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<document id="169A691EBB7423FBFD81A261C91EFC57" ID-CLB-Dataset="28431" ID-DOI="10.11646/zootaxa.4564.1.2" ID-GBIF-Dataset="d3a56aac-92aa-48f2-b9fe-9576a4c2a0ec" ID-ISSN="1175-5326" ID-Zenodo-Dep="2588621" ID-ZooBank="27F70B5F-24DD-4C9E-BBA7-2B87476BD49C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1552162070752" checkinUser="plazi" docAuthor="Gunter, Nicole L. &amp; Weir, Thomas A." docDate="2019" docId="03832713FF8B8D53A3DAFBD254419257" docLanguage="en" docName="zootaxa.4564.1.2.pdf" docOrigin="Zootaxa 4564 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Lepanus podocarp Gunter &amp; Weir 2019, new species" docType="treatment" docUuid="6B923B27-2007-4C2F-BB0C-1C28E2D2BB53" docUuidSource="ZooBank" docVersion="9" lastPageNumber="74" masterDocId="FFBA5F6BFFAB8D72A34DFFB65569910A" masterDocTitle="Revision of Australian species of the dung beetle genus Lepanus (Coleoptera: Scarabaeidae: Scarabaeinae): key to species groups and description of 14 new species from the L. pygmaeus species group" masterLastPageNumber="80" masterPageNumber="41" pageNumber="73" updateTime="1698700752634" updateUser="ExternalLinkService">
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<mods:title id="E65FF4618DEC82508A63619C3C76BF1F">Revision of Australian species of the dung beetle genus Lepanus (Coleoptera: Scarabaeidae: Scarabaeinae): key to species groups and description of 14 new species from the L. pygmaeus species group</mods:title>
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<mods:namePart id="9F6B0579578CF70845EDFC25627A35B5">Gunter, Nicole L.</mods:namePart>
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<mods:namePart id="BB3F05C7368988F1EB3D7F1962DA1225">Weir, Thomas A.</mods:namePart>
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<mods:title id="73072195ECB5CBE1EBDD9D6BB5D5F5BE">Zootaxa</mods:title>
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<mods:date id="B485B6C50D4EA3E193F3D2B26C5D9CE4">2019</mods:date>
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<mods:number id="C73B4779A83673E5080403697E92C55E">2019-03-05</mods:number>
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<mods:identifier id="B7241BFA19E3E750802E0EC6799A1A9A" type="CLB-Dataset">28431</mods:identifier>
<mods:identifier id="E39EEF6CCD2CA640788D94338DDAD5B9" type="DOI">10.11646/zootaxa.4564.1.2</mods:identifier>
<mods:identifier id="3B3CE7D6A3EB2EACE08CA6F020808D88" type="GBIF-Dataset">d3a56aac-92aa-48f2-b9fe-9576a4c2a0ec</mods:identifier>
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<treatment id="03832713FF8B8D53A3DAFBD254419257" ID-DOI="http://doi.org/10.5281/zenodo.5922828" ID-GBIF-Taxon="154366361" ID-Zenodo-Dep="5922828" LSID="urn:lsid:zoobank.org:act:6B923B27-2007-4C2F-BB0C-1C28E2D2BB53" httpUri="http://treatment.plazi.org/id/03832713FF8B8D53A3DAFBD254419257" lastPageId="33" lastPageNumber="74" pageId="32" pageNumber="73">
<subSubSection id="C330C58EFF8B8D52A3DAFBD257B39574" box="[151,730,1124,1150]" pageId="32" pageNumber="73" type="nomenclature">
<paragraph id="8B959605FF8B8D52A3DAFBD257B39574" blockId="32.[151,730,1124,1183]" box="[151,730,1124,1150]" pageId="32" pageNumber="73">
<heading id="D0DD2169FF8B8D52A3DAFBD257B39574" bold="true" box="[151,730,1124,1150]" fontSize="11" level="1" pageId="32" pageNumber="73" reason="1">
<emphasis id="B95E4A17FF8B8D52A3DAFBD257B39574" bold="true" box="[151,730,1124,1150]" pageId="32" pageNumber="73">
<taxonomicName id="4C2AED86FF8B8D52A3DAFBD257549574" ID-CoL="3T8VF" authority="Gunter &amp; Weir" authorityName="Gunter &amp; Weir" authorityYear="2019" box="[151,573,1124,1150]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="73" phylum="Arthropoda" rank="species" species="podocarp" status="sp. nov.">
<emphasis id="B95E4A17FF8B8D52A3DAFBD2541F9574" bold="true" box="[151,374,1124,1150]" italics="true" pageId="32" pageNumber="73">Lepanus podocarp</emphasis>
Gunter &amp; Weir
</taxonomicName>
,
<taxonomicNameLabel id="A26DF76CFF8B8D52A107FBD257B39574" box="[586,730,1124,1150]" pageId="32" pageNumber="73" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C330C58EFF8B8D52A3DAFB3054F79595" box="[151,414,1158,1183]" pageId="32" pageNumber="73" type="description">
<paragraph id="8B959605FF8B8D52A3DAFB3054F79595" blockId="32.[151,730,1124,1183]" box="[151,414,1158,1183]" pageId="32" pageNumber="73">
(
<figureCitation id="13118A80FF8B8D52A3D2FB3054699595" box="[159,256,1158,1183]" captionStart="FIGURE 6" captionStartId="16.[151,250,1948,1970]" captionTargetBox="[235,1333,202,1834]" captionTargetId="figure@16.[213,1373,193,1926]" captionTargetPageId="16" captionText="FIGURE 6. Habitus of species within the Lepanus pygmaeus species group. A, Lepanus lingziae; B, Lepanus mckenziei; C, Lepanus menendezae; D, Lepanus pecki; E: Lepanus podocarp; F, Lepanus pungalina. All specimens in comparative scale. Scale bar 1 mm." httpUri="https://zenodo.org/record/2588633/files/figure.png" pageId="32" pageNumber="73">Figs. 6E</figureCitation>
,
<figureCitation id="13118A80FF8B8D52A241FB3154479595" box="[268,302,1159,1183]" captionStart="FIGURE 9" captionStartId="19.[151,250,1201,1223]" captionTargetBox="[205,1310,206,1167]" captionTargetId="figure@19.[194,1392,193,1180]" captionTargetPageId="19" captionText="FIGURE 9. Pygidial configuration of species within the Lepanus pygmaeus species group. A, Lepanus pecki; B, Lepanus podocarp; C: Lepanus pungalina; D, Lepanus pygmaeus; E, Lepanus sauroni; F, Lepanus tozerensis. All specimens in comparative scale. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/2588639/files/figure.png" pageId="32" pageNumber="73">9B</figureCitation>
,
<figureCitation id="13118A80FF8B8D52A277FB3154059595" box="[314,364,1159,1183]" captionStart="FIGURE 12" captionStartId="25.[151,250,1905,1927]" captionTargetBox="[231,1341,208,1744]" captionTargetId="figure@25.[230,1357,193,1890]" captionTargetPageId="25" captionText="FIGURE 12. Left, dorsal, and right view of aedeagi of species within the Lepanus pygmaeus species group. A, Lepanus podocarp; B, Lepanus pungalina; C, Lepanus pygmaeus; D, Lepanus sauroni; E: Lepanus tozerensis. Not in comparative scale, given size difference aedegi are roughly grouped in three size classes as per lower left schematic and scaled accordingly. Size class indicated by shaded silhouette and 0.2 mm scale bar." httpUri="https://zenodo.org/record/2588645/files/figure.png" pageId="32" pageNumber="73">12A</figureCitation>
,
<figureCitation id="13118A80FF8B8D52A235FB3154FC9595" box="[376,405,1159,1183]" captionStart="FIGURE 14" captionStartId="31.[151,250,1220,1242]" captionTargetBox="[151,1436,375,1198]" captionTargetId="figure@31.[151,1436,375,1198]" captionTargetPageId="31" captionText="FIGURE 14. Map of known records for 10 species within the Lepanus pygmaeus species group overlaid with predicted distribution boundary of L. pygmaeus (dashed line) taken from Fig. 13 map." httpUri="https://zenodo.org/record/2588649/files/figure.png" pageId="32" pageNumber="73">14</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C330C58EFF8B8D52A3DAFB7857CB9491" pageId="32" pageNumber="73" type="materials_examined">
<paragraph id="8B959605FF8B8D52A3DAFB7857CB9491" blockId="32.[151,1437,1230,2011]" pageId="32" pageNumber="73">
<emphasis id="B95E4A17FF8B8D52A3DAFB7854F695ED" bold="true" box="[151,415,1230,1255]" pageId="32" pageNumber="73">
Type series.
<materialsCitation id="3B429C58FF8B8D52A264FB7854F695ED" ID-GBIF-Occurrence="2012745311" box="[297,415,1230,1255]" collectingDate="1993-12-15" collectingDateMax="1993-12-23" collectingDateMin="1993-12-15" collectionCode="AUST, NATURE, ANIC" collectorName="S. B. &amp; J. Peck" county="Arnhemland" latitude="-12.6663885" location="Rainforest" longLatPrecision="21" longitude="133.15833" municipality="Podocarp Canyon" pageId="32" pageNumber="73" specimenCode="CMNEN-00030225, CMNEN-00030226, CMNEN-00030230, ANIC 2, CMNEN- 00030227-00030229, CMNC 1" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="Northern Territory" typeStatus="holotype">
<typeStatus id="549128A7FF8B8D52A264FB7854F395ED" box="[297,410,1230,1255]" pageId="32" pageNumber="73" type="holotype">Holotype</typeStatus>
:
</materialsCitation>
</emphasis>
<specimenCount id="9D2C5D8CFF8B8D52A2EBFB7954D295EC" box="[422,443,1231,1254]" pageId="32" pageNumber="73" type="male">♂</specimenCount>
“AUST: NT;
<collectingCounty id="62F4EE89FF8B8D52A115FB78578D95ED" box="[600,740,1230,1255]" pageId="32" pageNumber="73">Arnhemland</collectingCounty>
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<quantity id="4CD23BE0FF8B8D52A1A6FB78564295ED" box="[747,811,1230,1255]" metricMagnitude="4" metricUnit="m" metricValue="3.3" pageId="32" pageNumber="73" unit="km" value="33.0">33km</quantity>
E Jabiru
</locationDeviation>
,
<collectingMunicipality id="6BF10C7FFF8B8D52A0D1FB78510E95ED" box="[924,1127,1230,1255]" pageId="32" pageNumber="73">Podocarp Canyon</collectingMunicipality>
,
<location id="8EF5C0DEFF8B8D52A73FFB78518395ED" LSID="urn:lsid:plazi:treatment:03832713FF8B8D53A3DAFBD254419257:8EF5C0DEFF8B8D52A73FFB78518395ED" box="[1138,1258,1230,1255]" county="Arnhemland" latitude="-12.6663885" longLatPrecision="21" longitude="133.15833" municipality="Podocarp Canyon" name="Rainforest" pageId="32" pageNumber="73" stateProvince="Northern Territory">Rainforest</location>
,
<date id="FF94B0C5FF8B8D52A7B8FB7850FE95ED" box="[1269,1431,1230,1255]" pageId="32" pageNumber="73" value="1993-12-15" valueMax="1993-12-23" valueMin="1993-12-15">
<collectingDate id="EFD0492DFF8B8D52A7B8FB7850FE95ED" box="[1269,1431,1230,1255]" pageId="32" pageNumber="73" value="1993-12-15" valueMax="1993-12-23" valueMin="1993-12-15">15-23. XII. 93</collectingDate>
</date>
, 5 FITs, S&amp;J Peck 93-130” [
<geoCoordinate id="EE1EF0C2FF8B8D52A2AEFB44570C9401" box="[483,613,1266,1291]" direction="south" orientation="latitude" pageId="32" pageNumber="73" precision="15" value="-12.6663885">12°39'59''S</geoCoordinate>
,
<geoCoordinate id="EE1EF0C2FF8B8D52A138FB45579E9401" box="[629,759,1267,1291]" direction="east" orientation="longitude" pageId="32" pageNumber="73" precision="15" value="133.15833">133°9'30''E</geoCoordinate>
] /
<location id="8EF5C0DEFF8B8D52A050FB44569C9400" LSID="urn:lsid:plazi:treatment:03832713FF8B8D53A3DAFBD254419257:8EF5C0DEFF8B8D52A050FB44569C9400" box="[797,1013,1266,1291]" county="Arnhemland" latitude="-12.6663885" longLatPrecision="21" longitude="133.15833" municipality="Podocarp Canyon" name="Canadian Museum" pageId="32" pageNumber="73" stateProvince="Northern Territory">Canadian Museum</location>
of NATURE
<specimenCode id="DB8C3E7EFF8B8D52A7D3FB4450179401" box="[1182,1406,1266,1291]" collectionCode="CMNEN-" pageId="32" pageNumber="73">CMNEN-00030225</specimenCode>
” / “ANIC Database No. 25-065550” / “
<taxonomicName id="4C2AED86FF8B8D52A105FAA157C09425" authorityName=", Matthews" authorityYear="1974" box="[584,681,1303,1327]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="79" phylum="Arthropoda" rank="genus">Lepanus</taxonomicName>
sp.1” / “
<taxonomicName id="4C2AED86FF8B8D52A05AFAA156119425" authorityName=", Matthews" authorityYear="1974" box="[791,888,1303,1327]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="79" phylum="Arthropoda" rank="genus">Lepanus</taxonomicName>
NT7 det.
<bibRefCitation id="EFBBEBF4FF8B8D52A0BAFAA051CD9425" author="Gunter, N. L. &amp; Monteith, G. B. &amp; Cameron, S. L. &amp; Weir, T." box="[1015,1188,1302,1327]" pageId="32" pageNumber="73" pagination="1 - 27" refId="ref29280" refString="Gunter, N. L., Monteith, G. B., Cameron, S. L. &amp; Weir, T. A (2018) Evidence from Australian mesic zone dung beetles support Gondwanan vicariance and Mesozoic origin of the Scarabaeinae (Coleoptera: Scarabaeidae). Invertebrate Systematics and Evolution, 1 - 27. [Advance online publication] https: // doi. org / 10.1163 / 1876312 X- 00002171" type="book chapter" year="2018">T.A.Weir 2018</bibRefCitation>
” / “ANIC Image” in ANIC.
<emphasis id="B95E4A17FF8B8D52A3BBFA8C57EE9459" bold="true" box="[246,647,1338,1363]" pageId="32" pageNumber="73">
<typeStatus id="549128A7FF8B8D52A3BBFA8C541B9459" box="[246,370,1338,1363]" pageId="32" pageNumber="73" type="paratype">Paratypes</typeStatus>
:
<collectingRegion id="49EE58E7FF8B8D52A2C6FA8C57EB9459" box="[395,642,1338,1363]" country="Australia" name="Northern Territory" pageId="32" pageNumber="73">Northern Territory</collectingRegion>
:
</emphasis>
<location id="8EF5C0DEFF8B8D52A1D6FA8C564E9459" LSID="urn:lsid:plazi:treatment:03832713FF8B8D53A3DAFBD254419257:8EF5C0DEFF8B8D52A1D6FA8C564E9459" box="[667,807,1338,1363]" county="Arnhemland" latitude="-12.6663885" longLatPrecision="21" longitude="133.15833" municipality="Podocarp Canyon" name="Arnhemland" pageId="32" pageNumber="73" stateProvince="Northern Territory">Arnhemland</location>
<locationDeviation id="9A278F27FF8B8D52A076FA8D51679459" box="[827,1038,1338,1363]" location="Jabiru" pageId="32" pageNumber="73">
<quantity id="4CD23BE0FF8B8D52A076FA8D56E79459" box="[827,910,1338,1363]" metricMagnitude="4" metricUnit="m" metricValue="3.3" pageId="32" pageNumber="73" unit="km" value="33.0">33 km</quantity>
E Jabiru
</locationDeviation>
, podocarp canyon, [
<geoCoordinate id="EE1EF0C2FF8B8D52A658FA8C50FE9459" box="[1301,1431,1338,1363]" direction="south" orientation="latitude" pageId="32" pageNumber="73" precision="15" value="-12.6663885">12°39'59''S</geoCoordinate>
,
<geoCoordinate id="EE1EF0C2FF8B8D52A3DAFAE95472947D" box="[151,283,1375,1399]" direction="east" orientation="longitude" pageId="32" pageNumber="73" precision="15" value="133.15833">133°9'30''E</geoCoordinate>
],
<collectingDate id="EFD0492DFF8B8D52A263FAE854B3947D" box="[302,474,1374,1399]" pageId="32" pageNumber="73" value="1993-12-15" valueMax="1993-12-23" valueMin="1993-12-15">
<date id="FF94B0C5FF8B8D52A263FAE854B3947D" box="[302,474,1374,1399]" pageId="32" pageNumber="73" value="1993-12-15" valueMax="1993-12-23" valueMin="1993-12-15">15–23.xii.1993</date>
</collectingDate>
,
<collectorName id="26DFF3D3FF8B8D52A2AAFAE957E4947D" box="[487,653,1374,1399]" pageId="32" pageNumber="73">S.B. &amp; J. Peck</collectorName>
(
<specimenCode id="DB8C3E7EFF8B8D52A1D1FAE856E9947D" box="[668,896,1374,1399]" collectionCode="CMNEN-" pageId="32" pageNumber="73">CMNEN-00030226</specimenCode>
,
<specimenCode id="DB8C3E7EFF8B8D52A0C1FAE95102947D" box="[908,1131,1375,1399]" collectionCode="CMNEN-" pageId="32" pageNumber="73">CMNEN-00030230</specimenCode>
in
<specimenCode id="DB8C3E7EFF8B8D52A7DDFAE95183947D" box="[1168,1258,1375,1399]" collectionCode="ANIC" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="32" pageNumber="73">ANIC 2</specimenCode>
×
<specimenCount id="9D2C5D8CFF8B8D52A647FAE95076947C" box="[1290,1311,1375,1398]" pageId="32" pageNumber="73" type="female">♀</specimenCount>
,
<specimenCode id="DB8C3E7EFF8B8D52A663FAE954EC9491" collectionCode="CMNEN-" pageId="32" pageNumber="73">CMNEN- 00030227–00030229</specimenCode>
in
<specimenCode id="DB8C3E7EFF8B8D52A2E4FA3557799491" box="[425,528,1411,1435]" collectionCode="CMNC" pageId="32" pageNumber="73">CMNC 1</specimenCode>
×
<specimenCount id="9D2C5D8CFF8B8D52A163FA35572C9490" box="[558,581,1411,1434]" pageId="32" pageNumber="73" type="male">♂</specimenCount>
, 2 ×
<specimenCount id="9D2C5D8CFF8B8D52A130FA3557F89491" box="[637,657,1411,1435]" pageId="32" pageNumber="73" type="female">♀</specimenCount>
).
</paragraph>
</subSubSection>
<subSubSection id="C330C58EFF8B8D53A38AFA105596906F" lastPageId="33" lastPageNumber="74" pageId="32" pageNumber="73" type="description">
<paragraph id="8B959605FF8B8D52A38AFA10502094B5" blockId="32.[151,1437,1230,2011]" box="[199,1353,1446,1471]" pageId="32" pageNumber="73">
<emphasis id="B95E4A17FF8B8D52A38AFA10543094B5" bold="true" box="[199,345,1446,1471]" pageId="32" pageNumber="73">Description.</emphasis>
Either uniformly dark brown or with red patches on elytral humeri. Antennal clubs black.
</paragraph>
<paragraph id="8B959605FF8B8D52A38AFA7C519B94E8" blockId="32.[151,1437,1230,2011]" box="[199,1266,1482,1507]" pageId="32" pageNumber="73">
Total length:
<quantity id="4CD23BE0FF8B8D52A211FA7C548F94E8" box="[348,486,1482,1507]" metricMagnitude="-3" metricUnit="m" metricValue="2.45" metricValueMax="2.6" metricValueMin="2.3" pageId="32" pageNumber="73" unit="mm" value="2.45" valueMax="2.6" valueMin="2.3">2.3–2.6 mm</quantity>
.
<typeStatus id="549128A7FF8B8D52A2BDFA7C573194E9" box="[496,600,1482,1507]" pageId="32" pageNumber="73" type="holotype">Holotype</typeStatus>
measurements: Total length
<quantity id="4CD23BE0FF8B8D52A0D1FA7D569D94E8" box="[924,1012,1483,1507]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="32" pageNumber="73" unit="mm" value="2.3">2.3 mm</quantity>
, elytral width
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.
</paragraph>
<paragraph id="8B959605FF8B8D52A38AFA5851A09779" blockId="32.[151,1437,1230,2011]" pageId="32" pageNumber="73">Male: Head: U-shaped between clypeal teeth, which are short and upturned. Margin slightly angulate at clypeogenal suture. Genal angle weakly defined. Basal carina present and narrowly interrupted. Punctation on the clypeus is finer than on rest of head. Dorsal part of eyes moderate, separated by interocular space approximately 15 times eye width (37: 2.5); eye canthus not dividing the eye. Head 1.2 × wider than long (45:37).</paragraph>
<paragraph id="8B959605FF8B8D52A38AF9C8567E97B1" blockId="32.[151,1437,1230,2011]" pageId="32" pageNumber="73">Prothorax: Hypomeral surface very finely reticulate without punctation. Hypomeral striae at least half length of hypomeron. Pronotum 1.65 × wider than long (74:45).</paragraph>
<paragraph id="8B959605FF8B8D52A38AF97050F39609" blockId="32.[151,1437,1230,2011]" pageId="32" pageNumber="73">Elytra: Surface smooth and striae slightly more impressed towards the apex. Striae 1–5 double with very fine punctate edges, striae 6–7 singular; elytral flange and groove relatively wide; Elytra 1.2 × wider than long (90:74).</paragraph>
<paragraph id="8B959605FF8B8D52A38AF8B9559A9665" blockId="32.[151,1437,1230,2011]" pageId="32" pageNumber="73">Legs: Front edge of protibiae deeply angulate; apical digit somewhat elongate and sharp; 2 teeth on outer edge. Inner edge of metatibiae almost straight. Metatarsi with 2nd tarsomere subequal to the 5th. Tarsal claws small and toothed.</paragraph>
<paragraph id="8B959605FF8B8D52A38AF8CC51F396D0" blockId="32.[151,1437,1230,2011]" pageId="32" pageNumber="73">Abdomen: Pygidium with wide pear shaped depression containing a small, slightly elongate tubercle in upper third. Depression occupies at least half size of disc and just extending under the slightly angulate upper edge of pygidium. Abdominal surface very finely reticulate right across. Segment 6 finely punctate.</paragraph>
<paragraph id="8B959605FF8A8D53A38AFF21575B9017" blockId="33.[151,1437,151,861]" pageId="33" pageNumber="74">Pterothorax: Medial lobe of metaventrite finely punctate and broadly margined between mesocoxae. Lateral lobe of metaventrite smooth and punctate with setae. Meso-metaventral suture very slightly arched. Mesoventrite with slight notch anteriorly. Mesoventrite smooth, with discernible punctures at least at sides. Mesepimeron smooth. Metanepisternum reticulate.</paragraph>
<paragraph id="8B959605FF8A8D53A38AFE915596906F" blockId="33.[151,1437,151,861]" pageId="33" pageNumber="74">
Female: As per the
<taxonomicName id="4C2AED86FF8A8D53A2F8FE9F572E904A" authorityName="in Matthews" authorityYear="1974" box="[437,583,296,320]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="74" phylum="Arthropoda" rank="species" species="pygmaeus">
<emphasis id="B95E4A17FF8A8D53A2F8FE9F54AD904A" box="[437,452,297,320]" italics="true" pageId="33" pageNumber="74">L</emphasis>
.
<emphasis id="B95E4A17FF8A8D53A29AFE9F572E904A" box="[471,583,297,320]" italics="true" pageId="33" pageNumber="74">pygmaeus</emphasis>
</taxonomicName>
species group but basal margin of abdominal ventrite 6 slightly convex medially.
</paragraph>
</subSubSection>
<subSubSection id="C330C58EFF8A8D53A38AFED951F89082" box="[199,1169,367,392]" pageId="33" pageNumber="74" type="etymology">
<paragraph id="8B959605FF8A8D53A38AFED951F89082" blockId="33.[151,1437,151,861]" box="[199,1169,367,392]" pageId="33" pageNumber="74">
<emphasis id="B95E4A17FF8A8D53A38AFED954279082" bold="true" box="[199,334,367,392]" pageId="33" pageNumber="74">Etymology.</emphasis>
Named for the only known locality, Podocarp Canyon, West Arnhemland.
</paragraph>
</subSubSection>
<subSubSection id="C330C58EFF8A8D53A38AFE22500390A6" box="[199,1386,404,429]" pageId="33" pageNumber="74" type="distribution">
<paragraph id="8B959605FF8A8D53A38AFE22500390A6" blockId="33.[151,1437,151,861]" box="[199,1386,404,429]" pageId="33" pageNumber="74">
<emphasis id="B95E4A17FF8A8D53A38AFE22540990A7" bold="true" box="[199,352,404,429]" pageId="33" pageNumber="74">Distribution.</emphasis>
Known only from Podocarp Canyon, West Arnhemland, in the ARP IBRA region (
<figureCitation id="13118A80FF8A8D53A644FE22503590A6" box="[1289,1372,404,429]" captionStart="FIGURE 14" captionStartId="31.[151,250,1220,1242]" captionTargetBox="[151,1436,375,1198]" captionTargetId="figure@31.[151,1436,375,1198]" captionTargetPageId="31" captionText="FIGURE 14. Map of known records for 10 species within the Lepanus pygmaeus species group overlaid with predicted distribution boundary of L. pygmaeus (dashed line) taken from Fig. 13 map." httpUri="https://zenodo.org/record/2588649/files/figure.png" pageId="33" pageNumber="74">Fig. 14</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C330C58EFF8A8D53A38AFE0154419257" pageId="33" pageNumber="74" type="discussion">
<paragraph id="8B959605FF8A8D53A38AFE0151FA93A2" blockId="33.[151,1437,151,861]" pageId="33" pageNumber="74">
<emphasis id="B95E4A17FF8A8D53A38AFE01542690DA" bold="true" box="[199,335,439,464]" pageId="33" pageNumber="74">Comments.</emphasis>
This species was originally reported as
<taxonomicName id="4C2AED86FF8A8D53A05AFE0F561F90DA" authorityName=", Matthews" authorityYear="1974" box="[791,886,441,464]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="74" phylum="Arthropoda" rank="genus">
<emphasis id="B95E4A17FF8A8D53A05AFE0F561F90DA" box="[791,886,441,464]" italics="true" pageId="33" pageNumber="74">Lepanus</emphasis>
</taxonomicName>
sp.
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<bibRefCitation id="EFBBEBF4FF8A8D53A097FE01510D90DA" author="Peck, S. B." box="[986,1124,439,464]" pageId="33" pageNumber="74" pagination="181 - 192" refId="ref30028" refString="Peck, S. B. (2002) Australian rainforest biogeography: is there a relictual beetle fauna in an Allosyncarpia rainforest refugium, Arnhemland, Northern Territory? Memoirs of the Queensland Museum, 48, 181 - 192." type="journal article" year="2002">Peck (2002)</bibRefCitation>
that documented the beetle fauna of this canyon.
<bibRefCitation id="EFBBEBF4FF8A8D53A2EFFE6A575890FE" author="Peck, S. B." box="[418,561,476,501]" pageId="33" pageNumber="74" pagination="181 - 192" refId="ref30028" refString="Peck, S. B. (2002) Australian rainforest biogeography: is there a relictual beetle fauna in an Allosyncarpia rainforest refugium, Arnhemland, Northern Territory? Memoirs of the Queensland Museum, 48, 181 - 192." type="journal article" year="2002">Peck (2002)</bibRefCitation>
lists erroneous coordinates 12°87'73&quot;S, 133°26'73&quot;E in this study, if the coordinates instead represent
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,
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they would place the site ~
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SE of Jabiru, close to but not in forested area as visualised though satellite images. The coordinates
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,
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more broadly match the description of the locality ~
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from Jabiru on the labels. Digital distributional records for
<taxonomicName id="4C2AED86FF8A8D53A3DAFDDA5477938F" box="[151,286,620,645]" class="Pinopsida" family="Podocarpaceae" genus="Podocarpus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="33" pageNumber="74" phylum="Tracheophyta" rank="genus">
<emphasis id="B95E4A17FF8A8D53A3DAFDDA5477938F" box="[151,286,620,645]" italics="true" pageId="33" pageNumber="74">Podocarpus</emphasis>
</taxonomicName>
in the Australian
<bibRefCitation id="EFBBEBF4FF8A8D53A2A9FDDA5101938E" author="Australasian Virtual Herbarium &amp; AVH" box="[484,1128,620,645]" pageId="33" pageNumber="74" refId="ref28857" refString="Australasian Virtual Herbarium [AVH] (2018) Genus Podocarpus, Northern Territory. Available from: https: // avh. ala. org. au / occurrences / search? q = lsid: http: // id. biodiversity. org. au / node / apni / 8905690 &amp; fq = cl 927: % 22 Northern % 20 Territory % 20 (including % 20 Coastal % 20 Waters) % 22 (accessed 15 October 2018)" type="url" year="2018">Virtual Herbarium (Australasian Virtual Herbarium 2018)</bibRefCitation>
indicate 25 records in West Arnhem to the east of Jabiru, however no records are congruent with the 1993 field survey.
</paragraph>
<paragraph id="8B959605FF8A8D53A38AFD0254419257" blockId="33.[151,1437,151,861]" pageId="33" pageNumber="74">
<bibRefCitation id="EFBBEBF4FF8A8D53A38AFD02543993C6" author="Peck, S. B." box="[199,336,692,717]" pageId="33" pageNumber="74" pagination="181 - 192" refId="ref30028" refString="Peck, S. B. (2002) Australian rainforest biogeography: is there a relictual beetle fauna in an Allosyncarpia rainforest refugium, Arnhemland, Northern Territory? Memoirs of the Queensland Museum, 48, 181 - 192." type="journal article" year="2002">Peck (2002)</bibRefCitation>
discussed and rejected the likelihood of this rainforested canyon being an invertebrate refugium. This species is distinct from all others known from the region and has only been collected on one occasion by flight intercept trap.
<taxonomicName id="4C2AED86FF8A8D53A209FD4B5772921F" authorityName="Gunter &amp; Weir" authorityYear="2019" box="[324,539,764,789]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="74" phylum="Arthropoda" rank="species" species="podocarp">
<emphasis id="B95E4A17FF8A8D53A209FD4B5772921F" box="[324,539,764,789]" italics="true" pageId="33" pageNumber="74">Lepanus podocarp</emphasis>
</taxonomicName>
was unknown at the time when the informal nomenclature for undescribed species of
<taxonomicName id="4C2AED86FF8A8D53A240FC9754059232" authorityName=", Matthews" authorityYear="1974" box="[269,364,801,824]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="74" phylum="Arthropoda" rank="genus">
<emphasis id="B95E4A17FF8A8D53A240FC9754059232" box="[269,364,801,824]" italics="true" pageId="33" pageNumber="74">Lepanus</emphasis>
</taxonomicName>
was reported (
<bibRefCitation id="EFBBEBF4FF8A8D53A15CFC9657B59232" author="Yeates, D. &amp; Gunter, N. L. &amp; Weir, T. &amp; Cameron, S. L." box="[529,732,799,824]" pageId="33" pageNumber="74" pagination="92 - 99" refId="ref30201" refString="Yeates, D. K, Gunter, N. L., Weir, T. &amp; Cameron, S. L. (2011) Developing a benchmark invertebrate group for biodiversity modelling. In: West, J. (Ed.), 21 st Century Taxonomy: accelerating research and discovery of Australia's biodiversity, Taxonomy Research and Identification Network Final Report- March, 2011, pp. 92 - 99. Available from: https: // www. environment. gov. au / system / files / pages / 56 c 08 f 6 f- 7 bf 7 - 4594 - 8175 - b 4 c 143110156 / files / hub-taxonomy. pdf (accessed 12 December 2018)" type="journal article" year="2011">
Yeates
<emphasis id="B95E4A17FF8A8D53A129FC9757F59232" box="[612,668,799,824]" italics="true" pageId="33" pageNumber="74">et al.</emphasis>
2011
</bibRefCitation>
) but was designated the informal name “
<taxonomicName id="4C2AED86FF8A8D53A7EAFC97506F9232" authorityName=", Matthews" authorityYear="1974" box="[1191,1286,801,824]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="74" phylum="Arthropoda" rank="genus">
<emphasis id="B95E4A17FF8A8D53A7EAFC97506F9232" box="[1191,1286,801,824]" italics="true" pageId="33" pageNumber="74">Lepanus</emphasis>
</taxonomicName>
NT7” within this revision.
</paragraph>
</subSubSection>
</treatment>
</document>
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