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<document ID-DOI="10.5252/geodiversitas2022v44a27" ID-ISSN="1638-9395" ID-ZooBank="urn:lsid:zoobank.org:pub:E4896081-9312-4EA6-AE33-AAC44201748E" checkinTime="1664917095605" checkinUser="admin" docAuthor="Sandoval, José" docDate="2022" docId="0394878DFF8B7B121A98F9F4FEB2DD4E" docLanguage="en" docName="Geodiversitas.44.27.801-851.pdf.imf" docOrigin="Geodiversitas 44 (27)" docStyle="DocumentStyle:F830B10FF475E64C1F1601E3B32DDC00.4:Geodiversitas.2018-.journal_article" docStyleId="F830B10FF475E64C1F1601E3B32DDC00" docStyleName="Geodiversitas.2018-.journal_article" docStyleVersion="4" docTitle="Dorsetensia Buckman 1892" docType="treatment" docVersion="5" lastPageNumber="841" masterDocId="FFADFFF5FFA27B381A52FFC7FFE0D81E" masterDocTitle="Sonniniidae Ammonitina, Middle Jurassic from Southern Spain: taxonomic, biostratigraphical and palaeobiogeographical analysis" masterLastPageNumber="851" masterPageNumber="801" pageNumber="840" updateTime="1665000364905" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Sonniniidae Ammonitina, Middle Jurassic from Southern Spain: taxonomic, biostratigraphical and palaeobiogeographical analysis</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sandoval, José</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Geodiversitas</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-09-15</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>44</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>27</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>801</mods:start>
<mods:end>851</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.5252/geodiversitas2022v44a27</mods:identifier>
<mods:identifier type="ISSN">1638-9395</mods:identifier>
<mods:identifier type="ZooBank">urn:lsid:zoobank.org:pub:E4896081-9312-4EA6-AE33-AAC44201748E</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.7145666" ID-Zenodo-Dep="7145666" LSID="urn:lsid:plazi:treatment:0394878DFF8B7B121A98F9F4FEB2DD4E" httpUri="http://treatment.plazi.org/id/0394878DFF8B7B121A98F9F4FEB2DD4E" lastPageId="42" lastPageNumber="841" pageId="41" pageNumber="840">
<subSubSection box="[202,705,1586,1613]" pageId="41" pageNumber="840" type="nomenclature">
<paragraph blockId="41.[202,705,1586,1613]" box="[202,705,1586,1613]" pageId="41" pageNumber="840">
<heading box="[202,705,1586,1613]" centered="true" fontSize="11" level="2" pageId="41" pageNumber="840" reason="2">
Genus
<taxonomicName authority="Buckman, 1892" authorityName="Buckman" authorityYear="1892" box="[279,578,1586,1613]" class="Cephalopoda" family="Sonniniidae" genus="Dorsetensia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ammonoidea" pageId="41" pageNumber="840" phylum="Mollusca" rank="genus">
<emphasis box="[279,397,1587,1612]" italics="true" pageId="41" pageNumber="840">Dorsetensia</emphasis>
Buckman, 1892
</taxonomicName>
[M] &amp; [m]
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="41" pageNumber="840" type="materials_examined">
<paragraph blockId="41.[132,776,1643,1694]" pageId="41" pageNumber="840">
<typeStatus box="[132,184,1644,1667]" pageId="41" pageNumber="840">
<collectionCode box="[132,184,1644,1667]" pageId="41" pageNumber="840">TYPE</collectionCode>
</typeStatus>
<collectionCode box="[192,266,1649,1666]" pageId="41" pageNumber="840">SPECIES</collectionCode>
. —
<taxonomicName box="[316,747,1643,1667]" pageId="41" pageNumber="840">
<emphasis box="[316,564,1643,1667]" italics="true" pageId="41" pageNumber="840">Ammonites edouardianus</emphasis>
d’Orbigny,
<quantity box="[695,776,1643,1667]" metricMagnitude="1" metricUnit="m" metricValue="4.68884" pageId="41" pageNumber="840" unit="in" value="1846.0">1846 in</quantity>
</taxonomicName>
d’Orbigny (1842-1851: 393, pl. 130, figs 3-5) (
<collectionCode box="[599,642,1670,1694]" pageId="41" pageNumber="840">OD</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection pageId="41" pageNumber="840" type="description">
<paragraph blockId="41.[132,776,1746,2028]" box="[132,277,1746,1772]" pageId="41" pageNumber="840">DESCRIPTION</paragraph>
<paragraph blockId="41.[132,776,1746,2028]" lastBlockId="41.[811,1455,757,1007]" pageId="41" pageNumber="840">
Macroconch [M] of medium-sized to large shells. The inner whorls are evolute with simple or, rarely, bifurcate (only in the earliest forms), radial or slightly flexuous ribs and no tubercles. The middle and outer whorls are more involute and compressed, with a sharp, sometimes undercut, umbilical edge, more or less flattened flanks, and a narrow venter with a full keel at all the stages. The ribbing usually weakens throughout ontogeny becoming smooth on the outer whorls. The mouth has plain borders.In some species, ribs are replaced by striation in the BC. The dimorphic microconch [m], traditionally classified with the name
<taxonomicName authorityName="Buckman" authorityYear="1927" box="[1178,1273,822,847]" genus="Nannina" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="41" pageNumber="840" phylum="Mollusca" rank="genus">
<emphasis box="[1178,1273,822,847]" italics="true" pageId="41" pageNumber="840">Nannina</emphasis>
</taxonomicName>
(gr. of
<taxonomicName authority="Haug, 1893" authorityName="Haug" authorityYear="1893" class="Cephalopoda" family="Sonniniidae" genus="Witchellia" kingdom="Animalia" order="Ammonoidea" pageId="41" pageNumber="840" phylum="Mollusca" rank="species" species="regrediens">
<emphasis italics="true" pageId="41" pageNumber="840">Witchellia regrediens</emphasis>
<bibRefCitation author="HAUG E." box="[918,1048,853,880]" pageId="41" pageNumber="840" pagination="277 - 333" refId="ref43527" refString="HAUG E. 1893. - Etude sur les ammonites des etages moyens du systeme Jurassique. Bulletin de la Societe geologique de France, ser. 3, 20: 277 - 333. https: // www. biodiversitylibrary. org / page / 31030302" type="journal article" year="1893">Haug, 1893</bibRefCitation>
</taxonomicName>
) are small and evolute, have ribs to the end of the BC and have lateral lappets on the mouth border. The septal suture, though relatively simple, is slightly more complex than in
<taxonomicName authorityName="Buckman" authorityYear="1889" box="[1002,1108,949,975]" class="Cephalopoda" family="Sonniniidae" genus="Witchellia" kingdom="Animalia" order="Ammonoidea" pageId="41" pageNumber="840" phylum="Mollusca" rank="genus">
<emphasis box="[1002,1108,949,975]" italics="true" pageId="41" pageNumber="840">Witchellia</emphasis>
</taxonomicName>
with wide and a rather shallow L, and the umbilical lobe is barely retracted.
</paragraph>
</subSubSection>
<subSubSection lastPageId="42" lastPageNumber="841" pageId="41" pageNumber="840" type="discussion">
<paragraph blockId="41.[811,1457,1045,2027]" box="[813,913,1045,1071]" pageId="41" pageNumber="840">REMARKS</paragraph>
<paragraph blockId="41.[811,1457,1045,2027]" pageId="41" pageNumber="840">
<taxonomicName authorityName="Buckman" authorityYear="1892" box="[813,934,1077,1102]" class="Cephalopoda" family="Sonniniidae" genus="Dorsetensia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ammonoidea" pageId="41" pageNumber="840" phylum="Mollusca" rank="genus">
<emphasis box="[813,934,1077,1102]" italics="true" pageId="41" pageNumber="840">Dorsetensia</emphasis>
</taxonomicName>
, included in the subfamily
<taxonomicName box="[1247,1380,1076,1102]" class="Cephalopoda" family="Sonniniidae" kingdom="Animalia" order="Ammonoidea" pageId="41" pageNumber="840" phylum="Mollusca" rank="subFamily" subFamily="Sonniniinae">Sonniniinae</taxonomicName>
in the
<emphasis box="[815,898,1109,1134]" italics="true" pageId="41" pageNumber="840">Treatise</emphasis>
(
<bibRefCitation author="HOWARTH M. K." box="[915,1084,1108,1134]" pageId="41" pageNumber="840" refId="ref43758" refString="HOWARTH M. K. 2013. - Treatease Online No. 57, Part L, Revised, Volume 3 B, Chapter 4: Psiloceratoidea, Eodoceratoidea, Hildoceratoidea. The University of Kansas, Lawrence, Kansas, 139 p. https: // doi. org / 10.17161 / to. v 0 i 0.4441" type="book" year="2013">Howarth 2013</bibRefCitation>
), is here formally transferred to
<taxonomicName box="[815,963,1140,1166]" class="Cephalopoda" family="Sonniniidae" kingdom="Animalia" order="Ammonoidea" pageId="41" pageNumber="822" phylum="Mollusca" rank="subFamily" subFamily="Witchelliinae">Witchelliinae</taxonomicName>
, because it shares many characters with
<taxonomicName authorityName="Buckman" authorityYear="1889" class="Cephalopoda" family="Sonniniidae" genus="Witchellia" kingdom="Animalia" order="Ammonoidea" pageId="41" pageNumber="840" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="41" pageNumber="840">Witchellia</emphasis>
</taxonomicName>
, such as: coiling; whorl section; ribbing; keel; and
<typeStatus box="[1409,1455,1172,1198]" pageId="41" pageNumber="840">type</typeStatus>
of dimorphism with medium-sized macroconchs that have a simple aperture, and much smaller microconchs that have a complex aperture and well-developed lateral lappets. The primary differences are that
<taxonomicName authorityName="Buckman" authorityYear="1892" box="[1124,1242,1300,1325]" class="Cephalopoda" family="Sonniniidae" genus="Dorsetensia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ammonoidea" pageId="41" pageNumber="840" phylum="Mollusca" rank="genus">
<emphasis box="[1124,1242,1300,1325]" italics="true" pageId="41" pageNumber="840">Dorsetensia</emphasis>
</taxonomicName>
never has tubercles and the venter is not tabulate, whereas
<taxonomicName authorityName="Buckman" authorityYear="1889" box="[1234,1340,1331,1357]" class="Cephalopoda" family="Sonniniidae" genus="Witchellia" kingdom="Animalia" order="Ammonoidea" pageId="41" pageNumber="840" phylum="Mollusca" rank="genus">
<emphasis box="[1234,1340,1331,1357]" italics="true" pageId="41" pageNumber="840">Witchellia</emphasis>
</taxonomicName>
has a relatively simpler septal suture. Some forms of
<taxonomicName authorityName="Buckman" authorityYear="1923" box="[1259,1365,1364,1390]" class="Cephalopoda" family="Sonniniidae" genus="Sonninites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ammonoidea" pageId="41" pageNumber="840" phylum="Mollusca" rank="genus">
<emphasis box="[1259,1365,1364,1390]" italics="true" pageId="41" pageNumber="840">Sonninites</emphasis>
</taxonomicName>
can also be quite similar to
<taxonomicName authorityName="Buckman" authorityYear="1892" box="[1019,1136,1396,1421]" class="Cephalopoda" family="Sonniniidae" genus="Dorsetensia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ammonoidea" pageId="41" pageNumber="840" phylum="Mollusca" rank="genus">
<emphasis box="[1019,1136,1396,1421]" italics="true" pageId="41" pageNumber="840">Dorsetensia</emphasis>
</taxonomicName>
, but
<taxonomicName authorityName="Buckman" authorityYear="1923" box="[1196,1303,1395,1421]" class="Cephalopoda" family="Sonniniidae" genus="Sonninites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ammonoidea" pageId="41" pageNumber="840" phylum="Mollusca" rank="genus">
<emphasis box="[1196,1303,1395,1421]" italics="true" pageId="41" pageNumber="840">Sonninites</emphasis>
</taxonomicName>
, usually more involute, has different ribbing with several furcate ribs in the inner whorls, may have tubercles in inner whorls, and has a more complex suture line.
</paragraph>
<paragraph blockId="41.[811,1457,1045,2027]" lastBlockId="42.[131,777,629,975]" lastPageId="42" lastPageNumber="841" pageId="41" pageNumber="840">
Some authors (e.g. Buckman 1927;
<bibRefCitation author="HUF W." box="[1231,1386,1523,1549]" pageId="41" pageNumber="840" pagination="1 - 126" refId="ref43817" refString="HUF W. 1968. - Uber Sonninien und Dorsetensien as dem Bajocium von Nordwestdeutschland. Beihefte zum Geologischen Jahrbuch 64: 1 - 126." type="journal article" year="1968">Huf 1968: 78</bibRefCitation>
,
<bibRefCitation author="PAVIA G." pageId="41" pageNumber="840" pagination="1 - 254" refId="ref45580" refString="PAVIA G. 1983. - Ammoniti e stratigrafia del Bajociano inferiore di Digne (Francia SE, dip. Alpes-Haute-Provence). Museo Regionale di Scienze Naturalli, Torino, Monografie II: 1 - 254. https: // archive. org / details / ammonitiebiostra 1983 pavi" type="journal article" year="1983">Pavia 1983: 62</bibRefCitation>
; Fernández-López 1985: 21; Fernández-López &amp; Mouterde 1994: 124;
<bibRefCitation author="SADKI D." box="[1065,1251,1586,1613]" pageId="41" pageNumber="840" pagination="1 - 245" refId="ref46296" refString="SADKI D. 1996. - Le Haut-Atlas Central (Maroc). Stratigraphie et paleontologie du Lias Superieur et du Dogger Inferieur. Documents des Laboratoires de Geologie de la Faculte des Sciences de Lyon 142: 1 - 245. https: // www. persee. fr / doc / geoly _ 0750 - 6635 _ 1996 _ mon _ 142 _ 1" type="journal article" year="1996">Sadki 1996: 170</bibRefCitation>
;
<bibRefCitation author="MYCZYNSKI R." pageId="41" pageNumber="840" pagination="7 - 31" refId="ref45059" refString="MYCZYNSKI R. 2004. - Toarcian, Aalenian and Early Bajocian (Jurassic) ammonite faunas and biostratigraphy in the Pieniny Klippen Belt and the Tatra Mts, West Carpathians. Studia Geologica Polonica 123: 7 - 31." type="journal article" year="2004">Myczynski 2004: 111</bibRefCitation>
) have considered
<taxonomicName authorityName="Buckman" authorityYear="1927" box="[1041,1135,1619,1644]" genus="Nannina" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="41" pageNumber="840" phylum="Mollusca" rank="genus">
<emphasis box="[1041,1135,1619,1644]" italics="true" pageId="41" pageNumber="840">Nannina</emphasis>
</taxonomicName>
to be the dimorphous partner [m] of
<taxonomicName authorityName="Buckman" authorityYear="1892" box="[882,998,1651,1676]" class="Cephalopoda" family="Sonniniidae" genus="Dorsetensia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ammonoidea" pageId="41" pageNumber="840" phylum="Mollusca" rank="genus">
<emphasis box="[882,998,1651,1676]" italics="true" pageId="41" pageNumber="840">Dorsetensia</emphasis>
</taxonomicName>
[M]. However,
<bibRefCitation author="HOWARTH M. K." box="[1164,1331,1650,1676]" pageId="41" pageNumber="840" refId="ref43758" refString="HOWARTH M. K. 2013. - Treatease Online No. 57, Part L, Revised, Volume 3 B, Chapter 4: Psiloceratoidea, Eodoceratoidea, Hildoceratoidea. The University of Kansas, Lawrence, Kansas, 139 p. https: // doi. org / 10.17161 / to. v 0 i 0.4441" type="book" year="2013">Howarth (2013</bibRefCitation>
, in
<emphasis box="[1369,1446,1651,1676]" italics="true" pageId="41" pageNumber="840">Treatise</emphasis>
: 119) deemed
<taxonomicName authority="Buckman, 1927" authorityName="Buckman" authorityYear="1927" box="[959,1234,1682,1708]" genus="Nannina" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="41" pageNumber="840" phylum="Mollusca" rank="genus">
<emphasis box="[959,1054,1683,1708]" italics="true" pageId="41" pageNumber="840">Nannina</emphasis>
Buckman, 1927
</taxonomicName>
as a junior synonym of
<taxonomicName authority="Buckman, 1902" authorityName="Buckman" authorityYear="1902" box="[842,1151,1714,1740]" class="Cephalopoda" family="Sonniniidae" genus="Fontannesia" kingdom="Animalia" order="Ammonoidea" pageId="41" pageNumber="840" phylum="Mollusca" rank="genus">
<emphasis box="[842,967,1715,1740]" italics="true" pageId="41" pageNumber="840">Fontannesia</emphasis>
Buckman, 1902
</taxonomicName>
. Clearly,
<taxonomicName authorityName="Buckman" authorityYear="1927" box="[1252,1347,1715,1740]" genus="Nannina" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="41" pageNumber="840" phylum="Mollusca" rank="genus">
<emphasis box="[1252,1347,1715,1740]" italics="true" pageId="41" pageNumber="840">Nannina</emphasis>
</taxonomicName>
is not the dimorphic microconch of
<taxonomicName authorityName="Buckman" authorityYear="1892" box="[1081,1195,1747,1772]" class="Cephalopoda" family="Sonniniidae" genus="Dorsetensia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ammonoidea" pageId="41" pageNumber="840" phylum="Mollusca" rank="genus">
<emphasis box="[1081,1195,1747,1772]" italics="true" pageId="41" pageNumber="840">Dorsetensia</emphasis>
</taxonomicName>
because the inner whorls of the
<typeStatus box="[883,929,1778,1804]" pageId="41" pageNumber="840">type</typeStatus>
species of the two genera are different. Furthermore,
<taxonomicName authorityName="Buckman" authorityYear="1927" box="[884,979,1810,1835]" genus="Nannina" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="41" pageNumber="840" phylum="Mollusca" rank="genus">
<emphasis box="[884,979,1810,1835]" italics="true" pageId="41" pageNumber="840">Nannina</emphasis>
</taxonomicName>
has a small coronate stage, a character that makes it closer to
<taxonomicName box="[1009,1139,1841,1867]" class="Cephalopoda" family="Sonniniidae" kingdom="Animalia" order="Ammonoidea" pageId="41" pageNumber="840" phylum="Mollusca" rank="subFamily" subFamily="Sonniniinae">Sonniniinae</taxonomicName>
and farther from
<taxonomicName authorityName="Buckman" authorityYear="1892" box="[1330,1448,1842,1867]" class="Cephalopoda" family="Sonniniidae" genus="Dorsetensia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ammonoidea" pageId="41" pageNumber="840" phylum="Mollusca" rank="genus">
<emphasis box="[1330,1448,1842,1867]" italics="true" pageId="41" pageNumber="840">Dorsetensia</emphasis>
</taxonomicName>
. Nor is
<taxonomicName authorityName="Buckman" authorityYear="1927" box="[889,984,1874,1899]" genus="Nannina" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="41" pageNumber="840" phylum="Mollusca" rank="genus">
<emphasis box="[889,984,1874,1899]" italics="true" pageId="41" pageNumber="840">Nannina</emphasis>
</taxonomicName>
synonymous with
<taxonomicName authorityName="Buckman" authorityYear="1902" box="[1190,1315,1874,1899]" class="Cephalopoda" family="Sonniniidae" genus="Fontannesia" kingdom="Animalia" order="Ammonoidea" pageId="41" pageNumber="840" phylum="Mollusca" rank="genus">
<emphasis box="[1190,1315,1874,1899]" italics="true" pageId="41" pageNumber="840">Fontannesia</emphasis>
</taxonomicName>
, as
<bibRefCitation author="HOWARTH M. K." pageId="41" pageNumber="840" refId="ref43758" refString="HOWARTH M. K. 2013. - Treatease Online No. 57, Part L, Revised, Volume 3 B, Chapter 4: Psiloceratoidea, Eodoceratoidea, Hildoceratoidea. The University of Kansas, Lawrence, Kansas, 139 p. https: // doi. org / 10.17161 / to. v 0 i 0.4441" type="book" year="2013">Howarth (2013)</bibRefCitation>
postulated, because the
<typeStatus box="[1138,1184,1906,1932]" pageId="41" pageNumber="840">type</typeStatus>
species of the two taxa are quite distinct:
<taxonomicName authorityName="Buckman" authorityYear="1902" box="[964,1088,1938,1963]" class="Cephalopoda" family="Sonniniidae" genus="Fontannesia" kingdom="Animalia" order="Ammonoidea" pageId="41" pageNumber="840" phylum="Mollusca" rank="genus">
<emphasis box="[964,1088,1938,1963]" italics="true" pageId="41" pageNumber="840">Fontannesia</emphasis>
</taxonomicName>
has inner whorls with well-defined simple ribs that are in no case tuberculate, differentiating this species from
<taxonomicName authorityName="Buckman" authorityYear="1927" box="[963,1060,2002,2027]" genus="Nannina" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="41" pageNumber="840" phylum="Mollusca" rank="genus">
<emphasis box="[963,1060,2002,2027]" italics="true" pageId="41" pageNumber="840">Nannina</emphasis>
</taxonomicName>
(
<typeStatus box="[1075,1124,2001,2027]" pageId="41" pageNumber="840">type</typeStatus>
species
<taxonomicName authority="Buckman" authorityName="Buckman" authorityYear="1927" box="[1220,1450,2001,2027]" class="Cephalopoda" family="Sonniniidae" genus="Witchellia" kingdom="Animalia" order="Ammonoidea" pageId="41" pageNumber="840" phylum="Mollusca" rank="species" species="evoluta">
<emphasis box="[1220,1334,2001,2027]" italics="true" pageId="41" pageNumber="840">N. evoluta</emphasis>
Buckman
</taxonomicName>
, 1927 [Buckman 1927:
<collectionCode box="[389,407,630,656]" name="Tavera, Department of Geology and Geophysics" pageId="42" pageNumber="841">T</collectionCode>
.A. 7, pl. 752]). Furthermore, the stratigraphic ranges also substantially differ, as
<taxonomicName authorityName="Buckman" authorityYear="1902" box="[651,777,662,687]" class="Cephalopoda" family="Sonniniidae" genus="Fontannesia" kingdom="Animalia" order="Ammonoidea" pageId="42" pageNumber="841" phylum="Mollusca" rank="genus">
<emphasis box="[651,777,662,687]" italics="true" pageId="42" pageNumber="841">Fontannesia</emphasis>
</taxonomicName>
is restricted to the interval between the Concavum (Upper Aalenian) and Ovale zones, whereas the first stratigraphic records for
<taxonomicName authorityName="Buckman" authorityYear="1927" box="[260,355,758,783]" genus="Nannina" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="42" pageNumber="841" phylum="Mollusca" rank="genus">
<emphasis box="[260,355,758,783]" italics="true" pageId="42" pageNumber="841">Nannina</emphasis>
</taxonomicName>
do not appear until the upper part of the Propinquans Zone (seePavia 1983: 62;Fernández-López 1985: 115). The above mentioned shows that the taxonomic status of the HT of
<taxonomicName box="[338,512,853,879]" genus="Nannina" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="42" pageNumber="841" phylum="Mollusca" rank="species" species="evoluta">
<emphasis box="[338,512,853,879]" italics="true" pageId="42" pageNumber="841">Nannina evoluta</emphasis>
</taxonomicName>
is not clear, but given its coiling, the coronate stage on the inner whorls and its stratigraphic distribution (Propinquans Zone), it could represent a dimorphic microconch of
<emphasis box="[435,455,949,975]" italics="true" pageId="42" pageNumber="841">S.</emphasis>
(
<taxonomicName box="[471,566,949,975]" class="Cephalopoda" family="Sonniniidae" genus="Sonninia" kingdom="Animalia" order="Ammonoidea" pageId="42" pageNumber="841" phylum="Mollusca" rank="genus">
<emphasis box="[471,566,949,975]" italics="true" pageId="42" pageNumber="841">Sonninia</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<caption ID-Table-UUID="DF426613FF887B121BB9FF1CFBA8D8F2" box="[491,1096,219,236]" httpUri="http://table.plazi.org/id/DF426613FF887B121BB9FF1CFBA8D8F2" pageId="42" pageNumber="841" startId="42.[491,539,219,236]" targetBox="[134,1436,295,549]" targetIsTable="true" targetPageId="42">
<paragraph blockId="42.[491,1096,219,236]" box="[491,1096,219,236]" pageId="42" pageNumber="841">
TABLE 25. — Measurements of
<taxonomicName authority="Buckman, 1892" authorityName="Buckman" authorityYear="1892" box="[749,1059,219,236]" class="Cephalopoda" family="Sonniniidae" genus="Dorsetensia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ammonoidea" pageId="42" pageNumber="841" phylum="Mollusca" rank="species" species="liostraca">
<emphasis box="[749,922,219,236]" italics="true" pageId="42" pageNumber="841">Dorsetensia liostraca</emphasis>
Buckman, 1892
</taxonomicName>
[M].
</paragraph>
</caption>
<paragraph pageId="42" pageNumber="841">
<table box="[134,1436,295,549]" gridcols="10" gridrows="6" pageId="42" pageNumber="841">
<tr box="[134,1436,295,315]" gridrow="0" pageId="42" pageNumber="841">
<th box="[134,243,295,315]" gridcol="0" gridrow="0" pageId="42" pageNumber="841">Specimen</th>
<th box="[333,387,295,315]" gridcol="1" gridrow="0" pageId="42" pageNumber="841">D</th>
<th box="[442,484,295,315]" gridcol="2" gridrow="0" pageId="42" pageNumber="841">U</th>
<th box="[545,598,295,315]" gridcol="3" gridrow="0" pageId="42" pageNumber="841">u</th>
<th box="[635,677,295,315]" gridcol="4" gridrow="0" pageId="42" pageNumber="841">H</th>
<th box="[737,779,295,315]" gridcol="5" gridrow="0" pageId="42" pageNumber="841">h</th>
<th box="[833,875,295,315]" gridcol="6" gridrow="0" pageId="42" pageNumber="841">W</th>
<th box="[930,972,295,315]" gridcol="7" gridrow="0" pageId="42" pageNumber="841">w</th>
<th box="[1050,1141,295,315]" gridcol="8" gridrow="0" pageId="42" pageNumber="841">Remarks</th>
<th box="[1212,1436,295,315]" gridcol="9" gridrow="0" pageId="42" pageNumber="841">
<emphasis bold="true" box="[1245,1401,295,315]" pageId="42" pageNumber="841">Biostratigraphy</emphasis>
</th>
</tr>
<tr box="[134,1436,330,376]" gridrow="1" pageId="42" pageNumber="841" rowspan-9="1">
<th box="[134,243,330,376]" gridcol="0" gridrow="1" pageId="42" pageNumber="841">JGa7.32.2</th>
<td box="[333,387,330,376]" gridcol="1" gridrow="1" pageId="42" pageNumber="841">74.0 51.0</td>
<td box="[442,484,330,376]" gridcol="2" gridrow="1" pageId="42" pageNumber="841">15.0 11.5</td>
<td box="[545,598,330,376]" gridcol="3" gridrow="1" pageId="42" pageNumber="841">0.20 0.22</td>
<td box="[635,677,330,376]" gridcol="4" gridrow="1" pageId="42" pageNumber="841">38.0 24.4</td>
<td box="[737,779,330,376]" gridcol="5" gridrow="1" pageId="42" pageNumber="841">0.51 0.44</td>
<td box="[833,875,330,376]" gridcol="6" gridrow="1" pageId="42" pageNumber="841">19.0 14.0</td>
<td box="[930,972,330,376]" gridcol="7" gridrow="1" pageId="42" pageNumber="841">0.26 0.27</td>
<td box="[1050,1141,330,376]" gridcol="8" gridrow="1" pageId="42" pageNumber="841">
<collectingCountry box="[1081,1110,343,363]" name="Philippines" pageId="42" pageNumber="841">PH</collectingCountry>
</td>
</tr>
<tr box="[134,1436,380,475]" gridrow="2" pageId="42" pageNumber="841">
<th box="[134,243,380,475]" gridcol="0" gridrow="2" pageId="42" pageNumber="841">JGa7.32.3 JGa7.32.4</th>
<td box="[333,387,380,475]" gridcol="1" gridrow="2" pageId="42" pageNumber="841">128.0 85.0 95.0 66.0</td>
<td box="[442,484,380,475]" gridcol="2" gridrow="2" pageId="42" pageNumber="841">16.0 12.0 13.0 10.0</td>
<td box="[545,598,380,475]" gridcol="3" gridrow="2" pageId="42" pageNumber="841">0.125 0.14 0.14 0.15</td>
<td box="[635,677,380,475]" gridcol="4" gridrow="2" pageId="42" pageNumber="841">69.0 45.0 49.0 33.5</td>
<td box="[737,779,380,475]" gridcol="5" gridrow="2" pageId="42" pageNumber="841">0.54 0.53 0.52 0.51</td>
<td box="[833,875,380,475]" gridcol="6" gridrow="2" pageId="42" pageNumber="841">30.0 23.0 24.0 17.5</td>
<td box="[930,972,380,475]" gridcol="7" gridrow="2" pageId="42" pageNumber="841">0.23 0.27 0.25 0.25</td>
<td box="[1050,1141,380,475]" gridcol="8" gridrow="2" pageId="42" pageNumber="841">
<collectingCountry box="[1054,1083,393,413]" name="Philippines" pageId="42" pageNumber="841">PH</collectingCountry>
&amp; BC
</td>
<td box="[1212,1436,380,475]" gridcol="9" gridrow="2" pageId="42" pageNumber="841">Humphriesianum Zone, Romani Subzone</td>
</tr>
<tr box="[134,1436,479,499]" gridrow="3" pageId="42" pageNumber="841" rowspan-9="1">
<th box="[134,243,479,499]" gridcol="0" gridrow="3" pageId="42" pageNumber="841">JGa7.32.5</th>
<td box="[333,387,479,499]" gridcol="1" gridrow="3" pageId="42" pageNumber="841">70.0</td>
<td box="[442,484,479,499]" gridcol="2" gridrow="3" pageId="42" pageNumber="841">14.0</td>
<td box="[545,598,479,499]" gridcol="3" gridrow="3" pageId="42" pageNumber="841">0.20</td>
<td box="[635,677,479,499]" gridcol="4" gridrow="3" pageId="42" pageNumber="841">35.0</td>
<td box="[737,779,479,499]" gridcol="5" gridrow="3" pageId="42" pageNumber="841">0.50</td>
<td box="[833,875,479,499]" gridcol="6" gridrow="3" pageId="42" pageNumber="841">24.0</td>
<td box="[930,972,479,499]" gridcol="7" gridrow="3" pageId="42" pageNumber="841">0.34</td>
<td box="[1050,1141,479,499]" gridcol="8" gridrow="3" pageId="42" pageNumber="841">PHs</td>
</tr>
<tr box="[134,1436,505,525]" gridrow="4" pageId="42" pageNumber="841" rowspan-0="1" rowspan-8="1" rowspan-9="1">
<td box="[333,387,505,525]" gridcol="1" gridrow="4" pageId="42" pageNumber="841">52.0</td>
<td box="[442,484,505,525]" gridcol="2" gridrow="4" pageId="42" pageNumber="841">12.0</td>
<td box="[545,598,505,525]" gridcol="3" gridrow="4" pageId="42" pageNumber="841">0.23</td>
<td box="[635,677,505,525]" gridcol="4" gridrow="4" pageId="42" pageNumber="841">25.5</td>
<td box="[737,779,505,525]" gridcol="5" gridrow="4" pageId="42" pageNumber="841">0.49</td>
<td box="[833,875,505,525]" gridcol="6" gridrow="4" pageId="42" pageNumber="841">18.3</td>
<td box="[930,972,505,525]" gridcol="7" gridrow="4" pageId="42" pageNumber="841">0.35</td>
</tr>
<tr box="[134,1436,529,549]" gridrow="5" pageId="42" pageNumber="841" rowspan-8="1" rowspan-9="1">
<th box="[134,243,529,549]" gridcol="0" gridrow="5" pageId="42" pageNumber="841">JGa7.32.10</th>
<td box="[333,387,529,549]" gridcol="1" gridrow="5" pageId="42" pageNumber="841">43.0</td>
<td box="[442,484,529,549]" gridcol="2" gridrow="5" pageId="42" pageNumber="841">9.0</td>
<td box="[545,598,529,549]" gridcol="3" gridrow="5" pageId="42" pageNumber="841">0.21</td>
<td box="[635,677,529,549]" gridcol="4" gridrow="5" pageId="42" pageNumber="841">20.0</td>
<td box="[737,779,529,549]" gridcol="5" gridrow="5" pageId="42" pageNumber="841">0.48</td>
<td box="[833,875,529,549]" gridcol="6" gridrow="5" pageId="42" pageNumber="841">12.0</td>
<td box="[930,972,529,549]" gridcol="7" gridrow="5" pageId="42" pageNumber="841">0.28</td>
</tr>
</table>
</paragraph>
<subSubSection pageId="42" pageNumber="841" type="distribution">
<paragraph blockId="42.[130,776,1014,1360]" box="[132,288,1014,1040]" pageId="42" pageNumber="841">DISTRIBUTION</paragraph>
<paragraph blockId="42.[130,776,1014,1360]" pageId="42" pageNumber="841">
Lower Bajocian (uppermost Propinquans Zone-Humphriesianum Zone): Europe, North Africa (
<taxonomicName authority="Ranges" authorityName="Ranges" box="[539,682,1078,1105]" genus="Atlas" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="42" pageNumber="841" phylum="Mollusca" rank="genus">Atlas Ranges</taxonomicName>
),
<collectingCountry box="[700,771,1079,1105]" name="Kenya" pageId="42" pageNumber="841">Kenya</collectingCountry>
,
<collectingCountry box="[132,260,1110,1136]" name="Madagascar" pageId="42" pageNumber="841">Madagascar</collectingCountry>
,
<collectingCountry box="[268,406,1110,1136]" name="Saudi Arabia" pageId="42" pageNumber="841">Saudi Arabia</collectingCountry>
,
<collectingCountry box="[416,484,1110,1136]" name="Russia" pageId="42" pageNumber="841">Russia</collectingCountry>
(south-eastern Pamirs), the Caucasus,
<collectingCountry box="[245,312,1142,1168]" name="China" pageId="42" pageNumber="841">China</collectingCountry>
(
<collectingRegion box="[328,392,1142,1169]" country="China" name="Xizang" pageId="42" pageNumber="841">Tibet</collectingRegion>
),
<collectingCountry box="[411,556,1142,1168]" name="United States of America" pageId="42" pageNumber="841">United States</collectingCountry>
(
<collectingRegion box="[571,646,1142,1168]" country="United States of America" name="Alaska" pageId="42" pageNumber="841">Alaska</collectingRegion>
,
<collectingRegion box="[657,769,1142,1168]" country="United States of America" name="California" pageId="42" pageNumber="841">California</collectingRegion>
,
<collectingRegion box="[133,223,1174,1200]" country="United States of America" name="Oregon" pageId="42" pageNumber="841">Oregon</collectingRegion>
),
<collectingCountry box="[244,308,1174,1200]" name="Chile" pageId="42" pageNumber="841">Chile</collectingCountry>
, and
<collectingCountry box="[372,486,1174,1200]" name="Argentina" pageId="42" pageNumber="841">Argentina</collectingCountry>
. In the Subbetic domain,
<taxonomicName authorityName="Buckman" authorityYear="1892" box="[134,252,1207,1232]" class="Cephalopoda" family="Sonniniidae" genus="Dorsetensia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ammonoidea" pageId="42" pageNumber="841" phylum="Mollusca" rank="genus">
<emphasis box="[134,252,1207,1232]" italics="true" pageId="42" pageNumber="841">Dorsetensia</emphasis>
</taxonomicName>
shares this stratigraphic distribution, appearing in the three Subbetic subdomains (External, Median, and Internal), but the genus is scarce except in the Pinguis horizon of the central sector of the Median Subbetic (Sierra de Alta Coloma area).
</paragraph>
</subSubSection>
</treatment>
</document>
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