214 lines
28 KiB
XML
214 lines
28 KiB
XML
<document id="41603FE01808EF0CB26C8DAB319990B3" ID-CLB-Dataset="26707" ID-DOI="10.11646/zootaxa.4609.2.8" ID-GBIF-Dataset="4b9fed80-8bb0-49eb-af85-7dbdbaf0d181" ID-ISSN="1175-5326" ID-Zenodo-Dep="3187667" ID-ZooBank="77EF20BE-A8AD-4B4D-8380-3B0E622CE4C5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1558594389829" checkinUser="plazi" docAuthor="Mielke, Carlos G. C., Grehan, John R. & Grados, Juan" docDate="2019" docId="03BA0223964FFFBBFF4EF73A4653334D" docLanguage="en" docName="zootaxa.4609.2.8.pdf" docOrigin="Zootaxa 4609 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Walkeriella Mielke & Grehan & Grados 2019, gen. nov." docType="treatment" docVersion="9" lastPageNumber="337" masterDocId="FF837A5B964EFFB8FFD9F1274474322A" masterDocTitle="Description of Walkeriella miraculosa, gen. n. et sp. n. from South-East Peru (Lepidoptera: Hepialoidea: Hepialidae)" masterLastPageNumber="342" masterPageNumber="335" pageNumber="336" updateTime="1698731842222" updateUser="ExternalLinkService">
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<mods:title id="8B2066DE4175F6465AD13A1BA2554DCC">Description of Walkeriella miraculosa, gen. n. et sp. n. from South-East Peru (Lepidoptera: Hepialoidea: Hepialidae)</mods:title>
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<mods:namePart id="DA855ADEEE01AB0E382B5188C03C477C">Mielke, Carlos G. C.</mods:namePart>
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<mods:namePart id="04281431FB35EAE5599DC7DAAE63C057">Grehan, John R.</mods:namePart>
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<mods:name id="AAD68C5F8FC40D4C206DA7C1B27A3CB9" type="personal">
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<mods:roleTerm id="A3747FC5C2D435042A252FCC62F2E1D6">Author</mods:roleTerm>
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<mods:namePart id="2C9E2565C84F3444C8A057476AA248C1">Grados, Juan</mods:namePart>
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<mods:typeOfResource id="D993E361C7E92FBF13BDF90A79E0B526">text</mods:typeOfResource>
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<mods:title id="5B560C4AD5ACA8CA170BA35E40F129E3">Zootaxa</mods:title>
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<mods:part id="15FB01C6315A7E07BE2EF99B19AFB77E">
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<mods:date id="B7B55E23F520F796C404CE402944FE77">2019</mods:date>
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<mods:number id="B688F0957BCE91B12FA5FFCCB7FF7F42">2019-05-23</mods:number>
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<treatment id="03BA0223964FFFBBFF4EF73A4653334D" ID-DOI="http://doi.org/10.5281/zenodo.5678533" ID-GBIF-Taxon="157251209" ID-Zenodo-Dep="5678533" LSID="urn:lsid:plazi:treatment:03BA0223964FFFBBFF4EF73A4653334D" httpUri="http://treatment.plazi.org/id/03BA0223964FFFBBFF4EF73A4653334D" lastPageId="3" lastPageNumber="337" pageId="1" pageNumber="336">
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<subSubSection id="C309E0BE964FFFB9FF4EF73A45E2341D" box="[151,406,1565,1592]" pageId="1" pageNumber="336" type="nomenclature">
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<paragraph id="8BACB335964FFFB9FF4EF73A45E2341D" blockId="1.[151,406,1565,1592]" box="[151,406,1565,1592]" pageId="1" pageNumber="336">
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<heading id="D0E40459964FFFB9FF4EF73A45E2341D" bold="true" box="[151,406,1565,1592]" fontSize="11" level="1" pageId="1" pageNumber="336" reason="1">
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<emphasis id="B9676F27964FFFB9FF4EF73A45E2341D" bold="true" box="[151,406,1565,1592]" pageId="1" pageNumber="336">
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<taxonomicName id="4C13C8B6964FFFB9FF4EF73A45553412" authority="Mielke & Grehan & Grados, 2019" authorityName="Mielke & Grehan & Grados" authorityYear="2019" box="[151,289,1565,1592]" class="Insecta" family="Hepialidae" genus="Walkeriella" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="336" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B9676F27964FFFB9FF4EF73A45553412" bold="true" box="[151,289,1565,1592]" italics="true" pageId="1" pageNumber="336">Walkeriella</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A254D25C964FFFB9FEF1F73A45E2341D" box="[296,406,1565,1591]" pageId="1" pageNumber="336" rank="genus">gen. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C309E0BE964FFFB9FF4EF74547A13453" box="[151,981,1632,1658]" pageId="1" pageNumber="336" type="materials_examined">
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<paragraph id="8BACB335964FFFB9FF4EF74547A13453" blockId="1.[151,981,1632,1658]" box="[151,981,1632,1658]" pageId="1" pageNumber="336">
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<typeStatus id="54A80D97964FFFB9FF4EF74544BD3450" box="[151,201,1634,1658]" pageId="1" pageNumber="336">Type</typeStatus>
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species:
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<taxonomicName id="4C13C8B6964FFFB9FEFFF74646673453" authority="Mielke & Grehan & Grados, 2019" authorityName="Mielke & Grehan & Grados" authorityYear="2019" box="[294,531,1633,1657]" class="Insecta" family="Hepialidae" genus="Walkeriella" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="336" phylum="Arthropoda" rank="species" species="miraculosa" status="sp. nov.">
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<emphasis id="B9676F27964FFFB9FEFFF74646673453" box="[294,531,1633,1657]" italics="true" pageId="1" pageNumber="336">Walkeriella miraculosa</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A254D25C964FFFB9FDC0F746461D3453" box="[537,617,1633,1657]" pageId="1" pageNumber="336" rank="species">
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<emphasis id="B9676F27964FFFB9FDC0F746461D3453" bold="true" box="[537,617,1633,1657]" pageId="1" pageNumber="336">sp. nov.</emphasis>
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</taxonomicNameLabel>
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, monotypic by present designation.
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</paragraph>
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</subSubSection>
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<subSubSection id="C309E0BE964FFFB9FF4EF780479A355F" pageId="1" pageNumber="336" type="diagnosis">
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<paragraph id="8BACB335964FFFB9FF4EF780479A355F" blockId="1.[151,1437,1703,1982]" pageId="1" pageNumber="336">
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<emphasis id="B9676F27964FFFB9FF4EF780457E34E8" bold="true" box="[151,266,1703,1730]" pageId="1" pageNumber="336">Diagnosis</emphasis>
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. Externally distinguished from all other Pan-American
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<taxonomicName id="4C13C8B6964FFFB9FCA0F780478634EB" box="[889,1010,1703,1729]" class="Insecta" family="Hepialidae" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="336" phylum="Arthropoda" rank="family">Hepialidae</taxonomicName>
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by a large white spot edged with dark brown at the base of each discal cell and by the large curved spot between Rs1 and M
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<subScript id="1797B170964FFFB9FBBDF7FA401834C1" attach="left" box="[1124,1132,1757,1771]" fontSize="6" pageId="1" pageNumber="336">1</subScript>
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at the outer anterior discal cell.
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<taxonomicName id="4C13C8B6964FFFB9FF17F7D7453A3520" authority="Mielke & Grehan & Grados, 2019" authorityName="Mielke & Grehan & Grados" authorityYear="2019" box="[206,334,1776,1802]" class="Insecta" family="Hepialidae" genus="Walkeriella" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="336" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B9676F27964FFFB9FF17F7D7453A3520" box="[206,334,1776,1802]" italics="true" pageId="1" pageNumber="336">Walkeriella</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A254D25C964FFFB9FE8CF7D745D03520" box="[341,420,1776,1802]" pageId="1" pageNumber="336" rank="genus">
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<emphasis id="B9676F27964FFFB9FE8CF7D7450B3520" bold="true" box="[341,383,1776,1802]" pageId="1" pageNumber="336">gen</emphasis>
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.
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<emphasis id="B9676F27964FFFB9FE54F7D745E93520" bold="true" box="[397,413,1776,1802]" pageId="1" pageNumber="336">n</emphasis>
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.
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</taxonomicNameLabel>
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is also recognized by the distinctive male genitalia, particularly the valva with its concave inner (medial) surface extending from the base to the apex in contrast to either a blade-like shape or lobular shape in the other American species, and by the ‘oxycanine’ venation on the forewing and ‘hepialine’ venation (
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<bibRefCitation id="EF82CEC4964FFFB9FAC5F61044A4355C" author="Dumbleton, L. J." pageId="1" pageNumber="336" pagination="920 - 981" refId="ref3611" refString="Dumbleton, L. J. (1966) Genitalia, classification and zoogeography of the New Zealand Hepialidae (Lepidoptera). New Zealand Journal of Science, 9, 920 - 981." type="journal article" year="1966">Dumbleton 1966</bibRefCitation>
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) on the hindwing with partial fusion of Rs1+Rs2 and Rs3 at their base.
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</paragraph>
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</subSubSection>
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<subSubSection id="C309E0BE964FFFBAFF1EF65845EB348E" lastPageId="2" lastPageNumber="337" pageId="1" pageNumber="336" type="description">
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<paragraph id="8BACB335964FFFB9FF1EF65847A33597" blockId="1.[151,1437,1703,1982]" pageId="1" pageNumber="336">
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<emphasis id="B9676F27964FFFB9FF1EF658452535B0" bold="true" box="[199,337,1919,1946]" pageId="1" pageNumber="336">Description</emphasis>
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.
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<emphasis id="B9676F27964FFFB9FE87F6A745EC35B0" box="[350,408,1920,1946]" italics="true" pageId="1" pageNumber="336">Male</emphasis>
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(
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<figureCitation id="1328AFB0964FFFB9FE7FF658467F35B0" box="[422,523,1919,1946]" captionStart-0="FIGURES 2−4" captionStart-1="FIGURES 5−9" captionStartId-0="3.[151,264,1478,1503]" captionStartId-1="4.[151,264,1830,1855]" captionTargetBox-0="[208,1378,683,1469]" captionTargetBox-1="[151,1435,304,1786]" captionTargetId-0="figure@3.[202,1386,676,1473]" captionTargetId-1="figure@4.[151,1435,294,1790]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="FIGURES 2−4. Walkeriella miraculosa sp. nov., male holotype habitus. 2, Dorsal view (a), ventral view (b). Scale bar = 1 cm. 3, Forewing ornamentation (adc = anterior discal cell, cdc = central discal cell, mb = marginal band, pdb = postdiscal band, pdc = posterior discal cell, pmb = premarginal band, smb = submarginal band). 4, Wing venation.Arrow indicates the fusion of Rs1 + Rs2 and Rs3." captionText-1="FIGURES 5−9. Walkeriella miraculosa sp. nov., male holotype morphological features. 5, Legs. Scale bar = 0.5 cm. 6, Tergosternal connection (ab = anterior break, db = dorsal brace, iz = intermediate zone, lb = lateral brace, SII = sternum II, TII = tergum II, tb = tergosternal bar, tl = tergal lobe). 7, Abdomen upper dorsal, lower ventral view. Scale bar = 0.5 mm. 8, Tergum (left) and sternum (right) VII and VIII. Arrow indicates the projections of tergite VII. Scale bar = 0.5 mm. 9, Genitalia: lateral view (9a), ventral view (9b; arrows indicates the fultura superior), anterior view (9c), phallus everted (9d). Scale bar = 0.5 mm." httpUri-0="https://zenodo.org/record/3187671/files/figure.png" httpUri-1="https://zenodo.org/record/3187673/files/figure.png" pageId="1" pageNumber="336">Figs 2–9</figureCitation>
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).
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<emphasis id="B9676F27964FFFB9FDC4F6A7462E35B0" box="[541,602,1920,1946]" italics="true" pageId="1" pageNumber="336">Head</emphasis>
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. Clypeus anteriorly glabrous and mesally projected, differentiated from the frons. Labial palpus with one tiny, rounded palpomere. Antenna lamellate.
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</paragraph>
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<caption id="DF6CE3BD964CFFBAFF4EF50046BA366A" box="[151,718,1063,1088]" httpUri="https://zenodo.org/record/3187669/files/figure.png" pageId="2" pageNumber="337" startId="2.[151,250,1063,1088]" targetBox="[151,1436,164,1020]" targetPageId="2">
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<paragraph id="8BACB335964CFFBAFF4EF50046BA366A" blockId="2.[151,718,1063,1088]" box="[151,718,1063,1088]" pageId="2" pageNumber="337">
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<emphasis id="B9676F27964CFFBAFF4EF5004579366A" bold="true" box="[151,269,1063,1088]" pageId="2" pageNumber="337">FIGURE 1</emphasis>
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. Biotope of
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<taxonomicName id="4C13C8B6964CFFBAFE52F500460C3615" authority="Mielke & Grehan & Grados, 2019" authorityName="Mielke & Grehan & Grados" authorityYear="2019" box="[395,632,1063,1087]" class="Insecta" family="Hepialidae" genus="Walkeriella" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="337" phylum="Arthropoda" rank="species" species="miraculosa" status="sp. nov.">
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<emphasis id="B9676F27964CFFBAFE52F500460C3615" box="[395,632,1063,1087]" italics="true" pageId="2" pageNumber="337">Walkeriella miraculosa</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A254D25C964CFFBAFDA7F50F46BA366A" box="[638,718,1064,1088]" pageId="2" pageNumber="337" rank="species">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</caption>
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<paragraph id="8BACB335964CFFBAFF1EF549469036FA" blockId="2.[151,1437,1134,2024]" pageId="2" pageNumber="337">
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<emphasis id="B9676F27964CFFBAFF1EF549456C36A2" box="[199,280,1134,1160]" italics="true" pageId="2" pageNumber="337">Thorax</emphasis>
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. Legs: Distitarsus without arolium. Fore- and hindwings: venation typical for cibyrine genera with Sc and R aligned very close together over much of the distal wing (
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<bibRefCitation id="EF82CEC4964CFFBAFCB3F5B540F03686" author="Grehan, J. R. & Rawlins, J. E." box="[874,1156,1170,1196]" pageId="2" pageNumber="337" pagination="165 - 192" refId="ref3926" refString="Grehan, J. R. & Rawlins, J. E. (2018) Taxonomic revision and vicariance biogeography of the Central and South American ghost moth genus Druceiella (Lepidoptera: Hepialidae). Annals of Carnegie Museum, 85 (2), 165 - 192." type="journal article" year="2018">Grehan & Rawlins 2018</bibRefCitation>
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). Forewing with Sc1 and hindwing with vestigial (barely visible) Sc1 (
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<figureCitation id="1328AFB0964CFFBAFD48F59146A336FA" box="[657,727,1206,1232]" captionStart="FIGURES 2−4" captionStartId="3.[151,264,1478,1503]" captionTargetBox="[208,1378,683,1469]" captionTargetId="figure@3.[202,1386,676,1473]" captionTargetPageId="3" captionText="FIGURES 2−4. Walkeriella miraculosa sp. nov., male holotype habitus. 2, Dorsal view (a), ventral view (b). Scale bar = 1 cm. 3, Forewing ornamentation (adc = anterior discal cell, cdc = central discal cell, mb = marginal band, pdb = postdiscal band, pdc = posterior discal cell, pmb = premarginal band, smb = submarginal band). 4, Wing venation.Arrow indicates the fusion of Rs1 + Rs2 and Rs3." httpUri="https://zenodo.org/record/3187671/files/figure.png" pageId="2" pageNumber="337">Fig. 4</figureCitation>
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).
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</paragraph>
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<paragraph id="8BACB335964CFFBAFF1EF5FD448637AF" blockId="2.[151,1437,1134,2024]" pageId="2" pageNumber="337">
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<emphasis id="B9676F27964CFFBAFF1EF5FD454536DE" box="[199,305,1242,1268]" italics="true" pageId="2" pageNumber="337">Abdomen</emphasis>
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(
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<figureCitation id="1328AFB0964CFFBAFE98F5FD45DC36DF" box="[321,424,1242,1269]" captionStart="FIGURES 5−9" captionStartId="4.[151,264,1830,1855]" captionTargetBox="[151,1435,304,1786]" captionTargetId="figure@4.[151,1435,294,1790]" captionTargetPageId="4" captionText="FIGURES 5−9. Walkeriella miraculosa sp. nov., male holotype morphological features. 5, Legs. Scale bar = 0.5 cm. 6, Tergosternal connection (ab = anterior break, db = dorsal brace, iz = intermediate zone, lb = lateral brace, SII = sternum II, TII = tergum II, tb = tergosternal bar, tl = tergal lobe). 7, Abdomen upper dorsal, lower ventral view. Scale bar = 0.5 mm. 8, Tergum (left) and sternum (right) VII and VIII. Arrow indicates the projections of tergite VII. Scale bar = 0.5 mm. 9, Genitalia: lateral view (9a), ventral view (9b; arrows indicates the fultura superior), anterior view (9c), phallus everted (9d). Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/3187673/files/figure.png" pageId="2" pageNumber="337">Figs 6–8</figureCitation>
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). Well sclerotized, tuberculate plates distinct; tergum I with lateral ridge meeting anteriolateral tuberculate plate; anterior margin with weakly sclerotized ridge, not fused to dorsal tergal brace of tergosternal connection. Tergosternal connection with short dorsal and lateral brace curving together next to broad, triangular intermediate zone with prominent posterior-ventral tergal knob; upper anterior margin with small space or lacuna (
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<figureCitation id="1328AFB0964CFFBAFF79F44D449037AF" box="[160,228,1386,1413]" captionStart="FIGURES 5−9" captionStartId="4.[151,264,1830,1855]" captionTargetBox="[151,1435,304,1786]" captionTargetId="figure@4.[151,1435,294,1790]" captionTargetPageId="4" captionText="FIGURES 5−9. Walkeriella miraculosa sp. nov., male holotype morphological features. 5, Legs. Scale bar = 0.5 cm. 6, Tergosternal connection (ab = anterior break, db = dorsal brace, iz = intermediate zone, lb = lateral brace, SII = sternum II, TII = tergum II, tb = tergosternal bar, tl = tergal lobe). 7, Abdomen upper dorsal, lower ventral view. Scale bar = 0.5 mm. 8, Tergum (left) and sternum (right) VII and VIII. Arrow indicates the projections of tergite VII. Scale bar = 0.5 mm. 9, Genitalia: lateral view (9a), ventral view (9b; arrows indicates the fultura superior), anterior view (9c), phallus everted (9d). Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/3187673/files/figure.png" pageId="2" pageNumber="337">Fig. 6</figureCitation>
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).
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</paragraph>
|
||
<paragraph id="8BACB335964CFFBAFF1EF4A9471E34AA" blockId="2.[151,1437,1134,2024]" pageId="2" pageNumber="337">
|
||
<emphasis id="B9676F27964CFFBAFF1EF4A945043782" box="[199,368,1422,1448]" italics="true" pageId="2" pageNumber="337">Male genitalia</emphasis>
|
||
(
|
||
<figureCitation id="1328AFB0964CFFBAFE5BF4A945BF3782" box="[386,459,1422,1448]" captionStart="FIGURES 5−9" captionStartId="4.[151,264,1830,1855]" captionTargetBox="[151,1435,304,1786]" captionTargetId="figure@4.[151,1435,294,1790]" captionTargetPageId="4" captionText="FIGURES 5−9. Walkeriella miraculosa sp. nov., male holotype morphological features. 5, Legs. Scale bar = 0.5 cm. 6, Tergosternal connection (ab = anterior break, db = dorsal brace, iz = intermediate zone, lb = lateral brace, SII = sternum II, TII = tergum II, tb = tergosternal bar, tl = tergal lobe). 7, Abdomen upper dorsal, lower ventral view. Scale bar = 0.5 mm. 8, Tergum (left) and sternum (right) VII and VIII. Arrow indicates the projections of tergite VII. Scale bar = 0.5 mm. 9, Genitalia: lateral view (9a), ventral view (9b; arrows indicates the fultura superior), anterior view (9c), phallus everted (9d). Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/3187673/files/figure.png" pageId="2" pageNumber="337">Fig. 9</figureCitation>
|
||
). Tegumen indistinct, likely fused to the pseudotegumen, ventral portion articulated with saccus. Saccus U-shaped anteriorly; posterior edge W-shaped with lateral expansions on either side, with indentation mesally. Tergal lobes forming narrow arch meeting anterior margin of pseudotegumen. Pseudotegumen compound of two conspicuous C-shaped lateral plates fused together on their anterior-ventral margins; its dorsal projection asymmetrical, ventral projection symmetrical with strongly sclerotized apex. Fultura inferior trapezoidal. Fultura superior comprising two reduced and narrow sclerotized longitudinal bands, one on either side. Valvae slightly curved, tapered and lightly setose. Phallus membranous.
|
||
</paragraph>
|
||
<paragraph id="8BACB335964CFFBAFF1EF7AD45EB348E" blockId="2.[151,1437,1134,2024]" box="[199,415,1674,1700]" pageId="2" pageNumber="337">
|
||
<emphasis id="B9676F27964CFFBAFF1EF7AD456F348E" box="[199,283,1674,1700]" italics="true" pageId="2" pageNumber="337">Female</emphasis>
|
||
. Unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C309E0BE964CFFBAFF1EF7894576351E" pageId="2" pageNumber="337" type="etymology">
|
||
<paragraph id="8BACB335964CFFBAFF1EF7894576351E" blockId="2.[151,1437,1134,2024]" pageId="2" pageNumber="337">
|
||
<emphasis id="B9676F27964CFFBAFF1EF789453C34E2" bold="true" box="[199,328,1710,1736]" pageId="2" pageNumber="337">Etymology</emphasis>
|
||
. This new genus is named for Francis Walker, a British entomologist, who described some Neotropical
|
||
<taxonomicName id="4C13C8B6964CFFBAFF2DF7F5451934C6" box="[244,365,1746,1772]" class="Insecta" family="Hepialidae" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="337" phylum="Arthropoda" rank="family">Hepialidae</taxonomicName>
|
||
species in the 19
|
||
<superScript id="7C661E7D964CFFBAFDEBF7F6464B34F5" attach="left" box="[562,575,1745,1759]" fontSize="6" pageId="2" pageNumber="337">th</superScript>
|
||
century. The name follows the tradition of
|
||
<taxonomicName id="4C13C8B6964CFFBAFBF0F7F540D534C6" authorityName="Viette" authorityYear="1949" box="[1065,1185,1746,1772]" class="Insecta" family="Hepialidae" genus="Druceiella" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="337" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9676F27964CFFBAFBF0F7F540D534C6" box="[1065,1185,1746,1772]" italics="true" pageId="2" pageNumber="337">Druceiella</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C13C8B6964CFFBAFB69F7F5455D353A" authority="Grehan & C. Mielke" authorityName="Grehan & C. Mielke" authorityYear="2018" class="Insecta" family="Hepialidae" genus="Dugdaleiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="337" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9676F27964CFFBAFB69F7F5413634C6" box="[1200,1346,1746,1772]" italics="true" pageId="2" pageNumber="337">Dugdaleiella</emphasis>
|
||
Grehan & C. Mielke
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C13C8B6964CFFBAFEECF7D145A6353A" authorityName="Viette" authorityYear="1949" box="[309,466,1782,1808]" class="Insecta" family="Limacodidae" genus="Hampsoniella" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="337" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9676F27964CFFBAFEECF7D145A6353A" box="[309,466,1782,1808]" italics="true" pageId="2" pageNumber="337">Hampsoniella</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C13C8B6964CFFBAFE38F7D14720353A" authority="Grehan & C. Mielke" authorityName="Grehan & C. Mielke" box="[481,852,1782,1808]" class="Bacillariophyceae" genus="Kozloviella" higherTaxonomySource="CoL" kingdom="Chromista" pageId="2" pageNumber="337" phylum="Ochrophyta" rank="genus">
|
||
<emphasis id="B9676F27964CFFBAFE38F7D14615353A" box="[481,609,1782,1808]" italics="true" pageId="2" pageNumber="337">Kozloviella</emphasis>
|
||
Grehan & C. Mielke
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C13C8B6964CFFBAFC48F7D140D4353A" authority="Viette. The" authorityName="Viette. The" box="[913,1184,1782,1808]" class="Insecta" family="Hepialidae" genus="Pfitzneriella" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="337" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9676F27964CFFBAFC48F7D1406F353A" box="[913,1051,1782,1808]" italics="true" pageId="2" pageNumber="337">Pfitzneriella</emphasis>
|
||
Viette. The
|
||
</taxonomicName>
|
||
gender of the name is feminine.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C309E0BE964CFFBBFF1EF6194653334D" lastPageId="3" lastPageNumber="338" pageId="2" pageNumber="337" type="discussion">
|
||
<paragraph id="8BACB335964CFFBBFF1EF6194653334D" blockId="2.[151,1437,1134,2024]" lastBlockId="3.[151,1436,153,359]" lastPageId="3" lastPageNumber="338" pageId="2" pageNumber="337">
|
||
<emphasis id="B9676F27964CFFBAFF1EF61945403572" bold="true" box="[199,308,1854,1880]" pageId="2" pageNumber="337">Remarks</emphasis>
|
||
. The presence of
|
||
<taxonomicName id="4C13C8B6964CFFBAFE2FF61946023572" authority="Mielke & Grehan & Grados, 2019" authorityName="Mielke & Grehan & Grados" authorityYear="2019" box="[502,630,1854,1880]" class="Insecta" family="Hepialidae" genus="Walkeriella" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="337" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B9676F27964CFFBAFE2FF61946023572" box="[502,630,1854,1880]" italics="true" pageId="2" pageNumber="337">Walkeriella</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A254D25C964CFFBAFDA5F61946BE3572" box="[636,714,1854,1880]" pageId="2" pageNumber="337" rank="genus">
|
||
<emphasis id="B9676F27964CFFBAFDA5F61946BE3572" bold="true" box="[636,714,1854,1880]" pageId="2" pageNumber="337">gen. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
in southern
|
||
<collectingCountry id="F304F3A5964CFFBAFC88F61947F33572" box="[849,903,1854,1880]" name="Peru" pageId="2" pageNumber="337">Peru</collectingCountry>
|
||
, a relatively well surveyed country, is surprising since no other similar taxon is so far known from the Neotropical region. The presence of a strongly developed posterior lateral knob, a broad intermediate connection between the tergosternal bar and the lateral ridge and a broken anterior margin at the tergosternal connection, along with a very close parallel position between the outer hindwing Sc and R veins support inclusion of
|
||
<taxonomicName id="4C13C8B6964CFFBAFDE9F6E946C435C2" authority="Mielke & Grehan & Grados, 2019" authorityName="Mielke & Grehan & Grados" authorityYear="2019" box="[560,688,1998,2024]" class="Insecta" family="Hepialidae" genus="Walkeriella" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="337" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B9676F27964CFFBAFDE9F6E946C435C2" box="[560,688,1998,2024]" italics="true" pageId="2" pageNumber="337">Walkeriella</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A254D25C964CFFBAFD61F6E9477335C2" box="[696,775,1998,2024]" pageId="2" pageNumber="337" rank="genus">
|
||
<emphasis id="B9676F27964CFFBAFD61F6E9477335C2" bold="true" box="[696,775,1998,2024]" pageId="2" pageNumber="337">gen. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
within the ‘cibyrine’ cluster of genera (
|
||
<bibRefCitation id="EF82CEC4964CFFBAFB1FF6E9412A35C2" author="Grehan, J. R." box="[1222,1374,1998,2024]" pageId="2" pageNumber="337" pagination="1 - 30" refId="ref3705" refString="Grehan, J. R. (2012) Morphological evidence for phylogenetic relationships within the Hepialidae (Lepidoptera: Exoporia). Bulletin of the Buffalo Society of Natural Sciences, 42, 1 - 30." type="journal article" year="2012">Grehan 2012</bibRefCitation>
|
||
). The configuration of Rs1+Rs2 and Rs3 is unique among all
|
||
<taxonomicName id="4C13C8B6964DFFBBFCC9F1BE47FD3299" box="[784,905,153,179]" class="Insecta" family="Hepialidae" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="338" phylum="Arthropoda" rank="family">Hepialidae</taxonomicName>
|
||
genera and this is the first case that both hepialine and oxycanine venations are found in the same species. In the
|
||
<typeStatus id="54A80D97964DFFBBFC7CF19A407332FD" box="[933,1031,189,215]" pageId="3" pageNumber="338" type="holotype">holotype</typeStatus>
|
||
the left forewing has an oxycanine arrangement (Rs3 joining the common stalk of Rs1+Rs2) whereas the right wing has a hepialine arrangement with Rs3 joining the base of Rs4. The hindwing arrangement is hepialine on both sides. The
|
||
<typeStatus id="54A80D97964DFFBBFB49F02240843335" box="[1168,1264,261,287]" pageId="3" pageNumber="338" type="paratype">paratype</typeStatus>
|
||
has oxycanine venation on both forewings and hepialine venation on both hindwings. Further investigation will be necessary for a better understanding of this pattern.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |