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<document id="30B8B17C4C4DE55E0FAB25316320EE57" ID="10.11646/zootaxa.4188.1.1" ID-DOI="10.11646/zootaxa.4188.1.1" ID-GBIF-Dataset="03a7cb89-747a-411a-855c-f46ad7df47b8" ID-ISSN="1175-5326" ID-Zenodo-Dep="165406" ID-ZooBank="AA85050E-7653-44BE-9330-AC617BFE6DF8" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1478593795583" checkinUser="plazi" docAuthor="Ng, Heok Hee &amp; Kottelat, Maurice" docDate="2016" docId="03E0BE0EFF8D5404FF58FF65FE53501B" docLanguage="en" docName="zootaxa.4188.1.1.pdf" docOrigin="Zootaxa 4188 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Glyptothorax famelicus Ng &amp; Kottelat, 2016, new species" docType="treatment" docVersion="11" lastPageNumber="24" masterDocId="FFD9C676FF995413FFCFFFF2FFE65323" masterDocTitle="The Glyptothorax of Sundaland: a revisionary study (Teleostei: Sisoridae)" masterLastPageNumber="92" masterPageNumber="1" pageNumber="21" updateTime="1698392689045" updateUser="plazi">
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<mods:title id="D3C6DC48184C0165A608CAE48B7F4CA5">The Glyptothorax of Sundaland: a revisionary study (Teleostei: Sisoridae)</mods:title>
</mods:titleInfo>
<mods:name id="55DB3AE7DB032E0FDB05A4DBD1751A9E" type="personal">
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<mods:namePart id="9D96373E02DCE898F2063D37219DB2DD">Ng, Heok Hee</mods:namePart>
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<mods:namePart id="F06533DC74868D3B3FFFA8F7DFDB0E61">Kottelat, Maurice</mods:namePart>
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<mods:title id="6E46C1F103C33F98421CB3304A1C16B3">Zootaxa</mods:title>
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<mods:date id="AA7E95466A093F0F0BC1129B62932A15">2016</mods:date>
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<mods:number id="D5AFD5206717A246C6F75831649E7132">4188</mods:number>
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<mods:identifier id="154379DB09E4F7F12A5045CED89A25DB" type="DOI">10.11646/zootaxa.4188.1.1</mods:identifier>
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<treatment id="03E0BE0EFF8D5404FF58FF65FE53501B" ID-DOI="http://doi.org/10.5281/zenodo.6063750" ID-GBIF-Taxon="124624014" ID-Zenodo-Dep="6063750" LSID="urn:lsid:plazi:treatment:03E0BE0EFF8D5404FF58FF65FE53501B" httpUri="http://treatment.plazi.org/id/03E0BE0EFF8D5404FF58FF65FE53501B" lastPageId="23" lastPageNumber="24" pageId="20" pageNumber="21">
<subSubSection id="C3535C93FF8D5407FF58FF65FF1E53F0" pageId="20" pageNumber="21" type="nomenclature">
<paragraph id="8BF60F18FF8D5407FF58FF65FDAC5391" blockId="20.[151,586,151,211]" box="[151,586,151,178]" pageId="20" pageNumber="21">
<heading id="D0BEB874FF8D5407FF58FF65FDAC5391" bold="true" box="[151,586,151,178]" fontSize="11" level="1" pageId="20" pageNumber="21" reason="1">
<emphasis id="B93DD30AFF8D5407FF58FF65FDAC5391" bold="true" box="[151,586,151,178]" pageId="20" pageNumber="21">
<taxonomicName id="4C49749BFF8D5407FF58FF65FE555392" box="[151,435,151,177]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="famelicus" status="sp. nov.">
<emphasis id="B93DD30AFF8D5407FF58FF65FE555392" bold="true" box="[151,435,151,177]" italics="true" pageId="20" pageNumber="21">Glyptothorax famelicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20E6E71FF8D5407FE75FF6AFDAC5391" box="[442,586,152,178]" pageId="20" pageNumber="21" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BF60F18FF8D5407FF58FF48FF1E53F0" blockId="20.[151,586,151,211]" box="[151,248,186,211]" pageId="20" pageNumber="21">
(
<figureCitation id="1372139DFF8D5407FF50FF48FF0953F0" box="[159,239,186,211]" captionStart="FIGURE 11" captionStartId="20.[151,250,1626,1648]" captionTargetBox="[151,1436,407,1604]" captionTargetId="figure@20.[151,1436,407,1604]" captionTargetPageId="20" captionText="FIGURE 11. Glyptothorax famelicus, USNM 193015, holotype, 73.3 mm SL; Sumatra: Danau Toba. Dorsal, lateral and ventral views." httpUri="https://zenodo.org/record/165417/files/figure.png" pageId="20" pageNumber="21">Fig. 11</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3535C93FF8D5407FF58FEF0FC075240" pageId="20" pageNumber="21" type="materials_examined">
<paragraph id="8BF60F18FF8D5407FF58FEF0FEF8521C" blockId="20.[151,1436,258,355]" pageId="20" pageNumber="21">
<materialsCitation id="3B210545FF8D5407FF58FEF0FC735238" ID-GBIF-Occurrence="1324948257" box="[151,917,258,283]" collectionCode="USNM" location="Sumatra" pageId="20" pageNumber="21" specimenCode="USNM 193015, 73.3" specimenCount="2" stateProvince="Sumatra" typeStatus="holotype">
<emphasis id="B93DD30AFF8D5407FF58FEF0FEAE5238" bold="true" box="[151,328,258,283]" pageId="20" pageNumber="21">
<typeStatus id="54F2B1BAFF8D5407FF58FEF0FF345238" box="[151,210,258,283]" pageId="20" pageNumber="21">Type</typeStatus>
material.
</emphasis>
<typeStatus id="54F2B1BAFF8D5407FE9CFEF0FE265238" box="[339,448,258,283]" pageId="20" pageNumber="21" type="holotype">Holotype</typeStatus>
:
<specimenCode id="DBEFA763FF8D5407FE02FEF1FD595238" box="[461,703,258,283]" pageId="20" pageNumber="21">
<collectionCode id="ED5897DDFF8D5407FE02FEF1FDF85238" box="[461,542,259,283]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="20" pageNumber="21">USNM</collectionCode>
193015,
<quantity id="4CB1A2FDFF8D5407FD41FEF1FD595238" box="[654,703,259,283]" metricMagnitude="-2" metricUnit="m" metricValue="7.33" pageId="20" pageNumber="21" unit="mm" value="73.3">73.3</quantity>
</specimenCode>
mm SL;
<collectingRegion id="498DC1FAFF8D5407FCFCFEF1FC735238" box="[819,917,259,283]" country="Indonesia" name="Sumatera Utara" pageId="20" pageNumber="21">Sumatra</collectingRegion>
</materialsCitation>
:
<materialsCitation id="3B210545FF8D5407FC6DFEF1FEF8521C" ID-GBIF-Occurrence="1324948271" collectorName="D. Silan" location="Sumatera Utara" pageId="20" pageNumber="21" specimenCount="1" stateProvince="Sumatera Utara" typeStatus="holotype">
<collectingRegion id="498DC1FAFF8D5407FC6DFEF1FBBE5239" box="[930,1112,259,283]" country="Indonesia" name="Sumatera Utara" pageId="20" pageNumber="21">Sumatera Utara</collectingRegion>
, Danau Toba at Parapat;
<collectorName id="26BC6ACEFF8D5407FA4EFEF1FF35521C" pageId="20" pageNumber="21">D. Silan</collectorName>
, 1937.
</materialsCitation>
</paragraph>
<paragraph id="8BF60F18FF8D5407FF08FEB9FC075240" blockId="20.[151,1436,258,355]" box="[199,993,330,355]" pageId="20" pageNumber="21">
<materialsCitation id="3B210545FF8D5407FF08FEB9FC075240" ID-GBIF-Occurrence="1324948261" box="[199,993,330,355]" collectionCode="USNM" pageId="20" pageNumber="21" specimenCode="USNM 410773" specimenCount="1" typeStatus="paratype">
<typeStatus id="54F2B1BAFF8D5407FF08FEB9FEDE5240" box="[199,312,331,355]" pageId="20" pageNumber="21" type="paratype">Paratypes</typeStatus>
.
<specimenCode id="DBEFA763FF8D5407FE8DFEB9FE085240" box="[322,494,331,355]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="20" pageNumber="21">
<collectionCode id="ED5897DDFF8D5407FE8DFEB9FE755240" box="[322,403,331,355]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="20" pageNumber="21">USNM</collectionCode>
410773
</specimenCode>
(6),
<quantity id="4CB1A2FDFF8D5407FDECFEB8FD205241" box="[547,710,330,355]" metricMagnitude="-2" metricUnit="m" metricValue="4.99" metricValueMax="7.43" metricValueMin="2.55" pageId="20" pageNumber="21" unit="mm" value="49.9" valueMax="74.3" valueMin="25.5">25.574.3 mm</quantity>
SL; data as for holotype.
</materialsCitation>
</paragraph>
</subSubSection>
<caption id="DF365F90FF8D5407FF58F9A8FEC055AC" httpUri="https://zenodo.org/record/165417/files/figure.png" pageId="20" pageNumber="21" targetBox="[151,1436,407,1604]" targetPageId="20">
<paragraph id="8BF60F18FF8D5407FF58F9A8FEC055AC" blockId="20.[151,1435,1626,1679]" pageId="20" pageNumber="21">
<emphasis id="B93DD30AFF8D5407FF58F9A8FEC4554C" bold="true" box="[151,290,1626,1648]" pageId="20" pageNumber="21">FIGURE 11.</emphasis>
<taxonomicName id="4C49749BFF8D5407FEE2F9A8FDF95553" box="[301,543,1626,1648]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="famelicus">
<emphasis id="B93DD30AFF8D5407FEE2F9A8FDF95553" box="[301,543,1626,1648]" italics="true" pageId="20" pageNumber="21">Glyptothorax famelicus</emphasis>
</taxonomicName>
, USNM 193015, holotype, 73.3 mm SL; Sumatra: Danau Toba. Dorsal, lateral and ventral views.
</paragraph>
</caption>
<subSubSection id="C3535C93FF8D5406FF08F944FAAB53DB" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21" type="diagnosis">
<paragraph id="8BF60F18FF8D5406FF08F944FAAB53DB" blockId="20.[151,1436,1718,2031]" lastBlockId="21.[151,1436,151,464]" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21">
<emphasis id="B93DD30AFF8D5407FF08F944FEA755EC" bold="true" box="[199,321,1718,1743]" pageId="20" pageNumber="21">Diagnosis.</emphasis>
<taxonomicName id="4C49749BFF8D5407FE85F944FDB555ED" box="[330,595,1718,1742]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="famelicus">
<emphasis id="B93DD30AFF8D5407FE85F944FDB555ED" box="[330,595,1718,1742]" italics="true" pageId="20" pageNumber="21">Glyptothorax famelicus</emphasis>
</taxonomicName>
differs from Sundaic congeners except for
<taxonomicName id="4C49749BFF8D5407FB89F945FB2555ED" box="[1094,1219,1718,1742]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="ketambe">
<emphasis id="B93DD30AFF8D5407FB89F945FB2555ED" box="[1094,1219,1718,1742]" italics="true" pageId="20" pageNumber="21">G. ketambe</emphasis>
</taxonomicName>
and
<taxonomicName id="4C49749BFF8D5407FB31F945FA9B55ED" box="[1278,1405,1718,1742]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="schmidti">
<emphasis id="B93DD30AFF8D5407FB31F945FA9B55ED" box="[1278,1405,1718,1742]" italics="true" pageId="20" pageNumber="21">G. schmidti</emphasis>
</taxonomicName>
in having a uniformly dark body with prominent pale mid-dorsal and midlateral stripes (vs. with either uniform body with or without dark spots or a mottled body, all without prominent pale mid-dorsal and midlateral stripes). It can be distinguished from
<taxonomicName id="4C49749BFF8D5407FE5AF8D1FDF75418" box="[405,529,1827,1851]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="ketambe">
<emphasis id="B93DD30AFF8D5407FE5AF8D1FDF75418" box="[405,529,1827,1851]" italics="true" pageId="20" pageNumber="21">G. ketambe</emphasis>
</taxonomicName>
in having a shorter head (22.623.6% SL vs. 24.025.5), a more slender caudal peduncle (depth 5.87.3% SL vs. 7.68.9) and more preanal vertebrae (1920 vs. 1617), and from
<taxonomicName id="4C49749BFF8D5407FACFF8B5FA99547D" box="[1280,1407,1862,1886]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="schmidti">
<emphasis id="B93DD30AFF8D5407FACFF8B5FA99547D" box="[1280,1407,1862,1886]" italics="true" pageId="20" pageNumber="21">G. schmidti</emphasis>
</taxonomicName>
in having a longer thoracic adhesive apparatus (reaching to or just beyond vs. not reaching the level of the last pectoral-fin-ray bases; 63.471.0% HL vs. 55.160.0). The following unique combination of the characters further distinguishes
<taxonomicName id="4C49749BFF8D5407FEFEF841FE5754E8" box="[305,433,1971,1995]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="famelicu">
<emphasis id="B93DD30AFF8D5407FEFEF841FE5754E8" box="[305,433,1971,1995]" italics="true" pageId="20" pageNumber="21">G. famelicu</emphasis>
</taxonomicName>
s from its Sundaic congeners: eye diameter 78% HL; interorbital distance 2831% HL; anteromedial striae in thoracic adhesive apparatus absent; predorsal length 32.135.2% SL; dorsal-fin spine length 8.914.0% SL; margin of dorsal fin concave, smooth posterior margin of dorsal-fin spine; pectoral-fin spine length 14.015.6% SL; adipose-fin base length 11.412.8% SL; straight dorsoposterior margin of adipose fin; body depth at anus 11.414.3% SL; post-adipose distance 19.122.0% SL; and caudal peduncle length 19.421.2% SL.
</paragraph>
</subSubSection>
<subSubSection id="C3535C93FF8C5405FF08FEF6FC5B57E6" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22" type="description">
<paragraph id="8BF60F18FF8C5406FF08FEF6FDD052F3" blockId="21.[151,1436,151,464]" pageId="21" pageNumber="22">
<emphasis id="B93DD30AFF8C5406FF08FEF6FEBF523E" bold="true" box="[199,345,260,285]" pageId="21" pageNumber="22">Description.</emphasis>
Morphometric data in
<tableCitation id="C6CB3AA3FF8C5406FD94FEF6FD57523F" box="[603,689,260,285]" captionStart="TABLE 4" captionStartId="21.[151,239,512,535]" captionTargetBox="[159,1282,562,1902]" captionTargetPageId="21" captionText="TABLE 4. Morphometric data for Glyptothorax famelicus (n = 7)." httpUri="http://table.plazi.org/id/DF365F90FF8C5406FF58FDF2FCA85134" pageId="21" pageNumber="22" tableUuid="DF365F90FF8C5406FF58FDF2FCA85134">Table 4</tableCitation>
. Head depressed; body very slender, subcylindrical. Dorsal profile rising evenly from tip of snout to origin of dorsal fin, then sloping gently ventrally from origin of dorsal fin to end of caudal peduncle. Ventral profile straight to anal-fin base, then sloping gently dorsally from anal-fin base to end of caudal peduncle. Anus and urogenital openings located at vertical through middle of adpressed pelvic fin. Skin tuberculate, with tubercles of even size on sides of body. Lateral line complete and midlateral. Vertebrae 19+18=37 (1), 19+19=38* (4) or 20+18=38 (2).
</paragraph>
<caption id="DF365F90FF8C5406FF58FDF2FCA85134" ID-Table-UUID="DF365F90FF8C5406FF58FDF2FCA85134" box="[151,846,511,535]" httpUri="http://table.plazi.org/id/DF365F90FF8C5406FF58FDF2FCA85134" pageId="21" pageNumber="22" targetBox="[159,1282,562,1902]" targetIsTable="true" targetPageId="21">
<paragraph id="8BF60F18FF8C5406FF58FDF2FCA85134" blockId="21.[151,846,511,535]" box="[151,846,511,535]" pageId="21" pageNumber="22">
<emphasis id="B93DD30AFF8C5406FF58FDF2FEEF5134" bold="true" box="[151,265,512,535]" pageId="21" pageNumber="22">TABLE 4.</emphasis>
Morphometric data for
<taxonomicName id="4C49749BFF8C5406FDC5FE0DFCE35134" box="[522,773,511,535]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="famelicus">
<emphasis id="B93DD30AFF8C5406FDC5FE0DFCE35134" box="[522,773,511,535]" italics="true" pageId="21" pageNumber="22">Glyptothorax famelicus</emphasis>
</taxonomicName>
(n=7).
</paragraph>
</caption>
<paragraph id="8BF60F18FF8C5406FD4AFDC0FB07544D" pageId="21" pageNumber="22">
<table id="F949FDB8FF8CABECFF50FDC0FAE4544D" box="[159,1282,562,1902]" gridcols="4" gridrows="12" pageId="21" pageNumber="22">
<tr id="35790D5AFF8CABECFF50FDC0FAE45144" box="[159,1282,562,615]" gridrow="0" pageId="21" pageNumber="22" rowspan-0="1">
<th id="76A86426FF8CABECFD4AFDC0FCC65144" box="[645,800,562,615]" gridcol="1" gridrow="0" pageId="21" pageNumber="22">Holotype USNM 193015</th>
<th id="76A86426FF8CABECFC5FFDC0FC105144" box="[912,1014,562,615]" gridcol="2" gridrow="0" pageId="21" pageNumber="22">Range</th>
<th id="76A86426FF8CABECFB54FDC0FAE45144" box="[1179,1282,562,615]" gridcol="3" gridrow="0" pageId="21" pageNumber="22">Mean±SD</th>
</tr>
<tr id="35790D5AFF8CABECFF50FD72FAE4519C" box="[159,1282,640,703]" gridrow="1" pageId="21" pageNumber="22" rowspan-3="1">
<th id="76A86426FF8CABECFF50FD72FE3B519C" box="[159,477,640,703]" gridcol="0" gridrow="1" pageId="21" pageNumber="22">Standard length (mm) %SL</th>
<td id="76A86426FF8CABECFD4AFD72FCC6519C" box="[645,800,640,703]" gridcol="1" gridrow="1" pageId="21" pageNumber="22">73.3</td>
<td id="76A86426FF8CABECFC5FFD72FC10519C" box="[912,1014,640,703]" gridcol="2" gridrow="1" pageId="21" pageNumber="22">25.574.3</td>
</tr>
<tr id="35790D5AFF8CABECFF50FD21FAE45018" box="[159,1282,723,827]" gridrow="2" pageId="21" pageNumber="22">
<th id="76A86426FF8CABECFF50FD21FE3B5018" box="[159,477,723,827]" gridcol="0" gridrow="2" pageId="21" pageNumber="22">Predorsal length Preanal length Prepelvic length</th>
<td id="76A86426FF8CABECFD4AFD21FCC65018" box="[645,800,723,827]" gridcol="1" gridrow="2" pageId="21" pageNumber="22">35.2 65.6 47.2</td>
<td id="76A86426FF8CABECFC5FFD21FC105018" box="[912,1014,723,827]" gridcol="2" gridrow="2" pageId="21" pageNumber="22">32.135.2 64.466.2 45.249.8</td>
<td id="76A86426FF8CABECFB54FD21FAE45018" box="[1179,1282,723,827]" gridcol="3" gridrow="2" pageId="21" pageNumber="22">34.1±1.38 65.1±0.76 46.7±1.87</td>
</tr>
<tr id="35790D5AFF8CABECFF50FCBDFAE45094" box="[159,1282,847,951]" gridrow="3" pageId="21" pageNumber="22">
<th id="76A86426FF8CABECFF50FCBDFE3B5094" box="[159,477,847,951]" gridcol="0" gridrow="3" pageId="21" pageNumber="22">Prepectoral length Length of dorsal-fin base Dorsal-fin spine length</th>
<td id="76A86426FF8CABECFD4AFCBDFCC65094" box="[645,800,847,951]" gridcol="1" gridrow="3" pageId="21" pageNumber="22">19.0 12.6 12.0</td>
<td id="76A86426FF8CABECFC5FFCBDFC105094" box="[912,1014,847,951]" gridcol="2" gridrow="3" pageId="21" pageNumber="22">18.921.8 10.913.2 8.914.0</td>
<td id="76A86426FF8CABECFB54FCBDFAE45094" box="[1179,1282,847,951]" gridcol="3" gridrow="3" pageId="21" pageNumber="22">20.1±1.24 12.3±0.91 12.0±2.07</td>
</tr>
<tr id="35790D5AFF8CABECFF50FC39FAE45710" box="[159,1282,971,1075]" gridrow="4" pageId="21" pageNumber="22">
<th id="76A86426FF8CABECFF50FC39FE3B5710" box="[159,477,971,1075]" gridcol="0" gridrow="4" pageId="21" pageNumber="22">Length of anal-fin base Pelvic-fin length Pectoral-fin length</th>
<td id="76A86426FF8CABECFD4AFC39FCC65710" box="[645,800,971,1075]" gridcol="1" gridrow="4" pageId="21" pageNumber="22">14.6 16.6 20.3</td>
<td id="76A86426FF8CABECFC5FFC39FC105710" box="[912,1014,971,1075]" gridcol="2" gridrow="4" pageId="21" pageNumber="22">14.616.9 15.117.6 20.322.1</td>
<td id="76A86426FF8CABECFB54FC39FAE45710" box="[1179,1282,971,1075]" gridcol="3" gridrow="4" pageId="21" pageNumber="22">15.5±1.12 16.4±0.89 21.1±0.65</td>
</tr>
<tr id="35790D5AFF8CABECFF50FBB5FAE4578C" box="[159,1282,1095,1199]" gridrow="5" pageId="21" pageNumber="22">
<th id="76A86426FF8CABECFF50FBB5FE3B578C" box="[159,477,1095,1199]" gridcol="0" gridrow="5" pageId="21" pageNumber="22">Pectoral-fin spine length Caudal-fin length Length of adipose-fin base</th>
<td id="76A86426FF8CABECFD4AFBB5FCC6578C" box="[645,800,1095,1199]" gridcol="1" gridrow="5" pageId="21" pageNumber="22">14.2 25.8 12.8</td>
<td id="76A86426FF8CABECFC5FFBB5FC10578C" box="[912,1014,1095,1199]" gridcol="2" gridrow="5" pageId="21" pageNumber="22">14.015.6 25.828.2 11.412.8</td>
<td id="76A86426FF8CABECFB54FBB5FAE4578C" box="[1179,1282,1095,1199]" gridcol="3" gridrow="5" pageId="21" pageNumber="22">15.0±0.79 26.9±0.97 12.1±0.63</td>
</tr>
<tr id="35790D5AFF8CABECFF50FB31FAE45608" box="[159,1282,1219,1323]" gridrow="6" pageId="21" pageNumber="22">
<th id="76A86426FF8CABECFF50FB31FE3B5608" box="[159,477,1219,1323]" gridcol="0" gridrow="6" pageId="21" pageNumber="22">Dorsal to adipose distance Post-adipose distance Length of caudal peduncle</th>
<td id="76A86426FF8CABECFD4AFB31FCC65608" box="[645,800,1219,1323]" gridcol="1" gridrow="6" pageId="21" pageNumber="22">25.0 19.9 20.1</td>
<td id="76A86426FF8CABECFC5FFB31FC105608" box="[912,1014,1219,1323]" gridcol="2" gridrow="6" pageId="21" pageNumber="22">20.027.3 19.122.0 19.421.2</td>
<td id="76A86426FF8CABECFB54FB31FAE45608" box="[1179,1282,1219,1323]" gridcol="3" gridrow="6" pageId="21" pageNumber="22">24.2±2.64 20.5±1.09 20.2±0.67</td>
</tr>
<tr id="35790D5AFF8CABECFF50FACDFAE45684" box="[159,1282,1343,1447]" gridrow="7" pageId="21" pageNumber="22">
<th id="76A86426FF8CABECFF50FACDFE3B5684" box="[159,477,1343,1447]" gridcol="0" gridrow="7" pageId="21" pageNumber="22">Depth of caudal peduncle Body depth at anus Body depth at dorsal-fin origin</th>
<td id="76A86426FF8CABECFD4AFACDFCC65684" box="[645,800,1343,1447]" gridcol="1" gridrow="7" pageId="21" pageNumber="22">6.5 11.9 14.0</td>
<td id="76A86426FF8CABECFC5FFACDFC105684" box="[912,1014,1343,1447]" gridcol="2" gridrow="7" pageId="21" pageNumber="22">5.87.3 11.414.3 14.017.8</td>
<td id="76A86426FF8CABECFB54FACDFAE45684" box="[1179,1282,1343,1447]" gridcol="3" gridrow="7" pageId="21" pageNumber="22">6.5±0.53 12.9±1.27 15.7±1.39</td>
</tr>
<tr id="35790D5AFF8CABECFF50FA49FAE45500" box="[159,1282,1467,1571]" gridrow="8" pageId="21" pageNumber="22">
<th id="76A86426FF8CABECFF50FA49FE3B5500" box="[159,477,1467,1571]" gridcol="0" gridrow="8" pageId="21" pageNumber="22">Head length Head width Head depth</th>
<td id="76A86426FF8CABECFD4AFA49FCC65500" box="[645,800,1467,1571]" gridcol="1" gridrow="8" pageId="21" pageNumber="22">23.1 18.4 13.2</td>
<td id="76A86426FF8CABECFC5FFA49FC105500" box="[912,1014,1467,1571]" gridcol="2" gridrow="8" pageId="21" pageNumber="22">22.623.6 16.720.0 12.514.0</td>
<td id="76A86426FF8CABECFB54FA49FAE45500" box="[1179,1282,1467,1571]" gridcol="3" gridrow="8" pageId="21" pageNumber="22">23.1±0.48 18.8±1.30 13.3±0.54</td>
</tr>
<tr id="35790D5AFF8CABECFF50F9C5FAE455BC" box="[159,1282,1591,1695]" gridrow="9" pageId="21" pageNumber="22">
<th id="76A86426FF8CABECFF50F9C5FE3B55BC" box="[159,477,1591,1695]" gridcol="0" gridrow="9" pageId="21" pageNumber="22">%HL Snout length Interorbital distance</th>
<td id="76A86426FF8CABECFD4AF9C5FCC655BC" box="[645,800,1591,1695]" gridcol="1" gridrow="9" pageId="21" pageNumber="22">49 28</td>
<td id="76A86426FF8CABECFC5FF9C5FC1055BC" box="[912,1014,1591,1695]" gridcol="2" gridrow="9" pageId="21" pageNumber="22">4552 2831</td>
<td id="76A86426FF8CABECFB54F9C5FAE455BC" box="[1179,1282,1591,1695]" gridcol="3" gridrow="9" pageId="21" pageNumber="22">49±2.5 29±1.0</td>
</tr>
<tr id="35790D5AFF8CABECFF50F941FAE45438" box="[159,1282,1715,1819]" gridrow="10" pageId="21" pageNumber="22">
<th id="76A86426FF8CABECFF50F941FE3B5438" box="[159,477,1715,1819]" gridcol="0" gridrow="10" pageId="21" pageNumber="22">Eye diameter Nasal barbel length Maxillary barbel length</th>
<td id="76A86426FF8CABECFD4AF941FCC65438" box="[645,800,1715,1819]" gridcol="1" gridrow="10" pageId="21" pageNumber="22">8 23 102</td>
<td id="76A86426FF8CABECFC5FF941FC105438" box="[912,1014,1715,1819]" gridcol="2" gridrow="10" pageId="21" pageNumber="22">78 1924 88102</td>
<td id="76A86426FF8CABECFB54F941FAE45438" box="[1179,1282,1715,1819]" gridcol="3" gridrow="10" pageId="21" pageNumber="22">8±0.5 22±2.1 95±6.0</td>
</tr>
<tr id="35790D5AFF8CABECFF50F8DDFAE4544D" box="[159,1282,1839,1902]" gridrow="11" pageId="21" pageNumber="22">
<th id="76A86426FF8CABECFF50F8DDFE3B544D" box="[159,477,1839,1902]" gridcol="0" gridrow="11" pageId="21" pageNumber="22">Inner mandibular barbel length Outer mandibular barbel length</th>
<td id="76A86426FF8CABECFD4AF8DDFCC6544D" box="[645,800,1839,1902]" gridcol="1" gridrow="11" pageId="21" pageNumber="22">33 54</td>
<td id="76A86426FF8CABECFC5FF8DDFC10544D" box="[912,1014,1839,1902]" gridcol="2" gridrow="11" pageId="21" pageNumber="22">3136 4457</td>
<td id="76A86426FF8CABECFB54F8DDFAE4544D" box="[1179,1282,1839,1902]" gridcol="3" gridrow="11" pageId="21" pageNumber="22">33±1.4 51±5.3</td>
</tr>
</table>
</paragraph>
<paragraph id="8BF60F18FF8C5405FF08F841FF3553DB" blockId="21.[151,1436,1971,2032]" lastBlockId="22.[151,1437,151,2013]" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22">Head depressed and moderately broad, triangular when viewed laterally. Snout prominent. Anterior and posterior nares large and separated only by base of nasal barbel. Gill opening broad, extending from ventral margin of post-temporal to isthmus. First branchial arch with 2+7 (3) or 2+8* (4) rakers. Bony elements of dorsal surface of head covered with thick, tuberculate skin. Eye ovoid, horizontal axis longest; located entirely in dorsal half of head.</paragraph>
<paragraph id="8BF60F18FF8F5405FF08FEF6FD9E52AB" blockId="22.[151,1437,151,2013]" pageId="22" pageNumber="23">Barbels in four pairs. Maxillary barbel long and slender, extending to middle of pectoral-fin base. Nasal barbel slender, extending to midway between its base and anterior orbital margin. Inner mandibular-barbel extending to midway between its base and that of pectoral spine. Outer mandibular barbel extending to two-thirds of distance between its base and that of pectoral spine.</paragraph>
<paragraph id="8BF60F18FF8F5405FF08FE66FF13513B" blockId="22.[151,1437,151,2013]" pageId="22" pageNumber="23">Mouth inferior, premaxillary tooth band partially (approximately half) exposed when mouth is closed. Oral teeth small and villiform, in irregular rows on all tooth-bearing surfaces. Premaxillary teeth appearing in single broad semilunate band. Dentary teeth in a single crescentic band, consisting of two separate halves tightly bound at midline.</paragraph>
<paragraph id="8BF60F18FF8F5405FF08FDD6FED851A6" blockId="22.[151,1437,151,2013]" pageId="22" pageNumber="23">
Thoracic adhesive apparatus consisting of keratinized striae in an ovate field extending from isthmus to or just beyond posterior limit of pectoral-fin base (
<figureCitation id="1372139DFF8F5405FD5EFDB5FD305143" box="[657,726,583,608]" captionStart="FIGURE 3" captionStartId="6.[151,250,1784,1806]" captionTargetBox="[228,1361,193,1761]" captionTargetId="figure@6.[228,1362,193,1762]" captionTargetPageId="6" captionText="FIGURE 3. Thoracic adhesive apparatus of: a. Glyptothorax amnestus, ZRC 54231, paratype, 51.2 mm SL; b. G. decussatus, MZB 17216, holotype, 74.5 mm SL; c. G. exodon, BMNH 1982.3. 29.190, paratype, 63.2 mm SL; d. G. famelicus, USNM 193015, holotype, 73.3 mm SL; e. G. f u s c u s, ZRC 42123, 54.3 mm SL; f. G. keluk, MZB 17217, holotype, 52.5 mm SL; g. G. ketambe, MZB 8694, holotype, 100.0 mm SL; h. G. major, ZRC 38761, 59.1 mm SL; i. G. nieuwenhuisi, MZB 16475, 61.0 mm SL; j. G. pictus, MZB 17218, holotype, 54.2 mm SL; k. G. platypogon, ZRC 48376, 56.3 mm SL; l. G. platypogonides, UF 166626, 52.4 mm SL; m. G. plectilis, ZRC 46420, paratype, 44.3 mm SL; n. G. prashadi, CMK 25369, 50.7 mm SL; o. G. robustus, ZRC 48377, 59.9 mm SL; p. G. schmidti, CMK 4536, 74.6 mm SL; q. G. stibaros, MZB 17215, holotype, 55.3 mm SL. Illustrations not to scale." httpUri="https://zenodo.org/record/266202/files/figure.png" pageId="22" pageNumber="23">Fig. 3</figureCitation>
d). Anteromedial striae absent. Chevron-shaped medial pit on posterior third.
</paragraph>
<paragraph id="8BF60F18FF8F5405FF08FD7DFEC850EB" blockId="22.[151,1437,151,2013]" pageId="22" pageNumber="23">Dorsal fin located above anterior third of body, with I,6 (7) rays; fin margin concave; spine short and straight, smooth on anterior and posterior margins. Adipose fin with anterior margin straight or slightly concave and posterior margin straight. Caudal fin strongly forked, with lower lobe slightly longer than upper lobe and i,7,8,i (7) principal rays. Procurrent rays symmetrical and extending only slightly anterior to fin base. Anal-fin base vertically opposite adipose-fin base. Anal fin with straight anterior margin and straight or slightly concave posterior margin; with iv,8* (6) or iv,9 (2) rays. Pelvic-fin origin at vertical through posterior limit of dorsal-fin base. Pelvic fin with slightly convex margin and i,5 (7) rays; tip of adpressed fin not reaching anal-fin origin. Pectoral fin with I,7,i* (4), I,8 (1) or I,8,i (2) rays; posterior fin margin slightly concave; anterior spine margin smooth, posterior margin with 10 serrations.</paragraph>
<paragraph id="8BF60F18FF8F5405FF08FC26FC5B57E6" blockId="22.[151,1437,151,2013]" pageId="22" pageNumber="23">
<emphasis id="B93DD30AFF8F5405FF08FC26FEA950CE" bold="true" box="[199,335,980,1005]" pageId="22" pageNumber="23">Coloration.</emphasis>
In 70% ethanol: Dorsal and lateral surfaces of head, and body brown, fading to beige on ventral surfaces. A faint thin, lighter brown mid-dorsal stripe extending from base of last dorsal-fin ray to origin of adipose fin. Laterosensory pores along lateral line rimmed in beige, imparting appearance of a diffuse lighter brown midlateral line. All fins with brown rays, and diffuse melanophores on membranes. Pectoral and pelvic fins with brown on base of fin rays and hyaline posterior margin. Anal fin with brown base and hyaline posteroventral corner. Adipose fin brown, with hyaline distal margin. Caudal fin brown, with tip of lobes hyaline. Maxillary and nasal barbels brown dorsally, beige ventrally. Mandibular barbels beige.
</paragraph>
</subSubSection>
<subSubSection id="C3535C93FF8F5404FF08FB3DFE53501B" lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23" type="distribution">
<paragraph id="8BF60F18FF8F5405FF08FB3DFDA6562E" blockId="22.[151,1437,151,2013]" pageId="22" pageNumber="23">
<emphasis id="B93DD30AFF8F5405FF08FB3DFE8657CB" bold="true" box="[199,352,1231,1256]" pageId="22" pageNumber="23">Distribution.</emphasis>
<taxonomicName id="4C49749BFF8F5405FEA7FB22FD9757CB" box="[360,625,1232,1256]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="famelicus">
<emphasis id="B93DD30AFF8F5405FEA7FB22FD9757CB" box="[360,625,1232,1256]" italics="true" pageId="22" pageNumber="23">Glyptothorax famelicus</emphasis>
</taxonomicName>
is only known from the
<typeStatus id="54F2B1BAFF8F5405FC44FB22FC5D57CB" box="[907,955,1232,1256]" pageId="22" pageNumber="23">type</typeStatus>
series collected in the Danau (Lake) Toba drainage in northern
<collectingRegion id="498DC1FAFF8F5405FE4EFB06FE38562F" box="[385,478,1268,1292]" country="Indonesia" name="Sumatera Utara" pageId="22" pageNumber="23">Sumatra</collectingRegion>
(
<figureCitation id="1372139DFF8F5405FE22FB06FDD4562E" box="[493,562,1268,1293]" captionStart="FIGURE 5" captionStartId="9.[151,250,1990,2012]" captionTargetBox="[170,1433,949,1972]" captionTargetId="figure@9.[170,1433,949,1972]" captionTargetPageId="9" captionText="FIGURE 5. Map showing collection localities of Glyptothorax amnestus, G. decussatus, G. exodon and G. famelicus." httpUri="https://zenodo.org/record/165411/files/figure.png" pageId="22" pageNumber="23">Fig. 5</figureCitation>
).
</paragraph>
<paragraph id="8BF60F18FF8F5405FF08FAE5FE1854FE" blockId="22.[151,1437,151,2013]" pageId="22" pageNumber="23">
<emphasis id="B93DD30AFF8F5405FF08FAE5FE8A5613" bold="true" box="[199,364,1303,1328]" pageId="22" pageNumber="23">Comparisons.</emphasis>
<taxonomicName id="4C49749BFF8F5405FEB8FAEAFD655613" box="[375,643,1304,1328]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="famelicus">
<emphasis id="B93DD30AFF8F5405FEB8FAEAFD655613" box="[375,643,1304,1328]" italics="true" pageId="22" pageNumber="23">Glyptothorax famelicus</emphasis>
</taxonomicName>
further differs from all Sundaic congeners except for
<taxonomicName id="4C49749BFF8F5405FB30FAEAFA955613" box="[1279,1395,1304,1328]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="exodon">
<emphasis id="B93DD30AFF8F5405FB30FAEAFA955613" box="[1279,1395,1304,1328]" italics="true" pageId="22" pageNumber="23">G. exodon</emphasis>
</taxonomicName>
,
<taxonomicName id="4C49749BFF8F5405FA4BFAEAFF345677" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="keluk">
<emphasis id="B93DD30AFF8F5405FA4BFAEAFF345677" italics="true" pageId="22" pageNumber="23">G. keluk</emphasis>
</taxonomicName>
,
<taxonomicName id="4C49749BFF8F5405FF2FFACEFEBA5677" box="[224,348,1340,1364]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="ketambe">
<emphasis id="B93DD30AFF8F5405FF2FFACEFEBA5677" box="[224,348,1340,1364]" italics="true" pageId="22" pageNumber="23">G. ketambe</emphasis>
</taxonomicName>
,
<taxonomicName id="4C49749BFF8F5405FEA5FACEFDD25677" box="[362,564,1340,1364]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="platypogonides">
<emphasis id="B93DD30AFF8F5405FEA5FACEFDD25677" box="[362,564,1340,1364]" italics="true" pageId="22" pageNumber="23">G. platypogonides</emphasis>
</taxonomicName>
and
<taxonomicName id="4C49749BFF8F5405FDA5FACEFD015677" box="[618,743,1340,1364]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="schmidti">
<emphasis id="B93DD30AFF8F5405FDA5FACEFD015677" box="[618,743,1340,1364]" italics="true" pageId="22" pageNumber="23">G. schmidti</emphasis>
</taxonomicName>
in having a more slender body (depth at anus 11.414.3% SL vs. 14.523.1) and from all Sundaic congeners except
<taxonomicName id="4C49749BFF8F5405FD32FA92FC8B565B" box="[765,877,1376,1400]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="exodon">
<emphasis id="B93DD30AFF8F5405FD32FA92FC8B565B" box="[765,877,1376,1400]" italics="true" pageId="22" pageNumber="23">G. exodon</emphasis>
</taxonomicName>
,
<taxonomicName id="4C49749BFF8F5405FCB3FA92FBC8565B" box="[892,1070,1376,1400]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="nieuwenhuisi">
<emphasis id="B93DD30AFF8F5405FCB3FA92FBC8565B" box="[892,1070,1376,1400]" italics="true" pageId="22" pageNumber="23">G. nieuwenhuisi</emphasis>
</taxonomicName>
,
<taxonomicName id="4C49749BFF8F5405FBF2FA92FAE1565B" box="[1085,1287,1376,1400]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="platypogonides">
<emphasis id="B93DD30AFF8F5405FBF2FA92FAE1565B" box="[1085,1287,1376,1400]" italics="true" pageId="22" pageNumber="23">G. platypogonides</emphasis>
</taxonomicName>
,
<emphasis id="B93DD30AFF8F5405FAD9FA92FA77565B" box="[1302,1425,1376,1400]" italics="true" pageId="22" pageNumber="23">G. ro b u s t u s</emphasis>
,
<taxonomicName id="4C49749BFF8F5405FF58FA76FEF256BF" box="[151,276,1412,1436]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="schmidti">
<emphasis id="B93DD30AFF8F5405FF58FA76FEF256BF" box="[151,276,1412,1436]" italics="true" pageId="22" pageNumber="23">G. schmidti</emphasis>
</taxonomicName>
and
<emphasis id="B93DD30AFF8F5405FE86FA76FE5A56BF" box="[329,444,1412,1436]" italics="true" pageId="22" pageNumber="23">G. s t i b a ro s</emphasis>
in having a more slender caudal peduncle (depth 5.87.3% SL vs. 7.411.4); it is further distinguished from
<taxonomicName id="4C49749BFF8F5405FEB9FA5AFE0F56E3" box="[374,489,1448,1472]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="exodon">
<emphasis id="B93DD30AFF8F5405FEB9FA5AFE0F56E3" box="[374,489,1448,1472]" italics="true" pageId="22" pageNumber="23">G. exodon</emphasis>
</taxonomicName>
and
<taxonomicName id="4C49749BFF8F5405FDEAFA5AFD1456E3" box="[549,754,1448,1472]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="platypogonides">
<emphasis id="B93DD30AFF8F5405FDEAFA5AFD1456E3" box="[549,754,1448,1472]" italics="true" pageId="22" pageNumber="23">G. platypogonides</emphasis>
</taxonomicName>
in having a smaller eye (diameter 78% HL vs. 1014), a shorter dorsal-fin spine (8.914.0% SL vs. 15.020.0) and a shorter pectoral-fin spine (14.015.6% SL vs. 16.2 22.7), and from
<taxonomicName id="4C49749BFF8F5405FE83FA02FDF0552B" box="[332,534,1520,1544]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="platypogonides">
<emphasis id="B93DD30AFF8F5405FE83FA02FDF0552B" box="[332,534,1520,1544]" italics="true" pageId="22" pageNumber="23">G. platypogonides</emphasis>
</taxonomicName>
in having a smaller predorsal length (32.135.2% SL vs. 36.539.1). Besides
<taxonomicName id="4C49749BFF8F5405FA4BFA02FF12550F" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="ketambe">
<emphasis id="B93DD30AFF8F5405FA4BFA02FF12550F" italics="true" pageId="22" pageNumber="23">G. ketambe</emphasis>
</taxonomicName>
,
<emphasis id="B93DD30AFF8F5405FEC4F9E6FE68550F" box="[267,398,1556,1580]" italics="true" pageId="22" pageNumber="23">G. s c h m i d t i</emphasis>
(whose differences with
<taxonomicName id="4C49749BFF8F5405FD00F9E6FC84550F" box="[719,866,1556,1580]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="famelicus">
<emphasis id="B93DD30AFF8F5405FD00F9E6FC84550F" box="[719,866,1556,1580]" italics="true" pageId="22" pageNumber="23">G. famelicus</emphasis>
</taxonomicName>
are already outlined in the diagnosis) and
<taxonomicName id="4C49749BFF8F5405FA4BF9E6FEA45573" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="platypogonides">
<emphasis id="B93DD30AFF8F5405FA4BF9E6FEA45573" italics="true" pageId="22" pageNumber="23">G. platypogonides</emphasis>
</taxonomicName>
(compared previously), there are five other congeners known from
<collectingRegion id="498DC1FAFF8F5405FBBDF9CAFB325573" box="[1138,1236,1592,1616]" country="Indonesia" name="Sumatera Utara" pageId="22" pageNumber="23">Sumatra</collectingRegion>
:
<taxonomicName id="4C49749BFF8F5405FB2BF9CAFA965573" box="[1252,1392,1592,1616]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="amnestus">
<emphasis id="B93DD30AFF8F5405FB2BF9CAFA965573" box="[1252,1392,1592,1616]" italics="true" pageId="22" pageNumber="23">G. amnestus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C49749BFF8F5405FA4BF9CAFF3B5557" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="fuscus">
<emphasis id="B93DD30AFF8F5405FA4BF9CAFF3B5557" italics="true" pageId="22" pageNumber="23">G. fuscus</emphasis>
</taxonomicName>
,
<emphasis id="B93DD30AFF8F5405FF23F9AEFEA25557" box="[236,324,1628,1652]" italics="true" pageId="22" pageNumber="23">G. ke l u k</emphasis>
, and
<taxonomicName id="4C49749BFF8F5405FE47F9AEFDE45557" box="[392,514,1628,1652]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="plectilis">
<emphasis id="B93DD30AFF8F5405FE47F9AEFDE45557" box="[392,514,1628,1652]" italics="true" pageId="22" pageNumber="23">G. plectilis</emphasis>
</taxonomicName>
.
<taxonomicName id="4C49749BFF8F5405FDDEF9AEFCFD5557" box="[529,795,1628,1652]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="famelicus">
<emphasis id="B93DD30AFF8F5405FDDEF9AEFCFD5557" box="[529,795,1628,1652]" italics="true" pageId="22" pageNumber="23">Glyptothorax famelicus</emphasis>
</taxonomicName>
further differs from all of these five species in having a smaller predorsal length (32.135.2% SL vs. 35.342.7), from
<taxonomicName id="4C49749BFF8F5405FCA4F972FC1255BB" box="[875,1012,1664,1688]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="amnestus">
<emphasis id="B93DD30AFF8F5405FCA4F972FC1255BB" box="[875,1012,1664,1688]" italics="true" pageId="22" pageNumber="23">G. amnestus</emphasis>
</taxonomicName>
in lacking (vs. having) anteromedial striae in the thoracic adhesive apparatus, and from
<emphasis id="B93DD30AFF8F5405FD1DF956FCD5559F" box="[722,819,1700,1724]" italics="true" pageId="22" pageNumber="23">G. f u s cu s</emphasis>
in having a concave (vs. straight) margin of the dorsal fin, a shorter dorsal-fin spine (8.914.0% SL vs. 14.620.1), a shorter pectoral-fin spine (14.015.6% SL vs. 17.0 22.7), a shorter adipose-fin base (11.412.8% SL vs. 13.121.0) and a shorter caudal fin (25.828.2% SL vs. 28.7 33.9). It is further distinguished from
<taxonomicName id="4C49749BFF8F5405FDF0F8E2FD7E540B" box="[575,664,1808,1832]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="keluk">
<emphasis id="B93DD30AFF8F5405FDF0F8E2FD7E540B" box="[575,664,1808,1832]" italics="true" pageId="22" pageNumber="23">G. keluk</emphasis>
</taxonomicName>
in having a shorter dorsal-fin spine (8.914.0% SL vs. 13.015.0), a shorter pectoral-fin spine (14.015.6% SL vs. 19.422.9), a shorter adipose-fin base (11.412.8% SL vs. 13.6 17.2), a longer post-adipose distance (19.122.0% SL vs. 15.216.7) and a straight (vs. convex) dorsoposterior margin of the adipose fin, and from
<taxonomicName id="4C49749BFF8F5405FDE1F88EFD4154B7" box="[558,679,1916,1940]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="plectilis">
<emphasis id="B93DD30AFF8F5405FDE1F88EFD4154B7" box="[558,679,1916,1940]" italics="true" pageId="22" pageNumber="23">G. plectilis</emphasis>
</taxonomicName>
in lacking (vs. having) anteromedial striae in the thoracic adhesive apparatus, having a shorter adipose-fin base (11.412.8% SL vs. 12.917.2) and a longer post-adipose distance (19.122.0% SL vs. 14.017.6).
</paragraph>
<paragraph id="8BF60F18FF8E5404FF08FF6AFE53501B" blockId="23.[151,1436,151,824]" pageId="23" pageNumber="24">
Among other Sundaic congeners,
<taxonomicName id="4C49749BFF8E5404FD96FF6AFD0E5393" box="[601,744,152,176]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="23" pageNumber="24" phylum="Chordata" rank="species" species="famelicus">
<emphasis id="B93DD30AFF8E5404FD96FF6AFD0E5393" box="[601,744,152,176]" italics="true" pageId="23" pageNumber="24">G. famelicus</emphasis>
</taxonomicName>
further differs from
<taxonomicName id="4C49749BFF8E5404FC2AFF6AFB655393" box="[997,1155,152,176]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="23" pageNumber="24" phylum="Chordata" rank="species" species="decussatus">
<emphasis id="B93DD30AFF8E5404FC2AFF6AFB655393" box="[997,1155,152,176]" italics="true" pageId="23" pageNumber="24">G. decussatus</emphasis>
</taxonomicName>
in lacking (vs. having) anteromedial striae in the thoracic adhesive apparatus, having a smaller predorsal length (32.135.2% SL vs. 35.8 40.4), a shorter dorsal-fin spine (8.914.0% SL vs. 16.017.3), and a longer post-adipose distance (19.122.0% SL vs. 16.217.5% SL), from
<taxonomicName id="4C49749BFF8E5404FE0AFEF6FDCE523F" box="[453,552,260,284]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="23" pageNumber="24" phylum="Chordata" rank="species" species="major">
<emphasis id="B93DD30AFF8E5404FE0AFEF6FDCE523F" box="[453,552,260,284]" italics="true" pageId="23" pageNumber="24">G. major</emphasis>
</taxonomicName>
in lacking (vs. having) anteromedial striae in the thoracic adhesive apparatus, having a smaller predorsal length (32.135.2% SL vs. 36.741.6), a shorter dorsal-fin spine (8.914.0% SL vs. 14.320.8) and a longer post-adipose distance (19.122.0% SL vs. 13.618.6), and from
<taxonomicName id="4C49749BFF8E5404FBB5FEBEFACA5247" box="[1146,1324,332,356]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="23" pageNumber="24" phylum="Chordata" rank="species" species="nieuwenhuisi">
<emphasis id="B93DD30AFF8E5404FBB5FEBEFACA5247" box="[1146,1324,332,356]" italics="true" pageId="23" pageNumber="24">G. nieuwenhuisi</emphasis>
</taxonomicName>
in having a smaller predorsal length (32.135.2% SL vs. 36.541.6), a shorter dorsal-fin spine (8.914.0% SL vs. 14.018.5) and a longer caudal peduncle (19.421.2% SL vs. 16.819.5). It is further distinguished from
<taxonomicName id="4C49749BFF8E5404FB7AFE66FAF1528F" box="[1205,1303,404,428]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="23" pageNumber="24" phylum="Chordata" rank="species" species="pictus">
<emphasis id="B93DD30AFF8E5404FB7AFE66FAF1528F" box="[1205,1303,404,428]" italics="true" pageId="23" pageNumber="24">G. pictus</emphasis>
</taxonomicName>
in having a smaller eye (diameter 78% HL vs. 813), a smaller predorsal length (32.135.2% SL vs. 35.140.4) and a shorter dorsal-fin spine (8.914.0% SL vs. 15.420.9), from
<taxonomicName id="4C49749BFF8E5404FD27FE2EFC6352D7" box="[744,901,476,500]" family="Sisoridae" genus="Glyptosternum" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageId="23" pageNumber="24" phylum="Chordata" rank="species" species="platypogon">
<emphasis id="B93DD30AFF8E5404FD27FE2EFC6352D7" box="[744,901,476,500]" italics="true" pageId="23" pageNumber="24">G. platypogon</emphasis>
</taxonomicName>
in having a straight (vs. convex) dorsoposterior margin of the adipose fin and a longer post-adipose distance (19.122.0% SL vs. 14.617.6), and from
<taxonomicName id="4C49749BFF8E5404FAD5FDF2FA7A513B" box="[1306,1436,512,536]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="23" pageNumber="24" phylum="Chordata" rank="species" species="prashadi">
<emphasis id="B93DD30AFF8E5404FAD5FDF2FA7A513B" box="[1306,1436,512,536]" italics="true" pageId="23" pageNumber="24">G. prashadi</emphasis>
</taxonomicName>
in having a narrower head (16.720.0% SL vs. 20.222.8), a smaller predorsal length (32.135.2% SL vs. 37.7 39.3) and a a concave (vs. straight) margin of the dorsal fin.
<taxonomicName id="4C49749BFF8E5404FCFCFDBAFBDD5143" box="[819,1083,584,608]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="23" pageNumber="24" phylum="Chordata" rank="species" species="famelicus">
<emphasis id="B93DD30AFF8E5404FCFCFDBAFBDD5143" box="[819,1083,584,608]" italics="true" pageId="23" pageNumber="24">Glyptothorax famelicus</emphasis>
</taxonomicName>
further differs from
<taxonomicName id="4C49749BFF8E5404FAD1FDBAFA7D5143" box="[1310,1435,584,608]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="23" pageNumber="24" phylum="Chordata" rank="species" species="robustus">
<emphasis id="B93DD30AFF8E5404FAD1FDBAFA7D5143" box="[1310,1435,584,608]" italics="true" pageId="23" pageNumber="24">G. robustus</emphasis>
</taxonomicName>
in having (vs. lacking) a medial pit in the thoracic adhesive apparatus, a smaller predorsal length (32.135.2% SL vs. 36.740.0) and a shorter pectoral-fin spine (14.015.6% SL vs. 16.819.8), and from
<emphasis id="B93DD30AFF8E5404FB57FD62FAE8518B" box="[1176,1294,656,680]" italics="true" pageId="23" pageNumber="24">G. s t i b a ro s</emphasis>
in having a smaller predorsal length (32.135.2% SL vs. 39.142.8), a shorter dorsal-fin spine (8.914.0% SL vs. 15.920.4), a shorter pectoral-fin spine (14.015.6% SL vs. 18.122.2), a shorter adipose-fin base (11.412.8% SL vs. 14.6 17.8), a smooth (vs. serrated) posterior margin of the dorsal-fin spine and a longer post-adipose distance (19.1 22.0% SL vs. 15.718.3).
</paragraph>
</subSubSection>
</treatment>
</document>