods:nameIdentifier id="36849D01E275B38A7A284A5D47E38FE1" type="ORCID">0000-0001-9890-2595</mods:nameIdentifier>
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<mods:nameIdentifier id="9393F9A878B8F9B4486E8DD0DA5CDF2A" type="email">harpal@zin.ru</mods:nameIdentifier>
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<mods:namePart id="193CBE3CA176BABF593C5A2FFABDF62E">Wrase, David W.</mods:namePart>
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<mods:nameIdentifier id="4FCC632EE82165D593EFDF648C0699DB" type="ORCID">0000-0003-1779-2684</mods:nameIdentifier>
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<mods:nameIdentifier id="329541B9D1BD2F9B6A2839EB1F892F68" type="email">carterus@gmx.de</mods:nameIdentifier>
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<mods:title id="8666E19D0DB2DC4C25089CBBD936E71D">Zootaxa</mods:title>
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<mods:part id="60F1B2EBF475B36B822DC6FF548B4CA5">
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<mods:date id="3BEA0042C71FD60BF16D90261A574370">2024</mods:date>
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<mods:number id="5FF807E6780A8980F6A8493BAEFFA8E0">2024-06-12</mods:number>
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<treatment id="039487C7FFB1BB66FF66FDA3989DFAAF" ID-DOI="http://doi.org/10.5281/zenodo.11621085" ID-GBIF-Taxon="232746601" ID-Zenodo-Dep="11621085" LSID="urn:lsid:plazi:treatment:039487C7FFB1BB66FF66FDA3989DFAAF" httpUri="http://treatment.plazi.org/id/039487C7FFB1BB66FF66FDA3989DFAAF" lastPageNumber="273" pageId="18" pageNumber="273">
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<subSubSection id="C327655AFFB1BB66FF66FDA39971FD64" pageId="18" pageNumber="273" type="nomenclature">
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<paragraph id="8B8236D1FFB1BB66FF66FDA39EB3FD41" blockId="18.[151,1136,632,695]" box="[151,946,632,659]" pageId="18" pageNumber="273">
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<heading id="D0CA81BDFFB1BB66FF66FDA39EB3FD41" bold="true" box="[151,946,632,659]" fontSize="11" level="1" pageId="18" pageNumber="273" reason="1">
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<emphasis id="B949EAC3FFB1BB66FF66FDA39EB3FD41" bold="true" box="[151,946,632,659]" pageId="18" pageNumber="273">
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Key to Western Palaearctic
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<taxonomicName id="4C3D4D52FFB1BB66FE02FDA39F7AFD40" authority="Chaudoir, 1838" authorityName="Chaudoir" authorityYear="1838" box="[499,635,632,659]" class="Insecta" family="Carabidae" genus="Lyrothorax" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="274" phylum="Arthropoda" rank="subGenus" subGenus="Lyrothorax">
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<emphasis id="B949EAC3FFB1BB66FE02FDA39F7AFD40" bold="true" box="[499,635,632,659]" italics="true" pageId="18" pageNumber="273">Lyrothorax</emphasis>
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</taxonomicName>
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species based on habitus
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B8236D1FFB1BB66FF66FD469971FD64" blockId="18.[151,1136,632,695]" box="[151,1136,668,695]" pageId="18" pageNumber="273">Because of variation between populations all characteristics must be taken into account.</paragraph>
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</subSubSection>
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<subSubSection id="C327655AFFB1BB66FF66FD3A989DFAAF" pageId="18" pageNumber="273" type="key">
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<paragraph id="8B8236D1FFB1BB66FF66FD3A989DFCBF" blockId="18.[151,1437,737,1405]" pageId="18" pageNumber="273">
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<keyLead id="30CC95E4FFB1BB66FF66FD3A989DFCBF" pageId="18" pageNumber="273">
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<specimenCount id="9D3BFD58FFB1BB66FF66FD3A9C4BFD24" box="[151,330,737,759]" count="1" pageId="18" pageNumber="273" type="generic">1 Specimens</specimenCount>
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from
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<collectingCountry id="F32A7641FFB1BB66FE72FD3A9F1FFD24" box="[387,542,737,759]" name="Czech Republic" pageId="18" pageNumber="273">Czech Republic</collectingCountry>
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.
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<location id="8EE2600AFFB1BB66FDD7FD3A9F70FD24" LSID="urn:lsid:plazi:treatment:039487C7FFB1BB66FF66FDA3989DFAAF:8EE2600AFFB1BB66FDD7FD3A9F70FD24" box="[550,625,737,759]" country="Czech Republic" county="Base" municipality="Basal" name="Habitus" pageId="18" pageNumber="273">Habitus</location>
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as in
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<figureCitation id="13062A54FFB1BB66FD56FD3A9FEDFD25" box="[679,748,737,759]" captionStart="FIGURE 2" captionStartId="6.[151,250,1236,1260]" captionTargetBox="[154,1435,182,1213]" captionTargetId="figure-29@6.[151,1435,181,1213]" captionTargetPageId="6" captionText="FIGURE 2. Habitus in dorsal view and pronotum increased, (a) P. caspius (Ménétriés, 1832) from Talysh, (b) P. fritzei sp. n. holotype, (c) P. vagus sp. n., from different localities, c1 holotype, c2 from Bylnice, Czech Republic. Scale bar =10 mm." figureDoi="http://doi.org/10.5281/zenodo.11615568" httpUri="https://zenodo.org/record/11615568/files/figure.png" pageId="18" pageNumber="273">Fig. 2c</figureCitation>
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.
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<location id="8EE2600AFFB1BB66FD05FD399E26FD24" LSID="urn:lsid:plazi:treatment:039487C7FFB1BB66FF66FDA3989DFAAF:8EE2600AFFB1BB66FD05FD399E26FD24" box="[756,807,738,759]" country="Czech Republic" county="Base" municipality="Basal" name="Inner" pageId="18" pageNumber="273">Inner</location>
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and outer basal foveae of pronotum joined, outer foveae normally separated from lateral margin by a long keel, inner foveae deep and long, longer than outer foveae.
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<location id="8EE2600AFFB1BB66FB65FD2499C5FCC6" LSID="urn:lsid:plazi:treatment:039487C7FFB1BB66FF66FDA3989DFAAF:8EE2600AFFB1BB66FB65FD2499C5FCC6" box="[1172,1220,767,789]" country="Czech Republic" county="Base" municipality="Basal" name="Hind" pageId="18" pageNumber="273">Hind</location>
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angles of pronotum as a rule right-angled, or weakly obtuse-angled, blunt at tip.
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<location id="8EE2600AFFB1BB66FCF4FCC79E57FCE1" LSID="urn:lsid:plazi:treatment:039487C7FFB1BB66FF66FDA3989DFAAF:8EE2600AFFB1BB66FCF4FCC79E57FCE1" box="[773,854,796,818]" country="Czech Republic" county="Base" municipality="Basal" name="Pronotal" pageId="18" pageNumber="273">Pronotal</location>
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sides converging towards base.
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<location id="8EE2600AFFB1BB66FB7BFCC799C4FCE1" LSID="urn:lsid:plazi:treatment:039487C7FFB1BB66FF66FDA3989DFAAF:8EE2600AFFB1BB66FB7BFCC799C4FCE1" box="[1162,1221,796,818]" country="Czech Republic" county="Base" municipality="Basal" name="Elytra" pageId="18" pageNumber="273">Elytra</location>
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and pronotum slightly shiny.
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<collectingMunicipality id="6BE6ACABFFB1BB66FED0FCE29C57FC9C" box="[289,342,825,847]" pageId="18" pageNumber="273">Basal</collectingMunicipality>
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margin of elytra more or less angularly merging into lateral margin.
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<collectingCounty id="62E34E5DFFB1BB66FC07FCE19924FC9C" box="[1014,1061,826,847]" pageId="18" pageNumber="273">Base</collectingCounty>
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of elytra more or less hooked towards lateral margins. BL: 10.0–
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<quantity id="4CC59B34FFB1BB66FE2EFC8D9F32FCBF" box="[479,563,854,876]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" pageId="18" pageNumber="273" unit="mm" value="12.5">12.5 mm</quantity>
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............................................................
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<taxonomicName id="4C3D4D52FFB1BB66FAE0FC8C9861FCBF" authority="Muilwijk & Kataev & Wrase, 2024" authorityName="Muilwijk & Kataev & Wrase" authorityYear="2024" box="[1297,1376,855,876]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="273" phylum="Arthropoda" rank="species" species="vagus" status="sp. n.">
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<emphasis id="B949EAC3FFB1BB66FAE0FC8C9861FCBF" box="[1297,1376,855,876]" italics="true" pageId="18" pageNumber="273">P. vagus</emphasis>
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</taxonomicName>
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<emphasis id="B949EAC3FFB1BB66FA97FC8C9899FCBF" bold="true" box="[1382,1432,855,876]" pageId="18" pageNumber="273">sp. n</emphasis>
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</taxonomicNameLabel>
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<emphasis id="B949EAC3FFB1BB66FA66FC8C989DFCBF" bold="true" box="[1431,1436,855,876]" pageId="18" pageNumber="273">.</emphasis>
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</keyLead>
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</paragraph>
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<paragraph id="8B8236D1FFB1BB66FF66FCAF989DFC5A" blockId="18.[151,1437,737,1405]" box="[151,1436,884,906]" pageId="18" pageNumber="273">
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<keyLead id="30CC95E4FFB1BB66FF66FCAF989DFC5A" box="[151,1436,884,906]" pageId="18" pageNumber="273">
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- Specimens from
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<collectingCountry id="F32A7641FFB1BB66FE75FCAF9CAAFC5A" box="[388,427,884,905]" name="Iran" pageId="18" pageNumber="273">Iran</collectingCountry>
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and
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<collectingCountry id="F32A7641FFB1BB66FE29FCAF9F40FC59" box="[472,577,884,906]" name="Azerbaijan" pageId="18" pageNumber="273">Azerbaijan</collectingCountry>
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...................................................................... 2
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</keyLead>
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</paragraph>
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||
</keyStep>
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||
<keyStep id="30C92E74FFB1BB66FF66FC4A989DFB42" pageId="18" pageNumber="273">
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<paragraph id="8B8236D1FFB1BB66FF66FC4A989DFB85" blockId="18.[151,1437,737,1405]" pageId="18" pageNumber="273">
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<keyLead id="30CC95E4FFB1BB66FF66FC4A989DFB85" pageId="18" pageNumber="273">
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||
2 Basal foveae of pronotum without punctures or only with fine punctures. Inner and outer basal foveae of pronotum joined, outer foveae normally separated from lateral margin by a long keel, inner foveae deep and long, longer than outer foveae. Pronotal hind angles as a rule right-angled, or weakly obtuse-angled, blunt at tip. Pronotal sides converging towards base (
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||
<figureCitation id="13062A54FFB1BB66FAB9FC179890FC32" box="[1352,1425,972,994]" captionStart="FIGURE 2" captionStartId="6.[151,250,1236,1260]" captionTargetBox="[154,1435,182,1213]" captionTargetId="figure-29@6.[151,1435,181,1213]" captionTargetPageId="6" captionText="FIGURE 2. Habitus in dorsal view and pronotum increased, (a) P. caspius (Ménétriés, 1832) from Talysh, (b) P. fritzei sp. n. holotype, (c) P. vagus sp. n., from different localities, c1 holotype, c2 from Bylnice, Czech Republic. Scale bar =10 mm." figureDoi="http://doi.org/10.5281/zenodo.11615568" httpUri="https://zenodo.org/record/11615568/files/figure.png" pageId="18" pageNumber="273">Fig. 2c</figureCitation>
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). Elytra and pronotum slightly shiny. Basal margin of elytra more or less angularly merging into lateral margin. Punctation of metepisternum clearly less dense than that of metasternum, punctures even coarse (
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<figureCitation id="13062A54FFB1BB66FC09FBDC993FFBCF" box="[1016,1086,1031,1053]" captionStart="FIGURE 4" captionStartId="7.[151,250,1790,1814]" captionTargetBox="[155,1429,1227,1759]" captionTargetId="figure-52@7.[151,1435,1227,1765]" captionTargetPageId="7" captionText="FIGURE 4. Metepisterna and metasternum, (a) P. caspius (Ménétriés, 1832) from Gilan, South Siahkal, (b) P. fritzei sp. n. holotype, (c) P. vagus sp. n. holotype. c1 holotype, c2 from Asålem Pass." figureDoi="http://doi.org/10.5281/zenodo.11615578" httpUri="https://zenodo.org/record/11615578/files/figure.png" pageId="18" pageNumber="273">Fig. 4c</figureCitation>
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). On average larger; specimens from
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<collectingRegion id="49F9F833FFB1BB66FF13FBFF9C36FBE9" box="[226,311,1060,1082]" country="Iran" name="Golestan" pageId="18" pageNumber="273">Golestan</collectingRegion>
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||
clearly larger than specimens from
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||
<collectingRegion id="49F9F833FFB1BB66FD61FBFF9E0BFBE9" box="[656,778,1060,1082]" country="Iran" name="Mazandaran" pageId="18" pageNumber="273">Māzandarān</collectingRegion>
|
||
. BL:
|
||
<quantity id="4CC59B34FFB1BB66FCCFFBFF9ECDFBEA" box="[830,972,1060,1082]" metricMagnitude="-2" metricUnit="m" metricValue="1.43" metricValueMax="1.61" metricValueMin="1.25" pageId="18" pageNumber="273" unit="mm" value="14.3" valueMax="16.1" valueMin="12.5">12.5–16.1 mm</quantity>
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||
.
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||
<collectingCountry id="F32A7641FFB1BB66FC24FBFF9EFFFBEA" box="[981,1022,1060,1081]" name="Iran" pageId="18" pageNumber="273">Iran</collectingCountry>
|
||
,
|
||
<collectingRegion id="49F9F833FFB1BB66FBF9FBFF997EFBE9" box="[1032,1151,1060,1082]" country="Iran" name="Mazandaran" pageId="18" pageNumber="273">Māzandarān</collectingRegion>
|
||
and
|
||
<collectingRegion id="49F9F833FFB1BB66FB5CFBFF9803FBE9" box="[1197,1282,1060,1082]" country="Iran" name="Golestan" pageId="18" pageNumber="273">Golestan</collectingRegion>
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||
....................................................................................................
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<taxonomicName id="4C3D4D52FFB1BB66FAE0FB999861FB84" authority="Muilwijk & Kataev & Wrase, 2024" authorityName="Muilwijk & Kataev & Wrase" authorityYear="2024" box="[1297,1376,1090,1111]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="273" phylum="Arthropoda" rank="species" species="vagus" status="sp. n.">
|
||
<emphasis id="B949EAC3FFB1BB66FAE0FB999861FB84" box="[1297,1376,1090,1111]" italics="true" pageId="18" pageNumber="273">P. vagus</emphasis>
|
||
</taxonomicName>
|
||
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|
||
<emphasis id="B949EAC3FFB1BB66FA97FB9A9899FB85" bold="true" box="[1382,1432,1089,1110]" pageId="18" pageNumber="273">sp. n</emphasis>
|
||
</taxonomicNameLabel>
|
||
<emphasis id="B949EAC3FFB1BB66FA66FB9A989DFB85" bold="true" box="[1431,1436,1089,1110]" pageId="18" pageNumber="273">.</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B8236D1FFB1BB66FF66FB84989DFB42" blockId="18.[151,1437,737,1405]" pageId="18" pageNumber="273">
|
||
<keyLead id="30CC95E4FFB1BB66FF66FB84989DFB42" pageId="18" pageNumber="273">
|
||
- Basal foveae of pronotum with at least some distinct punctures. Inner foveae less deep, shorter (
|
||
<figureCitation id="13062A54FFB1BB66FB73FB8499F0FBA6" box="[1154,1265,1119,1141]" captionStart="FIGURE 2" captionStartId="6.[151,250,1236,1260]" captionTargetBox="[154,1435,182,1213]" captionTargetId="figure-29@6.[151,1435,181,1213]" captionTargetPageId="6" captionText="FIGURE 2. Habitus in dorsal view and pronotum increased, (a) P. caspius (Ménétriés, 1832) from Talysh, (b) P. fritzei sp. n. holotype, (c) P. vagus sp. n., from different localities, c1 holotype, c2 from Bylnice, Czech Republic. Scale bar =10 mm." figureDoi="http://doi.org/10.5281/zenodo.11615568" httpUri="https://zenodo.org/record/11615568/files/figure.png" pageId="18" pageNumber="273">Figs 2a, 2b</figureCitation>
|
||
). Basal margin of elytra continuing fluently into lateral margin. On average smaller. BL:
|
||
<quantity id="4CC59B34FFB1BB66FC78FBA79915FB42" box="[905,1044,1148,1170]" metricMagnitude="-2" metricUnit="m" metricValue="1.27" metricValueMax="1.49" metricValueMin="1.05" pageId="18" pageNumber="273" unit="mm" value="12.7" valueMax="14.9" valueMin="10.5">10.5–14.9 mm</quantity>
|
||
............................... 3
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30C92E74FFB1BB66FF66FB42989DFAAF" pageId="18" pageNumber="273">
|
||
<paragraph id="8B8236D1FFB1BB66FF66FB42989DFAD5" blockId="18.[151,1437,737,1405]" pageId="18" pageNumber="273">
|
||
<keyLead id="30CC95E4FFB1BB66FF66FB42989DFAD5" pageId="18" pageNumber="273">
|
||
3 Pronotal hind angles as a rule right-angled. Pronotal sides converging towards base. Elytra in comparison more elongated, less convex, with inner intervals more convex (
|
||
<figureCitation id="13062A54FFB1BB66FD56FB6C9FECFB1F" box="[679,749,1207,1229]" captionStart="FIGURE 2" captionStartId="6.[151,250,1236,1260]" captionTargetBox="[154,1435,182,1213]" captionTargetId="figure-29@6.[151,1435,181,1213]" captionTargetPageId="6" captionText="FIGURE 2. Habitus in dorsal view and pronotum increased, (a) P. caspius (Ménétriés, 1832) from Talysh, (b) P. fritzei sp. n. holotype, (c) P. vagus sp. n., from different localities, c1 holotype, c2 from Bylnice, Czech Republic. Scale bar =10 mm." figureDoi="http://doi.org/10.5281/zenodo.11615568" httpUri="https://zenodo.org/record/11615568/files/figure.png" pageId="18" pageNumber="273">Fig. 2a</figureCitation>
|
||
). Head in comparison wide. Punctation on pronotum as a rule restricted to basal foveae. Punctation of metepisternum less coarse than that of metasternum (
|
||
<figureCitation id="13062A54FFB1BB66FBEEFB0F9966FB3A" box="[1055,1127,1236,1258]" captionStart="FIGURE 4" captionStartId="7.[151,250,1790,1814]" captionTargetBox="[155,1429,1227,1759]" captionTargetId="figure-52@7.[151,1435,1227,1765]" captionTargetPageId="7" captionText="FIGURE 4. Metepisterna and metasternum, (a) P. caspius (Ménétriés, 1832) from Gilan, South Siahkal, (b) P. fritzei sp. n. holotype, (c) P. vagus sp. n. holotype. c1 holotype, c2 from Asålem Pass." figureDoi="http://doi.org/10.5281/zenodo.11615578" httpUri="https://zenodo.org/record/11615578/files/figure.png" pageId="18" pageNumber="273">Fig. 4a</figureCitation>
|
||
).
|
||
<collectingRegion id="49F9F833FFB1BB66FB88FB0F99E3FB39" box="[1145,1250,1236,1258]" country="Iran" name="Azarbayjan-e Sharqi" pageId="18" pageNumber="273">Azerbaijan</collectingRegion>
|
||
and
|
||
<collectingCountry id="F32A7641FFB1BB66FAE7FB0F983CFB3A" box="[1302,1341,1236,1257]" name="Iran" pageId="18" pageNumber="273">Iran</collectingCountry>
|
||
(
|
||
<collectingRegion id="49F9F833FFB1BB66FABAFB0F9899FB39" box="[1355,1432,1236,1258]" country="Iran" name="Ardabil" pageId="18" pageNumber="273">Ardebil</collectingRegion>
|
||
,
|
||
<collectingRegion id="49F9F833FFB1BB66FF13FB2A9C18FAD4" box="[226,281,1265,1287]" country="Iran" name="Gilan" pageId="18" pageNumber="273">Gilan</collectingRegion>
|
||
).................................................................................
|
||
<taxonomicName id="4C3D4D52FFB1BB66FB04FB29989DFAD5" ID-CoL="4Q3WG" authority="(Men.)" baseAuthorityName="Menetries" baseAuthorityYear="1832" box="[1269,1436,1265,1287]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="273" phylum="Arthropoda" rank="species" species="caspius">
|
||
<emphasis id="B949EAC3FFB1BB66FB04FB299855FAD4" box="[1269,1364,1266,1287]" italics="true" pageId="18" pageNumber="273">P. caspius</emphasis>
|
||
(Mén.)
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B8236D1FFB1BB66FF66FAD4989DFAAF" blockId="18.[151,1437,737,1405]" pageId="18" pageNumber="273">
|
||
<keyLead id="30CC95E4FFB1BB66FF66FAD4989DFAAF" pageId="18" pageNumber="273">
|
||
- Pronotal hind angles as a rule acute-angled, distinctly directed outwards. Pronotal sides strongly converging towards base. Elytra in comparison more oval and more convex, with inner intervals flatter (
|
||
<figureCitation id="13062A54FFB1BB66FC2AFAF79925FA91" box="[987,1060,1324,1346]" captionStart="FIGURE 2" captionStartId="6.[151,250,1236,1260]" captionTargetBox="[154,1435,182,1213]" captionTargetId="figure-29@6.[151,1435,181,1213]" captionTargetPageId="6" captionText="FIGURE 2. Habitus in dorsal view and pronotum increased, (a) P. caspius (Ménétriés, 1832) from Talysh, (b) P. fritzei sp. n. holotype, (c) P. vagus sp. n., from different localities, c1 holotype, c2 from Bylnice, Czech Republic. Scale bar =10 mm." figureDoi="http://doi.org/10.5281/zenodo.11615568" httpUri="https://zenodo.org/record/11615568/files/figure.png" pageId="18" pageNumber="273">Fig. 2b</figureCitation>
|
||
). Head in comparison less wide. Basal foveae of pronotum as a rule densely punctured, also between them with scattered punctures. Punctation of metepisternum even coarser than that of metasternum (
|
||
<figureCitation id="13062A54FFB1BB66FDDBFABC9F73FAAE" box="[554,626,1383,1405]" captionStart="FIGURE 4" captionStartId="7.[151,250,1790,1814]" captionTargetBox="[155,1429,1227,1759]" captionTargetId="figure-52@7.[151,1435,1227,1765]" captionTargetPageId="7" captionText="FIGURE 4. Metepisterna and metasternum, (a) P. caspius (Ménétriés, 1832) from Gilan, South Siahkal, (b) P. fritzei sp. n. holotype, (c) P. vagus sp. n. holotype. c1 holotype, c2 from Asålem Pass." figureDoi="http://doi.org/10.5281/zenodo.11615578" httpUri="https://zenodo.org/record/11615578/files/figure.png" pageId="18" pageNumber="273">Fig. 4b</figureCitation>
|
||
).
|
||
<collectingCountry id="F32A7641FFB1BB66FD73FABC9FAAFAAF" box="[642,683,1383,1404]" name="Iran" pageId="18" pageNumber="273">Iran</collectingCountry>
|
||
, from
|
||
<collectingRegion id="49F9F833FFB1BB66FD1BFABC9E1EFAAE" box="[746,799,1383,1405]" country="Iran" name="Gilan" pageId="18" pageNumber="273">Gilan</collectingRegion>
|
||
to East
|
||
<collectingRegion id="49F9F833FFB1BB66FC9CFABC9EC3FAAE" box="[877,962,1383,1405]" country="Iran" name="Golestan" pageId="18" pageNumber="273">Golestan</collectingRegion>
|
||
..........................
|
||
<taxonomicName id="4C3D4D52FFB1BB66FAE0FABC9861FAAF" authority="Muilwijk & Kataev & Wrase, 2024" authorityName="Muilwijk & Kataev & Wrase" authorityYear="2024" box="[1297,1376,1383,1404]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="273" phylum="Arthropoda" rank="species" species="fritzei" status="sp. n.">
|
||
<emphasis id="B949EAC3FFB1BB66FAE0FABC9861FAAF" box="[1297,1376,1383,1404]" italics="true" pageId="18" pageNumber="273">P. fritzei</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27A57B8FFB1BB66FA97FABC9899FAAF" box="[1382,1432,1383,1404]" pageId="18" pageNumber="273" rank="species">
|
||
<emphasis id="B949EAC3FFB1BB66FA97FABC9899FAAF" bold="true" box="[1382,1432,1383,1404]" pageId="18" pageNumber="273">sp. n</emphasis>
|
||
</taxonomicNameLabel>
|
||
<emphasis id="B949EAC3FFB1BB66FA66FABC989DFAAF" bold="true" box="[1431,1436,1383,1404]" pageId="18" pageNumber="273">.</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |