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<document id="70004C77A61BAEBD49A6B55AD1059CB8" ID-CLB-Dataset="99514" ID-DOI="10.11646/zootaxa.5165.4.4" ID-GBIF-Dataset="87b9b049-2fe8-4121-babf-83a1058d40f3" ID-ISSN="1175-5326" ID-Zenodo-Dep="6854000" ID-ZooBank="A6E36593-8BDD-4381-8769-459CFFCC2E77" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1658127221786" checkinUser="plazi" docAuthor="Davidian, Elena M., Manukyan, Andranik R. &amp; Belokobylskij, Sergey A." docDate="2022" docId="03E55F686C74CA57EAC804B0FA37FCFB" docLanguage="en" docName="zootaxa.5165.4.4.pdf" docOrigin="Zootaxa 5165 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Protoctonus Davidian &amp; Manukyan &amp; Belokobylskij 2022, gen. nov." docType="treatment" docVersion="5" lastPageNumber="525" masterDocId="FFDC27106C75CA52EA5F073AFFABFFC1" masterDocTitle="A new genus and species of the aphid parasitoids (Hymenoptera: Braconidae Aphidiinae) from Baltic amber" masterLastPageNumber="526" masterPageNumber="520" pageNumber="521" updateTime="1699360171250" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="8392C68F8CD82F51B6D2E827B0636BD0">A new genus and species of the aphid parasitoids (Hymenoptera: Braconidae Aphidiinae) from Baltic amber</mods:title>
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<mods:namePart id="C627B114728E49D07779BC445C2F134E">Davidian, Elena M.</mods:namePart>
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<mods:affiliation id="A206ECAED3B8EB1CA38F3230DCCCB46C">All-Russian Institute of Plant Protection (FSBSI VIZR), Podbelskogo, 3, St Petersburg - Pushkin 196608, Russia GDavidian @ yandex. ru; https: // orcid. org / 0000 - 0003 - 3804 - 4618</mods:affiliation>
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<mods:namePart id="4380EC72D3BD6195B0963CCE2F8FC40B">Manukyan, Andranik R.</mods:namePart>
<mods:affiliation id="CF1FA111BD9420B4C7A85311EDF52384">Kaliningrad Amber Museum, Marshal Vasilevskiy Square, 1, Kaliningrad 236016, Russia</mods:affiliation>
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<mods:namePart id="955BED086FEDA41DA7426440B0B488C9">Belokobylskij, Sergey A.</mods:namePart>
<mods:affiliation id="C39281232CBD6F90BAFCD4EF0CE56E45">Zoological Institute of the Russian Academy of Sciences, St Petersburg 199034, Russia</mods:affiliation>
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<treatment id="03E55F686C74CA57EAC804B0FA37FCFB" ID-DOI="http://doi.org/10.5281/zenodo.6857543" ID-GBIF-Taxon="197819119" ID-Zenodo-Dep="6857543" LSID="urn:lsid:plazi:treatment:03E55F686C74CA57EAC804B0FA37FCFB" httpUri="http://treatment.plazi.org/id/03E55F686C74CA57EAC804B0FA37FCFB" lastPageId="5" lastPageNumber="525" pageId="1" pageNumber="521">
<subSubSection id="C356BDF56C74CA53EAC804B0FDDCFC64" box="[151,631,906,933]" pageId="1" pageNumber="521" type="nomenclature">
<paragraph id="8BF3EE7E6C74CA53EAC804B0FDDCFC64" blockId="1.[151,631,906,933]" box="[151,631,906,933]" pageId="1" pageNumber="521">
<heading id="D0BB59126C74CA53EAC804B0FDDCFC64" bold="true" box="[151,631,906,933]" fontSize="11" level="1" pageId="1" pageNumber="521" reason="1">
<emphasis id="B938326C6C74CA53EAC804B0FDDCFC64" bold="true" box="[151,631,906,933]" pageId="1" pageNumber="521">
Genus
<taxonomicName id="4C4C95FD6C74CA53EAB104B1FE57FC65" authority="Davidian" authorityName="Davidian &amp; Manukyan &amp; Belokobylskij" authorityYear="2022" box="[238,508,906,933]" class="Insecta" family="Braconidae" genus="Protoctonus" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="521" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B938326C6C74CA53EAB104B1FE2AFC64" bold="true" box="[238,385,907,933]" italics="true" pageId="1" pageNumber="521">Protoctonus</emphasis>
Davidian
</taxonomicName>
,
<taxonomicNameLabel id="A20B8F176C74CA53E85604B1FDDCFC64" box="[521,631,907,933]" pageId="1" pageNumber="521" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C356BDF56C74CA53EAC804F5FD18FC26" box="[151,691,975,999]" pageId="1" pageNumber="521" type="type_taxon">
<paragraph id="8BF3EE7E6C74CA53EAC804F5FD18FC26" blockId="1.[151,691,975,999]" box="[151,691,975,999]" pageId="1" pageNumber="521">
<emphasis id="B938326C6C74CA53EAC804F5FEB5FC26" bold="true" box="[151,286,975,999]" pageId="1" pageNumber="521">
<typeStatus id="54F750DC6C74CA53EAC804F5FF67FC26" box="[151,204,975,999]" pageId="1" pageNumber="521">Type</typeStatus>
species
</emphasis>
<taxonomicName id="4C4C95FD6C74CA53EB7C04EAFDF3FC26" authority="Davidian" authorityName="Davidian &amp; Manukyan &amp; Belokobylskij" authorityYear="2022" box="[291,600,975,999]" class="Insecta" family="Braconidae" genus="Protoctonus" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="521" phylum="Arthropoda" rank="species" species="groehni" status="sp. nov.">
<emphasis id="B938326C6C74CA53EB7C04EAFE58FC26" box="[291,499,975,999]" italics="true" pageId="1" pageNumber="521">Protoctonus groehni</emphasis>
Davidian
</taxonomicName>
,
<emphasis id="B938326C6C74CA53E83D04F5FD18FC26" bold="true" box="[610,691,975,999]" pageId="1" pageNumber="521">
<taxonomicNameLabel id="A20B8F176C74CA53E83D04F5FD18FC26" box="[610,691,975,999]" pageId="1" pageNumber="521" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C356BDF56C74CA53EAC8032FFC04FB92" pageId="1" pageNumber="521" type="etymology">
<paragraph id="8BF3EE7E6C74CA53EAC8032FFC04FB92" blockId="1.[151,1437,1045,2007]" pageId="1" pageNumber="521">
<emphasis id="B938326C6C74CA53EAC8032FFEB3FBEE" bold="true" box="[151,280,1045,1071]" pageId="1" pageNumber="521">Etymology</emphasis>
. The new genus name is composed from the
<collectingCountry id="F35BAEEE6C74CA53E949032FFCCEFBEE" box="[790,869,1045,1071]" name="Greece" pageId="1" pageNumber="521">Greece</collectingCountry>
word “protos” (= first, pioneering) and part of the related aphidiine genus name “
<taxonomicName id="4C4C95FD6C74CA53EBAD0300FDD7FB92" authorityName="Haliday" authorityYear="1833" box="[498,636,1082,1107]" class="Insecta" family="Braconidae" genus="Monoctonus" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="521" phylum="Arthropoda" rank="genus">
<emphasis id="B938326C6C74CA53EBAD0300FDD7FB92" box="[498,636,1082,1107]" italics="true" pageId="1" pageNumber="521">Monoctonus</emphasis>
</taxonomicName>
”. The gender is masculine.
</paragraph>
</subSubSection>
<subSubSection id="C356BDF56C74CA53EA980367FD7BF9C2" pageId="1" pageNumber="521" type="diagnosis">
<paragraph id="8BF3EE7E6C74CA53EA980367FD7BF9C2" blockId="1.[151,1437,1045,2007]" pageId="1" pageNumber="521">
<emphasis id="B938326C6C74CA53EA980367FE91FBB6" bold="true" box="[199,314,1117,1143]" pageId="1" pageNumber="521">Diagnosis</emphasis>
. Mandible bidentate; maxillary palpus 4-segmented, labial palpus 3-segmented (
<figureCitation id="1377F2FB6C74CA53EE8F0367FA82FBB6" box="[1232,1321,1117,1143]" captionStart="FIGURES 1" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[151,1436,179,1913]" captionTargetId="figure-22@3.[151,1436,179,1917]" captionTargetPageId="3" captionText="FIGURES 1. Photos of Protoctonus groehni gen. et sp. nov. (female, holotype). A. Habitus, left side. B. Habitus, right side. C. Head and basal antennomeres, front view. D. Antenna. E. Head and mesoscutum, dorsolateral view. F. Metasoma and ovipositor, dorsolateral view. G. Wings. H. Distal part of fore wing." figureDoi="http://doi.org/10.5281/zenodo.6854002" httpUri="https://zenodo.org/record/6854002/files/figure.png" pageId="1" pageNumber="521">Fig. 1C</figureCitation>
). Antenna (
<figureCitation id="1377F2FB6C74CA53EAC003BBFF5CFB5A" box="[159,247,1153,1179]" captionStart="FIGURES 1" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[151,1436,179,1913]" captionTargetId="figure-22@3.[151,1436,179,1917]" captionTargetPageId="3" captionText="FIGURES 1. Photos of Protoctonus groehni gen. et sp. nov. (female, holotype). A. Habitus, left side. B. Habitus, right side. C. Head and basal antennomeres, front view. D. Antenna. E. Head and mesoscutum, dorsolateral view. F. Metasoma and ovipositor, dorsolateral view. G. Wings. H. Distal part of fore wing." figureDoi="http://doi.org/10.5281/zenodo.6854002" httpUri="https://zenodo.org/record/6854002/files/figure.png" pageId="1" pageNumber="521">Fig. 1D</figureCitation>
) with 11 antennomeres, widened toward apex, densely pubescent. Mesoscutum with notauli reaching its middle (
<figureCitation id="1377F2FB6C74CA53EAAC039FFEECFB7E" box="[243,327,1189,1215]" captionStart="FIGURES 1" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[151,1436,179,1913]" captionTargetId="figure-22@3.[151,1436,179,1917]" captionTargetPageId="3" captionText="FIGURES 1. Photos of Protoctonus groehni gen. et sp. nov. (female, holotype). A. Habitus, left side. B. Habitus, right side. C. Head and basal antennomeres, front view. D. Antenna. E. Head and mesoscutum, dorsolateral view. F. Metasoma and ovipositor, dorsolateral view. G. Wings. H. Distal part of fore wing." figureDoi="http://doi.org/10.5281/zenodo.6854002" httpUri="https://zenodo.org/record/6854002/files/figure.png" pageId="1" pageNumber="521">Fig. 1E</figureCitation>
). Propodeum perhaps with median areola delineated by weak carinae. In fore wing (
<figureCitation id="1377F2FB6C74CA53EEB7039FFACDFB01" box="[1256,1382,1189,1216]" captionStart="FIGURES 1" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[151,1436,179,1913]" captionTargetId="figure-22@3.[151,1436,179,1917]" captionTargetPageId="3" captionText="FIGURES 1. Photos of Protoctonus groehni gen. et sp. nov. (female, holotype). A. Habitus, left side. B. Habitus, right side. C. Head and basal antennomeres, front view. D. Antenna. E. Head and mesoscutum, dorsolateral view. F. Metasoma and ovipositor, dorsolateral view. G. Wings. H. Distal part of fore wing." figureDoi="http://doi.org/10.5281/zenodo.6854002" httpUri="https://zenodo.org/record/6854002/files/figure.png" pageId="1" pageNumber="521">Figs 1G, H</figureCitation>
;
<figureCitation id="1377F2FB6C74CA53EF2E039FFA3EFB7E" box="[1393,1429,1189,1215]" captionStart="FIGURES 2" captionStartId="5.[151,264,1386,1410]" captionTargetBox="[164,1421,868,1356]" captionTargetId="figure-758@5.[151,1435,850,1363]" captionTargetPageId="5" captionText="FIGURES 2. Figures of Protoctonus groehni gen. et sp. nov. (female, holotype). A. Fore wing. B. Ovipositor sheaths." figureDoi="http://doi.org/10.5281/zenodo.6854004" httpUri="https://zenodo.org/record/6854004/files/figure.png" pageId="1" pageNumber="521">2A</figureCitation>
) pterostigma triangular. Metacarp (1-R1), radial (r, 3-SR, SR1) and parallel (CU1a) veins of fore wing long and almost reaching distal margin of wing; medial (1-SR+M, 2-M, 3-M) vein short and not reaching distal margin of wing. First and second radiomedial (submarginal) cells fused together with discoidal (discal) cell and formed united large central cell. Also present closed medial (basal), submedial (subbasal) and brachial (subdiscal) cells. Hind wing without closed cells. Legs (
<figureCitation id="1377F2FB6C74CA53EBC60263FDBFFAB2" box="[409,532,1369,1395]" captionStart="FIGURES 1" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[151,1436,179,1913]" captionTargetId="figure-22@3.[151,1436,179,1917]" captionTargetPageId="3" captionText="FIGURES 1. Photos of Protoctonus groehni gen. et sp. nov. (female, holotype). A. Habitus, left side. B. Habitus, right side. C. Head and basal antennomeres, front view. D. Antenna. E. Head and mesoscutum, dorsolateral view. F. Metasoma and ovipositor, dorsolateral view. G. Wings. H. Distal part of fore wing." figureDoi="http://doi.org/10.5281/zenodo.6854002" httpUri="https://zenodo.org/record/6854002/files/figure.png" pageId="1" pageNumber="521">Figs 1A, F</figureCitation>
) covered by dense and semi-erect setae (as those on body). Petiole of metasoma (
<figureCitation id="1377F2FB6C74CA53EAC00247FF5BFA56" box="[159,240,1405,1431]" captionStart="FIGURES 1" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[151,1436,179,1913]" captionTargetId="figure-22@3.[151,1436,179,1917]" captionTargetPageId="3" captionText="FIGURES 1. Photos of Protoctonus groehni gen. et sp. nov. (female, holotype). A. Habitus, left side. B. Habitus, right side. C. Head and basal antennomeres, front view. D. Antenna. E. Head and mesoscutum, dorsolateral view. F. Metasoma and ovipositor, dorsolateral view. G. Wings. H. Distal part of fore wing." figureDoi="http://doi.org/10.5281/zenodo.6854002" httpUri="https://zenodo.org/record/6854002/files/figure.png" pageId="1" pageNumber="521">Fig. 1F</figureCitation>
) rather long, slightly widened towards apex, about 1.6 × longer than its width medially at level of spiracles. Ovipositor sheaths (
<figureCitation id="1377F2FB6C74CA53EB27029BFE7AFA7A" box="[376,465,1441,1467]" captionStart="FIGURES 1" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[151,1436,179,1913]" captionTargetId="figure-22@3.[151,1436,179,1917]" captionTargetPageId="3" captionText="FIGURES 1. Photos of Protoctonus groehni gen. et sp. nov. (female, holotype). A. Habitus, left side. B. Habitus, right side. C. Head and basal antennomeres, front view. D. Antenna. E. Head and mesoscutum, dorsolateral view. F. Metasoma and ovipositor, dorsolateral view. G. Wings. H. Distal part of fore wing." figureDoi="http://doi.org/10.5281/zenodo.6854002" httpUri="https://zenodo.org/record/6854002/files/figure.png" pageId="1" pageNumber="521">Figs 1F</figureCitation>
;
<figureCitation id="1377F2FB6C74CA53EB82029BFE54FA7A" box="[477,511,1441,1467]" captionStart="FIGURES 2" captionStartId="5.[151,264,1386,1410]" captionTargetBox="[164,1421,868,1356]" captionTargetId="figure-758@5.[151,1435,850,1363]" captionTargetPageId="5" captionText="FIGURES 2. Figures of Protoctonus groehni gen. et sp. nov. (female, holotype). A. Fore wing. B. Ovipositor sheaths." figureDoi="http://doi.org/10.5281/zenodo.6854004" httpUri="https://zenodo.org/record/6854004/files/figure.png" pageId="1" pageNumber="521">2B</figureCitation>
) not long, weakly curved down, acuminate, 2.6 × longer than its maximum width, 0.15 × as long as metasoma, gradually tapered from base towards apex, narrowly rounded apically, with long setae on dorsal and ventral margins; setae apically short.
</paragraph>
</subSubSection>
<subSubSection id="C356BDF56C74CA50EA980137FB5CFE10" lastPageId="2" lastPageNumber="522" pageId="1" pageNumber="521" type="discussion">
<paragraph id="8BF3EE7E6C74CA53EA980137FB7AF976" blockId="1.[151,1437,1045,2007]" pageId="1" pageNumber="521">
<emphasis id="B938326C6C74CA53EA980137FE90F9E6" bold="true" box="[199,315,1549,1575]" pageId="1" pageNumber="521">Remarks.</emphasis>
The new aphidiine genus
<taxonomicName id="4C4C95FD6C74CA53E83E0134FD42F9E6" authority="Davidian &amp; Manukyan &amp; Belokobylskij, 2022" authorityName="Davidian &amp; Manukyan &amp; Belokobylskij" authorityYear="2022" box="[609,745,1550,1575]" class="Insecta" family="Braconidae" genus="Protoctonus" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="521" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B938326C6C74CA53E83E0134FD42F9E6" box="[609,745,1550,1575]" italics="true" pageId="1" pageNumber="521">Protoctonus</emphasis>
</taxonomicName>
<emphasis id="B938326C6C74CA53E8B00137FCF3F9E6" bold="true" box="[751,856,1549,1575]" pageId="1" pageNumber="521">
<taxonomicNameLabel id="A20B8F176C74CA53E8B00137FCF3F9E6" box="[751,856,1549,1575]" pageId="1" pageNumber="521" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
is most similar to the fossil genera
<taxonomicName id="4C4C95FD6C74CA53EEBB0137FEE7F98A" authority="Mackauer, 1961" authorityName="Mackauer" authorityYear="1961" class="Insecta" family="Braconidae" genus="Protacanthoides" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="521" phylum="Arthropoda" rank="genus">
<emphasis id="B938326C6C74CA53EEBB0137FA37F9E6" box="[1252,1436,1549,1575]" italics="true" pageId="1" pageNumber="521">Protacanthoides</emphasis>
Mackauer, 1961
</taxonomicName>
(with
<typeStatus id="54F750DC6C74CA53EBC90108FE6DF98D" box="[406,454,1586,1612]" pageId="1" pageNumber="521">type</typeStatus>
species
<taxonomicName id="4C4C95FD6C74CA53E8790108FC06F98A" authority="Quilis, 1940" authorityName="Quilis" authorityYear="1940" box="[550,941,1585,1611]" class="Insecta" family="Braconidae" genus="Trioxys" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="521" phylum="Arthropoda" rank="species" species="obscuriformis">
<emphasis id="B938326C6C74CA53E8790108FCB7F98A" box="[550,796,1585,1611]" italics="true" pageId="1" pageNumber="521">Trioxys obscuriformis</emphasis>
Quilis, 1940
</taxonomicName>
=
<taxonomicName id="4C4C95FD6C74CA53E9930108FA9FF98D" authority="Mackauer, 1959" authorityName="Mackauer" authorityYear="1959" box="[972,1332,1585,1612]" class="Insecta" family="Braconidae" genus="Trioxys" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="521" phylum="Arthropoda" rank="species" species="fossilis">
<emphasis id="B938326C6C74CA53E9930108FBDFF98A" box="[972,1140,1585,1611]" italics="true" pageId="1" pageNumber="521">Trioxys fossilis</emphasis>
Mackauer, 1959
</taxonomicName>
),
<taxonomicName id="4C4C95FD6C74CA53EF170108FE2FF9AE" authority="Stary, 1973" authorityName="Stary" authorityYear="1973" class="Insecta" family="Braconidae" genus="Promonoctonia" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="521" phylum="Arthropoda" rank="genus">
<emphasis id="B938326C6C74CA53EF170108FF52F9AE" italics="true" pageId="1" pageNumber="521">Promonoctonia</emphasis>
<bibRefCitation id="EFDD938F6C74CA53EB5C016FFE2FF9AE" author="Stary, P." box="[259,388,1621,1647]" pageId="1" pageNumber="521" pagination="1 - 22" refId="ref5097" refString="Stary, P. (1973) A revision of the fossil Aphidiidae (Hymenoptera). Annotationes Zoologicae &amp; Botanicae, 87, 1 - 22." type="journal article" year="1973">Stary, 1973</bibRefCitation>
</taxonomicName>
(with
<typeStatus id="54F750DC6C74CA53EB8D016CFDA9F9B1" box="[466,514,1622,1648]" pageId="1" pageNumber="521">type</typeStatus>
species
<taxonomicName id="4C4C95FD6C74CA53E839016FFC48F9AE" authority="Quilis, 1940" authorityName="Quilis" authorityYear="1940" box="[614,995,1621,1647]" class="Insecta" family="Braconidae" genus="Aphidius" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="521" phylum="Arthropoda" rank="species" species="quievreuxi">
<emphasis id="B938326C6C74CA53E839016FFCE3F9AE" box="[614,840,1621,1647]" italics="true" pageId="1" pageNumber="521">Aphidius quievreuxi</emphasis>
Quilis, 1940
</taxonomicName>
), and
<emphasis id="B938326C6C74CA53EE74016FFB4FF9AE" box="[1067,1252,1621,1647]" italics="true" pageId="1" pageNumber="521">Sakhalinoctonus</emphasis>
Davidian, 2021 (with
<typeStatus id="54F750DC6C74CA53EA860140FEA2F955" box="[217,265,1658,1684]" pageId="1" pageNumber="521">type</typeStatus>
species
<emphasis id="B938326C6C74CA53EB380143FD68F952" box="[359,707,1657,1683]" italics="true" pageId="1" pageNumber="521">Sakhalinoctonus alexrasnitsyni</emphasis>
Davidian, 2021) by the fewer number of antennomeres, presence the closed central cell in fore wing and curved downwards and taped apically ovipositor sheaths.
</paragraph>
<paragraph id="8BF3EE7E6C74CA53EA9801FBFD58F872" blockId="1.[151,1437,1045,2007]" pageId="1" pageNumber="521">
Genera
<taxonomicName id="4C4C95FD6C74CA53EB4101FBFE7DF91A" authorityName="Mackauer" authorityYear="1961" box="[286,470,1729,1755]" class="Insecta" family="Braconidae" genus="Protacanthoides" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="521" phylum="Arthropoda" rank="genus">
<emphasis id="B938326C6C74CA53EB4101FBFE7DF91A" box="[286,470,1729,1755]" italics="true" pageId="1" pageNumber="521">Protacanthoides</emphasis>
</taxonomicName>
and
<taxonomicName id="4C4C95FD6C74CA53E85401F8FD13F91A" authorityName="Stary" authorityYear="1973" box="[523,696,1730,1755]" class="Insecta" family="Braconidae" genus="Promonoctonia" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="521" phylum="Arthropoda" rank="genus">
<emphasis id="B938326C6C74CA53E85401F8FD13F91A" box="[523,696,1730,1755]" italics="true" pageId="1" pageNumber="521">Promonoctonia</emphasis>
</taxonomicName>
are characterised by a 13 antennomeres in thick antenna, short and not reaching distal margin of the wing the metacarp (1-R1), radial (r, 3-SR, SR1) and medial (2-M, 3-M) veins, and curved down the ovipositor sheaths. Besides those,
<taxonomicName id="4C4C95FD6C74CA53E8BA0033FC5FF8E2" baseAuthorityName="Mackauer" baseAuthorityYear="1957" box="[741,1012,1801,1827]" class="Insecta" family="Braconidae" genus="Protacanthoides" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="521" phylum="Arthropoda" rank="species" species="fossilis">
<emphasis id="B938326C6C74CA53E8BA0033FC5FF8E2" box="[741,1012,1801,1827]" italics="true" pageId="1" pageNumber="521">Protacanthoides fossilis</emphasis>
</taxonomicName>
has metasomal petiole short, smooth and without protruding spiracular tubercles, length of petiole approximately equal to its maximum width; ovipositor sheath evenly curved downwards, strongly tapered towards its apex and acuminate apically, about 0.15 × as long as metasoma. Unfortunately the information about condition of notauli and propodeum and pubescence of the ovipositor sheath are absent in the description of this species.
</paragraph>
<paragraph id="8BF3EE7E6C74CA50EA980084FDAFFF38" blockId="1.[151,1437,1045,2007]" lastBlockId="2.[151,1437,151,393]" lastPageId="2" lastPageNumber="522" pageId="1" pageNumber="521">
<taxonomicName id="4C4C95FD6C74CA53EA980084FE5AF816" baseAuthorityName="Quilis" baseAuthorityYear="1940" box="[199,497,1982,2007]" class="Insecta" family="Braconidae" genus="Promonoctonia" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="521" phylum="Arthropoda" rank="species" species="quievreuxi">
<emphasis id="B938326C6C74CA53EA980084FE5AF816" box="[199,497,1982,2007]" italics="true" pageId="1" pageNumber="521">Promonoctonia quievreuxi</emphasis>
</taxonomicName>
is additionally characterised by the following features: notauli missing; propodeum smooth and without delineated areola; metasomal petiole very short and wide; ovipositor sheaths claw-shaped, with long and wide its basal part and short and narrow apical part, approximately 3.0 × longer than its maximum width and 0.15 × as long as metasoma.
</paragraph>
<paragraph id="8BF3EE7E6C77CA50EA980639FE2CFE48" blockId="2.[151,1437,151,393]" pageId="2" pageNumber="522">
The genus
<emphasis id="B938326C6C77CA50EB1C0639FE57FEDC" box="[323,508,259,285]" italics="true" pageId="2" pageNumber="522">Sakhalinoctonus</emphasis>
has 12 antennomeres; relatively long radial (r, 3-SR, SR1) and medial (2-M, 3-M) veins, which do not reach the distal margin of fore wing; ovipositor sheath long, almost 6.0 × longer than its maximum width, weakly curved down and gradually tapered towards apex, very narrowly rounded apically, and covered by mainly long setae.
</paragraph>
<paragraph id="8BF3EE7E6C77CA50EAC8068DFB5CFE10" blockId="2.[151,1271,439,465]" box="[151,1271,439,465]" pageId="2" pageNumber="522">
The differences between
<taxonomicName id="4C4C95FD6C77CA50EBEE068DFD92FE11" authority="Davidian &amp; Manukyan &amp; Belokobylskij, 2022" authorityName="Davidian &amp; Manukyan &amp; Belokobylskij" authorityYear="2022" box="[433,569,439,464]" class="Insecta" family="Braconidae" genus="Protoctonus" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="522" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B938326C6C77CA50EBEE068DFD92FE11" box="[433,569,439,464]" italics="true" pageId="2" pageNumber="522">Protoctonus</emphasis>
</taxonomicName>
<emphasis id="B938326C6C77CA50E81F068DFD02FE10" bold="true" box="[576,681,439,465]" pageId="2" pageNumber="522">
<taxonomicNameLabel id="A20B8F176C77CA50E81F068DFD02FE10" box="[576,681,439,465]" pageId="2" pageNumber="522" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
and similar fossil genera are given in the key below.
</paragraph>
</subSubSection>
<subSubSection id="C356BDF56C77CA50EAC806C6FA37FBFE" pageId="2" pageNumber="522" type="key">
<key id="FFDDB94F6C77CA50EAC806C6FA37FBFE" pageId="2" pageNumber="522">
<keyStep id="30B8F6DB6C77CA50EAC806C6FA37FD62" pageId="2" pageNumber="522">
<paragraph id="8BF3EE7E6C77CA50EAC806C6FA37FD47" blockId="2.[151,1437,508,1087]" pageId="2" pageNumber="522">
<keyLead id="30BD4D4B6C77CA50EAC806C6FA37FD47" pageId="2" pageNumber="522">
1. Antenna with 11 antennomeres (
<figureCitation id="1377F2FB6C77CA50E84706C6FDF7FDD3" box="[536,604,508,530]" captionStart="FIGURES 1" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[151,1436,179,1913]" captionTargetId="figure-22@3.[151,1436,179,1917]" captionTargetPageId="3" captionText="FIGURES 1. Photos of Protoctonus groehni gen. et sp. nov. (female, holotype). A. Habitus, left side. B. Habitus, right side. C. Head and basal antennomeres, front view. D. Antenna. E. Head and mesoscutum, dorsolateral view. F. Metasoma and ovipositor, dorsolateral view. G. Wings. H. Distal part of fore wing." figureDoi="http://doi.org/10.5281/zenodo.6854002" httpUri="https://zenodo.org/record/6854002/files/figure.png" pageId="2" pageNumber="522">Fig. 1D</figureCitation>
)—Flagellomeres from fifth moniliform (
<figureCitation id="1377F2FB6C77CA50E9B306C6FB93FDD0" box="[1004,1080,508,530]" captionStart="FIGURES 1" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[151,1436,179,1913]" captionTargetId="figure-22@3.[151,1436,179,1917]" captionTargetPageId="3" captionText="FIGURES 1. Photos of Protoctonus groehni gen. et sp. nov. (female, holotype). A. Habitus, left side. B. Habitus, right side. C. Head and basal antennomeres, front view. D. Antenna. E. Head and mesoscutum, dorsolateral view. F. Metasoma and ovipositor, dorsolateral view. G. Wings. H. Distal part of fore wing." figureDoi="http://doi.org/10.5281/zenodo.6854002" httpUri="https://zenodo.org/record/6854002/files/figure.png" pageId="2" pageNumber="522">Fig. 1D</figureCitation>
). Metacarp (1-R1) of fore wing 1.3 × longer than pterostigma. Radial (r, 3-SR, SR1) vein and metacarp (1-R1) almost reaching distal margin of fore wing (
<figureCitation id="1377F2FB6C77CA50EF130523FF5DFD8D" captionStart="FIGURES 1" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[151,1436,179,1913]" captionTargetId="figure-22@3.[151,1436,179,1917]" captionTargetPageId="3" captionText="FIGURES 1. Photos of Protoctonus groehni gen. et sp. nov. (female, holotype). A. Habitus, left side. B. Habitus, right side. C. Head and basal antennomeres, front view. D. Antenna. E. Head and mesoscutum, dorsolateral view. F. Metasoma and ovipositor, dorsolateral view. G. Wings. H. Distal part of fore wing." figureDoi="http://doi.org/10.5281/zenodo.6854002" httpUri="https://zenodo.org/record/6854002/files/figure.png" pageId="2" pageNumber="522">Fig. 1G, H</figureCitation>
;
<figureCitation id="1377F2FB6C77CA50EB5F050CFE8AFD8A" box="[256,289,566,587]" captionStart="FIGURES 2" captionStartId="5.[151,264,1386,1410]" captionTargetBox="[164,1421,868,1356]" captionTargetId="figure-758@5.[151,1435,850,1363]" captionTargetPageId="5" captionText="FIGURES 2. Figures of Protoctonus groehni gen. et sp. nov. (female, holotype). A. Fore wing. B. Ovipositor sheaths." figureDoi="http://doi.org/10.5281/zenodo.6854004" httpUri="https://zenodo.org/record/6854004/files/figure.png" pageId="2" pageNumber="522">2A</figureCitation>
). Ovipositor sheaths (
<figureCitation id="1377F2FB6C77CA50EBAB050CFDEBFD8A" box="[500,576,566,588]" captionStart="FIGURES 1" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[151,1436,179,1913]" captionTargetId="figure-22@3.[151,1436,179,1917]" captionTargetPageId="3" captionText="FIGURES 1. Photos of Protoctonus groehni gen. et sp. nov. (female, holotype). A. Habitus, left side. B. Habitus, right side. C. Head and basal antennomeres, front view. D. Antenna. E. Head and mesoscutum, dorsolateral view. F. Metasoma and ovipositor, dorsolateral view. G. Wings. H. Distal part of fore wing." figureDoi="http://doi.org/10.5281/zenodo.6854002" httpUri="https://zenodo.org/record/6854002/files/figure.png" pageId="2" pageNumber="522">Figs 1F</figureCitation>
;
<figureCitation id="1377F2FB6C77CA50E814050CFDC3FD8A" box="[587,616,566,587]" captionStart="FIGURES 2" captionStartId="5.[151,264,1386,1410]" captionTargetBox="[164,1421,868,1356]" captionTargetId="figure-758@5.[151,1435,850,1363]" captionTargetPageId="5" captionText="FIGURES 2. Figures of Protoctonus groehni gen. et sp. nov. (female, holotype). A. Fore wing. B. Ovipositor sheaths." figureDoi="http://doi.org/10.5281/zenodo.6854004" httpUri="https://zenodo.org/record/6854004/files/figure.png" pageId="2" pageNumber="522">2B</figureCitation>
) curved downwards, subtriangular in shape, taped towards apex, entirely covered by dense setae: long setae arranged along dorsal and ventral margins and short setae located on apex of sheaths. Ovipositor sheath 2.6 × longer than its maximum basal width. Baltic amber.....................................
<taxonomicName id="4C4C95FD6C77CA50EE970548FA97FD46" authority="Davidian &amp; Manukyan &amp; Belokobylskij, 2022" authorityName="Davidian &amp; Manukyan &amp; Belokobylskij" authorityYear="2022" box="[1224,1340,626,647]" class="Insecta" family="Braconidae" genus="Protoctonus" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="522" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B938326C6C77CA50EE970548FA97FD46" box="[1224,1340,626,647]" italics="true" pageId="2" pageNumber="522">Protoctonus</emphasis>
</taxonomicName>
<emphasis id="B938326C6C77CA50EF1D054BFA37FD47" bold="true" box="[1346,1436,625,646]" pageId="2" pageNumber="522">
<taxonomicNameLabel id="A20B8F176C77CA50EF1D054BFA37FD47" box="[1346,1436,625,646]" pageId="2" pageNumber="522" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BF3EE7E6C77CA50EAC805B5FA37FD62" blockId="2.[151,1437,508,1087]" box="[151,1436,654,676]" pageId="2" pageNumber="522">
<keyLead id="30BD4D4B6C77CA50EAC805B5FA37FD62" box="[151,1436,654,676]" pageId="2" pageNumber="522">- Antenna with 1213 antennomeres....................................................................... 2</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B8F6DB6C77CA50EAC80596FA37FC4F" pageId="2" pageNumber="522">
<paragraph id="8BF3EE7E6C77CA50EAC80596FA37FCDB" blockId="2.[151,1437,508,1087]" pageId="2" pageNumber="522">
<keyLead id="30BD4D4B6C77CA50EAC80596FA37FCDB" pageId="2" pageNumber="522">
2. Antenna with 12 antennomeres, relatively slender. Metacarp (1-R1), radial (r, 3-SR, SR1) and medial (2 М, 3-M) veins of fore wing long and almost reaching distal margin of wing. Ovipositor sheath relatively long, curved down, gradually tapered towards apex, narrow curved apically, about 6.0 × longer than its maximum width and 0.4 × as long as metasoma. Sakhalinian amber...........................................................................
<emphasis id="B938326C6C77CA50EEFF043EFA94FCD8" box="[1184,1343,772,793]" italics="true" pageId="2" pageNumber="522">Sakhalinoctonus</emphasis>
Davidian
</keyLead>
</paragraph>
<paragraph id="8BF3EE7E6C77CA50EAC8041BFA37FC4F" blockId="2.[151,1437,508,1087]" pageId="2" pageNumber="522">
<keyLead id="30BD4D4B6C77CA50EAC8041BFA37FC4F" pageId="2" pageNumber="522">- Antenna with 13 antennomeres, thick. Metacarp (1-R1), radial (r, 3-SR, SR1) and medial (2 М, 3-M) veins of fore wing short and far not reaching distal margin of wing. Ovipositor sheaths short, claw-shaped, with long widened basal part and short narrow apical part, or evenly tapered to apex and acuminate apically, about 3.0 × longer than its maximum width and 0.150.20 × as long as metasoma.................................................................................. 3</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B8F6DB6C77CA50EAC804ACFA37FBFE" pageId="2" pageNumber="522">
<paragraph id="8BF3EE7E6C77CA50EAC804ACFA36FC26" blockId="2.[151,1437,508,1087]" pageId="2" pageNumber="522">
<keyLead id="30BD4D4B6C77CA50EAC804ACFA36FC26" pageId="2" pageNumber="522">
3. Ovipositor sheaths claw-shaped, with long and wide their basal part and short and narrow apically, perhaps narrowly rounded apically.—Mesoscutum without notauli. Propodeum without delineated areola. Petiole of metasoma very short and wide.
<collectingCountry id="F35BAEEE6C77CA50EABD04EBFE8DFC27" box="[226,294,977,998]" name="France" pageId="2" pageNumber="522">France</collectingCountry>
, Haut-Rhin, potassium layers......................................................
<taxonomicName id="4C4C95FD6C77CA50EE9004E8FA36FC26" ID-CoL="6WS2" authority="Stary" authorityName="Stary" authorityYear="1973" box="[1231,1437,977,999]" class="Insecta" family="Braconidae" genus="Promonoctonia" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="522" phylum="Arthropoda" rank="genus">
<emphasis id="B938326C6C77CA50EE9004E8FAC8FC26" box="[1231,1379,978,999]" italics="true" pageId="2" pageNumber="522">Promonoctonia</emphasis>
Starý
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BF3EE7E6C77CA50EAC804D5FA37FBFE" blockId="2.[151,1437,508,1087]" pageId="2" pageNumber="522">
<keyLead id="30BD4D4B6C77CA50EAC804D5FA37FBFE" pageId="2" pageNumber="522">
- Ovipositor sheaths weakly curved downwards, gradually tapered from base to apex and acuminate apically.—Petiole of metasoma subtriangular, smooth, without projecting spiracular tubercles, about as long as its maximum width.
<collectingCountry id="F35BAEEE6C77CA50EEBC0336FA8CFBE0" box="[1251,1319,1036,1057]" name="France" pageId="2" pageNumber="522">France</collectingCountry>
, Haut-Rhin, potassium layers.................................................................
<taxonomicName id="4C4C95FD6C77CA50EEC60313FA37FBFE" ID-CoL="6XBT" authority="Mackauer" authorityName="Mackauer" authorityYear="1961" box="[1177,1436,1065,1087]" class="Insecta" family="Braconidae" genus="Protacanthoides" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="522" phylum="Arthropoda" rank="genus">
<emphasis id="B938326C6C77CA50EEC60313FA9CFBFF" box="[1177,1335,1065,1086]" italics="true" pageId="2" pageNumber="522">Protacanthoides</emphasis>
Mackauer
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
<caption id="DF33BEF66C76CA51EAC800ADFD7FF836" ID-DOI="http://doi.org/10.5281/zenodo.6854002" ID-Zenodo-Dep="6854002" httpUri="https://zenodo.org/record/6854002/files/figure.png" pageId="3" pageNumber="523" startId="3.[151,264,1943,1967]" targetBox="[151,1436,179,1913]" targetPageId="3">
<paragraph id="8BF3EE7E6C76CA51EAC800ADFD7FF836" blockId="3.[151,1437,1943,2039]" pageId="3" pageNumber="523">
<emphasis id="B938326C6C76CA51EAC800ADFE8AF86E" bold="true" box="[151,289,1943,1967]" pageId="3" pageNumber="523">FIGURES 1.</emphasis>
Photos of
<taxonomicName id="4C4C95FD6C76CA51EBD200A2FDF0F86E" authority="Davidian &amp; Manukyan &amp; Belokobylskij, 2022" authorityName="Davidian &amp; Manukyan &amp; Belokobylskij" authorityYear="2022" box="[397,603,1943,1967]" class="Insecta" family="Braconidae" genus="Protoctonus" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="523" phylum="Arthropoda" rank="species" species="groehni" status="gen. et sp. nov.">
<emphasis id="B938326C6C76CA51EBD200A2FDF0F86E" box="[397,603,1943,1967]" italics="true" pageId="3" pageNumber="523">Protoctonus groehni</emphasis>
</taxonomicName>
<emphasis id="B938326C6C76CA51E83E00ADFD56F86E" bold="true" box="[609,765,1943,1967]" pageId="3" pageNumber="523">
<taxonomicNameLabel id="A20B8F176C76CA51E83E00ADFD56F86E" box="[609,765,1943,1967]" pageId="3" pageNumber="523" rank="species">gen. et sp. nov.</taxonomicNameLabel>
</emphasis>
(female, holotype).
<emphasis id="B938326C6C76CA51E99500ADFC48F86E" bold="true" box="[970,995,1943,1967]" pageId="3" pageNumber="523">A.</emphasis>
Habitus, left side.
<emphasis id="B938326C6C76CA51EEFE00ADFB13F86E" bold="true" box="[1185,1208,1943,1967]" pageId="3" pageNumber="523">B.</emphasis>
Habitus, right side.
<emphasis id="B938326C6C76CA51EFDB00ADFA37F86E" bold="true" box="[1412,1436,1943,1967]" pageId="3" pageNumber="523">C.</emphasis>
Head and basal antennomeres, front view.
<emphasis id="B938326C6C76CA51E8600081FDFCF812" bold="true" box="[575,599,1979,2003]" pageId="3" pageNumber="523">D.</emphasis>
Antenna.
<emphasis id="B938326C6C76CA51E8E30081FD7FF812" bold="true" box="[700,724,1979,2003]" pageId="3" pageNumber="523">E.</emphasis>
Head and mesoscutum, dorsolateral view.
<emphasis id="B938326C6C76CA51EE200081FB39F812" bold="true" box="[1151,1170,1979,2003]" pageId="3" pageNumber="523">F.</emphasis>
Metasoma and ovipositor, dorsolateral view.
<emphasis id="B938326C6C76CA51EB0E00E5FEC0F836" bold="true" box="[337,363,2015,2039]" pageId="3" pageNumber="523">G.</emphasis>
Wings.
<emphasis id="B938326C6C76CA51EBE000E5FE72F836" bold="true" box="[447,473,2015,2039]" pageId="3" pageNumber="523">H.</emphasis>
Distal part of fore wing.
</paragraph>
</caption>
<subSubSection id="C356BDF56C70CA57EAC8061DFC19FE80" box="[151,946,294,321]" pageId="5" pageNumber="525" type="discussion">
<paragraph id="8BF3EE7E6C70CA57EAC8061DFC19FE80" blockId="5.[151,946,294,321]" box="[151,946,294,321]" pageId="5" pageNumber="525">
<heading id="D0BB59126C70CA57EAC8061DFC19FE80" bold="true" box="[151,946,294,321]" fontSize="11" level="1" pageId="5" pageNumber="525" reason="1">
<emphasis id="B938326C6C70CA57EAC8061DFC19FE80" bold="true" box="[151,946,294,321]" pageId="5" pageNumber="525">
Key to
<taxonomicName id="4C4C95FD6C70CA57EAAE061DFE2FFE80" authority="Davidian &amp; Manukyan &amp; Belokobylskij, 2022" authorityName="Davidian &amp; Manukyan &amp; Belokobylskij" authorityYear="2022" box="[241,388,295,321]" class="Insecta" family="Braconidae" genus="Protoctonus" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="525" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B938326C6C70CA57EAAE061DFE2FFE80" bold="true" box="[241,388,295,321]" italics="true" pageId="5" pageNumber="525">Protoctonus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20B8F176C70CA57EBD4061DFE52FE80" box="[395,505,295,321]" pageId="5" pageNumber="525" rank="genus">gen. nov.</taxonomicNameLabel>
and the most similar recent genera
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C356BDF56C70CA57EAC80656FA37FCFB" pageId="5" pageNumber="525" type="key">
<keyStep id="30B8F6DB6C70CA57EAC80656FA37FE37" pageId="5" pageNumber="525">
<paragraph id="8BF3EE7E6C70CA57EAC80656FA37FE18" blockId="5.[151,1437,364,826]" pageId="5" pageNumber="525">
<keyLead id="30BD4D4B6C70CA57EAC80656FA37FE18" pageId="5" pageNumber="525">
1. Antenna with 11 antennomeres (
<figureCitation id="1377F2FB6C70CA57E8450656FDF4FE43" box="[538,607,364,386]" captionStart="FIGURES 1" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[151,1436,179,1913]" captionTargetId="figure-22@3.[151,1436,179,1917]" captionTargetPageId="3" captionText="FIGURES 1. Photos of Protoctonus groehni gen. et sp. nov. (female, holotype). A. Habitus, left side. B. Habitus, right side. C. Head and basal antennomeres, front view. D. Antenna. E. Head and mesoscutum, dorsolateral view. F. Metasoma and ovipositor, dorsolateral view. G. Wings. H. Distal part of fore wing." figureDoi="http://doi.org/10.5281/zenodo.6854002" httpUri="https://zenodo.org/record/6854002/files/figure.png" pageId="5" pageNumber="525">Fig. 1D</figureCitation>
)—Flagellomeres starting from fifth moniliform (
<figureCitation id="1377F2FB6C70CA57EE600656FB27FE40" box="[1087,1164,364,386]" captionStart="FIGURES 1" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[151,1436,179,1913]" captionTargetId="figure-22@3.[151,1436,179,1917]" captionTargetPageId="3" captionText="FIGURES 1. Photos of Protoctonus groehni gen. et sp. nov. (female, holotype). A. Habitus, left side. B. Habitus, right side. C. Head and basal antennomeres, front view. D. Antenna. E. Head and mesoscutum, dorsolateral view. F. Metasoma and ovipositor, dorsolateral view. G. Wings. H. Distal part of fore wing." figureDoi="http://doi.org/10.5281/zenodo.6854002" httpUri="https://zenodo.org/record/6854002/files/figure.png" pageId="5" pageNumber="525">Fig. 1D</figureCitation>
). Radial (r, 3-SR, SR1) vein and metacarp (1-R1) almost reaching distal margin of fore wing (
<figureCitation id="1377F2FB6C70CA57E91006B3FC34FE5F" box="[847,927,393,415]" captionStart="FIGURES 1" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[151,1436,179,1913]" captionTargetId="figure-22@3.[151,1436,179,1917]" captionTargetPageId="3" captionText="FIGURES 1. Photos of Protoctonus groehni gen. et sp. nov. (female, holotype). A. Habitus, left side. B. Habitus, right side. C. Head and basal antennomeres, front view. D. Antenna. E. Head and mesoscutum, dorsolateral view. F. Metasoma and ovipositor, dorsolateral view. G. Wings. H. Distal part of fore wing." figureDoi="http://doi.org/10.5281/zenodo.6854002" httpUri="https://zenodo.org/record/6854002/files/figure.png" pageId="5" pageNumber="525">Figs 1G</figureCitation>
;
<figureCitation id="1377F2FB6C70CA57E9F606B3FC61FE5F" box="[937,970,393,414]" captionStart="FIGURES 2" captionStartId="5.[151,264,1386,1410]" captionTargetBox="[164,1421,868,1356]" captionTargetId="figure-758@5.[151,1435,850,1363]" captionTargetPageId="5" captionText="FIGURES 2. Figures of Protoctonus groehni gen. et sp. nov. (female, holotype). A. Fore wing. B. Ovipositor sheaths." figureDoi="http://doi.org/10.5281/zenodo.6854004" httpUri="https://zenodo.org/record/6854004/files/figure.png" pageId="5" pageNumber="525">2A</figureCitation>
). Ovipositor sheaths (
<figureCitation id="1377F2FB6C70CA57EEC606B3FB4EFE5F" box="[1177,1253,393,415]" captionStart="FIGURES 1" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[151,1436,179,1913]" captionTargetId="figure-22@3.[151,1436,179,1917]" captionTargetPageId="3" captionText="FIGURES 1. Photos of Protoctonus groehni gen. et sp. nov. (female, holotype). A. Habitus, left side. B. Habitus, right side. C. Head and basal antennomeres, front view. D. Antenna. E. Head and mesoscutum, dorsolateral view. F. Metasoma and ovipositor, dorsolateral view. G. Wings. H. Distal part of fore wing." figureDoi="http://doi.org/10.5281/zenodo.6854002" httpUri="https://zenodo.org/record/6854002/files/figure.png" pageId="5" pageNumber="525">Figs 1F</figureCitation>
;
<figureCitation id="1377F2FB6C70CA57EEB006B3FAA7FE5F" box="[1263,1292,393,414]" captionStart="FIGURES 2" captionStartId="5.[151,264,1386,1410]" captionTargetBox="[164,1421,868,1356]" captionTargetId="figure-758@5.[151,1435,850,1363]" captionTargetPageId="5" captionText="FIGURES 2. Figures of Protoctonus groehni gen. et sp. nov. (female, holotype). A. Fore wing. B. Ovipositor sheaths." figureDoi="http://doi.org/10.5281/zenodo.6854004" httpUri="https://zenodo.org/record/6854004/files/figure.png" pageId="5" pageNumber="525">2B</figureCitation>
) subtriangular, tapered towards apex and curved downwards, entirely covered by dense and short setae; sheath 2.6 × longer than its maximum basal width..........................................................................
<taxonomicName id="4C4C95FD6C70CA57EE9706FEFA97FE18" authority="Davidian &amp; Manukyan &amp; Belokobylskij, 2022" authorityName="Davidian &amp; Manukyan &amp; Belokobylskij" authorityYear="2022" box="[1224,1340,452,473]" class="Insecta" family="Braconidae" genus="Protoctonus" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="525" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B938326C6C70CA57EE9706FEFA97FE18" box="[1224,1340,452,473]" italics="true" pageId="5" pageNumber="525">Protoctonus</emphasis>
</taxonomicName>
<emphasis id="B938326C6C70CA57EF1D06FEFA37FE18" bold="true" box="[1346,1436,452,473]" pageId="5" pageNumber="525">
<taxonomicNameLabel id="A20B8F176C70CA57EF1D06FEFA37FE18" box="[1346,1436,452,473]" pageId="5" pageNumber="525" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BF3EE7E6C70CA57EAC806DBFA37FE37" blockId="5.[151,1437,364,826]" box="[151,1436,481,503]" pageId="5" pageNumber="525">
<keyLead id="30BD4D4B6C70CA57EAC806DBFA37FE37" box="[151,1436,481,503]" pageId="5" pageNumber="525">
- Antenna with 12
<emphasis id="B938326C6C70CA57EBDD06D8FE25FE36" box="[386,398,482,503]" italics="true" pageId="5" pageNumber="525"></emphasis>
16 antennomeres....................................................................... 2
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B8F6DB6C70CA57EAC806C4FA37FD67" pageId="5" pageNumber="525">
<paragraph id="8BF3EE7E6C70CA57EAC806C4FA36FDAD" blockId="5.[151,1437,364,826]" pageId="5" pageNumber="525">
<keyLead id="30BD4D4B6C70CA57EAC806C4FA36FDAD" pageId="5" pageNumber="525">
2. Petiole of metasoma short and wide, its length almost equal to width at spiracle level. Propodeum without areola and keels. —Antenna with 1415 antennomeres, moniliform. Ovipositor sheath straight, subtriangular shape, strongly tapered towards apex.........................................................................................................................................................................................
<taxonomicName id="4C4C95FD6C70CA57EF5A056CFA36FDAD" ID-CoL="N29" authority="Foerster" authorityName="Foerster" authorityYear="1862" box="[1285,1437,598,620]" class="Insecta" family="Braconidae" genus="Aclitus" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="525" phylum="Arthropoda" rank="genus">
<emphasis id="B938326C6C70CA57EF5A056CFAE3FDAA" box="[1285,1352,598,619]" italics="true" pageId="5" pageNumber="525">Aclitus</emphasis>
Foerster
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BF3EE7E6C70CA57EAC8054EFA37FD67" blockId="5.[151,1437,364,826]" pageId="5" pageNumber="525">
<keyLead id="30BD4D4B6C70CA57EAC8054EFA37FD67" pageId="5" pageNumber="525">- Petiole of metasoma elongate, its length distinctly larger than width at level of spiracles. Propodeum with delineated medial areola.............................................................................................. 3</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B8F6DB6C70CA57EAC80594FA37FCFB" pageId="5" pageNumber="525">
<paragraph id="8BF3EE7E6C70CA57EAC80594FA37FD23" blockId="5.[151,1437,364,826]" pageId="5" pageNumber="525">
<keyLead id="30BD4D4B6C70CA57EAC80594FA37FD23" pageId="5" pageNumber="525">
3. Ovipositor sheaths curved downwards, gradually tapered towards acuminate apex. Petiole of mesosoma almost parallel-sided behind prominent spiracular tubercles. Antenna with 1214 antennomeres, evenly thick.......... ..
<taxonomicName id="4C4C95FD6C70CA57EE9E05F6FA37FD23" ID-CoL="62J6T" authority="Mackauer" authorityName="Mackauer" authorityYear="1961" box="[1217,1436,716,738]" class="Insecta" family="Braconidae" genus="Calaphidius" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="525" phylum="Arthropoda" rank="genus">
<emphasis id="B938326C6C70CA57EE9E05F6FA9DFD20" box="[1217,1334,716,737]" italics="true" pageId="5" pageNumber="525">Calaphidius</emphasis>
Mackauer
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BF3EE7E6C70CA57EAC805D3FA37FCFB" blockId="5.[151,1437,364,826]" pageId="5" pageNumber="525">
<keyLead id="30BD4D4B6C70CA57EAC805D3FA37FCFB" pageId="5" pageNumber="525">
- Ovipositor sheaths straight or very weakly curved downwards, gradually tapered towards obtuse apex. Petiole of mesosoma distinctly concave behind prominent spiracular tubercles. Antenna with 1316 antennomeres, slender but weakly thickened apically......................................................................... ....
<taxonomicName id="4C4C95FD6C70CA57EE93041EFA37FCFB" authority="Mackauer" authorityName="Mackauer" authorityYear="1959" box="[1228,1436,804,826]" class="Insecta" family="Braconidae" genus="Falciconus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="525" phylum="Arthropoda" rank="genus">
<emphasis id="B938326C6C70CA57EE93041EFA9CFCF8" box="[1228,1335,804,825]" italics="true" pageId="5" pageNumber="525">Falciconus</emphasis>
Mackauer
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>