213 lines
19 KiB
XML
213 lines
19 KiB
XML
<document id="6369A65DD50BE81D5C80AAFB54621C99" ENCODING="UTF-8" ID-GBIF-Dataset="68b0d7de-1651-4945-8576-b7328a26175b" ModsDocID="z00249p001" checkinTime="1246893442886" checkinUser="thomas" docAuthor="Jonathan W. Armbruster" docDate="2003" docId="43B23634594A5946DD4890010756F37F" docLanguage="en" docName="2003_Armbruster_2_gg1.xml" docOrigin="Zootaxa 249" docSource="http://www.zoobank.org/urn:lsid:zoobank.org:pub:DDFAA9D6-E4FA-4C3C-9749-CF0313D30F3C" docTitle="Hypostomus ericius Armbruster, 2003, new species" docType="treatment" docUuid="C7C2629A-1686-4E4E-9E15-59B1D1162CBE" docUuidSource="ZooBank" docVersion="14" lastPageNumber="28" masterDocId="A29EE709B7587AF1AD73A093A03CF987" masterDocTitle="The species of the Hypostomus cochliodon group (Siluriformes: Loricariidae)." masterLastPageNumber="60" masterPageNumber="1" pageNumber="25" updateTime="1701313868015" updateUser="plazi">
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<mods:titleInfo id="E472643718FD2B2FCB361490EED62D35">
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<mods:title id="78640133C1DCA9FFF6E5B73A03D2A12C">The species of the Hypostomus cochliodon group (Siluriformes: Loricariidae).</mods:title>
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</mods:titleInfo>
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<mods:name id="F08DB5607D97003AD8E192CC6444B917" type="personal">
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<mods:roleTerm id="0637EEC4AD1B100995D7D6EBF477179F">Author</mods:roleTerm>
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<mods:namePart id="8B975FB32BF40FAC551DA69783AE46B0">Jonathan W. Armbruster</mods:namePart>
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<mods:typeOfResource id="F345E3BFD1DB0FB2AC9125239F29D58F">text</mods:typeOfResource>
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<mods:titleInfo id="63C5F0661532A1C0113C1CDBA24A11AB">
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<mods:title id="4D6797329F8E36C5528679AF70482DB7">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="30E65D7B081F0A2AED8641D9582B8571">
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<mods:date id="8D1932ECCB687996686489687280B294">2003</mods:date>
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<mods:detail id="78E4F31DA681121E61B26B65368B66E2" type="volume">
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<mods:number id="7CC3B9B9CB8677D5587DFF075983BBFD">249</mods:number>
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<mods:start id="CA2CD25D95498565F54885C792A1FFE9">1</mods:start>
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<mods:end id="E24CD43C01768D19C7081734F5949BA1">60</mods:end>
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<mods:location id="17F9CF45B4ADDFEF2D1EF691F43101D7">
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<mods:url id="E2ED46A4F251D064F11C090CE2EEDCB0">http://www.zoobank.org/urn:lsid:zoobank.org:pub:DDFAA9D6-E4FA-4C3C-9749-CF0313D30F3C</mods:url>
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<mods:classification id="D801DE125AA2427B86A19B549BA51AB6">journal article</mods:classification>
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<mods:identifier id="DBD16BB5C82CBA2A8BEAD68886846A0E" type="Plazi-Custom">z00249p001</mods:identifier>
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<mods:identifier id="AF2E5C48FA8FC34DBADC36D1A8F69C6C" type="ZooBank">DDFAA9D6-E4FA-4C3C-9749-CF0313D30F3C</mods:identifier>
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<treatment id="43B23634594A5946DD4890010756F37F" ID-DOI="http://doi.org/10.5281/zenodo.6274705" ID-GBIF-Taxon="100125131" ID-Zenodo-Dep="6274705" LSID="urn:lsid:zoobank.org:act:C7C2629A-1686-4E4E-9E15-59B1D1162CBE" httpUri="http://treatment.plazi.org/id/43B23634594A5946DD4890010756F37F" lastPageNumber="28" pageNumber="25">
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<subSubSection id="7708E7C352C33379B4A576BA8AF831E6" type="nomenclature">
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<paragraph id="F00B1D7BC2C1597BC4240E962098D939" pageNumber="25">
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<taxonomicName id="6BE49F1C2FAAE70E1AEE190A28D81E04" ID-CoL="3NZ7J" LSID-ZBK="urn:lsid:zoobank.org:act:C7C2629A-1686-4E4E-9E15-59B1D1162CBE" family="Loricariidae" genus="Hypostomus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Hypostomus ericius Armbruster 2003:25" order="Siluriformes" pageNumber="25" phylum="Chordata" rank="species" species="ericius">Hypostomus ericius</taxonomicName>
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<taxonomicNameLabel id="1B431BF7F56AB76CA9CE908B7F4E1ABE" rank="species">new species</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="41BE739CBD09440F4505C5FCC9B2CD6E" type="description">
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<paragraph id="B65EBE4943AAFB7407C8F25D1EE109D8" pageNumber="25">(Fig. 9)</paragraph>
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</subSubSection>
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<subSubSection id="D6478E1911DE18E9CA5CD5BEC6D08924" type="nomenclature">
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<paragraph id="BA8B21FE046D465B9BD83C7D145F8015" pageNumber="25">
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<materialsCitation id="D8A4C7A297E98C5C1CA9E610AA64716C" ID-GBIF-Occurrence="1058481063" country="PERU" stateProvince="Amazonas">
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<typeStatus id="7F6CCEB08F6B5F201984338F3F79935F">Holotype</typeStatus>
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:
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<collectingCountry id="EFE3C65C0193FE7C02C37272F704E654">PERU</collectingCountry>
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.
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<collectingRegion id="8C191D7673A97F63D699BAE8CAE86C56">Amazonas</collectingRegion>
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||
:
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||
<normalizedToken id="C21C8379C77A912B708D83BB9A1064BD" originalValue="Río">Rio</normalizedToken>
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<normalizedToken id="C0E7C81AA5087B07760C76243AA0CCB6" originalValue="Marañon">Maranon</normalizedToken>
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dr. 1 km upstream from Caterpiza,
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<collectionCode id="66A2E9454DCCE2D6469DC19D93038A97" collectionName="Peru, Lima, Universidad Nacional Mayor de San Marcos, Museo de Historia Natural">MUSM</collectionCode>
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||
17441, 1, (186.8 mm SL).
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</materialsCitation>
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</paragraph>
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<paragraph id="650BEFC447EC2FC0BDD919EFD6898CDF" pageNumber="25">
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<materialsCitation id="28A9FCC8C59909E1B79F1389A3BB9922" ID-GBIF-Occurrence="1058481036" country="PERU" stateProvince="Amazonas">
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<typeStatus id="E1C689560ADEE53F1FC1239E6B0DFEAF">Paratypes</typeStatus>
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:
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<collectingCountry id="BA9869A43F49119EDBD56B68B5262999">PERU</collectingCountry>
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.
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<collectingRegion id="C325782C0226518AD4F91F818179F6FD">Amazonas</collectingRegion>
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||
:
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<normalizedToken id="A33E18B0541929302FB84926976583A0" originalValue="Río">Rio</normalizedToken>
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<normalizedToken id="3B50023AF55BCA1AF4A92D361A32B6AC" originalValue="Marañon">Maranon</normalizedToken>
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dr. 1 km upstream from Caterpiza,
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<collectionCode id="55FC91FB991695FC8F8138F1916058F7" collectionName="Peru, Lima, Universidad Nacional Mayor de San Marcos, Museo de Historia Natural">MUSM</collectionCode>
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17441, 1, (186.8)
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||
</materialsCitation>
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.
