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<document id="620E8F4A4D8E94B37591F880C8E578FD" ID-DOI="10.11646/zootaxa.4105.1.1" ID-GBIF-Dataset="cf0a97f6-504a-4de4-a51c-a11c5c80bca3" ID-ISSN="1175-5326" ID-Zenodo-Dep="271230" ID-ZooBank="621E2759-DDBF-4ADC-A1EC-3CA8F581C336" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1461046449829" checkinUser="plazi" docAuthor="Samimi-Namin, Kaveh &amp; Van Ofwegen, Leen P." docDate="2016" docId="038E8793991AD965D4C09AE0855C31F3" docLanguage="en" docName="zootaxa.4105.1.1.pdf.imf" docOrigin="Zootaxa 4105 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Trimuricea caledonica Grasshoff 1999" docType="treatment" docVersion="6" lastPageNumber="11" masterDocId="FFB7FFEB9910D96FD4579B7480603729" masterDocTitle="A revision of Trimuricea Gordon, 1926 (Cnidaria: Octocorallia: Plexauridae) with the description of six new species" masterLastPageNumber="44" masterPageNumber="1" pageNumber="11" updateTime="1698577331484" updateUser="plazi">
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<mods:title id="84B83F38A8B21E05671F36E7918D9818">A revision of Trimuricea Gordon, 1926 (Cnidaria: Octocorallia: Plexauridae) with the description of six new species</mods:title>
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<heading id="D0D081E9991AD965D4C09AE082153687" bold="true" box="[151,629,404,430]" fontSize="11" level="1" pageId="10" pageNumber="11" reason="1">
<taxonomicName id="4C274D06991AD965D4C09AE082153687" ID-CoL="58VCH" authority="Grasshoff, 1999" authorityName="Grasshoff" authorityYear="1999" box="[151,629,404,430]" class="Anthozoa" family="Plexauridae" genus="Trimuricea" kingdom="Animalia" order="Alcyonacea" pageId="10" pageNumber="11" phylum="Cnidaria" rank="species" species="caledonica">
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<emphasis id="B953EA97991AD965D4C09AE081C93687" bold="true" box="[151,425,404,430]" italics="true" pageId="10" pageNumber="11">Trimuricea caledonica</emphasis>
<bibRefCitation id="EFB64B74991AD965D5F89AE082153687" author="Grasshoff" box="[431,629,404,430]" pageId="10" pageNumber="11" refString="Grasshoff, M. (1999) The shallow water gorgonians of New Caledonia and adjacent islands (Coelenterata: Octocorallia). Senckenbergiana biologica, 78 (1 / 2), 1 - 245." type="journal article" year="1999">Grasshoff, 1999</bibRefCitation>
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(
<figureCitation id="131C2A00991AD965D4C89AC2808E36E6" box="[159,238,438,463]" captionStart="FIGURE 4" captionStartId="7.[151,250,1951,1973]" captionTargetBox="[151,1436,193,1927]" captionTargetId="figure@7.[151,1436,193,1929]" captionTargetPageId="7" captionText="FIGURE 4. Holotype colonies: a, T. africana, Gordon, 1926, BMNH 1933.5.3.102; b, T. bicolor n. sp., RMNH Coel. 39860; c, T. caledonica SMF 6752; d, T. flava n. sp., RMNH Coel. 39861." httpUri="https://zenodo.org/record/271233/files/figure.png" pageId="10" pageNumber="11">Figs. 4</figureCitation>
c, 78)
</paragraph>
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<taxonomicName id="4C274D06991AD965D4C09974822F353D" ID-CoL="58VCH" authority="Grasshoff, 1999: 48" authorityName="Grasshoff" authorityPageNumber="48" authorityYear="1999" box="[151,591,510,533]" class="Anthozoa" family="Plexauridae" genus="Trimuricea" kingdom="Animalia" order="Alcyonacea" pageId="10" pageNumber="11" phylum="Cnidaria" rank="species" species="caledonica">
<emphasis id="B953EA97991AD965D4C09974811D353D" box="[151,381,510,533]" italics="true" pageId="10" pageNumber="11">Trimuricea caledonica</emphasis>
<treatmentCitation id="0A861094991AD965D5D49A8A822F353D" author="Grasshoff" box="[387,591,510,532]" page="48" pageId="10" pageNumber="11" year="1999">
<bibRefCitation id="EFB64B74991AD965D5D49A8A824A353D" author="Grasshoff" box="[387,554,510,532]" pageId="10" pageNumber="11" refString="Grasshoff, M. (1999) The shallow water gorgonians of New Caledonia and adjacent islands (Coelenterata: Octocorallia). Senckenbergiana biologica, 78 (1 / 2), 1 - 245." type="journal article" year="1999">Grasshoff, 1999</bibRefCitation>
: 48
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, figs. 8183;
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<bibRefCitation id="EFB64B74991AD965D68F9A8A839C353D" author="Grasshoff" box="[728,1020,510,532]" pageId="10" pageNumber="11" refString="Grasshoff, M. &amp; Bargibant, G. (2001) Coral Reef Gorgonians of New Caledonia. Institut de recherche pour le developpement, Paris, 335 pp." type="book" year="2001">Grasshoff &amp; Bargibant 2001</bibRefCitation>
: 183
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; Fabricius &amp; Alderslade 2001: 196.
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</paragraph>
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<paragraph id="8B983685991AD965D4C0993581FA358B" blockId="10.[151,1436,577,1754]" pageId="10" pageNumber="11">
<emphasis id="B953EA97991AD965D4C0993581683573" bold="true" box="[151,264,577,602]" pageId="10" pageNumber="11">Material:</emphasis>
<collectionCode id="ED36AE40991AD965D547993581283570" LSID="urn:lsid:biocol.org:col:34838" box="[272,328,577,601]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="10" pageNumber="11">SMF</collectionCode>
6752, part of
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,
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, W lagon, Off Nouméa, Ilot Canard, Stn 136, 23 m;
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6753, part of
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,
<collectingCountry id="F3307615991AD965D5F2991282383557" box="[421,600,613,638]" name="New Caledonia" pageId="10" pageNumber="11">New Caledonia</collectingCountry>
, W GRext, Passe de Koumac,
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;
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6754,
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, South Pacific Ocean,
<collectingCountry id="F3307615991AD965D4BB99FE81FA358B" box="[236,410,649,674]" name="New Caledonia" pageId="10" pageNumber="11">New Caledonia</collectingCountry>
</paragraph>
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Description (modified after
<bibRefCitation id="EFB64B74991AD965D64499D982AB35EF" author="Grasshoff" box="[531,715,685,710]" pageId="10" pageNumber="11" refString="Grasshoff, M. (1999) The shallow water gorgonians of New Caledonia and adjacent islands (Coelenterata: Octocorallia). Senckenbergiana biologica, 78 (1 / 2), 1 - 245." type="journal article" year="1999">Grasshoff 1999</bibRefCitation>
).
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The complete
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was a large fan
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high, with many anastomoses; only a portion is shown in
<figureCitation id="131C2A00991AD965D60999A582C135C3" box="[606,673,721,746]" captionStart="FIGURE 4" captionStartId="7.[151,250,1951,1973]" captionTargetBox="[151,1436,193,1927]" captionTargetId="figure@7.[151,1436,193,1929]" captionTargetPageId="7" captionText="FIGURE 4. Holotype colonies: a, T. africana, Gordon, 1926, BMNH 1933.5.3.102; b, T. bicolor n. sp., RMNH Coel. 39860; c, T. caledonica SMF 6752; d, T. flava n. sp., RMNH Coel. 39861." httpUri="https://zenodo.org/record/271233/files/figure.png" pageId="10" pageNumber="11">Fig. 4</figureCitation>
c. The polyps are situated all around the branches, densely crowded, leaving little free coenenchyme in between. Calyces are dome shaped and up to
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high and
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wide.
</paragraph>
<paragraph id="8B983685991AD965D490986D81C334EB" blockId="10.[151,1436,577,1754]" pageId="10" pageNumber="11">
The points have 24 triradiates (
<figureCitation id="131C2A00991AD965D679986D8210341B" box="[558,624,793,818]" captionStart="FIGURE 7" captionStartId="11.[151,250,1965,1987]" captionTargetBox="[151,1436,193,1943]" captionTargetId="figure@11.[151,1436,193,1944]" captionTargetPageId="11" captionText="FIGURE 7. The polyp sclerites of T. caledonica Grasshoff, 1999, paratype, SMF 6753: a, point triradiates; b, point spindles; c, collaret spindles; d, tentacle scale." httpUri="https://zenodo.org/record/271236/files/figure.png" pageId="10" pageNumber="11">Fig. 7</figureCitation>
a) and 12 curved, hockeystick or boot-shaped sclerites, or spindles (
<figureCitation id="131C2A00991AD965D138986D80C6347F" captionStart="FIGURE 7" captionStartId="11.[151,250,1965,1987]" captionTargetBox="[151,1436,193,1943]" captionTargetId="figure@11.[151,1436,193,1944]" captionTargetPageId="11" captionText="FIGURE 7. The polyp sclerites of T. caledonica Grasshoff, 1999, paratype, SMF 6753: a, point triradiates; b, point spindles; c, collaret spindles; d, tentacle scale." httpUri="https://zenodo.org/record/271236/files/figure.png" pageId="10" pageNumber="11">Fig. 7</figureCitation>
b),
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long. The upper ray of the triradiates and the upper part of the spindles is smooth for up to
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, rather wide and blunt ended. The collaret spindles are
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long (
<figureCitation id="131C2A00991AD965D0629815841A3453" box="[1077,1146,865,890]" captionStart="FIGURE 7" captionStartId="11.[151,250,1965,1987]" captionTargetBox="[151,1436,193,1943]" captionTargetId="figure@11.[151,1436,193,1944]" captionTargetPageId="11" captionText="FIGURE 7. The polyp sclerites of T. caledonica Grasshoff, 1999, paratype, SMF 6753: a, point triradiates; b, point spindles; c, collaret spindles; d, tentacle scale." httpUri="https://zenodo.org/record/271236/files/figure.png" pageId="10" pageNumber="11">Fig. 7</figureCitation>
c); the middle part is less tuberculate than both ends. These polyp sclerites have very few tubercles. A few tentacle scales are present, up to
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long (
<figureCitation id="131C2A00991AD965D51498DD81E634EB" box="[323,390,937,962]" captionStart="FIGURE 7" captionStartId="11.[151,250,1965,1987]" captionTargetBox="[151,1436,193,1943]" captionTargetId="figure@11.[151,1436,193,1944]" captionTargetPageId="11" captionText="FIGURE 7. The polyp sclerites of T. caledonica Grasshoff, 1999, paratype, SMF 6753: a, point triradiates; b, point spindles; c, collaret spindles; d, tentacle scale." httpUri="https://zenodo.org/record/271236/files/figure.png" pageId="10" pageNumber="11">Fig. 7</figureCitation>
d).
</paragraph>
<paragraph id="8B983685991AD965D49098B980A13323" blockId="10.[151,1436,577,1754]" pageId="10" pageNumber="11">
The calyces have thornscales,
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long, with a short, smooth, blunt thorn up to
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long (
<figureCitation id="131C2A00991AD965D13898B980C53323" captionStart="FIGURE 8" captionStartId="12.[151,250,1953,1975]" captionTargetBox="[151,1436,193,1931]" captionTargetId="figure@12.[151,1436,193,1931]" captionTargetPageId="12" captionText="FIGURE 8. Sclerites of T. caledonica Grasshoff, 1999, paratype, SMF 6753: a, calicular thornscales; b, coenenchymal spindles; c, small, irregular shaped coenenchymal sclerites. Scale at c, only applies to c." httpUri="https://zenodo.org/record/271237/files/figure.png" pageId="10" pageNumber="11">Fig. 8</figureCitation>
a).
</paragraph>
<paragraph id="8B983685991AD965D4909F618181335F" blockId="10.[151,1436,577,1754]" pageId="10" pageNumber="11">
The coenenchyme has spindles and crosses,
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long, with simple tubercles (
<figureCitation id="131C2A00991AD965D0CF9F6184BA3307" box="[1176,1242,1045,1070]" captionStart="FIGURE 8" captionStartId="12.[151,250,1953,1975]" captionTargetBox="[151,1436,193,1931]" captionTargetId="figure@12.[151,1436,193,1931]" captionTargetPageId="12" captionText="FIGURE 8. Sclerites of T. caledonica Grasshoff, 1999, paratype, SMF 6753: a, calicular thornscales; b, coenenchymal spindles; c, small, irregular shaped coenenchymal sclerites. Scale at c, only applies to c." httpUri="https://zenodo.org/record/271237/files/figure.png" pageId="10" pageNumber="11">Fig. 8</figureCitation>
b). Some of these spindles have a side branch which maybe thorn-like. Furthermore, small, irregular shaped sclerites are present,
<quantity id="4CDF9B60991AD965D4C09F2A815A335C" box="[151,314,1117,1142]" metricMagnitude="-4" metricUnit="m" metricValue="1.3" metricValueMax="1.6" metricValueMin="1.0" pageId="10" pageNumber="11" unit="mm" value="0.13" valueMax="0.16" valueMin="0.1">0.100.16 mm</quantity>
long (
<figureCitation id="131C2A00991AD965D5D69F2981A5335F" box="[385,453,1117,1142]" captionStart="FIGURE 8" captionStartId="12.[151,250,1953,1975]" captionTargetBox="[151,1436,193,1931]" captionTargetId="figure@12.[151,1436,193,1931]" captionTargetPageId="12" captionText="FIGURE 8. Sclerites of T. caledonica Grasshoff, 1999, paratype, SMF 6753: a, calicular thornscales; b, coenenchymal spindles; c, small, irregular shaped coenenchymal sclerites. Scale at c, only applies to c." httpUri="https://zenodo.org/record/271237/files/figure.png" pageId="10" pageNumber="11">Fig. 8</figureCitation>
c).
</paragraph>
<paragraph id="8B983685991AD965D4909FF585153397" blockId="10.[151,1436,577,1754]" pageId="10" pageNumber="11">
<emphasis id="B953EA97991AD965D4909FF5814033B3" bold="true" box="[199,288,1153,1178]" pageId="10" pageNumber="11">Colour.</emphasis>
The live colony was red (for live colony photo see
<bibRefCitation id="EFB64B74991AD965D73F9FF584CD33B3" author="Grasshoff" box="[872,1197,1153,1178]" pageId="10" pageNumber="11" refString="Grasshoff, M. &amp; Bargibant, G. (2001) Coral Reef Gorgonians of New Caledonia. Institut de recherche pour le developpement, Paris, 335 pp." type="book" year="2001">Grasshoff &amp; Bargibant 2001</bibRefCitation>
). When dry it is dark brown, but it is yellowish-grey when preserved in fluid. No information about the colour of polyps is available.
</paragraph>
</subSubSection>
<subSubSection id="C33D650E991AD965D4909FBD855C31F3" pageId="10" pageNumber="11" type="discussion">
<paragraph id="8B983685991AD965D4909FBD82ED32F7" blockId="10.[151,1436,577,1754]" pageId="10" pageNumber="11">
<emphasis id="B953EA97991AD965D4909FBD815B33CB" bold="true" box="[199,315,1225,1250]" pageId="10" pageNumber="11">Remarks.</emphasis>
This species differs from all other
<taxonomicName id="4C274D06991AD965D6909FBE832433C8" box="[711,836,1226,1249]" class="Anthozoa" family="Plexauridae" genus="Trimuricea" kingdom="Animalia" order="Alcyonacea" pageId="10" pageNumber="11" phylum="Cnidaria" rank="genus">
<emphasis id="B953EA97991AD965D6909FBE832433C8" box="[711,836,1226,1249]" italics="true" pageId="10" pageNumber="11">Trimuricea</emphasis>
</taxonomicName>
species by having small, irregular shaped sclerites in the coenenchyme (
<figureCitation id="131C2A00991AD965D53C9F9981CE322F" box="[363,430,1261,1286]" captionStart="FIGURE 8" captionStartId="12.[151,250,1953,1975]" captionTargetBox="[151,1436,193,1931]" captionTargetId="figure@12.[151,1436,193,1931]" captionTargetPageId="12" captionText="FIGURE 8. Sclerites of T. caledonica Grasshoff, 1999, paratype, SMF 6753: a, calicular thornscales; b, coenenchymal spindles; c, small, irregular shaped coenenchymal sclerites. Scale at c, only applies to c." httpUri="https://zenodo.org/record/271237/files/figure.png" pageId="10" pageNumber="11">Fig. 8</figureCitation>
c), which were not reported by Grasshoff. Noteworthy, we noticed irregular radiates, and derivatives of radiates, similar to those in
<figureCitation id="131C2A00991AD965D62D9E6582DE3203" box="[634,702,1297,1322]" captionStart="FIGURE 8" captionStartId="12.[151,250,1953,1975]" captionTargetBox="[151,1436,193,1931]" captionTargetId="figure@12.[151,1436,193,1931]" captionTargetPageId="12" captionText="FIGURE 8. Sclerites of T. caledonica Grasshoff, 1999, paratype, SMF 6753: a, calicular thornscales; b, coenenchymal spindles; c, small, irregular shaped coenenchymal sclerites. Scale at c, only applies to c." httpUri="https://zenodo.org/record/271237/files/figure.png" pageId="10" pageNumber="11">Fig. 8</figureCitation>
c, that occur in the basal parts of colonies, and in branches with relatively solid axis, in other
<taxonomicName id="4C274D06991AD965D5B79E42823B3264" box="[480,603,1334,1357]" class="Anthozoa" family="Plexauridae" genus="Trimuricea" kingdom="Animalia" order="Alcyonacea" pageId="10" pageNumber="11" phylum="Cnidaria" rank="genus">
<emphasis id="B953EA97991AD965D5B79E42823B3264" box="[480,603,1334,1357]" italics="true" pageId="10" pageNumber="11">Trimuricea</emphasis>
</taxonomicName>
species. Therefore, we do not exclude the possibility that the piece of the colony we received from SMF might have been cut from the basal parts or older side branches of the colony, not from the top of the colony. This might also explains why Grasshoff did not mention these
<typeStatus id="549C8827991AD965D0C19E0A84B132BF" box="[1174,1233,1406,1430]" pageId="10" pageNumber="11">types</typeStatus>
of sclerites in the original description of
<taxonomicName id="4C274D06991AD965D5F79ED682593293" box="[416,569,1441,1466]" class="Anthozoa" family="Plexauridae" genus="Trimuricea" kingdom="Animalia" order="Alcyonacea" pageId="10" pageNumber="11" phylum="Cnidaria" rank="species" species="caledonica">
<emphasis id="B953EA97991AD965D5F79ED682593293" box="[416,569,1441,1466]" italics="true" pageId="10" pageNumber="11">T. caledonica</emphasis>
</taxonomicName>
. On the other hand it is possible that these small sclerites were missed by Grasshoffs technician making the drawings.
</paragraph>
<paragraph id="8B983685991AD965D4909E9D83BD3144" blockId="10.[151,1436,577,1754]" pageId="10" pageNumber="11">
If we assume that the irregular radiates (
<figureCitation id="131C2A00991AD965D6D99E9D82B0312B" box="[654,720,1513,1538]" captionStart="FIGURE 8" captionStartId="12.[151,250,1953,1975]" captionTargetBox="[151,1436,193,1931]" captionTargetId="figure@12.[151,1436,193,1931]" captionTargetPageId="12" captionText="FIGURE 8. Sclerites of T. caledonica Grasshoff, 1999, paratype, SMF 6753: a, calicular thornscales; b, coenenchymal spindles; c, small, irregular shaped coenenchymal sclerites. Scale at c, only applies to c." httpUri="https://zenodo.org/record/271237/files/figure.png" pageId="10" pageNumber="11">Fig. 8</figureCitation>
c) are not from the top of the colony, then
<taxonomicName id="4C274D06991AD965D0E69E9E8528312B" box="[1201,1352,1513,1538]" class="Anthozoa" family="Plexauridae" genus="Trimuricea" kingdom="Animalia" order="Alcyonacea" pageId="10" pageNumber="11" phylum="Cnidaria" rank="species" species="caledonica">
<emphasis id="B953EA97991AD965D0E69E9E84DE3128" box="[1201,1214,1514,1537]" italics="true" pageId="10" pageNumber="11">T</emphasis>
.
<emphasis id="B953EA97991AD965D09A9E9D8528312B" box="[1229,1352,1513,1538]" italics="true" pageId="10" pageNumber="11">caledonica</emphasis>
</taxonomicName>
mostly resembles
<taxonomicName id="4C274D06991AD965D5599D7A811E310C" box="[270,382,1550,1573]" class="Anthozoa" family="Plexauridae" genus="Trimuricea" kingdom="Animalia" order="Alcyonacea" pageId="10" pageNumber="11" phylum="Cnidaria" rank="species" species="inermis">
<emphasis id="B953EA97991AD965D5599D7A811E310C" box="[270,382,1550,1573]" italics="true" pageId="10" pageNumber="11">T. inermis</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C274D06991AD965D5EB9D7A822F310C" box="[444,591,1550,1573]" class="Anthozoa" family="Plexauridae" genus="Trimuricea" kingdom="Animalia" order="Alcyonacea" pageId="10" pageNumber="11" phylum="Cnidaria" rank="species" species="omanensis" status="sp. nov.">
<emphasis id="B953EA97991AD965D5EB9D7A822F310C" box="[444,591,1550,1573]" italics="true" pageId="10" pageNumber="11">T. omanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26057EC991AD965D6019D7982F5310F" box="[598,661,1549,1574]" pageId="10" pageNumber="11" rank="species">
<emphasis id="B953EA97991AD965D6019D7982F5310F" bold="true" box="[598,661,1549,1574]" pageId="10" pageNumber="11">n. sp.</emphasis>
</taxonomicNameLabel>
, however, it differs in having many thornscales and triradiates in the polyp with very smooth, blunt, thorn and upper ray. In addition, these thornscales are slightly larger and have wider and more tuberculate base than those in
<taxonomicName id="4C274D06991AD965D60E9D22828A3144" box="[601,746,1622,1645]" class="Anthozoa" family="Plexauridae" genus="Trimuricea" kingdom="Animalia" order="Alcyonacea" pageId="10" pageNumber="11" phylum="Cnidaria" rank="species" species="omanensis" status="sp. nov.">
<emphasis id="B953EA97991AD965D60E9D22828A3144" box="[601,746,1622,1645]" italics="true" pageId="10" pageNumber="11">T. omanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26057EC991AD965D6A59D2183513147" box="[754,817,1621,1646]" pageId="10" pageNumber="11" rank="species">
<emphasis id="B953EA97991AD965D6A59D2183513147" bold="true" box="[754,817,1621,1646]" pageId="10" pageNumber="11">n. sp.</emphasis>
</taxonomicNameLabel>
and
<taxonomicName id="4C274D06991AD965D7309D2283B73144" box="[871,983,1622,1645]" class="Anthozoa" family="Plexauridae" genus="Trimuricea" kingdom="Animalia" order="Alcyonacea" pageId="10" pageNumber="11" phylum="Cnidaria" rank="species" species="inermis">
<emphasis id="B953EA97991AD965D7309D2283B73144" box="[871,983,1622,1645]" italics="true" pageId="10" pageNumber="11">T. inermis</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B983685991AD965D4909D0D855C31F3" blockId="10.[151,1436,577,1754]" pageId="10" pageNumber="11">
The specimens described by Grasshoff were later broken in parts and stored in both MNHN and SMF. There is an additional fragment of
<typeStatus id="549C8827991AD965D5979DEA8240319F" box="[448,544,1694,1718]" pageId="10" pageNumber="11" type="paratype">paratype</typeStatus>
stored in SMF (SMF 6754), which is not mentioned by
<bibRefCitation id="EFB64B74991AD965D0FF9DE985F8319F" author="Grasshoff" box="[1192,1432,1693,1718]" pageId="10" pageNumber="11" refString="Grasshoff, M. (1999) The shallow water gorgonians of New Caledonia and adjacent islands (Coelenterata: Octocorallia). Senckenbergiana biologica, 78 (1 / 2), 1 - 245." type="journal article" year="1999">Grasshoff (1999: 48)</bibRefCitation>
, and it could be a fragment of SMF 6753. Grasshoff did not mention the number of spindles in the collaret.
</paragraph>
</subSubSection>
</treatment>
</document>