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<document id="8769F119C6958F9E2F8288B210AFF06A" ID-CLB-Dataset="29109" ID-DOI="10.11646/zootaxa.4455.1.9" ID-GBIF-Dataset="e1e20b66-1c1a-4865-8ad9-b64aac7620db" ID-ISSN="1175-5326" ID-Zenodo-Dep="1457091" ID-ZooBank="5C4466D7-8D53-4824-B417-0657913BEA40" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1539260996745" checkinUser="plazi" docAuthor="Zhang, Shuqian &amp; Zhang, Suping" docDate="2018" docId="03E00B20FF141500F8D7FEA2FD96FDA3" docLanguage="en" docName="zootaxa.4455.1.9.pdf" docOrigin="Zootaxa 4455 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cocculina delphinicula Zhang &amp; Zhang 2018, sp. nov." docType="treatment" docVersion="7" lastPageNumber="193" masterDocId="FFD97358FF161504F840FFE3FFD1FFC3" masterDocTitle="Cocculina delphinicula sp. nov., a new cocculinid species from whale bone in the East China Sea (Gastropoda: Cocculiniformia)" masterLastPageNumber="195" masterPageNumber="189" pageNumber="191" updateTime="1698575302015" updateUser="ExternalLinkService">
<mods:mods id="997B64473310EBACDB3B5729E1A402E8" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="810634499160EA16F097E0E3876D758C">
<mods:title id="932B2C98508DDDC5B1BBD7AA90E7CC71">Cocculina delphinicula sp. nov., a new cocculinid species from whale bone in the East China Sea (Gastropoda: Cocculiniformia)</mods:title>
</mods:titleInfo>
<mods:name id="4AC27D70EE01939B4D13C12531410C4C" type="personal">
<mods:role id="9AC668C0C62EDAAE80C9B219164B3232">
<mods:roleTerm id="1F78751A8192E919BCCDC7F3682977E0">Author</mods:roleTerm>
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<mods:namePart id="7DA132BFD483EB6E5F64CCCE568B5FAE">Zhang, Shuqian</mods:namePart>
</mods:name>
<mods:name id="02AA4130DD8C18DD5F9CF89540C91378" type="personal">
<mods:role id="8365EDE16B4BFA9C1819CFE31775FA0A">
<mods:roleTerm id="8ABB76F1D80D3133EC1DB7BA0F3F86E6">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="24A6B70BF6A6E69D2FBFF3461E493077">Zhang, Suping</mods:namePart>
</mods:name>
<mods:typeOfResource id="F520C1D0C135488213A96736064428FF">text</mods:typeOfResource>
<mods:relatedItem id="D5F1C26B17D9D9E153E5AC68DEE9EED6" type="host">
<mods:titleInfo id="E9874038E28D27F0107DB1EA2DAF08C8">
<mods:title id="EC45C79156A1722485FC6C9598DB4BA8">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="90EDD736F5902183531BD8C62D3BD666">
<mods:date id="E010DCE218267FBE9826E0EDFF288A3D">2018</mods:date>
<mods:detail id="E5B1FDFD7E3CFACD83DA91FDA8B224CE" type="pubDate">
<mods:number id="738B637C687565E6D0CD6501C8034A2B">2018-08-01</mods:number>
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<mods:detail id="6190C25F36C0151C4DAE4B4E2A5FBD3D" type="volume">
<mods:number id="81A1E47063976506C9DB9919C14DBD00">4455</mods:number>
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<mods:detail id="6E9C0716F9343C27A722E07CD8D8FD72" type="issue">
<mods:number id="7A33006E4B37580CE0B24D1646E8FB47">1</mods:number>
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<mods:classification id="E34F68E184C6A7372B41FC26C76D9FF4">journal article</mods:classification>
<mods:identifier id="415C022B6AD25E299EB81E719655360F" type="CLB-Dataset">29109</mods:identifier>
<mods:identifier id="1BAB2F040D3923FE2881FC0FC6B820E3" type="DOI">10.11646/zootaxa.4455.1.9</mods:identifier>
<mods:identifier id="B96C4942A5D520D9B9793EA8090D49E4" type="GBIF-Dataset">e1e20b66-1c1a-4865-8ad9-b64aac7620db</mods:identifier>
<mods:identifier id="54F2738D228B85B37B703BCE448E372C" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="E0494FE7A08F1F84B4D5896C8BBD6F76" type="Zenodo-Dep">1457091</mods:identifier>
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</mods:mods>
<treatment id="03E00B20FF141500F8D7FEA2FD96FDA3" ID-DOI="http://doi.org/10.5281/zenodo.5959205" ID-GBIF-Taxon="148691181" ID-Zenodo-Dep="5959205" LSID="urn:lsid:plazi:treatment:03E00B20FF141500F8D7FEA2FD96FDA3" httpUri="http://treatment.plazi.org/id/03E00B20FF141500F8D7FEA2FD96FDA3" lastPageId="4" lastPageNumber="193" pageId="2" pageNumber="191">
<subSubSection id="C353E9BDFF141506F8D7FEA2FEDAFEBF" pageId="2" pageNumber="191" type="nomenclature">
<paragraph id="8BF6BA36FF141506F8D7FEA2FDC7FE98" blockId="2.[151,534,321,381]" box="[151,534,321,347]" pageId="2" pageNumber="191">
<heading id="D0BE0D5AFF141506F8D7FEA2FDC7FE98" bold="true" box="[151,534,321,347]" fontSize="11" level="1" pageId="2" pageNumber="191" reason="1">
<emphasis id="B93D6624FF141506F8D7FEA2FDC7FE98" bold="true" box="[151,534,321,347]" pageId="2" pageNumber="191">
<taxonomicName id="4C49C1B5FF141506F8D7FEA2FE60FE98" authority="Zhang &amp; Zhang, 2018" authorityName="Zhang &amp; Zhang" authorityYear="2018" box="[151,433,321,347]" class="Gastropoda" family="Cocculinidae" genus="Cocculina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Archaeogastropoda" pageId="2" pageNumber="191" phylum="Mollusca" rank="species" species="delphinicula" status="sp. nov.">
<emphasis id="B93D6624FF141506F8D7FEA2FE60FE98" bold="true" box="[151,433,321,347]" italics="true" pageId="2" pageNumber="191">Cocculina delphinicula</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20EDB5FFF141506F9F9FEA2FDC7FE98" box="[441,534,321,347]" pageId="2" pageNumber="191" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BF6BA36FF141506F8D7FE87FEDAFEBF" blockId="2.[151,534,321,381]" box="[151,267,356,381]" pageId="2" pageNumber="191">
(
<figureCitation id="1372A6B3FF141506F8DFFE87FED2FEBF" box="[159,259,356,381]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStartId-0="3.[151,250,1286,1308]" captionStartId-1="3.[151,250,1941,1963]" captionStartId-2="4.[151,250,1500,1522]" captionTargetBox-0="[428,1159,361,1271]" captionTargetBox-1="[340,1247,1405,1931]" captionTargetBox-2="[340,1247,663,1478]" captionTargetId-0="figure@3.[428,1159,361,1271]" captionTargetId-1="figure@3.[340,1247,1405,1931]" captionTargetId-2="figure@4.[340,1247,663,1478]" captionTargetPageId-0="3" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="FIGURE 2. Shells of Cocculina delphinicula sp. nov. A. Dorsal, and B. lateral view of holotype, 4.9 mm, MBM285103, scale bar=10 mm; C, D. Microsculpture of shell near margin; E. Dorsal, and F. lateral view of protoconch; G, H. Microsculpture of protoconch." captionText-1="FIGURE 3. Ventral view of soft part of Cocculina delphinicula sp. nov., holotype MBM285103. Abbreviations. (co) copulatory organ; (ept) epipodial tentacle; (flw) lateral wall of foot; (mm) mantle margin; (o) oral; (ol) oral lappet; (ps) pedal sole." captionText-2="FIGURE 4. Radular of holotype of Cocculina delphinicula sp. nov., MBM285103. A. Dorsal view of intact radular segment; B. Enlargement of central region; C. Marginal portion, showing marginal teeth, scale bar=100 µm; D. Inner marginal teeth, scale bar=50 µm; E. Outer marginal teeth, scale bar=10 µm." httpUri-0="https://zenodo.org/record/1457095/files/figure.png" httpUri-1="https://zenodo.org/record/1457097/files/figure.png" httpUri-2="https://zenodo.org/record/1457099/files/figure.png" pageId="2" pageNumber="191">Figs 2–4</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C353E9BDFF141506F8D7FE4FFBB9FDEC" pageId="2" pageNumber="191" type="materials_examined">
<paragraph id="8BF6BA36FF141506F8D7FE4FFAEEFE2B" blockId="2.[151,1437,428,1064]" pageId="2" pageNumber="191">
<emphasis id="B93D6624FF141506F8D7FE4FFE1CFE06" bold="true" box="[151,461,428,453]" pageId="2" pageNumber="191">
<typeStatus id="54F20494FF141506F8D7FE4FFF03FE06" box="[151,210,428,453]" pageId="2" pageNumber="191">Type</typeStatus>
material.
<typeStatus id="54F20494FF141506F917FE4FFE19FE06" box="[343,456,428,453]" pageId="2" pageNumber="191" type="holotype">Holotype</typeStatus>
:
</emphasis>
<collectionCode id="ED5822F3FF141506F99AFE4FFDCFFE06" box="[474,542,428,453]" country="0" httpUri="http://grbio.org/cool/fc7c-p25f" name="San Jose State University, Museum of Birds and Mammals" pageId="2" pageNumber="191">MBM</collectionCode>
285103 (length
<quantity id="4CB117D3FF141506FA9BFE4FFCE8FE07" box="[731,825,428,452]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" pageId="2" pageNumber="191" unit="mm" value="4.9">4.9 mm</quantity>
, width
<quantity id="4CB117D3FF141506FBD7FE4FFC26FE07" box="[919,1015,428,452]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="2" pageNumber="191" unit="mm" value="3.8">3.8 mm</quantity>
), East
<collectingCountry id="F35EFAA6FF141506FC0DFE4FFB41FE06" box="[1101,1168,428,453]" name="China" pageId="2" pageNumber="191">China</collectingCountry>
Sea,
<date id="FFF79CF6FF141506FC99FE4FFA49FE07" box="[1241,1432,428,453]" pageId="2" pageNumber="191" value="2017-11">November 2017</date>
;
<emphasis id="B93D6624FF141506F8D7FE2CFEC9FE2B" bold="true" box="[151,280,463,488]" pageId="2" pageNumber="191">
<typeStatus id="54F20494FF141506F8D7FE2CFEC2FE2B" box="[151,275,463,488]" pageId="2" pageNumber="191" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<collectionCode id="ED5822F3FF141506F95EFE2CFEB2FE2B" box="[286,355,463,488]" country="0" httpUri="http://grbio.org/cool/fc7c-p25f" name="San Jose State University, Museum of Birds and Mammals" pageId="2" pageNumber="191">MBM</collectionCode>
285104, six specimens, collected with the
<typeStatus id="54F20494FF141506FB7EFE2CFC71FE2B" box="[830,928,463,488]" pageId="2" pageNumber="191" type="holotype">holotype</typeStatus>
at the
<typeStatus id="54F20494FF141506FBABFE33FBCAFE2B" box="[1003,1051,464,488]" pageId="2" pageNumber="191">type</typeStatus>
locality,
<date id="FFF79CF6FF141506FCC3FE2CFAEDFE2B" box="[1155,1340,463,488]" pageId="2" pageNumber="191" value="2017-11">November 2017</date>
.
</paragraph>
<paragraph id="8BF6BA36FF141506F887FE17FCE6FDCE" blockId="2.[151,1437,428,1064]" box="[199,823,500,525]" pageId="2" pageNumber="191">Six additional immature specimens were also observed.</paragraph>
<paragraph id="8BF6BA36FF141506F887FDF4FBB9FDEC" blockId="2.[151,1437,428,1064]" box="[199,1128,535,560]" pageId="2" pageNumber="191">
<materialsCitation id="3B21B06BFF141506F887FDF4FBB9FDEC" ID-GBIF-Occurrence="1917075841" box="[199,1128,535,560]" country="China" elevation="350" location="China" pageId="2" pageNumber="191" specimenCount="1" typeStatus="holotype">
<emphasis id="B93D6624FF141506F887FDF4FEB4FDF3" bold="true" box="[199,357,535,560]" pageId="2" pageNumber="191">
<typeStatus id="54F20494FF141506F887FDF4FED3FDF3" box="[199,258,535,560]" pageId="2" pageNumber="191">Type</typeStatus>
locality.
</emphasis>
East
<collectingCountry id="F35EFAA6FF141506F9E3FDF4FE37FDF3" box="[419,486,535,560]" name="China" pageId="2" pageNumber="191">China</collectingCountry>
Sea, approximately 28°N 127°E, at depth of
<elevation id="00645D05FF141506FBA2FDFBFBB5FDEC" box="[994,1124,536,560]" metricMagnitude="2" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="2" pageNumber="191" unit="m" value="350.0" valueMax="400.0" valueMin="300.0">
<quantity id="4CB117D3FF141506FBA2FDFBFBB5FDEC" box="[994,1124,536,560]" metricMagnitude="2" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="2" pageNumber="191" unit="m" value="350.0" valueMax="400.0" valueMin="300.0">300–400 m</quantity>
</elevation>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C353E9BDFF141506F887FDDFFF04FDBB" pageId="2" pageNumber="191" type="etymology">
<paragraph id="8BF6BA36FF141506F887FDDFFF04FDBB" blockId="2.[151,1437,428,1064]" pageId="2" pageNumber="191">
<emphasis id="B93D6624FF141506F887FDDFFE9FFD96" bold="true" box="[199,334,572,597]" pageId="2" pageNumber="191">Etymology.</emphasis>
The species is named after dolphin family
<taxonomicName id="4C49C1B5FF141506FB7CFDDFFC14FD96" authorityName="Gray" authorityYear="1821" box="[828,965,572,597]" class="Mammalia" family="Delphinidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="2" pageNumber="191" phylum="Chordata" rank="family">Delphinidae</taxonomicName>
because it was found living on a dolphin skull.
</paragraph>
</subSubSection>
<subSubSection id="C353E9BDFF141506F887FD67FE0AFC8C" pageId="2" pageNumber="191" type="diagnosis">
<paragraph id="8BF6BA36FF141506F887FD67FE0AFC8C" blockId="2.[151,1437,428,1064]" pageId="2" pageNumber="191">
<emphasis id="B93D6624FF141506F887FD67FE90FD5E" bold="true" box="[199,321,644,669]" pageId="2" pageNumber="191">Diagnosis.</emphasis>
Shell medium-sized, depressed, shell width 77%–81% of shell length, height 31%–36% of length. Shell surface sparsely pitted. Protoconch slightly posterior to center, with reticulate sculpture. Teleoconch sculptured with regularly spaced concentric growth lines and even finer radial threads. Aperture oval, longer than wider. Radula has a formula of (ca. 20)+5+1+5+(ca. 20), rachidian vestigial. Lateral 1 weak but well defined. Laterals 2 and 3 similar, overhanging tip with 3–4 cusps. Lateral 4 smaller and slender with a large primary cusp and an outer secondary cusp.
</paragraph>
</subSubSection>
<subSubSection id="C353E9BDFF141507F887FCBFFE85FE83" lastPageId="3" lastPageNumber="192" pageId="2" pageNumber="191" type="description">
<paragraph id="8BF6BA36FF141506F887FCBFFAF9FBEB" blockId="2.[151,1437,428,1064]" pageId="2" pageNumber="191">
<emphasis id="B93D6624FF141506F887FCBFFE88FCB6" bold="true" box="[199,345,860,885]" pageId="2" pageNumber="191">Description.</emphasis>
Shell (
<figureCitation id="1372A6B3FF141506F9EDFCBFFE24FCB7" box="[429,501,860,885]" captionStart="FIGURE 2" captionStartId="3.[151,250,1286,1308]" captionTargetBox="[428,1159,361,1271]" captionTargetId="figure@3.[428,1159,361,1271]" captionTargetPageId="3" captionText="FIGURE 2. Shells of Cocculina delphinicula sp. nov. A. Dorsal, and B. lateral view of holotype, 4.9 mm, MBM285103, scale bar=10 mm; C, D. Microsculpture of shell near margin; E. Dorsal, and F. lateral view of protoconch; G, H. Microsculpture of protoconch." httpUri="https://zenodo.org/record/1457095/files/figure.png" pageId="2" pageNumber="191">Fig. 2</figureCitation>
;
<tableCitation id="C6CB8F8DFF141506FA43FCBFFD98FCB7" box="[515,585,860,885]" captionStart="TABLE 1" captionStartId="2.[151,239,1112,1135]" captionText="TABLE 1. Shell measurements (in mm) and ratios of Cocculina delphinicula sp. nov." pageId="2" pageNumber="191">Tab. 1</tableCitation>
) medium-sized for genus (maximum length
<quantity id="4CB117D3FF141506FC0FFCBFFB7AFCB7" box="[1103,1195,860,885]" metricMagnitude="-3" metricUnit="m" metricValue="5.9" pageId="2" pageNumber="191" unit="mm" value="5.9">5.9 mm</quantity>
), thin, fragile, white, semi-transparent, periostracum not observed. Shell depressed, height of
<typeStatus id="54F20494FF141506FB87FC9CFBF8FC5B" box="[967,1065,895,920]" pageId="2" pageNumber="191" type="holotype">holotype</typeStatus>
0.33 times length. Apex slightly posterior to center, situated at 56–62% of shell length from anterior end. Protoconch offset to left, deflected from longitudinal axis of adult teleoconch by up to 20°. Aperture oval, longer than wide (width/length ratio 0.77–0.81), anterior end slightly narrower than posterior end. Apertural margin not planar, shallowly concave from side to side. Anterior and posterior slopes faintly convex, lateral slopes at first slightly convex, then weakly concave.
</paragraph>
<caption id="DF36EABEFF141506F8D7FBBBFBFDFBAC" box="[151,1068,1111,1135]" pageId="2" pageNumber="191">
<paragraph id="8BF6BA36FF141506F8D7FBBBFBFDFBAC" blockId="2.[151,1068,1111,1135]" box="[151,1068,1111,1135]" pageId="2" pageNumber="191">
<emphasis id="B93D6624FF141506F8D7FBBBFEDBFBAC" bold="true" box="[151,266,1112,1135]" pageId="2" pageNumber="191">TABLE 1.</emphasis>
Shell measurements (in mm) and ratios of
<taxonomicName id="4C49C1B5FF141506FA96FBB4FC1EFBAC" authority="Zhang &amp; Zhang, 2018" authorityName="Zhang &amp; Zhang" authorityYear="2018" box="[726,975,1111,1135]" class="Gastropoda" family="Cocculinidae" genus="Cocculina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Archaeogastropoda" pageId="2" pageNumber="191" phylum="Mollusca" rank="species" species="delphinicula" status="sp. nov.">
<emphasis id="B93D6624FF141506FA96FBB4FC1EFBAC" box="[726,975,1111,1135]" italics="true" pageId="2" pageNumber="191">Cocculina delphinicula</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20EDB5FFF141506FB96FBBBFBFDFBAC" box="[982,1068,1112,1135]" pageId="2" pageNumber="191" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="8BF6BA36FF141506F907FB69FB67FA28" pageId="2" pageNumber="191">
<table id="F9494896FF14EAFBF8DCFB69FA82FA28" box="[156,1363,1162,1515]" gridcols="7" gridrows="9" pageId="2" pageNumber="191">
<tr id="3579B874FF14EAFBF8DCFB69FA82FB63" box="[156,1363,1162,1184]" gridrow="0" pageId="2" pageNumber="191" rowspan-0="1">
<th id="76A8D108FF14EAFBF907FB69FE67FB63" box="[327,438,1162,1184]" gridcol="1" gridrow="0" pageId="2" pageNumber="191">Length (L)</th>
<th id="76A8D108FF14EAFBF9B2FB69FD8EFB63" box="[498,607,1162,1184]" gridcol="2" gridrow="0" pageId="2" pageNumber="191">Width (W)</th>
<th id="76A8D108FF14EAFBFADCFB69FCDAFB63" box="[668,779,1162,1184]" gridcol="3" gridrow="0" pageId="2" pageNumber="191">Height (H)</th>
<th id="76A8D108FF14EAFBFB07FB69FC75FB63" box="[839,932,1162,1184]" gridcol="4" gridrow="0" pageId="2" pageNumber="191">H/L ratio</th>
<th id="76A8D108FF14EAFBFBB2FB69FB84FB63" box="[1010,1109,1162,1184]" gridcol="5" gridrow="0" pageId="2" pageNumber="191">W/L ratio</th>
<th id="76A8D108FF14EAFBFCDCFB69FA82FB63" box="[1180,1363,1162,1184]" gridcol="6" gridrow="0" pageId="2" pageNumber="191">Anterior length %</th>
</tr>
<tr id="3579B874FF14EAFBF8DCFB57FA82FB09" box="[156,1363,1204,1226]" gridrow="1" pageId="2" pageNumber="191">
<th id="76A8D108FF14EAFBF8DCFB57FF2EFB09" box="[156,255,1204,1226]" gridcol="0" gridrow="1" pageId="2" pageNumber="191">Holotype</th>
<td id="76A8D108FF14EAFBF907FB57FE67FB09" box="[327,438,1204,1226]" gridcol="1" gridrow="1" pageId="2" pageNumber="191">4.9</td>
<td id="76A8D108FF14EAFBF9B2FB57FD8EFB09" box="[498,607,1204,1226]" gridcol="2" gridrow="1" pageId="2" pageNumber="191">3.8</td>
<td id="76A8D108FF14EAFBFADCFB57FCDAFB09" box="[668,779,1204,1226]" gridcol="3" gridrow="1" pageId="2" pageNumber="191">1.6</td>
<td id="76A8D108FF14EAFBFB07FB57FC75FB09" box="[839,932,1204,1226]" gridcol="4" gridrow="1" pageId="2" pageNumber="191">0.33</td>
<td id="76A8D108FF14EAFBFBB2FB57FB84FB09" box="[1010,1109,1204,1226]" gridcol="5" gridrow="1" pageId="2" pageNumber="191">0.79</td>
<td id="76A8D108FF14EAFBFCDCFB57FA82FB09" box="[1180,1363,1204,1226]" gridcol="6" gridrow="1" pageId="2" pageNumber="191">61</td>
</tr>
<tr id="3579B874FF14EAFBF8DCFB3EFA82FB30" box="[156,1363,1245,1267]" gridrow="2" pageId="2" pageNumber="191">
<th id="76A8D108FF14EAFBF8DCFB3EFF2EFB30" box="[156,255,1245,1267]" gridcol="0" gridrow="2" pageId="2" pageNumber="191">Paratypes</th>
<td id="76A8D108FF14EAFBF907FB3EFE67FB30" box="[327,438,1245,1267]" gridcol="1" gridrow="2" pageId="2" pageNumber="191">5.6</td>
<td id="76A8D108FF14EAFBF9B2FB3EFD8EFB30" box="[498,607,1245,1267]" gridcol="2" gridrow="2" pageId="2" pageNumber="191">4.3</td>
<td id="76A8D108FF14EAFBFADCFB3EFCDAFB30" box="[668,779,1245,1267]" gridcol="3" gridrow="2" pageId="2" pageNumber="191">2.0</td>
<td id="76A8D108FF14EAFBFB07FB3EFC75FB30" box="[839,932,1245,1267]" gridcol="4" gridrow="2" pageId="2" pageNumber="191">0.35</td>
<td id="76A8D108FF14EAFBFBB2FB3EFB84FB30" box="[1010,1109,1245,1267]" gridcol="5" gridrow="2" pageId="2" pageNumber="191">0.77</td>
<td id="76A8D108FF14EAFBFCDCFB3EFA82FB30" box="[1180,1363,1245,1267]" gridcol="6" gridrow="2" pageId="2" pageNumber="191">61</td>
</tr>
<tr id="3579B874FF14EAFBF8DCFAE5FA82FADF" box="[156,1363,1286,1308]" gridrow="3" pageId="2" pageNumber="191" rowspan-0="1">
<td id="76A8D108FF14EAFBF907FAE5FE67FADF" box="[327,438,1286,1308]" gridcol="1" gridrow="3" pageId="2" pageNumber="191">5.2</td>
<td id="76A8D108FF14EAFBF9B2FAE5FD8EFADF" box="[498,607,1286,1308]" gridcol="2" gridrow="3" pageId="2" pageNumber="191">4.2</td>
<td id="76A8D108FF14EAFBFADCFAE5FCDAFADF" box="[668,779,1286,1308]" gridcol="3" gridrow="3" pageId="2" pageNumber="191">1.8</td>
<td id="76A8D108FF14EAFBFB07FAE5FC75FADF" box="[839,932,1286,1308]" gridcol="4" gridrow="3" pageId="2" pageNumber="191">0.35</td>
<td id="76A8D108FF14EAFBFBB2FAE5FB84FADF" box="[1010,1109,1286,1308]" gridcol="5" gridrow="3" pageId="2" pageNumber="191">0.81</td>
<td id="76A8D108FF14EAFBFCDCFAE5FA82FADF" box="[1180,1363,1286,1308]" gridcol="6" gridrow="3" pageId="2" pageNumber="191">62</td>
</tr>
<tr id="3579B874FF14EAFBF8DCFAD3FA82FA85" box="[156,1363,1328,1350]" gridrow="4" pageId="2" pageNumber="191" rowspan-0="1">
<td id="76A8D108FF14EAFBF907FAD3FE67FA85" box="[327,438,1328,1350]" gridcol="1" gridrow="4" pageId="2" pageNumber="191">5.9</td>
<td id="76A8D108FF14EAFBF9B2FAD3FD8EFA85" box="[498,607,1328,1350]" gridcol="2" gridrow="4" pageId="2" pageNumber="191">4.7</td>
<td id="76A8D108FF14EAFBFADCFAD3FCDAFA85" box="[668,779,1328,1350]" gridcol="3" gridrow="4" pageId="2" pageNumber="191">1.8</td>
<td id="76A8D108FF14EAFBFB07FAD3FC75FA85" box="[839,932,1328,1350]" gridcol="4" gridrow="4" pageId="2" pageNumber="191">0.31</td>
<td id="76A8D108FF14EAFBFBB2FAD3FB84FA85" box="[1010,1109,1328,1350]" gridcol="5" gridrow="4" pageId="2" pageNumber="191">0.80</td>
<td id="76A8D108FF14EAFBFCDCFAD3FA82FA85" box="[1180,1363,1328,1350]" gridcol="6" gridrow="4" pageId="2" pageNumber="191">56</td>
</tr>
<tr id="3579B874FF14EAFBF8DCFABAFA82FAAC" box="[156,1363,1369,1391]" gridrow="5" pageId="2" pageNumber="191" rowspan-0="1">
<td id="76A8D108FF14EAFBF907FABAFE67FAAC" box="[327,438,1369,1391]" gridcol="1" gridrow="5" pageId="2" pageNumber="191">5.6</td>
<td id="76A8D108FF14EAFBF9B2FABAFD8EFAAC" box="[498,607,1369,1391]" gridcol="2" gridrow="5" pageId="2" pageNumber="191">4.3</td>
<td id="76A8D108FF14EAFBFADCFABAFCDAFAAC" box="[668,779,1369,1391]" gridcol="3" gridrow="5" pageId="2" pageNumber="191">2.0</td>
<td id="76A8D108FF14EAFBFB07FABAFC75FAAC" box="[839,932,1369,1391]" gridcol="4" gridrow="5" pageId="2" pageNumber="191">0.36</td>
<td id="76A8D108FF14EAFBFBB2FABAFB84FAAC" box="[1010,1109,1369,1391]" gridcol="5" gridrow="5" pageId="2" pageNumber="191">0.77</td>
<td id="76A8D108FF14EAFBFCDCFABAFA82FAAC" box="[1180,1363,1369,1391]" gridcol="6" gridrow="5" pageId="2" pageNumber="191">61</td>
</tr>
<tr id="3579B874FF14EAFBF8DCFA61FA82FA5B" box="[156,1363,1410,1432]" gridrow="6" pageId="2" pageNumber="191" rowspan-0="1">
<td id="76A8D108FF14EAFBF907FA61FE67FA5B" box="[327,438,1410,1432]" gridcol="1" gridrow="6" pageId="2" pageNumber="191">5.4</td>
<td id="76A8D108FF14EAFBF9B2FA61FD8EFA5B" box="[498,607,1410,1432]" gridcol="2" gridrow="6" pageId="2" pageNumber="191">4.2</td>
<td id="76A8D108FF14EAFBFADCFA61FCDAFA5B" box="[668,779,1410,1432]" gridcol="3" gridrow="6" pageId="2" pageNumber="191">1.7</td>
<td id="76A8D108FF14EAFBFB07FA61FC75FA5B" box="[839,932,1410,1432]" gridcol="4" gridrow="6" pageId="2" pageNumber="191">0.31</td>
<td id="76A8D108FF14EAFBFBB2FA61FB84FA5B" box="[1010,1109,1410,1432]" gridcol="5" gridrow="6" pageId="2" pageNumber="191">0.78</td>
<td id="76A8D108FF14EAFBFCDCFA61FA82FA5B" box="[1180,1363,1410,1432]" gridcol="6" gridrow="6" pageId="2" pageNumber="191">61</td>
</tr>
<tr id="3579B874FF14EAFBF8DCFA4FFA82FA01" box="[156,1363,1452,1474]" gridrow="7" pageId="2" pageNumber="191" rowspan-0="1">
<td id="76A8D108FF14EAFBF907FA4FFE67FA01" box="[327,438,1452,1474]" gridcol="1" gridrow="7" pageId="2" pageNumber="191">5.3</td>
<td id="76A8D108FF14EAFBF9B2FA4FFD8EFA01" box="[498,607,1452,1474]" gridcol="2" gridrow="7" pageId="2" pageNumber="191">4.2</td>
<td id="76A8D108FF14EAFBFADCFA4FFCDAFA01" box="[668,779,1452,1474]" gridcol="3" gridrow="7" pageId="2" pageNumber="191">1.9</td>
<td id="76A8D108FF14EAFBFB07FA4FFC75FA01" box="[839,932,1452,1474]" gridcol="4" gridrow="7" pageId="2" pageNumber="191">0.36</td>
<td id="76A8D108FF14EAFBFBB2FA4FFB84FA01" box="[1010,1109,1452,1474]" gridcol="5" gridrow="7" pageId="2" pageNumber="191">0.79</td>
<td id="76A8D108FF14EAFBFCDCFA4FFA82FA01" box="[1180,1363,1452,1474]" gridcol="6" gridrow="7" pageId="2" pageNumber="191">62</td>
</tr>
<tr id="3579B874FF14EAFBF8DCFA36FA82FA28" box="[156,1363,1493,1515]" gridrow="8" pageId="2" pageNumber="191">
<th id="76A8D108FF14EAFBF8DCFA36FF2EFA28" box="[156,255,1493,1515]" gridcol="0" gridrow="8" pageId="2" pageNumber="191">Mean</th>
<td id="76A8D108FF14EAFBF907FA36FE67FA28" box="[327,438,1493,1515]" gridcol="1" gridrow="8" pageId="2" pageNumber="191">5.4</td>
<td id="76A8D108FF14EAFBF9B2FA36FD8EFA28" box="[498,607,1493,1515]" gridcol="2" gridrow="8" pageId="2" pageNumber="191">4.2</td>
<td id="76A8D108FF14EAFBFADCFA36FCDAFA28" box="[668,779,1493,1515]" gridcol="3" gridrow="8" pageId="2" pageNumber="191">1.8</td>
<td id="76A8D108FF14EAFBFB07FA36FC75FA28" box="[839,932,1493,1515]" gridcol="4" gridrow="8" pageId="2" pageNumber="191">0.34</td>
<td id="76A8D108FF14EAFBFBB2FA36FB84FA28" box="[1010,1109,1493,1515]" gridcol="5" gridrow="8" pageId="2" pageNumber="191">0.79</td>
<td id="76A8D108FF14EAFBFCDCFA36FA82FA28" box="[1180,1363,1493,1515]" gridcol="6" gridrow="8" pageId="2" pageNumber="191">60</td>
</tr>
</table>
</paragraph>
<paragraph id="8BF6BA36FF141506F887F9CFFEFCF973" blockId="2.[151,1436,1580,2037]" pageId="2" pageNumber="191">
Protoconch (
<figureCitation id="1372A6B3FF141506F916F9CFFE02F987" box="[342,467,1580,1605]" captionStart="FIGURE 2" captionStartId="3.[151,250,1286,1308]" captionTargetBox="[428,1159,361,1271]" captionTargetId="figure@3.[428,1159,361,1271]" captionTargetPageId="3" captionText="FIGURE 2. Shells of Cocculina delphinicula sp. nov. A. Dorsal, and B. lateral view of holotype, 4.9 mm, MBM285103, scale bar=10 mm; C, D. Microsculpture of shell near margin; E. Dorsal, and F. lateral view of protoconch; G, H. Microsculpture of protoconch." httpUri="https://zenodo.org/record/1457095/files/figure.png" pageId="2" pageNumber="191">Figs 2E–H</figureCitation>
) about 260 µm long and 145 µm wide, located slightly below the highest point of the shell. Tip of protoconch partly immersed in the posterior slope of the teleoconch. Protoconch surface sculptured with a reticulate pattern of flattened, broad ridges (
<figureCitation id="1372A6B3FF141506FA93F997FC9FF94F" box="[723,846,1652,1677]" captionStart="FIGURE 2" captionStartId="3.[151,250,1286,1308]" captionTargetBox="[428,1159,361,1271]" captionTargetId="figure@3.[428,1159,361,1271]" captionTargetPageId="3" captionText="FIGURE 2. Shells of Cocculina delphinicula sp. nov. A. Dorsal, and B. lateral view of holotype, 4.9 mm, MBM285103, scale bar=10 mm; C, D. Microsculpture of shell near margin; E. Dorsal, and F. lateral view of protoconch; G, H. Microsculpture of protoconch." httpUri="https://zenodo.org/record/1457095/files/figure.png" pageId="2" pageNumber="191">Figs 2E–F</figureCitation>
). Surface between ridges of protoconch finely pitted (
<figureCitation id="1372A6B3FF141506F8DFF974FEF0F973" box="[159,289,1687,1712]" captionStart="FIGURE 2" captionStartId="3.[151,250,1286,1308]" captionTargetBox="[428,1159,361,1271]" captionTargetId="figure@3.[428,1159,361,1271]" captionTargetPageId="3" captionText="FIGURE 2. Shells of Cocculina delphinicula sp. nov. A. Dorsal, and B. lateral view of holotype, 4.9 mm, MBM285103, scale bar=10 mm; C, D. Microsculpture of shell near margin; E. Dorsal, and F. lateral view of protoconch; G, H. Microsculpture of protoconch." httpUri="https://zenodo.org/record/1457095/files/figure.png" pageId="2" pageNumber="191">Figs 2G–H</figureCitation>
).
</paragraph>
<paragraph id="8BF6BA36FF141506F887F95FFC0CF8DF" blockId="2.[151,1436,1580,2037]" pageId="2" pageNumber="191">
Teleoconch sculpture at first with regularly spaced concentric growth lines, then becoming coarser and stronger toward the aperture, crossed by even finer radial threads. The lines and threads producing waved nodules at intersections (see
<figureCitation id="1372A6B3FF141506F93BF8E7FE2BF8DF" box="[379,506,1796,1821]" captionStart="FIGURE 2" captionStartId="3.[151,250,1286,1308]" captionTargetBox="[428,1159,361,1271]" captionTargetId="figure@3.[428,1159,361,1271]" captionTargetPageId="3" captionText="FIGURE 2. Shells of Cocculina delphinicula sp. nov. A. Dorsal, and B. lateral view of holotype, 4.9 mm, MBM285103, scale bar=10 mm; C, D. Microsculpture of shell near margin; E. Dorsal, and F. lateral view of protoconch; G, H. Microsculpture of protoconch." httpUri="https://zenodo.org/record/1457095/files/figure.png" pageId="2" pageNumber="191">Figs 2C–D</figureCitation>
). Shell surface sparsely pitted (see
<figureCitation id="1372A6B3FF141506FBC2F8E7FC01F8DF" box="[898,976,1796,1821]" captionStart="FIGURE 2" captionStartId="3.[151,250,1286,1308]" captionTargetBox="[428,1159,361,1271]" captionTargetId="figure@3.[428,1159,361,1271]" captionTargetPageId="3" captionText="FIGURE 2. Shells of Cocculina delphinicula sp. nov. A. Dorsal, and B. lateral view of holotype, 4.9 mm, MBM285103, scale bar=10 mm; C, D. Microsculpture of shell near margin; E. Dorsal, and F. lateral view of protoconch; G, H. Microsculpture of protoconch." httpUri="https://zenodo.org/record/1457095/files/figure.png" pageId="2" pageNumber="191">Fig 2F</figureCitation>
).
</paragraph>
<paragraph id="8BF6BA36FF141506F887F8C4FC8CF84B" blockId="2.[151,1436,1580,2037]" pageId="2" pageNumber="191">
Soft parts (
<figureCitation id="1372A6B3FF141506F90BF8C4FE42F883" box="[331,403,1831,1856]" captionStart="FIGURE 3" captionStartId="3.[151,250,1941,1963]" captionTargetBox="[340,1247,1405,1931]" captionTargetId="figure@3.[340,1247,1405,1931]" captionTargetPageId="3" captionText="FIGURE 3. Ventral view of soft part of Cocculina delphinicula sp. nov., holotype MBM285103. Abbreviations. (co) copulatory organ; (ept) epipodial tentacle; (flw) lateral wall of foot; (mm) mantle margin; (o) oral; (ol) oral lappet; (ps) pedal sole." httpUri="https://zenodo.org/record/1457097/files/figure.png" pageId="2" pageNumber="191">Fig. 3</figureCitation>
). Copulatory organ bifurcated, at tip of right oral lappet. Foot with rounded pedal sole. Posterior end of foot with a pair of epipodial tentacles. Detailed morphology of head area and dorsal side of animal (visceral mass) could not be observed due to poor preservation.
</paragraph>
<paragraph id="8BF6BA36FF141507F887F877FE85FE83" blockId="2.[151,1436,1580,2037]" lastBlockId="3.[151,1436,151,320]" lastPageId="3" lastPageNumber="192" pageId="2" pageNumber="191">
Radula (
<figureCitation id="1372A6B3FF141506F967F877FEBDF86F" box="[295,364,1940,1965]" captionStart="FIGURE 4" captionStartId="4.[151,250,1500,1522]" captionTargetBox="[340,1247,663,1478]" captionTargetId="figure@4.[340,1247,663,1478]" captionTargetPageId="4" captionText="FIGURE 4. Radular of holotype of Cocculina delphinicula sp. nov., MBM285103. A. Dorsal view of intact radular segment; B. Enlargement of central region; C. Marginal portion, showing marginal teeth, scale bar=100 µm; D. Inner marginal teeth, scale bar=50 µm; E. Outer marginal teeth, scale bar=10 µm." httpUri="https://zenodo.org/record/1457099/files/figure.png" pageId="2" pageNumber="191">Fig. 4</figureCitation>
) has a formula of (ca. 20)+5+1+5+(ca. 20). Rachidian vestigial, represented by a longitudinal ridge. Lateral teeth numbering
<quantity id="4CB117D3FF141506FA47F85BFD82F812" box="[519,595,1976,2001]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" metricValueMax="12.7" metricValueMin="2.54" pageId="2" pageNumber="191" unit="in" value="3.0" valueMax="5.0" valueMin="1.0">1–5 in</quantity>
each half row, lateral 1 weak but well defined, having an outline that frequently terminates in a small beaklike cusp. Laterals 2 and 3 similar to each other, with long, parallel-sided shafts, tip overhanging, with 3–4 cusps. Lateral 4 smaller and slender, with a large primary cusp and an outer secondary cusp. Lateral 5 transformed to a broad claw-like tooth, equipped with a large primary cusp and a smaller secondary cusp on either side, finely longitudinally striated on its posterior surface. Marginal teeth with long, laterally compressed shafts, tips overhanging, cutting edges weakly serrated at inner margins and strongly serrated at outer margins.
</paragraph>
</subSubSection>
<caption id="DF36EABEFF151507F8D7FAE5FEC1FA99" httpUri="https://zenodo.org/record/1457095/files/figure.png" pageId="3" pageNumber="192" startId="3.[151,250,1286,1308]" targetBox="[428,1159,361,1271]" targetPageId="3">
<paragraph id="8BF6BA36FF151507F8D7FAE5FEC1FA99" blockId="3.[151,1436,1286,1370]" pageId="3" pageNumber="192">
<emphasis id="B93D6624FF151507F8D7FAE5FEC3FADF" bold="true" box="[151,274,1286,1308]" pageId="3" pageNumber="192">FIGURE 2.</emphasis>
Shells of
<taxonomicName id="4C49C1B5FF151507F939FAE5FDB7FADF" authority="Zhang &amp; Zhang, 2018" authorityName="Zhang &amp; Zhang" authorityYear="2018" box="[377,614,1286,1308]" class="Gastropoda" family="Cocculinidae" genus="Cocculina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Archaeogastropoda" pageId="3" pageNumber="192" phylum="Mollusca" rank="species" species="delphinicula" status="sp. nov.">
<emphasis id="B93D6624FF151507F939FAE5FDB7FADF" box="[377,614,1286,1308]" italics="true" pageId="3" pageNumber="192">Cocculina delphinicula</emphasis>
</taxonomicName>
<emphasis id="B93D6624FF151507FA2CFAE5FD0DFADF" bold="true" box="[620,732,1286,1308]" pageId="3" pageNumber="192">
<taxonomicNameLabel id="A20EDB5FFF151507FA2CFAE5FD6DFADF" box="[620,700,1286,1308]" pageId="3" pageNumber="192" rank="species">sp. nov.</taxonomicNameLabel>
A.
</emphasis>
Dorsal, and
<emphasis id="B93D6624FF151507FB1DFAE5FCA5FADF" bold="true" box="[861,884,1286,1308]" pageId="3" pageNumber="192">B.</emphasis>
lateral view of holotype, 4.9 mm, MBM285103, scale bar=10 mm;
<emphasis id="B93D6624FF151507F95BFAC6FE82FAF8" bold="true" box="[283,339,1317,1339]" pageId="3" pageNumber="192">C, D.</emphasis>
Microsculpture of shell near margin;
<emphasis id="B93D6624FF151507FA97FAC6FD3FFAF8" bold="true" box="[727,750,1317,1339]" pageId="3" pageNumber="192">E.</emphasis>
Dorsal, and
<emphasis id="B93D6624FF151507FB31FAC6FC52FAF8" bold="true" box="[881,899,1317,1339]" pageId="3" pageNumber="192">F.</emphasis>
lateral view of protoconch;
<emphasis id="B93D6624FF151507FCE6FAC6FB0AFAF8" bold="true" box="[1190,1243,1317,1339]" pageId="3" pageNumber="192">G, H.</emphasis>
Microsculpture of protoconch.
</paragraph>
</caption>
<caption id="DF36EABEFF151507F8D7F876FF16F82A" httpUri="https://zenodo.org/record/1457097/files/figure.png" pageId="3" pageNumber="192" startId="3.[151,250,1941,1963]" targetBox="[340,1247,1405,1931]" targetPageId="3">
<paragraph id="8BF6BA36FF151507F8D7F876FF16F82A" blockId="3.[151,1436,1941,2025]" pageId="3" pageNumber="192">
<emphasis id="B93D6624FF151507F8D7F876FEC8F868" bold="true" box="[151,281,1941,1963]" pageId="3" pageNumber="192">FIGURE 3.</emphasis>
Ventral view of soft part of
<taxonomicName id="4C49C1B5FF151507FA25F876FC88F868" authority="Zhang &amp; Zhang, 2018" authorityName="Zhang &amp; Zhang" authorityYear="2018" box="[613,857,1941,1963]" class="Gastropoda" family="Cocculinidae" genus="Cocculina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Archaeogastropoda" pageId="3" pageNumber="192" phylum="Mollusca" rank="species" species="delphinicula" status="sp. nov.">
<emphasis id="B93D6624FF151507FA25F876FC88F868" box="[613,857,1941,1963]" italics="true" pageId="3" pageNumber="192">Cocculina delphinicula</emphasis>
</taxonomicName>
<emphasis id="B93D6624FF151507FB25F876FC12F868" bold="true" box="[869,963,1941,1963]" pageId="3" pageNumber="192">
<taxonomicNameLabel id="A20EDB5FFF151507FB25F876FC6EF868" box="[869,959,1941,1963]" pageId="3" pageNumber="192" rank="species">sp. nov.</taxonomicNameLabel>
,
</emphasis>
holotype MBM285103.
<emphasis id="B93D6624FF151507FC93F876FAB1F868" box="[1235,1376,1941,1963]" italics="true" pageId="3" pageNumber="192">Abbreviations</emphasis>
.
<emphasis id="B93D6624FF151507FD33F876FA4AF868" bold="true" box="[1395,1435,1941,1963]" pageId="3" pageNumber="192">(co)</emphasis>
copulatory organ;
<emphasis id="B93D6624FF151507F911F857FE55F809" bold="true" box="[337,388,1972,1994]" pageId="3" pageNumber="192">(ept)</emphasis>
epipodial tentacle;
<emphasis id="B93D6624FF151507FA0DF857FD51F809" bold="true" box="[589,640,1972,1994]" pageId="3" pageNumber="192">(flw)</emphasis>
lateral wall of foot;
<emphasis id="B93D6624FF151507FB12F857FC5CF809" bold="true" box="[850,909,1972,1994]" pageId="3" pageNumber="192">(mm)</emphasis>
mantle margin;
<emphasis id="B93D6624FF151507FC75F857FB82F809" bold="true" box="[1077,1107,1972,1994]" pageId="3" pageNumber="192">(o)</emphasis>
oral;
<emphasis id="B93D6624FF151507FCCFF857FB64F809" bold="true" box="[1167,1205,1972,1994]" pageId="3" pageNumber="192">(ol)</emphasis>
oral lappet;
<emphasis id="B93D6624FF151507FD75F857FA8EF809" bold="true" box="[1333,1375,1972,1994]" pageId="3" pageNumber="192">(ps)</emphasis>
pedal sole.
</paragraph>
</caption>
<subSubSection id="C353E9BDFF121500F887FF74FD96FDA3" pageId="4" pageNumber="193" type="discussion">
<paragraph id="8BF6BA36FF121500F887FF74FC54FE83" blockId="4.[151,1437,151,608]" pageId="4" pageNumber="193">
<emphasis id="B93D6624FF121500F887FF74FEEAFF73" bold="true" box="[199,315,151,176]" pageId="4" pageNumber="193">Remarks.</emphasis>
<taxonomicName id="4C49C1B5FF121500F905FF7BFD9EFF73" authority="Zhang &amp; Zhang, 2018" authorityName="Zhang &amp; Zhang" authorityYear="2018" box="[325,591,152,176]" class="Gastropoda" family="Cocculinidae" genus="Cocculina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Archaeogastropoda" pageId="4" pageNumber="193" phylum="Mollusca" rank="species" species="delphinicula" status="sp. nov.">
<emphasis id="B93D6624FF121500F905FF7BFD9EFF73" box="[325,591,152,176]" italics="true" pageId="4" pageNumber="193">Cocculina delphinicula</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20EDB5FFF121500FA19FF74FD67FF73" box="[601,694,151,176]" pageId="4" pageNumber="193" rank="species">
<emphasis id="B93D6624FF121500FA19FF74FD67FF73" bold="true" box="[601,694,151,176]" pageId="4" pageNumber="193">sp. nov.</emphasis>
</taxonomicNameLabel>
is superficially similar to
<taxonomicName id="4C49C1B5FF121500FBADFF7BFA4DFF73" authority="Schepman, 1908" authorityName="Schepman" authorityYear="1908" box="[1005,1436,151,176]" class="Gastropoda" family="Cocculinidae" genus="Cocculina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Archaeogastropoda" pageId="4" pageNumber="193" phylum="Mollusca" rank="species" species="alveolata">
<emphasis id="B93D6624FF121500FBADFF7BFB02FF73" box="[1005,1235,152,176]" italics="true" pageId="4" pageNumber="193">Cocculina alveolata</emphasis>
<bibRefCitation id="EFD8C7C7FF121500FC9EFF74FA4DFF73" box="[1246,1436,151,176]" pageId="4" pageNumber="193" refId="ref3437" refString="Schepman, M. M. (1908) The Prosobranchia of the Siboga Expedition. Part I. Rhipidoglossa and Docoglossa. Uitkomsten op Zoologisch, Botanisch, Oceanographisch en Geologisch Gebied verzameld in Nederlandsch Oost-Indie 1899 - 1900 aan boord H. M. Siboga, 49, 1 - 95." type="journal article">Schepman, 1908</bibRefCitation>
</taxonomicName>
from
<collectingCountry id="F35EFAA6FF121500F896FF5FFE92FF16" box="[214,323,188,213]" name="Indonesia" pageId="4" pageNumber="193">Indonesia</collectingCountry>
in shell size and sculpture. From that species, however, the new species differs in having a larger width/length ratio (0.77–0.81
<emphasis id="B93D6624FF121500F9ACFF02FDDBFF3B" box="[492,522,225,248]" italics="true" pageId="4" pageNumber="193">vs.</emphasis>
0.72), a less convex anterior slope and a concave rather than elevated apertural margin. Moreover,
<taxonomicName id="4C49C1B5FF121500F92EFEE7FDA5FEDF" authority="Zhang &amp; Zhang, 2018" authorityName="Zhang &amp; Zhang" authorityYear="2018" box="[366,628,260,284]" class="Gastropoda" family="Cocculinidae" genus="Cocculina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Archaeogastropoda" pageId="4" pageNumber="193" phylum="Mollusca" rank="species" species="delphinicula" status="sp. nov.">
<emphasis id="B93D6624FF121500F92EFEE7FDA5FEDF" box="[366,628,260,284]" italics="true" pageId="4" pageNumber="193">Cocculina delphinicula</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20EDB5FFF121500FA3BFEE7FD05FEDE" box="[635,724,260,285]" pageId="4" pageNumber="193" rank="species">
<emphasis id="B93D6624FF121500FA3BFEE7FD05FEDE" bold="true" box="[635,724,260,285]" pageId="4" pageNumber="193">sp. nov.</emphasis>
</taxonomicNameLabel>
can be separated from
<taxonomicName id="4C49C1B5FF121500FB96FEE7FAA8FEDF" authority="Schepman, 1908" authorityName="Schepman" authorityYear="1908" box="[982,1401,260,285]" class="Gastropoda" family="Cocculinidae" genus="Cocculina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Archaeogastropoda" pageId="4" pageNumber="193" phylum="Mollusca" rank="species" species="alveolata">
<emphasis id="B93D6624FF121500FB96FEE7FB69FEDF" box="[982,1208,260,284]" italics="true" pageId="4" pageNumber="193">Cocculina alveolata</emphasis>
<bibRefCitation id="EFD8C7C7FF121500FCFFFEE7FAA8FEDF" box="[1215,1401,260,285]" pageId="4" pageNumber="193" refId="ref3437" refString="Schepman, M. M. (1908) The Prosobranchia of the Siboga Expedition. Part I. Rhipidoglossa and Docoglossa. Uitkomsten op Zoologisch, Botanisch, Oceanographisch en Geologisch Gebied verzameld in Nederlandsch Oost-Indie 1899 - 1900 aan boord H. M. Siboga, 49, 1 - 95." type="journal article">Schepman, 1908</bibRefCitation>
</taxonomicName>
by having an outer secondary cusp on the first and fourth lateral teeth.
</paragraph>
<paragraph id="8BF6BA36FF121500F887FEAFFD96FDA3" blockId="4.[151,1437,151,608]" pageId="4" pageNumber="193">
Five species from
<collectingCountry id="F35EFAA6FF121500F9D4FEAEFE06FEA7" box="[404,471,333,356]" name="Japan" pageId="4" pageNumber="193">Japan</collectingCountry>
,
<taxonomicName id="4C49C1B5FF121500F9A2FEAFFCECFEA7" authority="Dall, 1907" authorityName="Dall" authorityYear="1907" box="[482,829,332,357]" class="Gastropoda" family="Cocculinidae" genus="Cocculina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Archaeogastropoda" pageId="4" pageNumber="193" phylum="Mollusca" rank="species" species="japonica">
<emphasis id="B93D6624FF121500F9A2FEAFFD6FFEA7" box="[482,702,332,356]" italics="true" pageId="4" pageNumber="193">Cocculina japonica</emphasis>
<bibRefCitation id="EFD8C7C7FF121500FA84FEAFFCECFEA7" box="[708,829,332,357]" pageId="4" pageNumber="193" refId="ref2738" refString="Dall, W. H. (1907) Descriptions of new species of shell, chiefly Buccinidae, from the dredgings of the U. S. S. &quot; Albatross &quot; during 1906, in the northwestern Pacific, Bering, Okhotsk and Japanese Seas. Smithsonian Miscellaneous Collections, 50 (2), 139 - 173." type="journal article">Dall, 1907</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C49C1B5FF121500FB08FEAFFACFFEA7" authority="Kuroda &amp; Habe, 1949" authorityName="Kuroda &amp; Habe" authorityYear="1949" box="[840,1310,332,357]" class="Gastropoda" family="Cocculinidae" genus="Cocculina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Archaeogastropoda" pageId="4" pageNumber="193" phylum="Mollusca" rank="species" species="pacifica">
<emphasis id="B93D6624FF121500FB08FEAFFBCDFEA7" box="[840,1052,332,356]" italics="true" pageId="4" pageNumber="193">Cocculina pacifica</emphasis>
<bibRefCitation id="EFD8C7C7FF121500FC63FEAFFACFFEA7" box="[1059,1310,332,357]" pageId="4" pageNumber="193" refId="ref3118" refString="Kuroda, T. &amp; Habe, T. (1949) On the gastropod genus Cocculina from Japan. Lenus, 15, 58 - 67." type="journal article">Kuroda &amp; Habe, 1949</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C49C1B5FF121500FD69FEAFFE36FE4B" authority="Hasegawa, 1997" authorityName="Hasegawa" authorityYear="1997" class="Gastropoda" family="Cocculinidae" genus="Cocculina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Archaeogastropoda" pageId="4" pageNumber="193" phylum="Mollusca" rank="species" species="surugaensis">
<emphasis id="B93D6624FF121500FD69FEAFFECCFE44" italics="true" pageId="4" pageNumber="193">Cocculina surugaensis</emphasis>
<bibRefCitation id="EFD8C7C7FF121500F968FE93FE36FE4B" box="[296,487,368,392]" pageId="4" pageNumber="193" refId="ref2995" refString="Hasegawa, K. (1997) Sunken wood-associated gastropods collected from Suruga Bay, Pacific side of the central Honshu, Japan, with descriptions of 12 new species. National Science Museum Monographs, 12, 59 - 123." type="journal article">Hasegawa, 1997</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C49C1B5FF121500F9B7FE93FC7AFE4B" authority="Hasegawa, 1997" authorityName="Hasegawa" authorityYear="1997" box="[503,939,368,392]" class="Gastropoda" family="Cocculinidae" genus="Cocculina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Archaeogastropoda" pageId="4" pageNumber="193" phylum="Mollusca" rank="species" species="tenuitesta">
<emphasis id="B93D6624FF121500F9B7FE93FD33FE44" box="[503,738,368,392]" italics="true" pageId="4" pageNumber="193">Cocculina tenuitesta</emphasis>
<bibRefCitation id="EFD8C7C7FF121500FAADFE93FC7AFE4B" box="[749,939,368,392]" pageId="4" pageNumber="193" refId="ref2995" refString="Hasegawa, K. (1997) Sunken wood-associated gastropods collected from Suruga Bay, Pacific side of the central Honshu, Japan, with descriptions of 12 new species. National Science Museum Monographs, 12, 59 - 123." type="journal article">Hasegawa, 1997</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C49C1B5FF121500FBAAFE93FF1EFE6F" authority="Kuroda &amp; Habe, 1994" authorityName="Kuroda &amp; Habe" authorityYear="1994" class="Gastropoda" family="Cocculinidae" genus="Cocculina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Archaeogastropoda" pageId="4" pageNumber="193" phylum="Mollusca" rank="species" species="tosaensis">
<emphasis id="B93D6624FF121500FBAAFE93FB1EFE44" box="[1002,1231,368,392]" italics="true" pageId="4" pageNumber="193">Cocculina tosaensis</emphasis>
Kuroda &amp; Habe, 1994
</taxonomicName>
as well as two species from
<collectingCountry id="F35EFAA6FF121500FA51FE77FDB7FE6E" box="[529,614,404,429]" name="Taiwan" pageId="4" pageNumber="193">Taiwan</collectingCountry>
,
<taxonomicName id="4C49C1B5FF121500FA31FE77FC1AFE6F" authority="Dall, 1907" authorityName="Dall" authorityYear="1907" box="[625,971,404,429]" class="Gastropoda" family="Cocculinidae" genus="Cocculina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Archaeogastropoda" pageId="4" pageNumber="193" phylum="Mollusca" rank="species" species="japonica">
<emphasis id="B93D6624FF121500FA31FE77FC9CFE6F" box="[625,845,404,428]" italics="true" pageId="4" pageNumber="193">Cocculina japonica</emphasis>
<bibRefCitation id="EFD8C7C7FF121500FB14FE77FC1AFE6F" box="[852,971,404,429]" pageId="4" pageNumber="193" refId="ref2738" refString="Dall, W. H. (1907) Descriptions of new species of shell, chiefly Buccinidae, from the dredgings of the U. S. S. &quot; Albatross &quot; during 1906, in the northwestern Pacific, Bering, Okhotsk and Japanese Seas. Smithsonian Miscellaneous Collections, 50 (2), 139 - 173." type="journal article">Dall, 1907</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C49C1B5FF121500FC41FE77FF1EFE13" authority="Schepman, 1908" authorityName="Schepman" authorityYear="1908" class="Gastropoda" family="Cocculinidae" genus="Cocculina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Archaeogastropoda" pageId="4" pageNumber="193" phylum="Mollusca" rank="species" species="subcompressa">
<emphasis id="B93D6624FF121500FC41FE77FACBFE6F" box="[1025,1306,404,428]" italics="true" pageId="4" pageNumber="193">Cocculina subcompressa</emphasis>
<bibRefCitation id="EFD8C7C7FF121500FD60FE77FF1EFE13" pageId="4" pageNumber="193" refId="ref3437" refString="Schepman, M. M. (1908) The Prosobranchia of the Siboga Expedition. Part I. Rhipidoglossa and Docoglossa. Uitkomsten op Zoologisch, Botanisch, Oceanographisch en Geologisch Gebied verzameld in Nederlandsch Oost-Indie 1899 - 1900 aan boord H. M. Siboga, 49, 1 - 95." type="journal article">Schepman, 1908</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="EFD8C7C7FF121500F89DFE5BFE47FE13" box="[221,406,439,464]" pageId="4" pageNumber="193" refId="ref3652" refString="Wu, W. L. &amp; Lee, Y. C. (2005) The Taiwan common mollusks in color. The Forestry Bureau, the Council of Agriculture, Taipei, 294 pp. [in Chinese]" type="book">Wu &amp; Lee 2005</bibRefCitation>
;
<bibRefCitation id="EFD8C7C7FF121500F9E2FE54FDF1FE13" box="[418,544,439,464]" pageId="4" pageNumber="193" refId="ref2599" refString="Chen, W. D. (2005) Shells from Donggang Township. 2. The Pei-yo, 31, 48 - 53. [in Chinese]" type="journal article">Chen 2005</bibRefCitation>
), may represent the geographically closest congeners to
<taxonomicName id="4C49C1B5FF121500FCD6FE5BFA4DFE13" authority="Zhang &amp; Zhang, 2018" authorityName="Zhang &amp; Zhang" authorityYear="2018" box="[1174,1436,440,464]" class="Gastropoda" family="Cocculinidae" genus="Cocculina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Archaeogastropoda" pageId="4" pageNumber="193" phylum="Mollusca" rank="species" species="delphinicula" status="sp. nov.">
<emphasis id="B93D6624FF121500FCD6FE5BFA4DFE13" box="[1174,1436,440,464]" italics="true" pageId="4" pageNumber="193">Cocculina delphinicula</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20EDB5FFF121500F8D7FE3FFF27FE36" box="[151,246,476,501]" pageId="4" pageNumber="193" rank="species">
<emphasis id="B93D6624FF121500F8D7FE3FFF27FE36" bold="true" box="[151,246,476,501]" pageId="4" pageNumber="193">sp. nov.</emphasis>
</taxonomicNameLabel>
Of those,
<taxonomicName id="4C49C1B5FF121500F93DFE3FFCE6FE37" authority="Hasegawa, 1997" authorityName="Hasegawa" authorityYear="1997" box="[381,823,476,501]" class="Gastropoda" family="Cocculinidae" genus="Cocculina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Archaeogastropoda" pageId="4" pageNumber="193" phylum="Mollusca" rank="species" species="tenuitesta">
<emphasis id="B93D6624FF121500F93DFE3FFDBBFE37" box="[381,618,476,500]" italics="true" pageId="4" pageNumber="193">Cocculina tenuitesta</emphasis>
<bibRefCitation id="EFD8C7C7FF121500FA38FE3EFCE6FE37" box="[632,823,476,501]" pageId="4" pageNumber="193" refId="ref2995" refString="Hasegawa, K. (1997) Sunken wood-associated gastropods collected from Suruga Bay, Pacific side of the central Honshu, Japan, with descriptions of 12 new species. National Science Museum Monographs, 12, 59 - 123." type="journal article">Hasegawa, 1997</bibRefCitation>
</taxonomicName>
resembles the new species in shell size. However,
<taxonomicName id="4C49C1B5FF121500F8D7FDE3FE72FDDB" authority="Zhang &amp; Zhang, 2018" authorityName="Zhang &amp; Zhang" authorityYear="2018" box="[151,419,512,536]" class="Gastropoda" family="Cocculinidae" genus="Cocculina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Archaeogastropoda" pageId="4" pageNumber="193" phylum="Mollusca" rank="species" species="delphinicula" status="sp. nov.">
<emphasis id="B93D6624FF121500F8D7FDE3FE72FDDB" box="[151,419,512,536]" italics="true" pageId="4" pageNumber="193">Cocculina delphinicula</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20EDB5FFF121500F9EFFE1CFDDEFDDB" box="[431,527,511,536]" pageId="4" pageNumber="193" rank="species">
<emphasis id="B93D6624FF121500F9EFFE1CFDDEFDDB" bold="true" box="[431,527,511,536]" pageId="4" pageNumber="193">sp. nov.</emphasis>
</taxonomicNameLabel>
differs from this species in having a depressed shell with regularly spaced, concentric growth lines and finer radial threads, a sparsely pitted shell surface and a protoconch that has wavy ridges and numerous microscopic pits.
</paragraph>
</subSubSection>
</treatment>
</document>
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