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<materialsCitation id="C91C26E3FECE868AFC1CD474193C4D0A" ID-GBIF-Occurrence="1058481015" country="Peru" stateProvince="Amazonas">
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||
<normalizedToken id="A5CF1770C56AB911CB7CF6BEF97EAD4F" originalValue="Río">Rio</normalizedToken>
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<normalizedToken id="FBDB7A62AA6C0467F8136AA2787A3038" originalValue="Marañon">Maranon</normalizedToken>
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dr. 200 m downstream from Caterpiza,
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<collectionCode id="659B780F60DFBA3E2444289F3D1773A3" collectionName="USA, California, Los Angeles, Los Angeles County Museum of Natural History">LACM</collectionCode>
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42004-9, 1, (148.3)
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||
</materialsCitation>
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||
.
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<materialsCitation id="DF1CE5FCD57912130AE893EB41D7D9CA" ID-GBIF-Occurrence="1058481075" country="Peru" stateProvince="Amazonas">
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||
<normalizedToken id="AA0EC93B8ED35819BE441D664FE7D7CB" originalValue="Río">Rio</normalizedToken>
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||
<normalizedToken id="FF93961B13A4CEC4995B75C9DDF311F8" originalValue="Marañon">Maranon</normalizedToken>
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||
dr. Caterpiza,
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<collectionCode id="C3F8EED85238E60D5A6A83A69BE1C5F0" collectionName="USA, California, Los Angeles, Los Angeles County Museum of Natural History">LACM</collectionCode>
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41893-6, 1, (80.5)
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||
</materialsCitation>
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<materialsCitation id="9F3FBE3F2CF5F601B8CF34C1445098DF" ID-GBIF-Occurrence="1058481037" country="Peru" stateProvince="Amazonas">
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and
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<collectionCode id="C5696F66AF984EF5DB3F8F8FFDAC13DC" collectionName="USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]">USNM</collectionCode>
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357392, 1, (93.6)
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||
</materialsCitation>
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.
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<materialsCitation id="04DB629C87F19632B1C79F59711F94D9" ID-GBIF-Occurrence="1058481073" country="Peru" stateProvince="Amazonas">
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||
Quebrada Pastazilla,
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<normalizedToken id="EA61CAB59E1E269DA149CFA839EF8894" originalValue="Río">Rio</normalizedToken>
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||
Santiago -
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<normalizedToken id="719470B261C8636CCE71581DECF21136" originalValue="Río">Rio</normalizedToken>
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||
<normalizedToken id="BEEB97D647E4268D0D187BED50578F99" originalValue="Marañon">Maranon</normalizedToken>
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dr.
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<collectionCode id="B32647401C916AEA445C66DF4D3619DA" collectionName="USA, California, Los Angeles, Los Angeles County Museum of Natural History">LACM</collectionCode>
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39947-7, 1, (120.9)
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||
</materialsCitation>
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.
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<materialsCitation id="D011BB0C0FACCF901A2412BFCF211C79" ID-GBIF-Occurrence="1058481082" country="Peru" stateProvince="Loreto">
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||
<collectingRegion id="375F9FCB78FCCBCA4400CAB65F79968B">Loreto</collectingRegion>
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:
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<normalizedToken id="C87152EED54145EE517B0134D32B7382" originalValue="Río">Rio</normalizedToken>
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Amazonas dr. Iquitos,
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<collectionCode id="1A6188749FA9B80E6361A2CF2B8AC0F2" collectionName="USA, New York, New York, American Museum of Natural History">AMNH</collectionCode>
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218035, 1, (85.1)
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||
</materialsCitation>
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.
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<materialsCitation id="3A49DEA73D31E3DE465A6D70E4205D85" ID-GBIF-Occurrence="1058481046" country="Peru" stateProvince="Loreto">
|
||
<normalizedToken id="EB44047C6F756B211D8C2553A773BB28" originalValue="Río">Rio</normalizedToken>
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||
Itaya,
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<normalizedToken id="087D742AA27CC35C294E5826985CA324" originalValue="Río">Rio</normalizedToken>
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||
Amazonas dr. approx. 10 km S Santa Clara,
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<collectionCode id="9C3CA18A925D87ABF1F86F500AD89D2D" collectionName="USA, Illinois, Champaign, Illinois Natural History Survey">INHS</collectionCode>
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36876, 1, (80.1)
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||
</materialsCitation>
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||
.
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<materialsCitation id="B033DF13D731335B33E0364B76F7F315" ID-GBIF-Occurrence="1058481087" country="Peru" stateProvince="Loreto">
|
||
<normalizedToken id="F54365FF47B609901ED76D7C0A4F6CE0" originalValue="Río">Rio</normalizedToken>
|
||
Itaya,
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<normalizedToken id="4F02AC5EAF4E737B9AF75DD565E8E48F" originalValue="Río">Rio</normalizedToken>
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||
Amazonas dr. upriver of
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||
<normalizedToken id="04BDEDC02D044ABFE381EF606ED08FA4" originalValue="Belém">Belem</normalizedToken>
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, approx. 4.5 km,
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<collectingRegion id="239548B0CEE62A751431141DFAC4660B">Loreto</collectingRegion>
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, Peru,
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<geoCoordinate id="D89A845188E25EE9C0A3FE04526130FE" direction="south" orientation="latitude" value="-3.7951667">3°47.71'S</geoCoordinate>
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,
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<geoCoordinate id="BB731ADE97BAFB89C9F6D241C533C3A3" direction="west" orientation="longitude" value="-73.28817">73°17.29'W</geoCoordinate>
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,
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<collectionCode id="6E3525054737AF1DCC9B447F22C4743D" collectionName="USA, Illinois, Carbondale, Southern Illinois University, Research Museum of Zoology">SIUC</collectionCode>
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29195, 1, (94.8)
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||
</materialsCitation>
|
||
.
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<materialsCitation id="0485177D9A5B2919DDA4F21E042C7389" ID-GBIF-Occurrence="1058481112" country="Peru" stateProvince="Loreto">
|
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<normalizedToken id="268E9739A9563AFC043AE044477DACD1" originalValue="Río">Rio</normalizedToken>
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Nanay,
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<normalizedToken id="CC45B4BD18169DF4436A6B2222D56E8B" originalValue="Río">Rio</normalizedToken>
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Amazonas dr. sand beach on right bank, 0.25 mi below and across river from Nina Rumi,
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<geoCoordinate id="480D4F981A6B790254FC5F627B00D3CD" direction="south" orientation="latitude" value="-3.7333333">03°44’S</geoCoordinate>
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,
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<geoCoordinate id="3EB0414EC005416ACDDCDE8A96C5B40E" direction="west" orientation="longitude" value="-73.333336">73°20’W</geoCoordinate>
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,
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<collectionCode id="EB8A679B7F2CEF987689A1BE69E0E065" collectionName="USA, Pennsylvania, Philadelphia, Academy of Natural Sciences">ANSP</collectionCode>
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176149, 1, (104.5)
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="242CA82EF55C4EA81F143EE9E4B41909" type="diagnosis">
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<paragraph id="FAF0F82429DFCC8643CD42DF3259ECD4" pageNumber="25">
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Diagnosis:
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<taxonomicName id="4DACAF87F7B6B7922E3DBFFDB3B406D3" LSID-ZBK="urn:lsid:zoobank.org:act:C7C2629A-1686-4E4E-9E15-59B1D1162CBE" family="Loricariidae" genus="Hypostomus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Hypostomus ericius Armbruster 2003:25" order="Siluriformes" pageNumber="25" phylum="Chordata" rank="species" species="ericius">Hypostomus ericius</taxonomicName>
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can be separated from all other members of the
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<taxonomicName id="B71906E0D072EE946841C88286999ED8" LSID-ZBK="urn:lsid:zoobank.org:act:7520B3CD-59C7-4425-A30E-1336A0B2AADD" family="Loricariidae" genus="Hypostomus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Hypostomus cochliodon Kner 1854:265 [15]" order="Siluriformes" pageNumber="25" phylum="Chordata" rank="species" species="cochliodon">H. cochliodon</taxonomicName>
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group by coloration: light tan-gray with well-separated spots and no stripes. In addition,
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<taxonomicName id="C8E3507835AAD84E92ACF6D024D44DBE" LSID-ZBK="urn:lsid:zoobank.org:act:C7C2629A-1686-4E4E-9E15-59B1D1162CBE" family="Loricariidae" genus="Hypostomus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Hypostomus ericius Armbruster 2003:25" order="Siluriformes" pageNumber="25" phylum="Chordata" rank="species" species="ericius">H. ericius</taxonomicName>
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differs from
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<taxonomicName id="A21F603204E0A0B2ACBADC923E2D43FB" LSID-ZBK="urn:lsid:zoobank.org:act:4125CE0B-B466-47CA-B90F-CF9073FEE6F8" family="Loricariidae" genus="Hypostomus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Hypostomus hemicochliodon Armbruster 2003:29" order="Siluriformes" pageNumber="25" phylum="Chordata" rank="species" species="hemicochliodon">H. hemicochliodon</taxonomicName>
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by a lack of a buccal papilla and from
|
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<taxonomicName id="53ECE832CFCF731D4A16BB19473C99C0" family="Loricariidae" genus="Hypostomus" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="25" phylum="Chordata" rank="species" species="pyrineusi">H. pyrineusi</taxonomicName>
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by the presence of strong, sharp keels on the lateral plates.
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<taxonomicName id="E52B833F221DFA9BA8905E0BBE34219C" LSID-ZBK="urn:lsid:zoobank.org:act:C7C2629A-1686-4E4E-9E15-59B1D1162CBE" family="Loricariidae" genus="Hypostomus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Hypostomus ericius Armbruster 2003:25" order="Siluriformes" pageNumber="25" phylum="Chordata" rank="species" species="ericius">Hypostomus ericius</taxonomicName>
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is most similar to
|
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<taxonomicName id="066A65F1D56FB124D6723149668766A1" family="Loricariidae" genus="Hypostomus" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="25" phylum="Chordata" rank="species" species="oculeus">H. oculeus</taxonomicName>
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from which it differs mainly in coloration and morphometrics(Fig. 10B).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C6ECEFA0D4E1C6E85A7AEB29FC71BF0C" type="description">
|
||
<paragraph id="DBCE9413F0E7AA9A0E7963DF21689C75" pageNumber="25">
|
||
Description: See
|
||
<taxonomicName id="4A60D6F9A48A7BBE1D94733E826C827D" LSID-ZBK="urn:lsid:zoobank.org:act:7520B3CD-59C7-4425-A30E-1336A0B2AADD" family="Loricariidae" genus="Hypostomus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Hypostomus cochliodon Kner 1854:265 [15]" order="Siluriformes" pageNumber="25" phylum="Chordata" rank="species" species="cochliodon">Hypostomus cochliodon</taxonomicName>
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group description for more details. Morphometric data given in Table 1. Coloration generally tan-gray with large, widely-spaced spots. Spots present over entire body and becoming larger posteriorly. Caudal-fin membrane and rays often with black wash distally, and lighter area proximally instead of being spotted. Spots on fin spines and rays occasionally large. Spots less pronounced in juveniles, spots may be absent on abdomen. Juveniles with four faint dorsal saddles: first below anterior rays of dorsal fin, second below posterior rays of dorsal fin and slightly posterior to dorsal fin, third slightly anterior to and ventral to adipose fin, and fourth at base of caudal fin.
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</paragraph>
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<paragraph id="86935225ED0EEFFD572CEDB479464D08" lastPageNumber="26" pageNumber="25">Dorsal fin moderately long, usually just barely reaching preadipose plate when depressed. Depressed pectoral-fin spine ventral to pelvic fin reaches beyond bases of pelvic-fin rays. Pectoral-fin spine supporting numerous stout, recurved, hypertrophied odontodes in nuptial males.</paragraph>
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<paragraph id="5693FF8994FE747E47631985A998AEF3" pageNumber="28">
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Keels sharp, very strongly developed. Orbits forming ridge distinctly raised above medial surface of head; ridges of dorsal and lateral aspect of head well-developed. Longitudinal ridge formed of raised bone and slightly larger odontodes absent on pterotic-supracleithrum beginning at postdorsal corner of orbit. Opercle usually not supporting odontodes, but sometimes up to 10 odontodes may be present on opercle (Fig. 2B). Nuptial body odontodes present (Fig. 2B). Plates in skin anterior to dorsal-fin spine usually absent or not numerous (Fig. 6A; Table 7). Cheek plates generally support several stout odontodes slightly larger than surrounding odontodes. Head appears wider and taller than in other species of
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<taxonomicName id="0C843FFA93949441C65035555AE51253" LSID-ZBK="urn:lsid:zoobank.org:act:E0BC8DAD-9CD9-4E32-920A-063D33AD0E80" family="Loricariidae" genus="Hypostomus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Hypostomus Lacépède 1803:144" order="Siluriformes" pageNumber="28" phylum="Chordata" rank="genus">Hypostomus</taxonomicName>
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.
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</paragraph>
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<paragraph id="E76057B8618EEC7EEED0E8EF906F4337" pageNumber="28">Each jaw with 5-9 teeth (mode = 7), teeth large and spoon-shaped. Average angle between dentaries 53° (SD = 9°; range: 40°-69°; N=7). Lateral line plates 27-28; dorsal plates 7-9; interdorsal plates 5-7; adipose caudal plates 9-10.</paragraph>
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</subSubSection>
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<subSubSection id="ABFC59B764623806ADF7B95020038DED" type="distribution">
|
||
<paragraph id="C733BAB0D0D63AB35C31D8F9489C39CB" pageNumber="28">
|
||
Range: Currently known from the
|
||
<normalizedToken id="8A2B04EEA5FF95C710FCED8589FC897E" originalValue="Ríos">Rios</normalizedToken>
|
||
<normalizedToken id="51CD65EAA1A1CE69A24AA37EBE16A9DA" originalValue="Marañon">Maranon</normalizedToken>
|
||
, Itaya, and Nanay of the Upper
|
||
<normalizedToken id="AA540EA54EF40F0C8088297CB56C3830" originalValue="Río">Rio</normalizedToken>
|
||
Amazonas drainage of Peru (Fig. 11).
|
||
<taxonomicName id="C0069FC290CB6CF2E04AEB86D7454928" LSID-ZBK="urn:lsid:zoobank.org:act:C7C2629A-1686-4E4E-9E15-59B1D1162CBE" family="Loricariidae" genus="Hypostomus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Hypostomus ericius Armbruster 2003:25" order="Siluriformes" pageNumber="28" phylum="Chordata" rank="species" species="ericius">Hypostomus ericius</taxonomicName>
|
||
is sympatric with
|
||
<taxonomicName id="AE111A704826BA9DE29427B586CF1920" LSID-ZBK="urn:lsid:zoobank.org:act:4125CE0B-B466-47CA-B90F-CF9073FEE6F8" family="Loricariidae" genus="Hypostomus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Hypostomus hemicochliodon Armbruster 2003:29" order="Siluriformes" pageNumber="28" phylum="Chordata" rank="species" species="hemicochliodon">H. hemicochliodon</taxonomicName>
|
||
,
|
||
<taxonomicName id="7C8B0F09F81F3059ACD2351D4E23CF0A" family="Loricariidae" genus="Hypostomus" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="28" phylum="Chordata" rank="species" species="oculeus">H. oculeus</taxonomicName>
|
||
, and
|
||
<taxonomicName id="DE960281884173B5A16995F91028FAD4" family="Loricariidae" genus="Hypostomus" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="28" phylum="Chordata" rank="species" species="pyrineusi">H. pyrineusi</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="01A64394863FC71429F52E1FC261B850" type="etymology">
|
||
<paragraph id="FEF1A8B988156A31C42536AE6F9C9FA6" pageNumber="28">Etymology: From the Latin, ericius, for hedgehog in reference to the many sharp odontodes on the keels.</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |