treatments-xml/data/03/B9/33/03B93371FFF5FFA6FF73FA28B6860506.xml

<document id="C550699F15275003C9F3A9D229982A4D" ID-DOI="10.5281/zenodo.196263" ID-GBIF-Dataset="3cd616a5-134b-46d2-898b-9dec696d1dfc" ID-ISSN="1175-5326" ID-Zenodo-Dep="196263" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460241135365" checkinUser="plazi" docAuthor="Fuangarworn, Marut" docDate="2010" docId="03B93371FFF5FFA6FF73FA28B6860506" docLanguage="en" docName="zt02521p036.pdf" docOrigin="Zootaxa 2521" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Phyllochthonius peniculus Fuangarworn, 2010, n. sp." docType="treatment" docVersion="9" lastPageNumber="35" masterDocId="FF804B09FFF1FFAFFFE4FFD5B306010A" masterDocTitle="Two new species of the oribatid mite genus Phyllochthonius Travé, 1967 (Acari: Oribatida: Phyllochthoniidae) from Thailand" masterLastPageNumber="36" masterPageNumber="26" pageNumber="30" updateTime="1698240851758" updateUser="plazi">
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<mods:title id="6F86E370FA8FC03539F00CE237FE31C0">Two new species of the oribatid mite genus Phyllochthonius Travé, 1967 (Acari: Oribatida: Phyllochthoniidae) from Thailand</mods:title>
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<mods:namePart id="40A47BA3B4029323BBE67490670A83D2">Fuangarworn, Marut</mods:namePart>
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<mods:date id="E76EB1A07A4F5785E135234C3F395CE9">2010</mods:date>
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<treatment id="03B93371FFF5FFA6FF73FA28B6860506" ID-DOI="http://doi.org/10.5281/zenodo.6197580" ID-GBIF-Taxon="119389768" ID-Zenodo-Dep="6197580" LSID="urn:lsid:plazi:treatment:03B93371FFF5FFA6FF73FA28B6860506" httpUri="http://treatment.plazi.org/id/03B93371FFF5FFA6FF73FA28B6860506" lastPageId="9" lastPageNumber="35" pageId="4" pageNumber="30">
<subSubSection id="C30AD1ECFFF5FFABFF73FA28B3E80730" pageId="4" pageNumber="30" type="nomenclature">
<paragraph id="8BAF8267FFF5FFABFF73FA28B122071D" blockId="4.[151,548,1533,1594]" box="[151,548,1533,1559]" pageId="4" pageNumber="30">
<heading id="D0E7350BFFF5FFABFF73FA28B122071D" bold="true" box="[151,548,1533,1559]" fontSize="11" level="1" pageId="4" pageNumber="30" reason="1">
<emphasis id="B9645E75FFF5FFABFF73FA28B122071D" bold="true" box="[151,548,1533,1559]" pageId="4" pageNumber="30">
<taxonomicName id="4C10F9E4FFF5FFABFF73FA28B2DC071D" box="[151,474,1533,1559]" class="Arachnida" family="Phyllochthoniidae" genus="Phyllochthonius" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="30" phylum="Arthropoda" rank="species" species="peniculus" status="sp. nov.">
<emphasis id="B9645E75FFF5FFABFF73FA28B2DC071D" bold="true" box="[151,474,1533,1559]" italics="true" pageId="4" pageNumber="30">Phyllochthonius peniculus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A257E30EFFF5FFABFE05FA28B122071D" box="[481,548,1533,1559]" pageId="4" pageNumber="30" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BAF8267FFF5FFABFF73F9F5B3E80730" blockId="4.[151,548,1533,1594]" box="[151,238,1568,1594]" pageId="4" pageNumber="30">
(
<figureCitation id="132B9EE2FFF5FFABFF7BF9F5B3E30730" box="[159,229,1568,1594]" captionStart="FIGURE 5" captionStartId="7.[151,256,1968,1992]" captionTargetBox="[202,1378,409,1895]" captionTargetId="figure@7.[196,1386,400,1940]" captionTargetPageId="7" captionText="FIGURE 5. Phyllochthonius peniculus n. sp., female. A, dorsal view; B, interlamellar seta; C, ventral view (without legs); D, lateral view; E, seta e 2 with imaginary cross section." httpUri="https://zenodo.org/record/196268/files/figure.png" pageId="4" pageNumber="30">Fig. 5</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30AD1ECFFF5FFA6FF73F9BFB152032D" lastPageId="9" lastPageNumber="35" pageId="4" pageNumber="30" type="description">
<paragraph id="8BAF8267FFF5FFABFF73F9BFB3EC07A0" blockId="4.[151,1437,1642,2004]" pageId="4" pageNumber="30">
<emphasis id="B9645E75FFF5FFABFF73F9BFB238078E" bold="true" box="[151,318,1642,1668]" pageId="4" pageNumber="30">Adult female</emphasis>
. 
<emphasis id="B9645E75FFF5FFABFEA9F9BFB2E7078E" bold="true" box="[333,481,1642,1668]" pageId="4" pageNumber="30">Dimensions</emphasis>
(n=1). Body length 250. Body width (at the level of setae 
<emphasis id="B9645E75FFF5FFABFB48F9BEB7CE078E" box="[1196,1224,1643,1668]" italics="true" pageId="4" pageNumber="30">cp</emphasis>
) 125. Color light brown.
</paragraph>
<paragraph id="8BAF8267FFF5FFABFF22F960B73106DE" blockId="4.[151,1437,1642,2004]" pageId="4" pageNumber="30">
<emphasis id="B9645E75FFF5FFABFF22F960B25507C5" bold="true" box="[198,339,1717,1743]" pageId="4" pageNumber="30">Prodorsum</emphasis>
. Triangular in outline, posterior end slightly narrower than anterior margin of notogaster. Integument weakly sclerotized, surface smooth. Rostrum extended, with irregular teeth laterally. Area between interlamellar setae (
<emphasis id="B9645E75FFF5FFABFE08F8D5B1050613" box="[492,515,1792,1817]" italics="true" pageId="4" pageNumber="30">in</emphasis>
) mostly elevated, except concave anteriorly. Rostral setae (
<emphasis id="B9645E75FFF5FFABFB22F8D5B7D80613" box="[1222,1246,1792,1817]" italics="true" pageId="4" pageNumber="30">ro</emphasis>
), lamellar setae (
<emphasis id="B9645E75FFF5FFABFF45F8F0B3B00634" box="[161,182,1829,1854]" italics="true" pageId="4" pageNumber="30">le</emphasis>
), and anterior exobothridial setae (
<emphasis id="B9645E75FFF5FFABFDBFF8F0B1820634" box="[603,644,1829,1854]" italics="true" pageId="4" pageNumber="30">exa</emphasis>
) broadly plate-like, with foveae; of these 
<emphasis id="B9645E75FFF5FFABFB9FF8F0B7950634" box="[1147,1171,1829,1854]" italics="true" pageId="4" pageNumber="30">ro</emphasis>
smallest with pointed process, 
<emphasis id="B9645E75FFF5FFABFF1BF89EB212066E" box="[255,276,1867,1892]" italics="true" pageId="4" pageNumber="30">le</emphasis>
largest, their shape depicted in 
<figureCitation id="132B9EE2FFF5FFABFD76F89FB1E4066E" box="[658,738,1866,1892]" captionStart="FIGURE 5" captionStartId="7.[151,256,1968,1992]" captionTargetBox="[202,1378,409,1895]" captionTargetId="figure@7.[196,1386,400,1940]" captionTargetPageId="7" captionText="FIGURE 5. Phyllochthonius peniculus n. sp., female. A, dorsal view; B, interlamellar seta; C, ventral view (without legs); D, lateral view; E, seta e 2 with imaginary cross section." httpUri="https://zenodo.org/record/196268/files/figure.png" pageId="4" pageNumber="30">Figs. 5</figureCitation>
A and D. Setae 
<emphasis id="B9645E75FFF5FFABFC46F89EB0BF066E" box="[930,953,1867,1892]" italics="true" pageId="4" pageNumber="30">in</emphasis>
sub-elliptical, basally narrower than the distal part, and covered by micro-barbs (
<figureCitation id="132B9EE2FFF5FFABFD98F8A5B1C30680" box="[636,709,1904,1930]" captionStart="FIGURE 5" captionStartId="7.[151,256,1968,1992]" captionTargetBox="[202,1378,409,1895]" captionTargetId="figure@7.[196,1386,400,1940]" captionTargetPageId="7" captionText="FIGURE 5. Phyllochthonius peniculus n. sp., female. A, dorsal view; B, interlamellar seta; C, ventral view (without legs); D, lateral view; E, seta e 2 with imaginary cross section." httpUri="https://zenodo.org/record/196268/files/figure.png" pageId="4" pageNumber="30">Fig. 5</figureCitation>
B). Setae 
<emphasis id="B9645E75FFF5FFABFCDEF8A5B0650683" box="[826,867,1904,1929]" italics="true" pageId="4" pageNumber="30">exa</emphasis>
located at normal position but expand dorsally, able to cover bothridium as well as area in front of setae 
<emphasis id="B9645E75FFF5FFABFCA9F840B06206A4" box="[845,868,1941,1966]" italics="true" pageId="4" pageNumber="30">in</emphasis>
. Posterior exobothridial setae (
<emphasis id="B9645E75FFF5FFABFB05F840B60C06A4" box="[1249,1290,1941,1966]" italics="true" pageId="4" pageNumber="30">exp</emphasis>
) spine-like. Bothridia prominent, with barbed, long flagelliform sensilli (
<figureCitation id="132B9EE2FFF5FFABFCBBF86FB0A806DE" box="[863,942,1978,2004]" captionStart="FIGURE 5" captionStartId="7.[151,256,1968,1992]" captionTargetBox="[202,1378,409,1895]" captionTargetId="figure@7.[196,1386,400,1940]" captionTargetPageId="7" captionText="FIGURE 5. Phyllochthonius peniculus n. sp., female. A, dorsal view; B, interlamellar seta; C, ventral view (without legs); D, lateral view; E, seta e 2 with imaginary cross section." httpUri="https://zenodo.org/record/196268/files/figure.png" pageId="4" pageNumber="30">Figs. 5</figureCitation>
A &amp; D, 
<emphasis id="B9645E75FFF5FFABFBEBF86EB72306DE" box="[1039,1061,1979,2004]" italics="true" pageId="4" pageNumber="30">ss</emphasis>
).
</paragraph>
<caption id="DF6FD2EFFFF4FFAAFF73F852B2C506D4" httpUri="https://zenodo.org/record/196266/files/figure.png" pageId="5" pageNumber="31" targetBox="[215,1388,211,1886]" targetPageId="5">
<paragraph id="8BAF8267FFF4FFAAFF73F852B2C506D4" blockId="5.[151,1436,1927,2014]" pageId="5" pageNumber="31">
<emphasis id="B9645E75FFF4FFAAFF73F852B21C0695" bold="true" box="[151,282,1927,1951]" pageId="5" pageNumber="31">FIGURE 3.</emphasis>
<taxonomicName id="4C10F9E4FFF4FFAAFEC6F852B1660694" box="[290,608,1927,1950]" class="Arachnida" family="Phyllochthoniidae" genus="Phyllochthonius" kingdom="Animalia" order="Sarcoptiformes" pageId="5" pageNumber="31" phylum="Arthropoda" rank="species" species="ovatosetosus" status="sp. nov.">
<emphasis id="B9645E75FFF4FFAAFEC6F852B1660694" box="[290,608,1927,1950]" italics="true" pageId="5" pageNumber="31">Phyllochthonius ovatosetosus</emphasis>
</taxonomicName>
<emphasis id="B9645E75FFF4FFAAFD83F852B1A50695" bold="true" box="[615,675,1927,1951]" pageId="5" pageNumber="31">
<taxonomicNameLabel id="A257E30EFFF4FFAAFD83F852B1A50695" box="[615,675,1927,1951]" pageId="5" pageNumber="31" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, female. A, leg I, lateral (abaxial) view; B, leg II, dorsal view (setae 
<emphasis id="B9645E75FFF4FFAAFA6AF852B69D0694" box="[1422,1435,1927,1950]" italics="true" pageId="5" pageNumber="31">d</emphasis>
on femur–tibia not illustrated in place, but having the form of the figure at right); C, leg III, dorsal view; D, leg IV, dorso-lateral (abaxial) view.
</paragraph>
</caption>
<caption id="DF6FD2EFFFF7FFA9FF73F8DCB0C8064A" httpUri="https://zenodo.org/record/196267/files/figure.png" pageId="6" pageNumber="32" targetBox="[168,1424,213,1742]" targetPageId="6">
<paragraph id="8BAF8267FFF7FFA9FF73F8DCB0C8064A" blockId="6.[151,1436,1801,1856]" pageId="6" pageNumber="32">
<emphasis id="B9645E75FFF7FFA9FF73F8DCB21A062B" bold="true" box="[151,284,1801,1825]" pageId="6" pageNumber="32">FIGURE 4.</emphasis>
<taxonomicName id="4C10F9E4FFF7FFA9FEC7F8DCB162062A" box="[291,612,1801,1824]" class="Arachnida" family="Phyllochthoniidae" genus="Phyllochthonius" kingdom="Animalia" order="Sarcoptiformes" pageId="6" pageNumber="32" phylum="Arthropoda" rank="species" species="ovatosetosus" status="sp. nov.">
<emphasis id="B9645E75FFF7FFA9FEC7F8DCB162062A" box="[291,612,1801,1824]" italics="true" pageId="6" pageNumber="32">Phyllochthonius ovatosetosus</emphasis>
</taxonomicName>
<emphasis id="B9645E75FFF7FFA9FD8FF8DCB1AE062B" bold="true" box="[619,680,1801,1825]" pageId="6" pageNumber="32">
<taxonomicNameLabel id="A257E30EFFF7FFA9FD8FF8DCB1AE062B" box="[619,680,1801,1825]" pageId="6" pageNumber="32" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, immatures. A–B, larva, dorsal and ventral views, respectively; C–E, anogenital region of protonymph, deutonymph, and tritonymph, respectively.
</paragraph>
</caption>
<paragraph id="8BAF8267FFF7FFA8FF22F8BBB05B0066" blockId="6.[151,1436,1902,2003]" lastBlockId="7.[151,1437,152,364]" lastPageId="7" lastPageNumber="33" pageId="6" pageNumber="32">
<emphasis id="B9645E75FFF7FFA9FF22F8BBB24B0682" bold="true" box="[198,333,1902,1928]" pageId="6" pageNumber="32">Notogaster</emphasis>
. Generally elliptical in dorsal aspect, posterior rounded (
<figureCitation id="132B9EE2FFF7FFA9FC0FF8BBB73C0682" box="[1003,1082,1902,1928]" captionStart="FIGURE 5" captionStartId="7.[151,256,1968,1992]" captionTargetBox="[202,1378,409,1895]" captionTargetId="figure@7.[196,1386,400,1940]" captionTargetPageId="7" captionText="FIGURE 5. Phyllochthonius peniculus n. sp., female. A, dorsal view; B, interlamellar seta; C, ventral view (without legs); D, lateral view; E, seta e 2 with imaginary cross section." httpUri="https://zenodo.org/record/196268/files/figure.png" pageId="6" pageNumber="32">Figs. 5</figureCitation>
A &amp; C); moderately arched in lateral aspect (
<figureCitation id="132B9EE2FFF7FFA9FEA3F841B29606A4" box="[327,400,1940,1966]" captionStart="FIGURE 5" captionStartId="7.[151,256,1968,1992]" captionTargetBox="[202,1378,409,1895]" captionTargetId="figure@7.[196,1386,400,1940]" captionTargetPageId="7" captionText="FIGURE 5. Phyllochthonius peniculus n. sp., female. A, dorsal view; B, interlamellar seta; C, ventral view (without legs); D, lateral view; E, seta e 2 with imaginary cross section." httpUri="https://zenodo.org/record/196268/files/figure.png" pageId="6" pageNumber="32">Fig. 5</figureCitation>
D). Integument weakly sclerotized, surface smooth. Two transverse scissures present, located behind setae row 
<emphasis id="B9645E75FFF7FFA9FE26F86CB2C906D8" box="[450,463,1977,2002]" italics="true" pageId="6" pageNumber="32">c</emphasis>
and just in front of setae row 
<emphasis id="B9645E75FFF7FFA9FCD5F86CB03806D8" box="[817,830,1977,2002]" italics="true" pageId="6" pageNumber="32">e</emphasis>
. Sixteen pairs of notogastral setae present; margins of adjacent setae usually meet. Setae 
<emphasis id="B9645E75FFF6FFA8FDA8FF4DB16101BB" box="[588,615,152,177]" italics="true" pageId="7" pageNumber="33">c1</emphasis>
, 
<emphasis id="B9645E75FFF6FFA8FD92FF4DB19401BB" box="[630,658,152,177]" italics="true" pageId="7" pageNumber="33">c2</emphasis>
, 
<emphasis id="B9645E75FFF6FFA8FD44FF4DB1BA01BB" box="[672,700,152,177]" italics="true" pageId="7" pageNumber="33">c3</emphasis>
, 
<emphasis id="B9645E75FFF6FFA8FD2FFF4DB1E001BB" box="[715,742,152,177]" italics="true" pageId="7" pageNumber="33">cp</emphasis>
, 
<emphasis id="B9645E75FFF6FFA8FD11FF4DB01401BB" box="[757,786,152,177]" italics="true" pageId="7" pageNumber="33">d1</emphasis>
, 
<emphasis id="B9645E75FFF6FFA8FCC5FF4DB03901BB" box="[801,831,152,177]" italics="true" pageId="7" pageNumber="33">d2</emphasis>
, 
<emphasis id="B9645E75FFF6FFA8FCA9FF4DB06E01BB" box="[845,872,152,177]" italics="true" pageId="7" pageNumber="33">e1</emphasis>
, 
<emphasis id="B9645E75FFF6FFA8FC93FF4DB08801BB" box="[887,910,152,177]" italics="true" pageId="7" pageNumber="33">f1</emphasis>
, and 
<emphasis id="B9645E75FFF6FFA8FC2AFF4DB0E301BB" box="[974,997,152,177]" italics="true" pageId="7" pageNumber="33">f2</emphasis>
broadly phylliform or plate-like with foveate pattern; of these, 
<emphasis id="B9645E75FFF6FFA8FE03FF68B10001DC" box="[487,518,189,214]" italics="true" pageId="7" pageNumber="33">c1</emphasis>
elongate, reaching posterior notogastral scissure, 
<emphasis id="B9645E75FFF6FFA8FB7CFF68B7B401DC" box="[1176,1202,189,214]" italics="true" pageId="7" pageNumber="33">f2</emphasis>
basally broad but dramatically tapered at about distal half, 
<emphasis id="B9645E75FFF6FFA8FD9EFF37B19001F1" box="[634,662,226,251]" italics="true" pageId="7" pageNumber="33">e1</emphasis>
originating on tubercles and permanently erect, 
<emphasis id="B9645E75FFF6FFA8FB36FF37B7E801F1" box="[1234,1262,226,251]" italics="true" pageId="7" pageNumber="33">e2</emphasis>
originating on tubercles, bottlebrush-like—having thickened, sub-cylindrical stalk and ciliated in distal third (
<figureCitation id="132B9EE2FFF6FFA8FACAFEDDB67E0028" box="[1326,1400,264,290]" captionStart="FIGURE 5" captionStartId="7.[151,256,1968,1992]" captionTargetBox="[202,1378,409,1895]" captionTargetId="figure@7.[196,1386,400,1940]" captionTargetPageId="7" captionText="FIGURE 5. Phyllochthonius peniculus n. sp., female. A, dorsal view; B, interlamellar seta; C, ventral view (without legs); D, lateral view; E, seta e 2 with imaginary cross section." httpUri="https://zenodo.org/record/196268/files/figure.png" pageId="7" pageNumber="33">Fig. 5</figureCitation>
E), permanently erect and directed posterio-laterally. Setal rows 
<emphasis id="B9645E75FFF6FFA8FC82FEF8B073004C" box="[870,885,301,326]" italics="true" pageId="7" pageNumber="33">h</emphasis>
and 
<emphasis id="B9645E75FFF6FFA8FC54FEF8B0CC004C" box="[944,970,301,326]" italics="true" pageId="7" pageNumber="33">ps</emphasis>
on venter; 
<emphasis id="B9645E75FFF6FFA8FBB5FEF8B768004C" box="[1105,1134,301,326]" italics="true" pageId="7" pageNumber="33">h2</emphasis>
, 
<emphasis id="B9645E75FFF6FFA8FB9AFEF8B79A004C" box="[1150,1180,301,326]" italics="true" pageId="7" pageNumber="33">h3</emphasis>
, and 
<emphasis id="B9645E75FFF6FFA8FB39FEF8B643004C" box="[1245,1349,301,326]" italics="true" pageId="7" pageNumber="33">ps1 –ps3</emphasis>
simple and setiform, 
<emphasis id="B9645E75FFF6FFA8FEDDFE87B2500061" box="[313,342,338,363]" italics="true" pageId="7" pageNumber="33">h1</emphasis>
basally broad but distally narrow and acute.
</paragraph>
<caption id="DF6FD2EFFFF6FFA8FF73F865B02D06ED" httpUri="https://zenodo.org/record/196268/files/figure.png" pageId="7" pageNumber="33" targetBox="[202,1378,409,1895]" targetPageId="7">
<paragraph id="8BAF8267FFF6FFA8FF73F865B02D06ED" blockId="7.[151,1436,1968,2023]" pageId="7" pageNumber="33">
<emphasis id="B9645E75FFF6FFA8FF73F865B21B06C2" bold="true" box="[151,285,1968,1992]" pageId="7" pageNumber="33">FIGURE 5.</emphasis>
<taxonomicName id="4C10F9E4FFF6FFA8FEC0F865B14206CD" box="[292,580,1968,1991]" class="Arachnida" family="Phyllochthoniidae" genus="Phyllochthonius" kingdom="Animalia" order="Sarcoptiformes" pageId="7" pageNumber="33" phylum="Arthropoda" rank="species" species="peniculus" status="sp. nov.">
<emphasis id="B9645E75FFF6FFA8FEC0F865B14206CD" box="[292,580,1968,1991]" italics="true" pageId="7" pageNumber="33">Phyllochthonius peniculus</emphasis>
</taxonomicName>
<emphasis id="B9645E75FFF6FFA8FDA8F865B18C06C2" bold="true" box="[588,650,1968,1992]" pageId="7" pageNumber="33">
<taxonomicNameLabel id="A257E30EFFF6FFA8FDA8F865B18C06C2" box="[588,650,1968,1992]" pageId="7" pageNumber="33" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, female. A, dorsal view; B, interlamellar seta; C, ventral view (without legs); D, lateral view; E, seta 
<emphasis id="B9645E75FFF6FFA8FE36F805B2ED06ED" box="[466,491,2000,2023]" italics="true" pageId="7" pageNumber="33">e2</emphasis>
with imaginary cross section.
</paragraph>
</caption>
<paragraph id="8BAF8267FFF9FFA7FF98FDE4B02D0361" blockId="8.[124,1708,554,619]" pageId="8" pageNumber="34">
<emphasis id="B9645E75FFF9FFA7FF98FDE4B3FD0340" bold="true" box="[124,251,561,586]" pageId="8" pageNumber="34">TABLE 1.</emphasis>
Ontogeny of leg setae and solenidia of 
<taxonomicName id="4C10F9E4FFF9FFA7FD20FDFAB71C0343" box="[708,1050,559,585]" class="Arachnida" family="Phyllochthoniidae" genus="Phyllochthonius" kingdom="Animalia" order="Sarcoptiformes" pageId="8" pageNumber="34" phylum="Arthropoda" rank="species" species="ovatosetosus" status="sp. nov.">
<emphasis id="B9645E75FFF9FFA7FD20FDFAB71C0343" box="[708,1050,559,585]" italics="true" pageId="8" pageNumber="34">Phyllochthonius ovatosetosus</emphasis>
</taxonomicName>
<emphasis id="B9645E75FFF9FFA7FBC5FDE4B7640340" bold="true" box="[1057,1122,561,586]" pageId="8" pageNumber="34">
<taxonomicNameLabel id="A257E30EFFF9FFA7FBC5FDE4B7640340" box="[1057,1122,561,586]" pageId="8" pageNumber="34" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(placement indicates instar of 1st appearance, dash indicates no change, and parentheses indicates a given pair)
</paragraph>
<paragraph id="8BAF8267FFF9FFA7FEA2FD40B7E904D7" pageId="8" pageNumber="34">
<table id="F91070C7FFF90050FF6EFD40B6B004D7" box="[138,1462,661,1501]" gridcols="5" gridrows="8" pageId="8" pageNumber="34">
<tr id="35208025FFF90050FF6EFD40B6B0027D" box="[138,1462,661,887]" gridrow="0" pageId="8" pageNumber="34">
<th id="76F1E959FFF90050FF6EFD40B2C1027D" box="[138,455,661,887]" gridcol="0" gridrow="0" pageId="8" pageNumber="34">
<emphasis id="B9645E75FFF9FFA7FEA2FD40B2C103A6" bold="true" box="[326,455,661,684]" pageId="8" pageNumber="34">Trochanter</emphasis>
<emphasis id="B9645E75FFF9FFA7FF6EFD6DB3B903C7" bold="true" box="[138,191,696,717]" pageId="8" pageNumber="34">Leg I</emphasis>
Larva - Protonymph - Deutonymph - Tritonymph - Adult -
</th>
<th id="76F1E959FFF90050FE1BFD40B14D027D" box="[511,587,661,887]" gridcol="1" gridrow="0" pageId="8" pageNumber="34">
<emphasis id="B9645E75FFF9FFA7FE1BFD40B14D03A6" bold="true" box="[511,587,661,684]" pageId="8" pageNumber="34">Femur</emphasis>
<emphasis id="B9645E75FFF9FFA7FDE3FD0CB11503E5" box="[519,531,729,751]" italics="true" pageId="8" pageNumber="34">d</emphasis>
, 
<emphasis id="B9645E75FFF9FFA7FDFBFD0CB14203E5" box="[543,580,729,751]" italics="true" pageId="8" pageNumber="34">bv”</emphasis>
- - 
<emphasis id="B9645E75FFF9FFA7FDFAFCEAB12B025F" box="[542,557,831,853]" italics="true" pageId="8" pageNumber="34">l’</emphasis>
-
</th>
<th id="76F1E959FFF90050FD74FD40B000027D" box="[656,774,661,887]" gridcol="2" gridrow="0" pageId="8" pageNumber="34">
<emphasis id="B9645E75FFF9FFA7FD48FD40B1EF03A6" bold="true" box="[684,745,661,684]" pageId="8" pageNumber="34">Genu</emphasis>
<emphasis id="B9645E75FFF9FFA7FD74FD0CB19A03E5" box="[656,668,729,751]" italics="true" pageId="8" pageNumber="34">d</emphasis>
, (
<emphasis id="B9645E75FFF9FFA7FD54FD0CB1B003E5" box="[688,694,729,751]" italics="true" pageId="8" pageNumber="34">l</emphasis>
), σ 
<emphasis id="B9645E75FFF9FFA7FD32FD0CB1E403E5" box="[726,738,729,751]" italics="true" pageId="8" pageNumber="34">1</emphasis>
, σ 
<emphasis id="B9645E75FFF9FFA7FD1EFD0CB1D2025E" italics="true" pageId="8" pageNumber="34">2 v” - v’</emphasis>
-
</th>
<th id="76F1E959FFF90050FCDCFD40B0E4027D" box="[824,994,661,887]" gridcol="3" gridrow="0" pageId="8" pageNumber="34">
<emphasis id="B9645E75FFF9FFA7FC8BFD40B0AA03A6" bold="true" box="[879,940,661,684]" pageId="8" pageNumber="34">Tibia</emphasis>
<emphasis id="B9645E75FFF9FFA7FCDCFD0EB04203FB" box="[824,836,731,753]" italics="true" pageId="8" pageNumber="34">d</emphasis>
, 
<emphasis id="B9645E75FFF9FFA7FCB4FD0EB06E03FB" box="[848,872,731,753]" italics="true" pageId="8" pageNumber="34">l’s</emphasis>
, 
<emphasis id="B9645E75FFF9FFA7FC90FD0EB08903FB" box="[884,911,731,753]" italics="true" pageId="8" pageNumber="34">l”i</emphasis>
, 
<emphasis id="B9645E75FFF9FFA7FC7FFD0EB0AF03FB" box="[923,937,731,753]" italics="true" pageId="8" pageNumber="34">l’</emphasis>
, 
<emphasis id="B9645E75FFF9FFA7FC51FD0EB0C103FA" box="[949,967,731,752]" italics="true" pageId="8" pageNumber="34">v’</emphasis>
, ϕ - 
<emphasis id="B9645E75FFF9FFA7FC65FCCBB09F0239" box="[897,921,798,819]" italics="true" pageId="8" pageNumber="34">v”</emphasis>
- -
</th>
<th id="76F1E959FFF90050FBC4FD40B6B0027D" box="[1056,1462,661,887]" gridcol="4" gridrow="0" pageId="8" pageNumber="34">
<emphasis id="B9645E75FFF9FFA7FB20FD40B61403A6" bold="true" box="[1220,1298,661,684]" pageId="8" pageNumber="34">Tarsus</emphasis>
ω 
<emphasis id="B9645E75FFF9FFA7FBD4FD0EB73A03FB" box="[1072,1084,731,753]" italics="true" pageId="8" pageNumber="34">1</emphasis>
, ε, (
<emphasis id="B9645E75FFF9FFA7FB83FD0EB77203FB" box="[1127,1140,731,753]" italics="true" pageId="8" pageNumber="34">ft</emphasis>
), (
<emphasis id="B9645E75FFF9FFA7FB75FD09B7A403FB" box="[1169,1186,732,753]" italics="true" pageId="8" pageNumber="34">tc</emphasis>
), (
<emphasis id="B9645E75FFF9FFA7FB5AFD09B7CC03FB" box="[1214,1226,732,753]" italics="true" pageId="8" pageNumber="34">p</emphasis>
), (
<emphasis id="B9645E75FFF9FFA7FB02FD09B7F403FB" box="[1254,1266,732,753]" italics="true" pageId="8" pageNumber="34">u</emphasis>
), (
<emphasis id="B9645E75FFF9FFA7FAEAFD09B61C03FB" box="[1294,1306,732,753]" italics="true" pageId="8" pageNumber="34">a</emphasis>
), 
<emphasis id="B9645E75FFF9FFA7FACAFD09B63103FB" box="[1326,1335,732,753]" italics="true" pageId="8" pageNumber="34">s</emphasis>
, 
<emphasis id="B9645E75FFF9FFA7FAA7FD09B65203FB" box="[1347,1364,732,753]" italics="true" pageId="8" pageNumber="34">m</emphasis>
, (
<emphasis id="B9645E75FFF9FFA7FA8CFD0EB67D03FB" box="[1384,1403,731,753]" italics="true" pageId="8" pageNumber="34">pl</emphasis>
), (
<emphasis id="B9645E75FFF9FFA7FA73FD09B6AB03FB" box="[1431,1453,732,753]" italics="true" pageId="8" pageNumber="34">pv</emphasis>
) ω 
<emphasis id="B9645E75FFF9FFA7FB22FD28B7D40219" box="[1222,1234,765,787]" italics="true" pageId="8" pageNumber="34">2</emphasis>
, (
<emphasis id="B9645E75FFF9FFA7FB03FD2BB7F20219" box="[1255,1268,766,787]" italics="true" pageId="8" pageNumber="34">it</emphasis>
), 
<emphasis id="B9645E75FFF9FFA7FAECFD2BB6260219" box="[1288,1312,766,787]" italics="true" pageId="8" pageNumber="34">v”</emphasis>
- ω 
<emphasis id="B9645E75FFF9FFA7FB3AFC94B7EC025D" box="[1246,1258,833,855]" italics="true" pageId="8" pageNumber="34">3</emphasis>
, 
<emphasis id="B9645E75FFF9FFA7FB12FC97B60F025D" box="[1270,1289,834,855]" italics="true" pageId="8" pageNumber="34">v’</emphasis>
-
</th>
</tr>
<tr id="35208025FFF90050FF6EFC50B6B002D7" box="[138,1462,901,989]" gridrow="1" pageId="8" pageNumber="34">
<th id="76F1E959FFF90050FF6EFC50B2C102D7" box="[138,455,901,989]" gridcol="0" gridrow="1" pageId="8" pageNumber="34">
<emphasis id="B9645E75FFF9FFA7FF6EFC50B3CF0290" bold="true" box="[138,201,901,922]" pageId="8" pageNumber="34">Leg II</emphasis>
Larva - Protonymph -
</th>
<td id="76F1E959FFF90050FE1BFC50B14D02D7" box="[511,587,901,989]" gridcol="1" gridrow="1" pageId="8" pageNumber="34">
<emphasis id="B9645E75FFF9FFA7FDE3FC70B11502B1" box="[519,531,933,955]" italics="true" pageId="8" pageNumber="34">d</emphasis>
, 
<emphasis id="B9645E75FFF9FFA7FDFBFC70B12902D7" italics="true" pageId="8" pageNumber="34">bv” l”</emphasis>
</td>
<td id="76F1E959FFF90050FD74FC50B00002D7" box="[656,774,901,989]" gridcol="2" gridrow="1" pageId="8" pageNumber="34">
<emphasis id="B9645E75FFF9FFA7FD43FC70B1B502B1" box="[679,691,933,955]" italics="true" pageId="8" pageNumber="34">d</emphasis>
, (
<emphasis id="B9645E75FFF9FFA7FD2CFC70B1C902B1" box="[712,719,933,955]" italics="true" pageId="8" pageNumber="34">l</emphasis>
), σ -
</td>
<td id="76F1E959FFF90050FCDCFC50B0E402D7" box="[824,994,901,989]" gridcol="3" gridrow="1" pageId="8" pageNumber="34">
<emphasis id="B9645E75FFF9FFA7FCB1FC72B06702B7" box="[853,865,935,957]" italics="true" pageId="8" pageNumber="34">d</emphasis>
, (
<emphasis id="B9645E75FFF9FFA7FC91FC72B07A02B7" box="[885,892,935,957]" italics="true" pageId="8" pageNumber="34">l</emphasis>
), (
<emphasis id="B9645E75FFF9FFA7FC7CFC7DB0A502B7" box="[920,931,936,957]" italics="true" pageId="8" pageNumber="34">v</emphasis>
), ϕ -
</td>
<td id="76F1E959FFF90050FBC4FC50B6B002D7" box="[1056,1462,901,989]" gridcol="4" gridrow="1" pageId="8" pageNumber="34">
ω 
<emphasis id="B9645E75FFF9FFA7FBAEFC72B75002B7" box="[1098,1110,935,957]" italics="true" pageId="8" pageNumber="34">1</emphasis>
, (
<emphasis id="B9645E75FFF9FFA7FB8FFC72B77E02B7" box="[1131,1144,935,957]" italics="true" pageId="8" pageNumber="34">ft</emphasis>
), (
<emphasis id="B9645E75FFF9FFA7FB70FC7DB7A302B7" box="[1172,1189,936,957]" italics="true" pageId="8" pageNumber="34">tc</emphasis>
), (
<emphasis id="B9645E75FFF9FFA7FB25FC7DB7CB02B7" box="[1217,1229,936,957]" italics="true" pageId="8" pageNumber="34">p</emphasis>
), (
<emphasis id="B9645E75FFF9FFA7FB0DFC7DB7F302B7" box="[1257,1269,936,957]" italics="true" pageId="8" pageNumber="34">u</emphasis>
), 
<emphasis id="B9645E75FFF9FFA7FAEDFC7DB61402B7" box="[1289,1298,936,957]" italics="true" pageId="8" pageNumber="34">s</emphasis>
, (
<emphasis id="B9645E75FFF9FFA7FAC3FC7DB63B02B7" box="[1319,1341,936,957]" italics="true" pageId="8" pageNumber="34">pv</emphasis>
), (
<emphasis id="B9645E75FFF9FFA7FABDFC7DB66302B7" box="[1369,1381,936,957]" italics="true" pageId="8" pageNumber="34">a</emphasis>
), (
<emphasis id="B9645E75FFF9FFA7FA65FC72B69202B7" box="[1409,1428,935,957]" italics="true" pageId="8" pageNumber="34">pl</emphasis>
) -
</td>
</tr>
<tr id="35208025FFF90050FF6EFC3FB6B0052B" box="[138,1462,1002,1057]" gridrow="2" pageId="8" pageNumber="34">
<th id="76F1E959FFF90050FF6EFC3FB2C1052B" box="[138,455,1002,1057]" gridcol="0" gridrow="2" pageId="8" pageNumber="34">
Deutonymph 
<emphasis id="B9645E75FFF9FFA7FE99FC3FB28902F5" box="[381,399,1002,1023]" italics="true" pageId="8" pageNumber="34">v’</emphasis>
Tritonymph -
</th>
<td id="76F1E959FFF90050FE1BFC3FB14D052B" box="[511,587,1002,1057]" gridcol="1" gridrow="2" pageId="8" pageNumber="34">- -</td>
<td id="76F1E959FFF90050FD74FC3FB000052B" box="[656,774,1002,1057]" gridcol="2" gridrow="2" pageId="8" pageNumber="34">
- 
<emphasis id="B9645E75FFF9FFA7FD5BFBD9B1D1052B" box="[703,727,1036,1057]" italics="true" pageId="8" pageNumber="34">v”</emphasis>
</td>
<td id="76F1E959FFF90050FCDCFC3FB0E4052B" box="[824,994,1002,1057]" gridcol="3" gridrow="2" pageId="8" pageNumber="34">- -</td>
<td id="76F1E959FFF90050FBC4FC3FB6B0052B" box="[1056,1462,1002,1057]" gridcol="4" gridrow="2" pageId="8" pageNumber="34">
ω 
<emphasis id="B9645E75FFF9FFA7FB3CFC39B7E20508" box="[1240,1252,1004,1026]" italics="true" pageId="8" pageNumber="34">2</emphasis>
, (
<emphasis id="B9645E75FFF9FFA7FB1DFC38B6000508" box="[1273,1286,1005,1026]" italics="true" pageId="8" pageNumber="34">it</emphasis>
) -
</td>
</tr>
<tr id="35208025FFF90050FF6EFBFAB6B0054E" box="[138,1462,1071,1092]" gridrow="3" pageId="8" pageNumber="34">
<th id="76F1E959FFF90050FF6EFBFAB2C1054E" box="[138,455,1071,1092]" gridcol="0" gridrow="3" pageId="8" pageNumber="34">Adult -</th>
<td id="76F1E959FFF90050FE1BFBFAB14D054E" box="[511,587,1071,1092]" gridcol="1" gridrow="3" pageId="8" pageNumber="34">-</td>
<td id="76F1E959FFF90050FD74FBFAB000054E" box="[656,774,1071,1092]" gridcol="2" gridrow="3" pageId="8" pageNumber="34">
<emphasis id="B9645E75FFF9FFA7FD25FBFAB1D2054E" box="[705,724,1071,1092]" italics="true" pageId="8" pageNumber="34">v’</emphasis>
</td>
<td id="76F1E959FFF90050FCDCFBFAB0E4054E" box="[824,994,1071,1092]" gridcol="3" gridrow="3" pageId="8" pageNumber="34">-</td>
<td id="76F1E959FFF90050FBC4FBFAB6B0054E" box="[1056,1462,1071,1092]" gridcol="4" gridrow="3" pageId="8" pageNumber="34">-</td>
</tr>
<tr id="35208025FFF90050FF6EFB84B6B005C6" box="[138,1462,1105,1228]" gridrow="4" pageId="8" pageNumber="34">
<th id="76F1E959FFF90050FF6EFB84B2C105C6" box="[138,455,1105,1228]" gridcol="0" gridrow="4" pageId="8" pageNumber="34">
<emphasis id="B9645E75FFF9FFA7FF6EFB84B3D4056C" bold="true" box="[138,210,1105,1126]" pageId="8" pageNumber="34">Leg III</emphasis>
Larva - Protonymph 
<emphasis id="B9645E75FFF9FFA7FE99FB40B28905A0" box="[381,399,1173,1194]" italics="true" pageId="8" pageNumber="34">v’</emphasis>
Deutonymph 
<emphasis id="B9645E75FFF9FFA7FE9BFB63B28805C6" box="[383,398,1206,1228]" italics="true" pageId="8" pageNumber="34">l’</emphasis>
</th>
<td id="76F1E959FFF90050FE1BFB84B14D05C6" box="[511,587,1105,1228]" gridcol="1" gridrow="4" pageId="8" pageNumber="34">
<emphasis id="B9645E75FFF9FFA7FDEFFBA7B1110582" box="[523,535,1138,1160]" italics="true" pageId="8" pageNumber="34">d</emphasis>
, 
<emphasis id="B9645E75FFF9FFA7FDC7FBA7B146058D" box="[547,576,1138,1159]" italics="true" pageId="8" pageNumber="34">ev’</emphasis>
- -
</td>
<td id="76F1E959FFF90050FD74FB84B00005C6" box="[656,774,1105,1228]" gridcol="2" gridrow="4" pageId="8" pageNumber="34">
<emphasis id="B9645E75FFF9FFA7FD48FBA7B1BE0582" box="[684,696,1138,1160]" italics="true" pageId="8" pageNumber="34">d</emphasis>
, 
<emphasis id="B9645E75FFF9FFA7FD20FBA7B1D40582" box="[708,722,1138,1160]" italics="true" pageId="8" pageNumber="34">l’</emphasis>
, σ - -
</td>
<td id="76F1E959FFF90050FCDCFB84B0E405C6" box="[824,994,1105,1228]" gridcol="3" gridrow="4" pageId="8" pageNumber="34">
<emphasis id="B9645E75FFF9FFA7FCBDFBA1B0630580" box="[857,869,1140,1162]" italics="true" pageId="8" pageNumber="34">d</emphasis>
, 
<emphasis id="B9645E75FFF9FFA7FC95FBA1B0860580" box="[881,896,1140,1162]" italics="true" pageId="8" pageNumber="34">l’</emphasis>
, (
<emphasis id="B9645E75FFF9FFA7FC70FBA0B0990580" box="[916,927,1141,1162]" italics="true" pageId="8" pageNumber="34">v</emphasis>
), ϕ - -
</td>
<td id="76F1E959FFF90050FBC4FB84B6B005C6" box="[1056,1462,1105,1228]" gridcol="4" gridrow="4" pageId="8" pageNumber="34">
(
<emphasis id="B9645E75FFF9FFA7FB90FBA7B7870582" box="[1140,1153,1138,1160]" italics="true" pageId="8" pageNumber="34">ft</emphasis>
), (
<emphasis id="B9645E75FFF9FFA7FB79FBA6B7A80582" box="[1181,1198,1139,1160]" italics="true" pageId="8" pageNumber="34">tc</emphasis>
), (
<emphasis id="B9645E75FFF9FFA7FB2EFBA6B7D00582" box="[1226,1238,1139,1160]" italics="true" pageId="8" pageNumber="34">p</emphasis>
), (
<emphasis id="B9645E75FFF9FFA7FB16FBA6B7F80582" box="[1266,1278,1139,1160]" italics="true" pageId="8" pageNumber="34">u</emphasis>
), 
<emphasis id="B9645E75FFF9FFA7FAF6FBA6B61D0582" box="[1298,1307,1139,1160]" italics="true" pageId="8" pageNumber="34">s</emphasis>
, (
<emphasis id="B9645E75FFF9FFA7FACBFBA6B63D0582" box="[1327,1339,1139,1160]" italics="true" pageId="8" pageNumber="34">a</emphasis>
), 
<emphasis id="B9645E75FFF9FFA7FAABFBA7B66C0582" box="[1359,1386,1138,1160]" italics="true" pageId="8" pageNumber="34">pl’</emphasis>
(
<emphasis id="B9645E75FFF9FFA7FB3BFB40B7F005A0" box="[1247,1270,1173,1194]" italics="true" pageId="8" pageNumber="34">pv</emphasis>
) (
<emphasis id="B9645E75FFF9FFA7FB01FB62B7F405C6" box="[1253,1266,1207,1228]" italics="true" pageId="8" pageNumber="34">it</emphasis>
)
</td>
</tr>
<tr id="35208025FFF90050FF6EFB0CB6B005E4" box="[138,1462,1241,1262]" gridrow="5" pageId="8" pageNumber="34">
<th id="76F1E959FFF90050FF6EFB0CB2C105E4" box="[138,455,1241,1262]" gridcol="0" gridrow="5" pageId="8" pageNumber="34">Tritonymph -</th>
<td id="76F1E959FFF90050FE1BFB0CB14D05E4" box="[511,587,1241,1262]" gridcol="1" gridrow="5" pageId="8" pageNumber="34">-</td>
<td id="76F1E959FFF90050FD74FB0CB00005E4" box="[656,774,1241,1262]" gridcol="2" gridrow="5" pageId="8" pageNumber="34">-</td>
<td id="76F1E959FFF90050FCDCFB0CB0E405E4" box="[824,994,1241,1262]" gridcol="3" gridrow="5" pageId="8" pageNumber="34">-</td>
<td id="76F1E959FFF90050FBC4FB0CB6B005E4" box="[1056,1462,1241,1262]" gridcol="4" gridrow="5" pageId="8" pageNumber="34">-</td>
</tr>
<tr id="35208025FFF90050FF6EFB2EB6B0041A" box="[138,1462,1275,1296]" gridrow="6" pageId="8" pageNumber="34">
<th id="76F1E959FFF90050FF6EFB2EB2C1041A" box="[138,455,1275,1296]" gridcol="0" gridrow="6" pageId="8" pageNumber="34">Adult -</th>
<td id="76F1E959FFF90050FE1BFB2EB14D041A" box="[511,587,1275,1296]" gridcol="1" gridrow="6" pageId="8" pageNumber="34">-</td>
<td id="76F1E959FFF90050FD74FB2EB000041A" box="[656,774,1275,1296]" gridcol="2" gridrow="6" pageId="8" pageNumber="34">
(
<emphasis id="B9645E75FFF9FFA7FD21FB2EB1D6041A" box="[709,720,1275,1296]" italics="true" pageId="8" pageNumber="34">v</emphasis>
)
</td>
<td id="76F1E959FFF90050FCDCFB2EB0E4041A" box="[824,994,1275,1296]" gridcol="3" gridrow="6" pageId="8" pageNumber="34">-</td>
<td id="76F1E959FFF90050FBC4FB2EB6B0041A" box="[1056,1462,1275,1296]" gridcol="4" gridrow="6" pageId="8" pageNumber="34">-</td>
</tr>
<tr id="35208025FFF90050FF6EFACBB6B004D7" box="[138,1462,1310,1501]" gridrow="7" pageId="8" pageNumber="34">
<th id="76F1E959FFF90050FF6EFACBB2C104D7" box="[138,455,1310,1501]" gridcol="0" gridrow="7" pageId="8" pageNumber="34">
<emphasis id="B9645E75FFF9FFA7FF6EFACBB3D60439" bold="true" box="[138,208,1310,1331]" pageId="8" pageNumber="34">Leg IV</emphasis>
Larva - Protonymph - Deutonymph 
<emphasis id="B9645E75FFF9FFA7FE99FA56B2890492" box="[381,399,1411,1432]" italics="true" pageId="8" pageNumber="34">v’</emphasis>
Tritonymph 
<emphasis id="B9645E75FFF9FFA7FE9BFA71B28804B0" box="[383,398,1444,1466]" italics="true" pageId="8" pageNumber="34">l’</emphasis>
Adult -
</th>
<td id="76F1E959FFF90050FE1BFACBB14D04D7" box="[511,587,1310,1501]" gridcol="1" gridrow="7" pageId="8" pageNumber="34">
- - 
<emphasis id="B9645E75FFF9FFA7FDFBFA57B13204B0" italics="true" pageId="8" pageNumber="34">d ev’</emphasis>
-
</td>
<td id="76F1E959FFF90050FD74FACBB00004D7" box="[656,774,1310,1501]" gridcol="2" gridrow="7" pageId="8" pageNumber="34">
- - 
<emphasis id="B9645E75FFF9FFA7FD48FA57B1BE0492" box="[684,696,1410,1432]" italics="true" pageId="8" pageNumber="34">d</emphasis>
, 
<emphasis id="B9645E75FFF9FFA7FD20FA57B1D40492" box="[708,722,1410,1432]" italics="true" pageId="8" pageNumber="34">l’</emphasis>
, σ - (
<emphasis id="B9645E75FFF9FFA7FD21FA1DB1D604D7" box="[709,720,1480,1501]" italics="true" pageId="8" pageNumber="34">v</emphasis>
)
</td>
<td id="76F1E959FFF90050FCDCFACBB0E404D7" box="[824,994,1310,1501]" gridcol="3" gridrow="7" pageId="8" pageNumber="34">
- 
<emphasis id="B9645E75FFF9FFA7FC62FAB5B0860492" italics="true" pageId="8" pageNumber="34">l’ d</emphasis>
, (
<emphasis id="B9645E75FFF9FFA7FC70FA56B0990492" box="[916,927,1411,1432]" italics="true" pageId="8" pageNumber="34">v</emphasis>
) ϕ -
</td>
<td id="76F1E959FFF90050FBC4FACBB6B004D7" box="[1056,1462,1310,1501]" gridcol="4" gridrow="7" pageId="8" pageNumber="34">
- 
<emphasis id="B9645E75FFF9FFA7FB78FAB5B7B0047C" box="[1180,1206,1376,1398]" italics="true" pageId="8" pageNumber="34">ft”</emphasis>
, (
<emphasis id="B9645E75FFF9FFA7FB2FFAB4B7D1047C" box="[1227,1239,1377,1398]" italics="true" pageId="8" pageNumber="34">p</emphasis>
), (
<emphasis id="B9645E75FFF9FFA7FB17FAB4B7F9047C" box="[1267,1279,1377,1398]" italics="true" pageId="8" pageNumber="34">u</emphasis>
), (
<emphasis id="B9645E75FFF9FFA7FAFFFAB4B637047C" box="[1307,1329,1377,1398]" italics="true" pageId="8" pageNumber="34">pv</emphasis>
) 
<emphasis id="B9645E75FFF9FFA7FB4FFA57B7C60492" box="[1195,1216,1410,1432]" italics="true" pageId="8" pageNumber="34">ft’</emphasis>
, (
<emphasis id="B9645E75FFF9FFA7FB30FA56B7E30492" box="[1236,1253,1411,1432]" italics="true" pageId="8" pageNumber="34">tc</emphasis>
), (
<emphasis id="B9645E75FFF9FFA7FAE5FA56B60B0492" box="[1281,1293,1411,1432]" italics="true" pageId="8" pageNumber="34">a</emphasis>
), 
<emphasis id="B9645E75FFF9FFA7FAC6FA56B7FE04B0" italics="true" pageId="8" pageNumber="34">s pl’</emphasis>
-
</td>
</tr>
</table>
</paragraph>
<paragraph id="8BAF8267FFF8FFA6FF22FF42B7920028" blockId="9.[151,1436,151,1036]" pageId="9" pageNumber="35">
<emphasis id="B9645E75FFF8FFA6FF22FF42B21101BB" bold="true" box="[198,279,151,177]" pageId="9" pageNumber="35">Venter</emphasis>
(
<figureCitation id="132B9EE2FFF8FFA6FECEFF4DB27701B8" box="[298,369,152,178]" captionStart="FIGURE 5" captionStartId="7.[151,256,1968,1992]" captionTargetBox="[202,1378,409,1895]" captionTargetId="figure@7.[196,1386,400,1940]" captionTargetPageId="7" captionText="FIGURE 5. Phyllochthonius peniculus n. sp., female. A, dorsal view; B, interlamellar seta; C, ventral view (without legs); D, lateral view; E, seta e 2 with imaginary cross section." httpUri="https://zenodo.org/record/196268/files/figure.png" pageId="9" pageNumber="35">Fig. 5</figureCitation>
C). Except for epimeres, ventral surface soft. Genital and anal valves delimited by folds of integument. Genital valves with seven pairs of setae. Three pairs of genital papillae present. Ovipositor with simple (uncounted) setae. Anal valves with three pairs of anal setae (
<emphasis id="B9645E75FFF8FFA6FC59FF37B0DC01F1" box="[957,986,226,251]" italics="true" pageId="9" pageNumber="35">an</emphasis>
), one pair of peranal setae (
<emphasis id="B9645E75FFF8FFA6FAC6FF37B63901F1" box="[1314,1343,226,251]" italics="true" pageId="9" pageNumber="35">pa</emphasis>
). Three pairs of adanal setae (
<emphasis id="B9645E75FFF8FFA6FE72FEDDB2B5002B" box="[406,435,264,289]" italics="true" pageId="9" pageNumber="35">ad</emphasis>
) present. Epimeral setation 3-1-4-3; all setae short and simple.
</paragraph>
<paragraph id="8BAF8267FFF8FFA6FF22FEF8B27400D6" blockId="9.[151,1436,151,1036]" pageId="9" pageNumber="35">
<emphasis id="B9645E75FFF8FFA6FF22FEF8B260004D" bold="true" box="[198,358,301,327]" pageId="9" pageNumber="35">Gnathosoma</emphasis>
. Generally similar to that of 
<taxonomicName id="4C10F9E4FFF8FFA6FD5DFEF8B06D004C" box="[697,875,301,326]" class="Arachnida" family="Phyllochthoniidae" genus="Phyllochthonius" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="35" phylum="Arthropoda" rank="species" species="ovatosetosus" status="sp. nov.">
<emphasis id="B9645E75FFF8FFA6FD5DFEF8B06D004C" box="[697,875,301,326]" italics="true" pageId="9" pageNumber="35">P. ovatosetosus</emphasis>
</taxonomicName>
<emphasis id="B9645E75FFF8FFA6FC96FEF8B0B3004D" bold="true" box="[882,949,301,327]" pageId="9" pageNumber="35">
<taxonomicNameLabel id="A257E30EFFF8FFA6FC96FEF8B0B3004D" box="[882,949,301,327]" pageId="9" pageNumber="35" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
Subcapitulum with 3 pairs of setae (
<emphasis id="B9645E75FFF8FFA6FA86FEF8B676004C" box="[1378,1392,301,326]" italics="true" pageId="9" pageNumber="35">a</emphasis>
, 
<emphasis id="B9645E75FFF8FFA6FA9BFEF8B692004C" box="[1407,1428,301,326]" italics="true" pageId="9" pageNumber="35">m</emphasis>
, 
<emphasis id="B9645E75FFF8FFA6FF73FE87B3A00061" box="[151,166,338,363]" italics="true" pageId="9" pageNumber="35">h</emphasis>
). Rutellum broad distally with 4 teeth; basal tooth absent. Three pairs of adoral setae present, similar to those of 
<taxonomicName id="4C10F9E4FFF8FFA6FF1BFEADB2B3009B" box="[255,437,376,401]" class="Arachnida" family="Phyllochthoniidae" genus="Phyllochthonius" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="35" phylum="Arthropoda" rank="species" species="ovatosetosus" status="sp. nov.">
<emphasis id="B9645E75FFF8FFA6FF1BFEADB2B3009B" box="[255,437,376,401]" italics="true" pageId="9" pageNumber="35">P. ovatosetosus</emphasis>
</taxonomicName>
<emphasis id="B9645E75FFF8FFA6FE59FEA2B104009B" bold="true" box="[445,514,375,401]" pageId="9" pageNumber="35">
<taxonomicNameLabel id="A257E30EFFF8FFA6FE59FEA2B104009B" box="[445,514,375,401]" pageId="9" pageNumber="35" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
Palpi five-segmented; setation 0-2-1-2-9; tarsi with baculiform solenidion ω and terminally with forked eupathidium. Chelicerae dentate with two cheliceral setae, 
<emphasis id="B9645E75FFF8FFA6FB4AFE48B7DF00BC" box="[1198,1241,413,438]" italics="true" pageId="9" pageNumber="35">cha</emphasis>
minute and 
<emphasis id="B9645E75FFF8FFA6FA95FE48B69A00BC" box="[1393,1436,413,438]" italics="true" pageId="9" pageNumber="35">chb</emphasis>
longer, with barbs.
</paragraph>
<paragraph id="8BAF8267FFF8FFA6FF22FE32B74B030B" blockId="9.[151,1436,151,1036]" box="[198,1101,487,514]" pageId="9" pageNumber="35">
<emphasis id="B9645E75FFF8FFA6FF22FE32B207030B" bold="true" box="[198,257,487,513]" pageId="9" pageNumber="35">Legs</emphasis>
. Leg I–IV bidactyl. Leg setation similar to that of 
<taxonomicName id="4C10F9E4FFF8FFA6FCB5FE3DB702030B" box="[849,1028,488,513]" class="Arachnida" family="Phyllochthoniidae" genus="Phyllochthonius" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="35" phylum="Arthropoda" rank="species" species="ovatosetosus" status="sp. nov.">
<emphasis id="B9645E75FFF8FFA6FCB5FE3DB702030B" box="[849,1028,488,513]" italics="true" pageId="9" pageNumber="35">P. ovatosetosus</emphasis>
</taxonomicName>
<emphasis id="B9645E75FFF8FFA6FBEFFE32B74B030B" bold="true" box="[1035,1101,487,513]" pageId="9" pageNumber="35">
<taxonomicNameLabel id="A257E30EFFF8FFA6FBEFFE32B74B030B" box="[1035,1101,487,513]" pageId="9" pageNumber="35" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8BAF8267FFF8FFA6FF22FDD8B152032D" blockId="9.[151,1436,151,1036]" box="[198,596,525,551]" pageId="9" pageNumber="35">
<emphasis id="B9645E75FFF8FFA6FF22FDD8B2CC032D" bold="true" box="[198,458,525,551]" pageId="9" pageNumber="35">Male and immatures</emphasis>
. Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C30AD1ECFFF8FFA6FF22FDE7B7A403B6" pageId="9" pageNumber="35" type="materials_examined">
<paragraph id="8BAF8267FFF8FFA6FF22FDE7B7A403B6" blockId="9.[151,1436,151,1036]" pageId="9" pageNumber="35">
<emphasis id="B9645E75FFF8FFA6FF22FDE7B2BC0346" bold="true" box="[198,442,562,588]" pageId="9" pageNumber="35">Material examined</emphasis>
. Female 
<typeStatus id="54AB3CC5FFF8FFA6FDD6FDE7B19E0346" box="[562,664,562,588]" pageId="9" pageNumber="35" type="holotype">holotype</typeStatus>
(in alcohol, CUMZ-AC2010.39); north entrance of Tharn-lod Cave, 
<quantity id="4CE82F82FFF8FFA6FF05FD8DB2290378" box="[225,303,600,626]" metricMagnitude="4" metricUnit="m" metricValue="8.0" pageId="9" pageNumber="35" unit="km" value="80.0">80 km</quantity>
. NW of Kanchanaburi Province, 
<collectingCountry id="F307C2F7FFF8FFA6FD57FD8DB01D0378" box="[691,795,600,626]" name="Thailand" pageId="9" pageNumber="35">Thailand</collectingCountry>
(
<geoCoordinate id="EE24E4A0FFF8FFA6FCCFFD8DB09B0378" box="[811,925,600,626]" direction="north" orientation="latitude" pageId="9" pageNumber="35" precision="925" value="14.65">14° 39' N</geoCoordinate>
, 
<geoCoordinate id="EE24E4A0FFF8FFA6FC4DFD8DB71E0378" box="[937,1048,600,626]" direction="east" orientation="longitude" pageId="9" pageNumber="35" precision="925" value="99.3">99° 18' E</geoCoordinate>
); ex. forest litter; 
<date id="FFAEA4A7FFF8FFA6FB0CFD8DB6630378" box="[1256,1381,600,626]" pageId="9" pageNumber="35" value="2008-03-03">3-III-2008</date>
; leg. M. Fuangarworn (Field No. MF 
<date id="FFAEA4A7FFF8FFA6FDCEFDA8B194039D" box="[554,658,637,663]" pageId="9" pageNumber="35" value="2008" valueMax="2015">2008-15</date>
). No other specimens known. 
<typeStatus id="54AB3CC5FFF8FFA6FBE8FDA8B779039D" box="[1036,1151,637,663]" pageId="9" pageNumber="35" type="holotype">Holotype</typeStatus>
deposited in the Acari collection at Chulalongkorn University Museum of Natural History, Bangkok. 
<collectingCountry id="F307C2F7FFF8FFA6FBD6FD77B79803B6" box="[1074,1182,674,700]" name="Thailand" pageId="9" pageNumber="35">Thailand</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C30AD1ECFFF8FFA6FF22FD12B62603E8" box="[198,1312,711,738]" pageId="9" pageNumber="35" type="distribution">
<paragraph id="8BAF8267FFF8FFA6FF22FD12B62603E8" blockId="9.[151,1436,151,1036]" box="[198,1312,711,738]" pageId="9" pageNumber="35">
<emphasis id="B9645E75FFF8FFA6FF22FD12B25903EB" bold="true" box="[198,351,711,737]" pageId="9" pageNumber="35">Distribution</emphasis>
. Known only from its 
<typeStatus id="54AB3CC5FFF8FFA6FD8EFD1DB19A03E8" box="[618,668,712,738]" pageId="9" pageNumber="35">type</typeStatus>
locality — Kanchanaburi Province, Western 
<collectingCountry id="F307C2F7FFF8FFA6FB54FD1DB61A03E8" box="[1200,1308,712,738]" name="Thailand" pageId="9" pageNumber="35">Thailand</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C30AD1ECFFF8FFA6FF22FD38B1870226" pageId="9" pageNumber="35" type="etymology">
<paragraph id="8BAF8267FFF8FFA6FF22FD38B1870226" blockId="9.[151,1436,151,1036]" pageId="9" pageNumber="35">
<emphasis id="B9645E75FFF8FFA6FF22FD38B24D020D" bold="true" box="[198,331,749,775]" pageId="9" pageNumber="35">Etymology</emphasis>
. The new species is named after the peculiar form of notogastral setae 
<emphasis id="B9645E75FFF8FFA6FB14FD38B60D020C" box="[1264,1291,749,774]" italics="true" pageId="9" pageNumber="35">e2</emphasis>
, which are bottlebrush-like (
<taxonomicName id="4C10F9E4FFF8FFA6FE84FCC7B1240226" authority="Latin" authorityName="Latin" box="[352,546,786,812]" class="Arachnida" family="Phyllochthoniidae" genus="Phyllochthonius" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="35" phylum="Arthropoda" rank="species" species="peniculus">peniculus: Latin</taxonomicName>
, brush).
</paragraph>
</subSubSection>
<subSubSection id="C30AD1ECFFF8FFA6FF22FCE2B6860506" pageId="9" pageNumber="35" type="discussion">
<paragraph id="8BAF8267FFF8FFA6FF22FCE2B6860506" blockId="9.[151,1436,151,1036]" pageId="9" pageNumber="35">
<emphasis id="B9645E75FFF8FFA6FF22FCE2B23E025B" bold="true" box="[198,312,823,849]" pageId="9" pageNumber="35">Remarks</emphasis>
. 
<taxonomicName id="4C10F9E4FFF8FFA6FEABFCEDB18B025B" box="[335,653,824,849]" class="Arachnida" family="Phyllochthoniidae" genus="Phyllochthonius" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="35" phylum="Arthropoda" rank="species" species="peniculus" status="sp. nov.">
<emphasis id="B9645E75FFF8FFA6FEABFCEDB18B025B" box="[335,653,824,849]" italics="true" pageId="9" pageNumber="35">Phyllochthonius peniculus</emphasis>
</taxonomicName>
<emphasis id="B9645E75FFF8FFA6FD78FCE2B1E1025B" bold="true" box="[668,743,823,849]" pageId="9" pageNumber="35">
<taxonomicNameLabel id="A257E30EFFF8FFA6FD78FCE2B1E1025B" box="[668,743,823,849]" pageId="9" pageNumber="35" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
has a set of characters that corresponds to the genus 
<taxonomicName id="4C10F9E4FFF8FFA6FF73FC88B250027C" box="[151,342,861,886]" class="Arachnida" family="Phyllochthoniidae" genus="Phyllochthonius" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B9645E75FFF8FFA6FF73FC88B250027C" box="[151,342,861,886]" italics="true" pageId="9" pageNumber="35">Phyllochthonius</emphasis>
</taxonomicName>
as redefined below. It exhibits a degree of setal expansion similar to that of 
<taxonomicName id="4C10F9E4FFF8FFA6FB04FC88B3D30296" authority="Trave, 1967" authorityName="Trave" authorityYear="1967" class="Arachnida" family="Phyllochthoniidae" genus="Phyllochthonius" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="35" phylum="Arthropoda" rank="species" species="aoutii">
<emphasis id="B9645E75FFF8FFA6FB04FC88B647027C" box="[1248,1345,861,886]" italics="true" pageId="9" pageNumber="35">P. aoutii</emphasis>
, 
<bibRefCitation id="EF81FF96FFF8FFA6FAB5FC88B3D30296" author="Trave" pageId="9" pageNumber="36" refString="Trave, J. (1967) Phyllochthonius aoutii nov. gen., nov. spec., un Enarthronota (Acarien, Oribatei) nouveau de Cote d'Ivoire, avec la creation d'une superfamille nouvelle, Phyllochthonoidea. Zoologische Mededelingen, 42, 83 - 105." type="journal article" year="1967">Travé, 1967</bibRefCitation>
</taxonomicName>
. However, these species can be distinguished easily by the shape of setae 
<emphasis id="B9645E75FFF8FFA6FBD8FC57B75E0291" box="[1084,1112,898,923]" italics="true" pageId="9" pageNumber="35">e2</emphasis>
, which are bottlebrush-like in 
<taxonomicName id="4C10F9E4FFF8FFA6FF52FC7DB24002CB" box="[182,326,936,961]" class="Arachnida" family="Phyllochthoniidae" genus="Phyllochthonius" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="35" phylum="Arthropoda" rank="species" species="peniculus" status="sp. nov.">
<emphasis id="B9645E75FFF8FFA6FF52FC7DB24002CB" box="[182,326,936,961]" italics="true" pageId="9" pageNumber="35">P. peniculus</emphasis>
</taxonomicName>
<emphasis id="B9645E75FFF8FFA6FEABFC72B29202CB" bold="true" box="[335,404,935,961]" pageId="9" pageNumber="35">
<taxonomicNameLabel id="A257E30EFFF8FFA6FEABFC72B29202CB" box="[335,404,935,961]" pageId="9" pageNumber="35" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
but plate-like in 
<taxonomicName id="4C10F9E4FFF8FFA6FD86FC7DB1C002CB" box="[610,710,936,961]" class="Arachnida" family="Phyllochthoniidae" genus="Phyllochthonius" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="35" phylum="Arthropoda" rank="species" species="aoutii">
<emphasis id="B9645E75FFF8FFA6FD86FC7DB1C002CB" box="[610,710,936,961]" italics="true" pageId="9" pageNumber="35">P. aoutii</emphasis>
</taxonomicName>
. In addition, the new species possesses shorter interlamellar setae, seven pairs of genital setae, and tarsi III lacking solenidia. In 
<emphasis id="B9645E75FFF8FFA6FC49FC18B71002EC" box="[941,1046,973,998]" italics="true" pageId="9" pageNumber="35">
<taxonomicName id="4C10F9E4FFF8FFA6FC49FC18B71702EC" box="[941,1041,973,998]" class="Arachnida" family="Phyllochthoniidae" genus="Phyllochthonius" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="35" phylum="Arthropoda" rank="species" species="aoutii">P. aoutii</taxonomicName>
,
</emphasis>
the interlamellar setae are longer than the distance between their bases, eight pairs of genital setae are present, and tarsi III bear a solenidion.
</paragraph>
</subSubSection>
</treatment>
</document>