375 lines
42 KiB
XML
375 lines
42 KiB
XML
<document id="A4D11F358C6A7B2AD65C4C30E2E05F28" ID-CLB-Dataset="53799" ID-DOI="10.1080/00222933.2023.2171820" ID-GBIF-Dataset="e3544c70-f9c9-4daa-a16e-a827022616bc" ID-ISSN="1464-5262" ID-Zenodo-Dep="7737268" ID-ZooBank="AA69FCBE-81ED-4B41-90D6-0D32EBE887CA" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1678694520066" checkinUser="plazi" docAuthor="Lee, Young June, Marshall, David C., Mohagan, Alma B., Hill, Kathy B. R. & Mohagan, Dave P." docDate="2023" docId="038E87BB9B4AFFC04B36FCA6FD0266A6" docLanguage="en" docName="JNATHIST.57.1-4.193-242.pdf" docOrigin="Journal of Natural History 57 (1 - 4)" docSource="http://dx.doi.org/10.1080/00222933.2023.2171820" docStyle="DocumentStyle:3C762F9783B768A90048B82ACD135A62.3:JNatHist.2017-.journal_article.0cover" docStyleId="3C762F9783B768A90048B82ACD135A62" docStyleName="JNatHist.2017-.journal_article.0cover" docStyleVersion="3" docTitle="Oncotympana obesa Lee and Marshall 2023, sp. nov." docType="treatment" docVersion="7" lastPageNumber="216" masterDocId="FFB7FFC39B59FFD74A52FFEEFFDF6525" masterDocTitle="Revised checklist of Cicadidae (Insecta: Hemiptera) of Mindanao, Philippines, with descriptions of a new genus and nine new species" masterLastPageNumber="242" masterPageNumber="193" pageNumber="212" updateTime="1699476474976" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="027F184B06637FBD70987A907061445A">
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<mods:title id="466CE71263AC04AA9FE32C6394B13824">Revised checklist of Cicadidae (Insecta: Hemiptera) of Mindanao, Philippines, with descriptions of a new genus and nine new species</mods:title>
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<mods:namePart id="53A395302C5E118D2CBCF28599763F82">Lee, Young June</mods:namePart>
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<mods:namePart id="018C01D462225B6D66B225F2B66F4A92">Marshall, David C.</mods:namePart>
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<mods:namePart id="CF0D868C15D64662ACCE3468872B5606">Mohagan, Alma B.</mods:namePart>
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<mods:namePart id="8553B136338E3752E89F6E4120D3E18E">Hill, Kathy B. R.</mods:namePart>
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<mods:namePart id="369B089694273B1B77183AAD6ADC9FA0">Mohagan, Dave P.</mods:namePart>
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<mods:title id="C5EC25F33FD78ABF2A74F0EAB8BBD247">Journal of Natural History</mods:title>
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<mods:part id="81169B9FB5141C488C06C71507C0577C">
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<mods:date id="1D83FD60061C36D67726735DD304BB62">2023</mods:date>
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<mods:number id="77DFA3B061C46C8AE51AD6812496D4E5">2023-03-06</mods:number>
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<mods:number id="0C5E60E56C8FF2404D7BC27D2AF45304">1 - 4</mods:number>
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<treatment id="038E87BB9B4AFFC04B36FCA6FD0266A6" ID-DOI="http://doi.org/10.5281/zenodo.7738570" ID-GBIF-Taxon="209450831" ID-Zenodo-Dep="7738570" LSID="urn:lsid:plazi:treatment:038E87BB9B4AFFC04B36FCA6FD0266A6" httpUri="http://treatment.plazi.org/id/038E87BB9B4AFFC04B36FCA6FD0266A6" lastPageId="23" lastPageNumber="216" pageId="19" pageNumber="212">
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<subSubSection id="C33D65269B4AFFC44B36FCA6FC606647" box="[356,959,840,866]" pageId="19" pageNumber="212" type="nomenclature">
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<paragraph id="8B9836AD9B4AFFC44B36FCA6FC606647" blockId="19.[356,959,840,866]" box="[356,959,840,866]" pageId="19" pageNumber="212">
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<heading id="D0D081C19B4AFFC44B36FCA6FC606647" box="[356,959,840,866]" centered="true" fontSize="11" level="2" pageId="19" pageNumber="212" reason="5">
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16.
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<taxonomicName id="4C274D2E9B4AFFC44BDDFCA6FC836647" ID-CoL="1467649b-119d-4671-a4a9-46c6a9e7ee27" authority="Lee and Marshall" authorityName="Lee and Marshall" authorityYear="2023" box="[399,860,840,866]" class="Insecta" family="Cicadidae" genus="Oncotympana" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="212" phylum="Arthropoda" rank="species" species="obesa" status="sp. nov.">
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<emphasis id="B953EABF9B4AFFC44BDDFCA6FD536647" bold="true" box="[399,652,840,866]" italics="true" pageId="19" pageNumber="212">Oncotympana obesa</emphasis>
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Lee and Marshall
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</taxonomicName>
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<taxonomicNameLabel id="A26057C49B4AFFC44936FCA6FC606647" box="[868,959,840,866]" pageId="19" pageNumber="212" rank="species">sp. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C33D65269B4AFFC4486DFC61FD3A668C" box="[575,741,911,937]" pageId="19" pageNumber="212" type="description">
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<paragraph id="8B9836AD9B4AFFC4486DFC61FD3A668C" blockId="19.[575,741,911,937]" box="[575,741,911,937]" pageId="19" pageNumber="212">
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(
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<figureCitation id="131C2A289B4AFFC44815FC61FD6B668C" box="[583,692,911,937]" captionStart="Figure 9" captionStartId="20.[160,225,1348,1371]" captionTargetBox="[162,1132,175,1297]" captionTargetId="figure-74@20.[162,1154,142,1317]" captionTargetPageId="20" captionText="Figure 9. Oncotympana obesa Lee and Marshall sp. nov., male, Mindanao, Philippines. (A) Dorsal habitus of holotype; (B) ventral habitus of holotype; (C) pygofer, slightly oblique ventral view." figureDoi="http://doi.org/10.5281/zenodo.7737288" httpUri="https://zenodo.org/record/7737288/files/figure.png" pageId="19" pageNumber="212">Figures 9</figureCitation>
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,
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<figureCitation id="131C2A289B4AFFC44892FC61FD02668C" box="[704,733,911,937]" captionStart="Figure 10" captionStartId="21.[160,225,894,917]" captionTargetBox="[164,1151,144,859]" captionTargetId="figure-233@21.[162,1154,142,862]" captionTargetPageId="21" captionText="Figure 10. Oncotympana obesa Lee and Marshall sp. nov., holotype male calling song. Waveforms (A, C) and spectrograms (B, D) are shown at two time scales." figureDoi="http://doi.org/10.5281/zenodo.7737290" httpUri="https://zenodo.org/record/7737290/files/figure.png" pageId="19" pageNumber="212">10</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C33D65269B4AFFC44AF2FC2AFC8F66F8" box="[160,848,964,989]" pageId="19" pageNumber="212" type="reference_group">
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<paragraph id="8B9836AD9B4AFFC44AF2FC2AFC8F66F8" blockId="19.[160,848,964,989]" box="[160,848,964,989]" pageId="19" pageNumber="212">
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<taxonomicName id="4C274D2E9B4AFFC44AF2FC2AFD8A66F8" ID-CoL="49JQV" authority=": Hill et al. 2021" authorityName="Hill" authorityYear="2021" box="[160,597,964,989]" class="Insecta" family="Cicadidae" genus="Oncotympana" isUncertain="true" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="212" phylum="Arthropoda" rank="species" species="averta">
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<emphasis id="B953EABF9B4AFFC44AF2FC2AFEE466F8" box="[160,315,964,989]" italics="true" pageId="19" pageNumber="212">Oncotympana</emphasis>
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cf.
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<emphasis id="B953EABF9B4AFFC44B31FC2AFE7766F8" box="[355,424,964,989]" italics="true" pageId="19" pageNumber="212">averta</emphasis>
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:
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<bibRefCitation id="EFB64B5C9B4AFFC44BE7FC2AFD8A66F8" author="Hill KBR & Marshall DC & Marathe K & Moulds M & Lee YJ & Pham TH & Mohagan AB & Sarkar V & Price BW & Duffels JP" box="[437,597,964,989]" pageId="19" pageNumber="212" pagination="570 - 601" refId="ref19849" refString="Hill KBR, Marshall DC, Marathe K, Moulds M, Lee YJ, Pham TH, Mohagan AB, Sarkar V, Price BW, Duffels JP, et al. 2021. The molecular systematics and diversification of a taxonomically unstable group of Asian cicada tribes related to Cicadini Latreille, 1802 (Hemiptera: Cicadidae). Invertebr Syst. 35 (5): 570 - 601. doi: 10.1071 / IS 20079." type="journal article" year="2021">Hill et al. 2021</bibRefCitation>
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</taxonomicName>
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: table 1, figs 4, 12, 14.
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</paragraph>
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</subSubSection>
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<subSubSection id="C33D65269B4AFFC44AF2FBFAFD706009" pageId="19" pageNumber="212" type="materials_examined">
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<emphasis id="B953EABF9B4AFFC44AF2FBFAFE9B610B" bold="true" box="[160,324,1044,1070]" italics="true" pageId="19" pageNumber="212">Type material</emphasis>
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</paragraph>
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<paragraph id="8B9836AD9B4AFFC44AF2FBD9FE1C61B3" blockId="19.[160,1155,1044,1174]" pageId="19" pageNumber="212">
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<emphasis id="B953EABF9B4AFFC44AF2FBD9FECD6174" bold="true" box="[160,274,1079,1105]" italics="true" pageId="19" pageNumber="212">
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<typeStatus id="549C880F9B4AFFC44AF2FBD9FED26174" box="[160,269,1079,1105]" pageId="19" pageNumber="212" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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<location id="8EF860769B4AFFC44B73FBD6FE856174" LSID="urn:lsid:plazi:treatment:038E87BB9B4AFFC04B36FCA6FD0266A6:8EF860769B4AFFC44B73FBD6FE856174" box="[289,346,1080,1105]" country="Philippines" county="Mt. Kitanglad" latitude="8.1635" longLatPrecision="13" longitude="124.9325" municipality="Mindanao" name="Male" pageId="19" pageNumber="212">Male</location>
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, specimen code 12.PH.
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<collectionCode id="ED36AE689B4AFFC4480EFBD6FD5B6174" box="[604,644,1080,1105]" country="Brazil" name="Museu Nacional, Universidade Federal do Rio de Janeiro" pageId="19" pageNumber="212">MN</collectionCode>
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.KGC.01,
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,
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<collectingMunicipality id="6BFCACD79B4AFFC4492FFBD6FC2F6174" box="[893,1008,1080,1105]" pageId="19" pageNumber="212">Mindanao</collectingMunicipality>
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, ca. 1/2 h up trail to
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<collectingCounty id="62F94E219B4AFFC44ABCFBB4FE5B6156" box="[238,388,1114,1139]" pageId="19" pageNumber="212">Mt. Kitanglad</collectingCounty>
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,
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<quantity id="4CDF9B489B4AFFC44BDFFBB4FE3E6156" box="[397,481,1114,1139]" metricMagnitude="3" metricUnit="m" metricValue="1.7759999999999998" pageId="19" pageNumber="212" unit="m" value="1776.0">
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</quantity>
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,
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08°09.81
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<emphasis id="B953EABF9B4AFFC44818FBB4FD8F6157" bold="true" box="[586,592,1114,1138]" pageId="19" pageNumber="212">ʹ</emphasis>
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N
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</geoCoordinate>
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,
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<geoCoordinate id="EE13506A9B4AFFC4483FFBB4FD2E6156" box="[621,753,1114,1139]" degrees="124" direction="east" minutes="55.95" orientation="longitude" pageId="19" pageNumber="212" precision="9" value="124.9325">
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124°55.95
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<emphasis id="B953EABF9B4AFFC44889FBB4FD3E6157" bold="true" box="[731,737,1114,1138]" pageId="19" pageNumber="212">ʹ</emphasis>
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E
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</geoCoordinate>
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,
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</date>
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,
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<collectorName id="26D2537B9B4AFFC449D1FBB4FC746156" box="[899,939,1114,1139]" name="Hill KBR" pageId="19" pageNumber="212">K.B.</collectorName>
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<collectionCode id="ED36AE689B4AFFC449F8FBB4FC646156" box="[938,955,1114,1139]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="19" pageNumber="212">R</collectionCode>
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.
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<collectorName id="26D2537B9B4AFFC44996FBB4FBC26156" box="[964,1053,1114,1139]" pageId="19" pageNumber="212">Hill, D.C</collectorName>
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. Marshall and
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<collectorName id="26D2537B9B4AFFC44A80FB93FE8461B3" box="[210,347,1149,1174]" pageId="19" pageNumber="212">D. Mohagan</collectorName>
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(
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<collectionCode id="ED36AE689B4AFFC44B3BFB93FE6661B3" box="[361,441,1149,1174]" pageId="19" pageNumber="212">NMPM</collectionCode>
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).
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</materialsCitation>
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</paragraph>
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<paragraph id="8B9836AD9B4AFFC44AF2FB23FD706009" blockId="19.[160,1156,1229,1324]" pageId="19" pageNumber="212">
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<materialsCitation id="3B4F3CF09B4AFFC44AF2FB23FC8A61C2" ID-GBIF-Occurrence="4058636342" box="[160,853,1229,1255]" country="Philippines" location="Mt" municipality="Mindanao" pageId="19" pageNumber="212" specimenCount="2" specimenCount-male="2" stateProvince="Bukidnon" typeStatus="paratype">
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<emphasis id="B953EABF9B4AFFC44AF2FB23FEC261C2" bold="true" box="[160,285,1229,1255]" italics="true" pageId="19" pageNumber="212">
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<typeStatus id="549C880F9B4AFFC44AF2FB23FEC761C2" box="[160,280,1229,1255]" pageId="19" pageNumber="212" type="paratype">Paratypes</typeStatus>
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.
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</emphasis>
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<specimenCount id="9D21FD249B4AFFC44B7FFB20FE5361C2" box="[301,396,1230,1255]" pageId="19" pageNumber="212" type="male">2 males</specimenCount>
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,
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<collectingCountry id="F330763D9B4AFFC44BC8FB20FDF961C2" box="[410,550,1230,1255]" name="Philippines" pageId="19" pageNumber="212">PHILIPPINES</collectingCountry>
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,
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<collectingMunicipality id="6BFCACD79B4AFFC44866FB20FD7761C2" box="[564,680,1230,1255]" pageId="19" pageNumber="212">Mindanao</collectingMunicipality>
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,
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<collectingRegion id="49E3F84F9B4AFFC448E5FB20FCF961C2" box="[695,806,1230,1255]" country="Philippines" name="Bukidnon" pageId="19" pageNumber="212">Bukidnon</collectingRegion>
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,
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<location id="8EF860769B4AFFC44966FB20FC8A61C2" LSID="urn:lsid:plazi:treatment:038E87BB9B4AFFC04B36FCA6FD0266A6:8EF860769B4AFFC44966FB20FC8A61C2" box="[820,853,1230,1255]" country="Philippines" municipality="Mindanao" name="Mt" pageId="19" pageNumber="212" stateProvince="Bukidnon">Mt</location>
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</materialsCitation>
|
||
. Kiamo,
|
||
<date id="FF99106D9B4AFFC449EEFB20FB9461C2" box="[956,1099,1230,1255]" pageId="19" pageNumber="212" value="2012-05-07">7 May 2012</date>
|
||
, A.B.
|
||
<materialsCitation id="3B4F3CF09B4AFFC44AF2FB1EFCC0602C" ID-GBIF-Occurrence="4058636307" box="[160,799,1264,1289]" collectionCode="UCONN" collectorName="Mohagan" country="Philippines" location="Mt" municipality="Mindanao" pageId="19" pageNumber="212" specimenCount="2" specimenCount-male="2" typeStatus="paratype">
|
||
<collectorName id="26D2537B9B4AFFC44AF2FB1EFED5602C" box="[160,266,1264,1289]" pageId="19" pageNumber="212">Mohagan</collectorName>
|
||
(
|
||
<collectionCode id="ED36AE689B4AFFC44B4AFB1EFEAA602C" box="[280,373,1264,1289]" country="USA" httpUri="http://grbio.org/cool/jrf7-y871" name="University of Connecticut Biodiversity Research Collections" pageId="19" pageNumber="212">UCONN</collectionCode>
|
||
);
|
||
<specimenCount id="9D21FD249B4AFFC44BD4FB1EFE3F602C" box="[390,480,1264,1289]" pageId="19" pageNumber="212" type="male">2 males</specimenCount>
|
||
,
|
||
<collectingCountry id="F330763D9B4AFFC44BB9FB1EFDA9602C" box="[491,630,1264,1289]" name="Philippines" pageId="19" pageNumber="212">PHILIPPINES</collectingCountry>
|
||
,
|
||
<collectingMunicipality id="6BFCACD79B4AFFC448D2FB1EFD2C602C" box="[640,755,1264,1289]" pageId="19" pageNumber="212">Mindanao</collectingMunicipality>
|
||
,
|
||
<location id="8EF860769B4AFFC448ADFB1EFCC0602C" LSID="urn:lsid:plazi:treatment:038E87BB9B4AFFC04B36FCA6FD0266A6:8EF860769B4AFFC448ADFB1EFCC0602C" box="[767,799,1264,1289]" country="Philippines" municipality="Mindanao" name="Mt" pageId="19" pageNumber="212">Mt</location>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B4F3CF09B4AFFC44979FB1EFE906009" ID-GBIF-Occurrence="4058636337" country="Philippines" elevation="1773" latitude="8.165167" location="PH" longLatPrecision="13" longitude="124.93266" pageId="19" pageNumber="212" specimenCount="1" typeStatus="paratype">
|
||
Kitanglad,
|
||
<quantity id="4CDF9B489B4AFFC449F1FB1EFC25602C" box="[931,1018,1264,1289]" metricMagnitude="3" metricUnit="m" metricValue="1.773" pageId="19" pageNumber="212" unit="m" value="1773.0">
|
||
<elevation id="000AD19E9B4AFFC449F1FB1EFC25602C" box="[931,1018,1264,1289]" metricMagnitude="3" metricUnit="m" metricValue="1.773" pageId="19" pageNumber="212" unit="m" value="1773.0">1773 m</elevation>
|
||
</quantity>
|
||
,
|
||
<geoCoordinate id="EE13506A9B4AFFC44E54FB1EFBA0602C" box="[1030,1151,1264,1289]" degrees="08" direction="north" minutes="09.91" orientation="latitude" pageId="19" pageNumber="212" precision="9" value="8.165167">
|
||
08°09.91
|
||
<emphasis id="B953EABF9B4AFFC44E34FB1EFBB3602D" bold="true" box="[1126,1132,1264,1288]" pageId="19" pageNumber="212">ʹ</emphasis>
|
||
N
|
||
</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE13506A9B4AFFC44AF2FAFDFEFB6009" box="[160,292,1299,1324]" degrees="124" direction="east" minutes="55.96" orientation="longitude" pageId="19" pageNumber="212" precision="9" value="124.93266">
|
||
124°55.96
|
||
<emphasis id="B953EABF9B4AFFC44B5CFAFDFECB600E" bold="true" box="[270,276,1299,1323]" pageId="19" pageNumber="212">ʹ</emphasis>
|
||
E
|
||
</geoCoordinate>
|
||
,
|
||
<collectingCountry id="F330763D9B4AFFC44B7CFAFDFE906009" box="[302,335,1299,1324]" name="Philippines" pageId="19" pageNumber="212">PH</collectingCountry>
|
||
</materialsCitation>
|
||
.
|
||
<collectionCode id="ED36AE689B4AFFC44B03FAFDFEA86009" box="[337,375,1299,1324]" country="Brazil" name="Museu Nacional, Universidade Federal do Rio de Janeiro" pageId="19" pageNumber="212">MN</collectionCode>
|
||
.
|
||
<materialsCitation id="3B4F3CF09B4AFFC44B28FAFDFD746009" ID-GBIF-Occurrence="4058636327" box="[378,683,1299,1324]" collectingDate="2012-05-03" collectionCode="UCONN" country="Philippines" pageId="19" pageNumber="212" specimenCount="1" typeStatus="paratype">
|
||
KGS,
|
||
<date id="FF99106D9B4AFFC44BEBFAFDFDE36009" box="[441,572,1299,1324]" pageId="19" pageNumber="212" value="2012-05-03">
|
||
<collectingDate id="EFDDE9859B4AFFC44BEBFAFDFDE36009" box="[441,572,1299,1324]" pageId="19" pageNumber="212" value="2012-05-03">3 May 2012</collectingDate>
|
||
</date>
|
||
(
|
||
<collectionCode id="ED36AE689B4AFFC4481BFAFDFD796009" box="[585,678,1299,1324]" country="USA" httpUri="http://grbio.org/cool/jrf7-y871" name="University of Connecticut Biodiversity Research Collections" pageId="19" pageNumber="212">UCONN</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33D65269B4AFFC44AF2FA8DFED860E7" pageId="19" pageNumber="212" type="etymology">
|
||
<paragraph id="8B9836AD9B4AFFC44AF2FA8DFEC26058" blockId="19.[160,1155,1379,1474]" box="[160,285,1379,1405]" pageId="19" pageNumber="212">
|
||
<emphasis id="B953EABF9B4AFFC44AF2FA8DFEC26058" bold="true" box="[160,285,1379,1405]" italics="true" pageId="19" pageNumber="212">Etymology</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B9836AD9B4AFFC44AF2FA68FED860E7" blockId="19.[160,1155,1379,1474]" pageId="19" pageNumber="212">
|
||
The specific name is the Latin feminine adjective
|
||
<taxonomicName id="4C274D2E9B4AFFC4489BFA68FCD660BA" authorityName="Lee and Marshall" authorityYear="2023" box="[713,777,1414,1439]" class="Insecta" family="Cicadidae" genus="Oncotympana" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="212" phylum="Arthropoda" rank="species" species="obesa">
|
||
<emphasis id="B953EABF9B4AFFC4489BFA68FCD660BA" box="[713,777,1414,1439]" italics="true" pageId="19" pageNumber="212">obesa</emphasis>
|
||
</taxonomicName>
|
||
, meaning ̍fat̾, ̍obese̾, ̍plump̾ or ̍swollen̾.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33D65269B4AFFC44AF2FA17FD1163E5" pageId="19" pageNumber="212" type="description">
|
||
<paragraph id="8B9836AD9B4AFFC44AF2FA17FDF86336" blockId="19.[160,1156,1529,1728]" box="[160,551,1529,1555]" pageId="19" pageNumber="212">
|
||
<emphasis id="B953EABF9B4AFFC44AF2FA17FDF86336" bold="true" box="[160,551,1529,1555]" italics="true" pageId="19" pageNumber="212">
|
||
Measurements of
|
||
<typeStatus id="549C880F9B4AFFC44B26FA17FE6D6336" box="[372,434,1529,1555]" pageId="19" pageNumber="212">types</typeStatus>
|
||
(
|
||
<specimenCount id="9D21FD249B4AFFC44B91FA17FDC16336" box="[451,542,1529,1555]" pageId="19" pageNumber="212" type="male">5 males</specimenCount>
|
||
)
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B9836AD9B4AFFC44AF2F9F3FD1163E5" blockId="19.[160,1156,1529,1728]" pageId="19" pageNumber="212">Length of body: 22.3 (21.3–23.3). Length of head and thorax together: 11.3 (11.0–11.6). Length of abdomen: 11.0 (10.2–11.7). Width of head including compound eyes: 6.3 (6.1– 6.4). Width of pronotum: 8.9 (8.6–9.4). Width of mesonotum: 8.0 (7.7–8.4). Width of abdominal tergite 3: 9.8 (9.7–10.1). Length of fore wing: 31.2 (30.6–32.3). Width of fore wing: 10.0 (9.5–10.2). Wing span: 68.6 (67.1–71.7).</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF5866259B4DFFC34AF2FAAAFB9D605C" ID-DOI="http://doi.org/10.5281/zenodo.7737288" ID-Zenodo-Dep="7737288" httpUri="https://zenodo.org/record/7737288/files/figure.png" pageId="20" pageNumber="213" startId="20.[160,225,1348,1371]" targetBox="[162,1132,175,1297]" targetPageId="20">
|
||
<paragraph id="8B9836AD9B4DFFC34AF2FAAAFB9D605C" blockId="20.[160,1155,1348,1401]" pageId="20" pageNumber="213">
|
||
<emphasis id="B953EABF9B4DFFC34AF2FAAAFF22607E" bold="true" box="[160,253,1348,1371]" pageId="20" pageNumber="213">Figure 9.</emphasis>
|
||
<taxonomicName id="4C274D2E9B4DFFC34B5AFAAAFD55607E" authority="Lee and Marshall" authorityName="Lee and Marshall" authorityYear="2023" box="[264,650,1348,1371]" class="Insecta" family="Cicadidae" genus="Oncotympana" kingdom="Animalia" order="Hemiptera" pageId="20" pageNumber="213" phylum="Arthropoda" rank="species" species="obesa" status="sp. nov.">
|
||
<emphasis id="B953EABF9B4DFFC34B5AFAAAFE0E607E" box="[264,465,1348,1371]" italics="true" pageId="20" pageNumber="213">Oncotympana obesa</emphasis>
|
||
Lee and Marshall
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26057C49B4DFFC348C6FAAAFD38607E" box="[660,743,1348,1371]" pageId="20" pageNumber="213" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, male, Mindanao, Philippines. (A) Dorsal habitus of holotype; (B) ventral habitus of holotype; (C) pygofer, slightly oblique ventral view.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C33D65269B4DFFC34AF2FA5AFBB26313" pageId="20" pageNumber="213" type="diagnosis">
|
||
<paragraph id="8B9836AD9B4DFFC34AF2FA5AFECB60EB" blockId="20.[160,1155,1460,1590]" box="[160,276,1460,1486]" pageId="20" pageNumber="213">
|
||
<emphasis id="B953EABF9B4DFFC34AF2FA5AFECB60EB" bold="true" box="[160,276,1460,1486]" italics="true" pageId="20" pageNumber="213">Diagnosis</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B9836AD9B4DFFC34AF2FA39FBB26313" blockId="20.[160,1155,1460,1590]" pageId="20" pageNumber="213">
|
||
This new species resembles
|
||
<taxonomicName id="4C274D2E9B4DFFC34B8AFA39FD6160D5" box="[472,702,1495,1520]" class="Insecta" family="Cicadidae" genus="Oncotympana" kingdom="Animalia" order="Hemiptera" pageId="20" pageNumber="213" phylum="Arthropoda" rank="species" species="averta">
|
||
<emphasis id="B953EABF9B4DFFC34B8AFA39FD6160D5" box="[472,702,1495,1520]" italics="true" pageId="20" pageNumber="213">Oncotympana averta</emphasis>
|
||
</taxonomicName>
|
||
but has a distinctively larger body, more distinct infuscations on the fore wing, a proportionally much larger male abdomen, and differently shaped uncal lobes of the male genitalia, as detailed in the Remarks below.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33D65269B4DFFC14AF2F98FFBEB61D8" lastPageId="22" lastPageNumber="215" pageId="20" pageNumber="213" type="description">
|
||
<paragraph id="8B9836AD9B4DFFC34AF2F98FFE57635E" blockId="20.[160,1156,1633,1728]" box="[160,392,1633,1659]" pageId="20" pageNumber="213">
|
||
<emphasis id="B953EABF9B4DFFC34AF2F98FFE57635E" bold="true" box="[160,392,1633,1659]" italics="true" pageId="20" pageNumber="213">Description of male</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B9836AD9B4DFFC24AF2F96AFCC66066" blockId="20.[160,1156,1633,1728]" lastBlockId="21.[160,1157,1045,1347]" lastPageId="21" lastPageNumber="214" pageId="20" pageNumber="213">
|
||
<emphasis id="B953EABF9B4DFFC34AF2F96AFF3A63BB" bold="true" box="[160,229,1668,1694]" italics="true" pageId="20" pageNumber="213">Head.</emphasis>
|
||
Vertex yellowish green with the following black to dark brown marks: median mark enclosing ocelli, reaching posterior margin of vertex; a pair of spots on supraantennal plates; a pair of obliquely longitudinal small spots on posterolateral corners of vertex; a pair of indistinct tiny spots on anterolateral parts of vertex; and irregularly shaped transverse fascia along anterior margin of vertex. Supra-antennal plates produced anteriad, covering scape. Compound eyes brown in life. Distance between lateral ocelli and compound eyes about twice the distance between lateral ocelli. Antenna mostly black but with dark brown to fuscous pedicel. Postclypeus moderately swollen; yellowish green with a few black to dark brown fasciae along anterior transverse grooves. Anteclypeus ochraceous with a pair of central fuscous spots. Rostrum ochraceous with black apex, passing posterior margin of hind trochanter.
|
||
</paragraph>
|
||
<caption id="DF5866259B4CFFC24AF2FC90FD246697" ID-DOI="http://doi.org/10.5281/zenodo.7737290" ID-Zenodo-Dep="7737290" httpUri="https://zenodo.org/record/7737290/files/figure.png" pageId="21" pageNumber="214" startId="21.[160,225,894,917]" targetBox="[164,1151,144,859]" targetPageId="21">
|
||
<paragraph id="8B9836AD9B4CFFC24AF2FC90FD246697" blockId="21.[160,1155,894,946]" pageId="21" pageNumber="214">
|
||
<emphasis id="B953EABF9B4CFFC24AF2FC90FED666B0" bold="true" box="[160,265,894,917]" pageId="21" pageNumber="214">Figure 10.</emphasis>
|
||
<taxonomicName id="4C274D2E9B4CFFC24B40FC90FD4F66B0" authority="Lee and Marshall" authorityName="Lee and Marshall" authorityYear="2023" box="[274,656,894,917]" class="Insecta" family="Cicadidae" genus="Oncotympana" kingdom="Animalia" order="Hemiptera" pageId="21" pageNumber="214" phylum="Arthropoda" rank="species" species="obesa" status="sp. nov.">
|
||
<emphasis id="B953EABF9B4CFFC24B40FC90FE0566B0" box="[274,474,894,917]" italics="true" pageId="21" pageNumber="214">Oncotympana obesa</emphasis>
|
||
Lee and Marshall
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26057C49B4CFFC248CAFC90FD3566B0" box="[664,746,894,917]" pageId="21" pageNumber="214" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype male calling song. Waveforms (A, C) and spectrograms (B, D) are shown at two time scales.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B9836AD9B4CFFC14AF2FA80FB926452" blockId="21.[160,1156,1390,1728]" lastBlockId="22.[160,1156,142,1034]" lastPageId="22" lastPageNumber="215" pageId="21" pageNumber="214">
|
||
<emphasis id="B953EABF9B4CFFC24AF2FA80FF2560AD" bold="true" box="[160,250,1390,1416]" italics="true" pageId="21" pageNumber="214">Thorax.</emphasis>
|
||
Pronotum greenish ochraceous. Inner area of pronotum with the following fuscous marks: a pair of medial longitudinal fasciae not reaching anterior margin of pronotum, broadened near posterior ends and meeting each other at posterior ends; a pair of marks along paramedian fissures; a pair of short longitudinal fasciae between median parts of paramedian fissures and posterior ends of lateral fissures; a pair of marks along lateral fissures; and a pair of curved fasciae along lateral margins of inner area. Pronotal collar with a pair of indistinct marks near posterolateral corners; posterior marginal area very narrow. Anterolateral pronotal collar not dentate but sinuate. Mesonotum greenish ochraceous with the following fuscous to black marks: medial longitudinal fascia reaching anterior margin of cruciform elevation but not reaching anterior margin of mesonotum; a pair of small roundish spots enclosing scutal depressions; a pair of large obconical paramedian marks on submedian sigilla, indistinct in their inner parts; a pair of long obconical sublateral marks on lateral sigilla, discontinued in the middle; and a pair of small spots on posterolateral corners of mesonotum. Cruciform elevation greenish ochraceous without distinct marks. Thoracic sternites greenish ochraceous. Legs greenish ochraceous to green with some brown parts and black to fuscous marks. Fore femur with primary and secondary spines and minute subapical spines.
|
||
</paragraph>
|
||
<paragraph id="8B9836AD9B4FFFC14A92FE6FFE3E674F" blockId="22.[160,1156,142,1034]" pageId="22" pageNumber="215">Wings hyaline, slightly tinged with brown. Fore wing costal vein mostly ochraceous but fuscous near node. Basal vein of apical cell 1 about one-third to one-fourth as long as longitudinal vein of the apical cell 1. Infuscation present at bases of apical cells 1–5 and 7– 8 and on cubitus anterior 2 vein. A distinct spot appearing on each hind margin of radius anterior 2, radius posterior, median 1–4 and cubitus anterior 1 veins, forming a series of spots on subapical margin of fore wing. Basal membrane and base of hind wing jugum greenish (bright blue in life).</paragraph>
|
||
<paragraph id="8B9836AD9B4FFFC14A92FD9DFE8367D1" blockId="22.[160,1156,142,1034]" pageId="22" pageNumber="215">Operculum light greenish ochraceous without distinct marks; much wider than long, with rounded posterior apex passing beyond posterior margin of abdominal sternite II and with posterolateral corner almost straight but slightly convex. Opercula separated from each other.</paragraph>
|
||
<paragraph id="8B9836AD9B4FFFC14A92FD10FB9F612F" blockId="22.[160,1156,142,1034]" pageId="22" pageNumber="215">Abdomen obconical in dorsal view, short, shorter than or about as long as head and thorax together. Tergites 2–8 mostly castaneous, with light green to green posterior margins (blue in life). Tergites 2–5 with a pair of fuscous paramedian patches. Tergites 4–8 with a pair of lateral fuscous patches. Posterior margin of tergite 3 much wider than pronotum or mesonotum. Timbal cover dull greenish ochraceous with black to fuscous patch along posterior margin; globose, wider than long, with anterior and posterior inner corners smoothly rounded. Timbal completely concealed by timbal cover in dorsal view. Abdominal sternites II–VIII fuscous, with green posterior margins of sternites III–VI.</paragraph>
|
||
<paragraph id="8B9836AD9B4FFFC14AF2FBD8FBEB61D8" blockId="22.[160,1156,1078,1277]" pageId="22" pageNumber="215">
|
||
<emphasis id="B953EABF9B4FFFC14AF2FBD8FECB6175" bold="true" box="[160,276,1078,1104]" italics="true" pageId="22" pageNumber="215">Genitalia.</emphasis>
|
||
Pygofer barrel-shaped in ventral view. Uncus bifurcate. Uncal lobes each divided into two parts: a long external part, and a shorter internal part, which is less than half as long as external part: in ventral view, external parts curved and extended obliquely laterad, and internal parts convexly curved with tips directed inward; in lateral view, external parts convexly curved with tips directed inward, and internal parts curved inward. Aedeagus slender. Dorsal beak long, narrow, about as long as anal styles.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33D65269B4FFFC04AF2FAC7FD6565C8" lastPageId="23" lastPageNumber="216" pageId="22" pageNumber="215" type="discussion">
|
||
<paragraph id="8B9836AD9B4FFFC14AF2FAC7FED96066" blockId="22.[160,1156,1321,1728]" box="[160,262,1321,1347]" pageId="22" pageNumber="215">
|
||
<emphasis id="B953EABF9B4FFFC14AF2FAC7FED96066" bold="true" box="[160,262,1321,1347]" italics="true" pageId="22" pageNumber="215">Remarks</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B9836AD9B4FFFC04AF2FAA3FD6565C8" blockId="22.[160,1156,1321,1728]" lastBlockId="23.[160,1156,142,237]" lastPageId="23" lastPageNumber="216" pageId="22" pageNumber="215">
|
||
This species is very similar to
|
||
<taxonomicName id="4C274D2E9B4FFFC14BB0FAA3FD186043" box="[482,711,1357,1382]" class="Insecta" family="Cicadidae" genus="Oncotympana" kingdom="Animalia" order="Hemiptera" pageId="22" pageNumber="215" phylum="Arthropoda" rank="species" species="averta">
|
||
<emphasis id="B953EABF9B4FFFC14BB0FAA3FD186043" box="[482,711,1357,1382]" italics="true" pageId="22" pageNumber="215">Oncotympana averta</emphasis>
|
||
</taxonomicName>
|
||
but can be distinguished from
|
||
<taxonomicName id="4C274D2E9B4FFFC14E73FAA3FB5C6043" box="[1057,1155,1357,1382]" class="Insecta" family="Cicadidae" genus="Oncotympana" kingdom="Animalia" order="Hemiptera" pageId="22" pageNumber="215" phylum="Arthropoda" rank="species" species="averta">
|
||
<emphasis id="B953EABF9B4FFFC14E73FAA3FB5C6043" box="[1057,1155,1357,1382]" italics="true" pageId="22" pageNumber="215">O. averta</emphasis>
|
||
</taxonomicName>
|
||
by the following characteristics: body distinctly larger; infuscation present at base of apical cells 4 and 8 (vs absent there in
|
||
<taxonomicName id="4C274D2E9B4FFFC14821FA7CFD05608E" box="[627,730,1426,1451]" class="Insecta" family="Cicadidae" genus="Oncotympana" kingdom="Animalia" order="Hemiptera" pageId="22" pageNumber="215" phylum="Arthropoda" rank="species" species="averta">
|
||
<emphasis id="B953EABF9B4FFFC14821FA7CFD05608E" box="[627,730,1426,1451]" italics="true" pageId="22" pageNumber="215">O. averta</emphasis>
|
||
</taxonomicName>
|
||
); basal membrane and base of hind wing jugum greenish (vs greyish in
|
||
<taxonomicName id="4C274D2E9B4FFFC1481CFA5BFD6960EB" box="[590,694,1461,1486]" class="Insecta" family="Cicadidae" genus="Oncotympana" kingdom="Animalia" order="Hemiptera" pageId="22" pageNumber="215" phylum="Arthropoda" rank="species" species="averta">
|
||
<emphasis id="B953EABF9B4FFFC1481CFA5BFD6960EB" box="[590,694,1461,1486]" italics="true" pageId="22" pageNumber="215">O. averta</emphasis>
|
||
</taxonomicName>
|
||
); operculum with posterolateral corner almost straight but slightly convex (vs slightly concave in
|
||
<taxonomicName id="4C274D2E9B4FFFC1497DFA39FC4B60D5" box="[815,916,1495,1520]" class="Insecta" family="Cicadidae" genus="Oncotympana" kingdom="Animalia" order="Hemiptera" pageId="22" pageNumber="215" phylum="Arthropoda" rank="species" species="averta">
|
||
<emphasis id="B953EABF9B4FFFC1497DFA39FC4B60D5" box="[815,916,1495,1520]" italics="true" pageId="22" pageNumber="215">O. averta</emphasis>
|
||
</taxonomicName>
|
||
); posterior margin of tergite 3 much wider than pronotum or mesonotum (vs about as wide as or slightly narrower than mesonotum in
|
||
<taxonomicName id="4C274D2E9B4FFFC14BBEF9F3FD916313" box="[492,590,1565,1590]" class="Insecta" family="Cicadidae" genus="Oncotympana" kingdom="Animalia" order="Hemiptera" pageId="22" pageNumber="215" phylum="Arthropoda" rank="species" species="averta">
|
||
<emphasis id="B953EABF9B4FFFC14BBEF9F3FD916313" box="[492,590,1565,1590]" italics="true" pageId="22" pageNumber="215">O. averta</emphasis>
|
||
</taxonomicName>
|
||
); abdominal sternites II–VIII fuscous (vs sternites II– V mostly ochraceous but sternites VI–VIII fuscous in
|
||
<taxonomicName id="4C274D2E9B4FFFC148BFF9D1FC8D637D" box="[749,850,1599,1624]" class="Insecta" family="Cicadidae" genus="Oncotympana" kingdom="Animalia" order="Hemiptera" pageId="22" pageNumber="215" phylum="Arthropoda" rank="species" species="averta">
|
||
<emphasis id="B953EABF9B4FFFC148BFF9D1FC8D637D" box="[749,850,1599,1624]" italics="true" pageId="22" pageNumber="215">O. averta</emphasis>
|
||
</taxonomicName>
|
||
); timbal cover globose but more or less flat (vs much more prominently globose in
|
||
<taxonomicName id="4C274D2E9B4FFFC14976F98CFC56635E" box="[804,905,1634,1659]" class="Insecta" family="Cicadidae" genus="Oncotympana" kingdom="Animalia" order="Hemiptera" pageId="22" pageNumber="215" phylum="Arthropoda" rank="species" species="averta">
|
||
<emphasis id="B953EABF9B4FFFC14976F98CFC56635E" box="[804,905,1634,1659]" italics="true" pageId="22" pageNumber="215">O. averta</emphasis>
|
||
</taxonomicName>
|
||
); dark-coloured patch absent on inner corner of timbal cover (vs present in
|
||
<taxonomicName id="4C274D2E9B4FFFC1495AF96BFCB263BB" box="[776,877,1669,1694]" class="Insecta" family="Cicadidae" genus="Oncotympana" kingdom="Animalia" order="Hemiptera" pageId="22" pageNumber="215" phylum="Arthropoda" rank="species" species="averta">
|
||
<emphasis id="B953EABF9B4FFFC1495AF96BFCB263BB" box="[776,877,1669,1694]" italics="true" pageId="22" pageNumber="215">O. averta</emphasis>
|
||
</taxonomicName>
|
||
); external parts of uncal lobes curved and extended obliquely laterad in ventral view (vs nearly parallel to each other but with curved tips extended obliquely laterad in
|
||
<taxonomicName id="4C274D2E9B4EFFC04976FF60FC576582" box="[804,904,142,167]" class="Insecta" family="Cicadidae" genus="Oncotympana" kingdom="Animalia" order="Hemiptera" pageId="23" pageNumber="216" phylum="Arthropoda" rank="species" species="averta">
|
||
<emphasis id="B953EABF9B4EFFC04976FF60FC576582" box="[804,904,142,167]" italics="true" pageId="23" pageNumber="216">O. averta</emphasis>
|
||
</taxonomicName>
|
||
); and internal parts of uncal lobes convexly curved with tips directed inward in ventral view (vs concavely curved with hooked tips extended laterad in
|
||
<taxonomicName id="4C274D2E9B4EFFC0481BFF3AFD7265C8" box="[585,685,212,237]" class="Insecta" family="Cicadidae" genus="Oncotympana" kingdom="Animalia" order="Hemiptera" pageId="23" pageNumber="216" phylum="Arthropoda" rank="species" species="averta">
|
||
<emphasis id="B953EABF9B4EFFC0481BFF3AFD7265C8" box="[585,685,212,237]" italics="true" pageId="23" pageNumber="216">O. averta</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33D65269B4EFFC04AF2FEF2FD0266A6" pageId="23" pageNumber="216" type="description">
|
||
<paragraph id="8B9836AD9B4EFFC04AF2FEF2FEB86413" blockId="23.[160,1156,284,899]" box="[160,359,284,310]" pageId="23" pageNumber="216">
|
||
<emphasis id="B953EABF9B4EFFC04AF2FEF2FEB86413" bold="true" box="[160,359,284,310]" pageId="23" pageNumber="216">
|
||
<emphasis id="B953EABF9B4EFFC04AF2FEF2FF036413" bold="true" box="[160,220,284,310]" italics="true" pageId="23" pageNumber="216">Song</emphasis>
|
||
(
|
||
<figureCitation id="131C2A289B4EFFC04ABEFEF2FE806413" box="[236,351,284,310]" captionStart="Figure 10" captionStartId="21.[160,225,894,917]" captionTargetBox="[164,1151,144,859]" captionTargetId="figure-233@21.[162,1154,142,862]" captionTargetPageId="21" captionText="Figure 10. Oncotympana obesa Lee and Marshall sp. nov., holotype male calling song. Waveforms (A, C) and spectrograms (B, D) are shown at two time scales." figureDoi="http://doi.org/10.5281/zenodo.7737290" httpUri="https://zenodo.org/record/7737290/files/figure.png" pageId="23" pageNumber="216">Figure 10</figureCitation>
|
||
)
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B9836AD9B4EFFC04AF2FED1FE266796" blockId="23.[160,1156,284,899]" pageId="23" pageNumber="216">
|
||
Recordings of male calling song totalling 320 s were taken at the type locality (the
|
||
<typeStatus id="549C880F9B4EFFC04AF2FE8CFEDB645E" box="[160,260,354,379]" pageId="23" pageNumber="216" type="holotype">holotype</typeStatus>
|
||
male is illustrated). Males were recorded producing song in phrases of 4–11 s duration (4–6 most common) comprised of two-syllable echemes repeated at about 5.2– 6.5 echemes/s. Note that such values are commonly dependent on temperature. The syllables within each echeme contain strongly contrasting sound frequencies. The first syllable contains most of its sound energy within the range 7–15 kHz, with a peak at about 10 kHz, while the second syllable, which is nearly an order of magnitude quieter than the first, contains sound energy with a peak of 4.5 kHz and a broad shoulder of quieter frequencies ranging from 2–16 kHz. The fundamental sound pulses are produced at about 225/s in the first syllable, which lasts
|
||
<geoCoordinate id="EE13506A9B4EFFC04867FD99FDA967B5" box="[565,630,631,656]" degrees="0.04" direction="south" orientation="latitude" pageId="23" pageNumber="216" precision="555" value="-0.04">0.04 s</geoCoordinate>
|
||
, and slightly faster at about 235/s in the second syllable which is twice as long.
|
||
</paragraph>
|
||
<paragraph id="8B9836AD9B4EFFC04A92FD53FD0266A6" blockId="23.[160,1156,284,899]" pageId="23" pageNumber="216">
|
||
This species sang from low stations on tree trunks in the forest understory in low light conditions, as early as 7.00am. Males could be attracted by mouth-clicks or finger-snaps produced a fraction of a second after the end of the complete song phrase. Immediately after hearing the response, the male would flick his wings with a synchronised timbal click, before turning or moving in the direction of the sound. Several specimens from the
|
||
<typeStatus id="549C880F9B4EFFC04AF2FC84FEDB66A6" box="[160,260,874,899]" pageId="23" pageNumber="216" type="paratype">paratype</typeStatus>
|
||
series were collected with this technique.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |