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<document id="D866E9CF3B0B379F855A4EBB66543658" ID-CLB-Dataset="29479" ID-DOI="10.11646/zootaxa.4471.2.3" ID-GBIF-Dataset="694f85ad-7913-43ea-89b9-8eb4e20f0821" ID-ISSN="1175-5326" ID-Zenodo-Dep="1439674" ID-ZooBank="CE4C6E73-09CB-4E74-91C5-771B3696DC52" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1538364447371" checkinUser="plazi" docAuthor="Smith, Andrew B. T. &amp; Evans, Arthur V." docDate="2018" docId="03AF87E48E4BFFDEFF49FAAAFE0081FC" docLanguage="en" docName="zootaxa.4471.2.3.pdf" docOrigin="Zootaxa 4471 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Apteroathlia translucida Smith &amp; Evans 2018, new species" docType="treatment" docUuid="D4F8FA09-1E22-41A0-91F1-CD2945005552" docUuidSource="ZooBank" docVersion="8" lastPageNumber="285" masterDocId="FF96FF9C8E4FFFD8FFDEFFC7FFE38255" masterDocTitle="Taxonomic review of Athliini (Coleoptera: Scarabaeidae: Melolonthinae), a new tribe of scarab beetles endemic to South America" masterLastPageNumber="308" masterPageNumber="279" pageNumber="283" updateTime="1698558211898" updateUser="ExternalLinkService">
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<mods:title id="B2F8365F6703C0343E0668EC9FAA07A2">Taxonomic review of Athliini (Coleoptera: Scarabaeidae: Melolonthinae), a new tribe of scarab beetles endemic to South America</mods:title>
</mods:titleInfo>
<mods:name id="67CD5F5FD06369893C1CCD45BA375FD3" type="personal">
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<mods:namePart id="4BD7F9ED046E30B42F2C7BEC6DF97D32">Smith, Andrew B. T.</mods:namePart>
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<mods:namePart id="8EE9AD3D40A93CE131B6535D3BDB3F46">Evans, Arthur V.</mods:namePart>
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<mods:title id="23D03B327EF8C3AD1D8E9D170F5FEA33">Zootaxa</mods:title>
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<mods:part id="118A0CA41A704BCB74E07243911F6653">
<mods:date id="677A2CA12367A075CA5677AF88F81B80">2018</mods:date>
<mods:detail id="4C157CDB2933A4F2340C0E048393A4AD" type="pubDate">
<mods:number id="04E6A90DC76C926FC3B257F635A5F162">2018-09-05</mods:number>
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<mods:number id="E28951382EDEFE7D90DC4210F0161522">4471</mods:number>
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<mods:classification id="46207110A4CC9BE1AF56562DE9EF0B19">journal article</mods:classification>
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<mods:identifier id="6A65539F32E8CDF0C2ACC22972595965" type="DOI">10.11646/zootaxa.4471.2.3</mods:identifier>
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<treatment id="03AF87E48E4BFFDEFF49FAAAFE0081FC" ID-DOI="http://doi.org/10.5281/zenodo.5958486" ID-GBIF-Taxon="148415679" ID-Zenodo-Dep="5958486" LSID="urn:lsid:zoobank.org:act:D4F8FA09-1E22-41A0-91F1-CD2945005552" httpUri="http://treatment.plazi.org/id/03AF87E48E4BFFDEFF49FAAAFE0081FC" lastPageId="6" lastPageNumber="285" pageId="4" pageNumber="283">
<subSubSection id="C31C65798E4BFFDCFF49FAAAFEDD87FC" pageId="4" pageNumber="283" type="nomenclature">
<paragraph id="8BB936F28E4BFFDCFF49FAAAFCC187D2" blockId="4.[151,1169,1389,1485]" box="[151,802,1389,1415]" pageId="4" pageNumber="283">
<heading id="D0F1819E8E4BFFDCFF49FAAAFCC187D2" bold="true" box="[151,802,1389,1415]" fontSize="11" level="1" pageId="4" pageNumber="283" reason="1">
<emphasis id="B972EAE08E4BFFDCFF49FAAAFCC187D2" bold="true" box="[151,802,1389,1415]" pageId="4" pageNumber="283">
<taxonomicName id="4C064D718E4BFFDCFF49FAAAFD6687D2" ID-CoL="FYHZ" authority="Smith &amp; Evans" authorityName="Smith &amp; Evans" authorityYear="2018" box="[151,645,1389,1415]" class="Insecta" family="Scarabaeidae" genus="Apteroathlia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="283" phylum="Arthropoda" rank="species" species="translucida" status="sp. nov.">
<emphasis id="B972EAE08E4BFFDCFF49FAAAFE5D87D2" bold="true" box="[151,446,1389,1415]" italics="true" pageId="4" pageNumber="283">Apteroathlia translucida</emphasis>
Smith &amp; Evans
</taxonomicName>
,
<taxonomicNameLabel id="A241579B8E4BFFDCFD4DFAAAFCC187D2" box="[659,802,1389,1415]" pageId="4" pageNumber="283" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BB936F28E4BFFDCFF49FA57FEDD87FC" blockId="4.[151,1169,1389,1485]" box="[151,318,1424,1449]" pageId="4" pageNumber="283">
<figureCitation id="133D2A778E4BFFDCFF49FA57FEF387FC" box="[151,272,1424,1449]" captionStart="FIGURES 610" captionStartId="5.[151,264,1714,1736]" captionTargetBox="[335,1263,553,1700]" captionTargetId="figure@5.[335,1263,553,1700]" captionTargetPageId="5" captionText="FIGURES 610. Apteroathlia translucida Smith &amp; Evans, new species, holotype. 6, dorsal habitus; 7, oblique habitus; 8, ventral habitus; 9, head and pronotum; 10, parameres." httpUri="https://zenodo.org/record/1439678/files/figure.png" pageId="4" pageNumber="283">Figs. 610</figureCitation>
,
<figureCitation id="133D2A778E4BFFDCFEC5FA57FED987FC" box="[283,314,1424,1449]" captionStart="FIGURE 66" captionStartId="21.[151,250,1961,1983]" captionTargetBox="[153,1433,499,1939]" captionTargetId="figure@21.[153,1433,499,1939]" captionTargetPageId="21" captionText="FIGURE 66. Distribution map of Apteroathlia nox Smith &amp; Evans, new species (black circles) and Apteroathlia translucida Smith &amp; Evans, new species (red circles)." httpUri="https://zenodo.org/record/1439706/files/figure.png" pageId="4" pageNumber="283">66</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C31C65798E4BFFDCFF49FA73FB728798" box="[151,1169,1460,1485]" pageId="4" pageNumber="283" type="description">
<paragraph id="8BB936F28E4BFFDCFF49FA73FB728798" blockId="4.[151,1169,1389,1485]" box="[151,1169,1460,1485]" pageId="4" pageNumber="283">http://zoobank.org/urn:lsid:zoobank.org:act:D4F8FA09-1E22-41A0-91F1-CD2945005552</paragraph>
</subSubSection>
<subSubSection id="C31C65798E4BFFDDFF49FA3BFD1583A0" lastPageId="5" lastPageNumber="284" pageId="4" pageNumber="283" type="materials_examined">
<paragraph id="8BB936F28E4BFFDCFF49FA3BFD648440" blockId="4.[151,1437,1532,2025]" box="[151,647,1532,1557]" pageId="4" pageNumber="283">
<materialsCitation id="3B6E3CAF8E4BFFDCFF49FA3BFD648440" ID-GBIF-Occurrence="1913888834" box="[151,647,1532,1557]" country="Uruguay" location="Cerro" pageId="4" pageNumber="283" specimenCount="1" stateProvince="Montevideo" typeStatus="holotype">
<emphasis id="B972EAE08E4BFFDCFF49FA3BFED68440" bold="true" box="[151,309,1532,1557]" pageId="4" pageNumber="283">
<typeStatus id="54BD88508E4BFFDCFF49FA3BFF318440" box="[151,210,1532,1557]" pageId="4" pageNumber="283">Type</typeStatus>
locality.
</emphasis>
<location id="8ED960298E4BFFDCFEE2FA3AFE9C8440" LSID="urn:lsid:plazi:treatment:03AF87E48E4BFFDEFF49FAAAFE0081FC:8ED960298E4BFFDCFEE2FA3AFE9C8440" box="[316,383,1533,1557]" country="Uruguay" name="Cerro" pageId="4" pageNumber="283" stateProvince="Montevideo">Cerro</location>
,
<collectingRegion id="49C2F8108E4BFFDCFE54FA3BFDF78440" box="[394,532,1532,1557]" country="Uruguay" name="Montevideo" pageId="4" pageNumber="283">Montevideo</collectingRegion>
,
<collectingCountry id="F31176628E4BFFDCFDC1FA3AFD608440" box="[543,643,1533,1557]" name="Uruguay" pageId="4" pageNumber="283">Uruguay</collectingCountry>
.
</materialsCitation>
</paragraph>
<paragraph id="8BB936F28E4BFFDDFF19F9E7FD1583A0" blockId="4.[151,1437,1532,2025]" lastBlockId="5.[151,1436,151,501]" lastPageId="5" lastPageNumber="284" pageId="4" pageNumber="283">
<emphasis id="B972EAE08E4BFFDCFF19F9E7FEB7846C" bold="true" box="[199,340,1568,1593]" pageId="4" pageNumber="283">Type series.</emphasis>
One male holotype, 14 male paratypes, and 14 female paratypes.
<materialsCitation id="3B6E3CAF8E4BFFDCFB98F9E7FC86850C" ID-GBIF-Occurrence="1913888848" collectingDate="1933-null-null" collectionCode="CMNC, USNM, USMN" collectorName="A. Martinez &amp; Apterops &amp; Cand. &amp; One &amp; Bs As &amp; Azul &amp; Daguerre &amp; Colln. &amp; J. Daguerre &amp; Four" country="Argentina" determinerName="Vialle" location="Buenos Aires" pageId="4" pageNumber="283" specimenCount="5" stateProvince="Buenos Aires" typeStatus="holotype">
<typeStatus id="54BD88508E4BFFDCFB98F9E7FB4D846C" box="[1094,1198,1568,1593]" pageId="4" pageNumber="283" type="holotype">Holotype</typeStatus>
male (
<figureCitation id="133D2A778E4BFFDCFB20F9E7FA94846C" box="[1278,1399,1568,1593]" captionStart="FIGURES 610" captionStartId="5.[151,264,1714,1736]" captionTargetBox="[335,1263,553,1700]" captionTargetId="figure@5.[335,1263,553,1700]" captionTargetPageId="5" captionText="FIGURES 610. Apteroathlia translucida Smith &amp; Evans, new species, holotype. 6, dorsal habitus; 7, oblique habitus; 8, ventral habitus; 9, head and pronotum; 10, parameres." httpUri="https://zenodo.org/record/1439678/files/figure.png" pageId="4" pageNumber="283">Figs. 610</figureCitation>
) at
<collectionCode id="ED17AE378E4BFFDCFF49F982FF098408" box="[151,234,1605,1629]" pageId="4" pageNumber="283">CMNC</collectionCode>
labeled a) “Montevideo / Cerro /
<collectingDate id="EFFCE9DA8E4BFFDCFDB4F982FD348408" box="[618,727,1605,1629]" pageId="4" pageNumber="283" value="1933-null-null">
<date id="FFB810328E4BFFDCFDB4F982FD348408" box="[618,727,1605,1629]" pageId="4" pageNumber="283" value="1933-null-null">1. X. 33</date>
</collectingDate>
” (handwritten), b) “H. &amp; A. HOWDEN / COLLECTION /
<emphasis id="B972EAE08E4BFFDCFAA2F982FA778409" box="[1404,1428,1605,1628]" italics="true" pageId="4" pageNumber="283">ex</emphasis>
.
<collectorName id="26F353248E4BFFDCFF49F9AFFEF484D5" box="[151,279,1640,1664]" pageId="4" pageNumber="283">A.Martínez</collectorName>
coll.” (typeset with black border), c) “
<collectorName id="26F353248E4BFFDCFD00F9AFFCA484D5" box="[734,839,1640,1664]" pageId="4" pageNumber="283">Apterops</collectorName>
/ translucidus / (
<collectorName id="26F353248E4BFFDCFBD1F9AFFBB084D4" box="[1039,1107,1640,1665]" pageId="4" pageNumber="283">Cand.</collectorName>
in litt.)” (handwritten).
<collectorName id="26F353248E4BFFDCFAB3F9AFFA7F84D5" box="[1389,1436,1640,1664]" pageId="4" pageNumber="283">One</collectorName>
<typeStatus id="54BD88508E4BFFDCFF49F94AFF1484F0" box="[151,247,1677,1701]" pageId="4" pageNumber="283" type="paratype">paratype</typeStatus>
male at
<collectionCode id="ED17AE378E4BFFDCFEBFF94AFE5184F0" box="[353,434,1677,1701]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="4" pageNumber="283">USNM</collectionCode>
labeled a) “
<collectorName id="26F353248E4BFFDCFD9DF94AFD6884F0" box="[579,651,1677,1701]" pageId="4" pageNumber="283">Bs As</collectorName>
/
<collectorName id="26F353248E4BFFDCFD76F94BFD3D84F0" box="[680,734,1676,1701]" pageId="4" pageNumber="283">Azul</collectorName>
/ V.956.
<collectorName id="26F353248E4BFFDCFC90F94AFC5A84F0" box="[846,953,1677,1701]" pageId="4" pageNumber="283">Daguerre</collectorName>
” (handwritten), b) “
<collectingCountry id="F31176628E4BFFDCFB7CF94AFAAB84F0" box="[1186,1352,1677,1701]" name="Argentina" pageId="4" pageNumber="283">ARGENTINA</collectingCountry>
/ 1968
<collectorName id="26F353248E4BFFDCFF49F977FF3E849C" box="[151,221,1712,1737]" pageId="4" pageNumber="283">Colln.</collectorName>
/
<collectorName id="26F353248E4BFFDCFF2DF977FE8C849D" box="[243,367,1712,1736]" pageId="4" pageNumber="283">J.Daguerre</collectorName>
” (typeset over red, white, and green background), c) “
<collectorName id="26F353248E4BFFDCFC0BF977FBDD849D" box="[981,1086,1712,1736]" pageId="4" pageNumber="283">Apterops</collectorName>
/ translucidus / (in lit.)
<collectorName id="26F353248E4BFFDCFA94F977FA6E849C" box="[1354,1421,1712,1737]" pageId="4" pageNumber="283">Cand.</collectorName>
/
<determinerName id="7E65DA368E4BFFDCFF49F913FF3984B8" box="[151,218,1748,1773]" pageId="4" pageNumber="283">Vialle</determinerName>
det” (handwritten).
<collectorName id="26F353248E4BFFDCFE1DF912FE1184B8" box="[451,498,1749,1773]" pageId="4" pageNumber="283">One</collectorName>
<typeStatus id="54BD88508E4BFFDCFE25F912FDB884B8" box="[507,603,1749,1773]" pageId="4" pageNumber="283" type="paratype">paratype</typeStatus>
male at
<collectionCode id="ED17AE378E4BFFDCFD1FF912FCF784B8" box="[705,788,1749,1773]" pageId="4" pageNumber="283">CMNC</collectionCode>
and one parartype female at
<collectionCode id="ED17AE378E4BFFDCFBB8F912FB5484B8" box="[1126,1207,1749,1773]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="4" pageNumber="283">USNM</collectionCode>
labeled a) “
<collectorName id="26F353248E4BFFDCFA9AF912FA6884B8" box="[1348,1419,1749,1773]" pageId="4" pageNumber="283">Bs As</collectorName>
/
<collectorName id="26F353248E4BFFDCFF49F93FFF2E8544" box="[151,205,1784,1809]" pageId="4" pageNumber="283">Azul</collectorName>
/ V.956.
<collectorName id="26F353248E4BFFDCFEECF93FFE7E8545" box="[306,413,1784,1808]" pageId="4" pageNumber="283">Daguerre</collectorName>
” (handwritten), b) “
<collectingCountry id="F31176628E4BFFDCFDA2F93FFCC18545" box="[636,802,1784,1808]" name="Argentina" pageId="4" pageNumber="283">ARGENTINA</collectingCountry>
/ 1968
<collectorName id="26F353248E4BFFDCFCA8F93FFC5F8544" box="[886,956,1784,1809]" pageId="4" pageNumber="283">Colln.</collectorName>
/
<collectorName id="26F353248E4BFFDCFC0FF93FFBAE8545" box="[977,1101,1784,1808]" pageId="4" pageNumber="283">J.Daguerre</collectorName>
” (typeset over red, white, and green background).
<collectorName id="26F353248E4BFFDCFEA9F8DAFE488561" box="[375,427,1821,1844]" pageId="4" pageNumber="283">Four</collectorName>
male
<typeStatus id="54BD88508E4BFFDCFE2CF8DAFDB88560" box="[498,603,1821,1845]" pageId="4" pageNumber="283" type="paratype">paratypes</typeStatus>
at
<collectionCode id="ED17AE378E4BFFDCFD5EF8DAFD328560" box="[640,721,1821,1845]" pageId="4" pageNumber="283">CMNC</collectionCode>
(2) and
<collectionCode id="ED17AE378E4BFFDCFCEAF8DAFC678560" box="[820,900,1821,1845]" pageId="4" pageNumber="283">USMN</collectionCode>
(2) and six female
<typeStatus id="54BD88508E4BFFDCFBBDF8DAFB2F8560" box="[1123,1228,1821,1845]" pageId="4" pageNumber="283" type="paratype">paratypes</typeStatus>
at
<collectionCode id="ED17AE378E4BFFDCFB2FF8DAFAA08560" box="[1265,1347,1821,1845]" pageId="4" pageNumber="283">CMNC</collectionCode>
(3) and
<collectionCode id="ED17AE378E4BFFDCFF49F887FF04850D" box="[151,231,1856,1880]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="4" pageNumber="283">USNM</collectionCode>
(3) labeled a) “
<collectingCountry id="F31176628E4BFFDCFE53F887FDD3850D" box="[397,560,1856,1880]" name="Argentina" pageId="4" pageNumber="283">ARGENTINA</collectingCountry>
/
<collectingRegion id="49C2F8108E4BFFDCFD9BF886FD39850C" box="[581,730,1856,1881]" country="Argentina" name="Ciudad Autonoma de Buenos Aires" pageId="4" pageNumber="283">Buenos Aires</collectingRegion>
Prov. /
<collectorName id="26F353248E4BFFDCFCF2F887FC86850C" box="[812,869,1856,1881]" pageId="4" pageNumber="283">Azul</collectorName>
</materialsCitation>
;
<date id="FFB810328E4BFFDCFCB1F887FC5D850C" box="[879,958,1856,1881]" pageId="4" pageNumber="283" value="1956-05">V.1956</date>
/ J.
<materialsCitation id="3B6E3CAF8E4BFFDDFC37F887FDB082AD" ID-GBIF-Occurrence="1913888851" collectingDate="1909" collectingDateMax="1974" collectingDateMin="1909" collectionCode="USNM, CMNC, BMNH" collectorName="A. Martinez &amp; One &amp; La Plata &amp; M Video &amp; Fry Coll." country="Uruguay" lastPageId="5" lastPageNumber="284" location="Montevideo" pageId="4" pageNumber="283" specimenCount="5" stateProvince="Montevideo" typeStatus="paratype">
Daguerre” (typeset). One male
<typeStatus id="54BD88508E4BFFDCFAE0F886FA7F850C" box="[1342,1436,1857,1881]" pageId="4" pageNumber="283" type="paratype">paratype</typeStatus>
at
<collectionCode id="ED17AE378E4BFFDCFF6AF8A2FEE78528" box="[180,260,1893,1917]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="4" pageNumber="283">USNM</collectionCode>
labeled a) “
<collectingRegion id="49C2F8108E4BFFDCFE53F8A3FDF78528" box="[397,532,1892,1917]" country="Uruguay" name="Montevideo" pageId="4" pageNumber="283">Montevideo</collectingRegion>
/ Cerro /
<collectingDate id="EFFCE9DA8E4BFFDCFD59F8A2FD0C8528" box="[647,751,1893,1917]" pageId="4" pageNumber="283" value="1934-09-12">
<date id="FFB810328E4BFFDCFD59F8A2FD0C8528" box="[647,751,1893,1917]" pageId="4" pageNumber="283" value="1934-09-12">12 IX 34</date>
</collectingDate>
” (handwritten), b) “Apterops / translucidus / (Chand. i. litt)” (handwritten). One
<typeStatus id="54BD88508E4BFFDCFEB1F84EFE2E85F4" box="[367,461,1929,1953]" pageId="4" pageNumber="283" type="paratype">paratype</typeStatus>
male at
<collectionCode id="ED17AE378E4BFFDCFDF0F84FFD6385F5" box="[558,640,1928,1952]" pageId="4" pageNumber="283">CMNC</collectionCode>
labeled a) “
<collectingRegion id="49C2F8108E4BFFDCFCDAF84FFC6885F4" box="[772,907,1928,1953]" country="Uruguay" name="Montevideo" pageId="4" pageNumber="283">Montevideo</collectingRegion>
/ Cerro /
<collectingDate id="EFFCE9DA8E4BFFDCFC29F84FFB8285F5" box="[1015,1121,1928,1952]" pageId="4" pageNumber="283" value="1933-null-null">
<date id="FFB810328E4BFFDCFC29F84FFB8285F5" box="[1015,1121,1928,1952]" pageId="4" pageNumber="283" value="1933-null-null">1. X. 33</date>
</collectingDate>
” (handwritten), b) “H. &amp; A. HOWDEN / COLLECTION /
<emphasis id="B972EAE08E4BFFDCFE37F86AFDE28591" box="[489,513,1965,1988]" italics="true" pageId="4" pageNumber="283">ex</emphasis>
.
<collectorName id="26F353248E4BFFDCFDD1F86AFD6E8590" box="[527,653,1965,1989]" pageId="4" pageNumber="283">A.Martínez</collectorName>
coll.” (typeset with black border). Four male and two female
<typeStatus id="54BD88508E4BFFDCFAEDF86AFA7F8590" box="[1331,1436,1965,1989]" pageId="4" pageNumber="283" type="paratype">paratypes</typeStatus>
at
<collectionCode id="ED17AE378E4BFFDCFF6FF817FEE185BD" box="[177,258,2000,2024]" pageId="4" pageNumber="283">CMNC</collectionCode>
labeled a) “R. O. del /
<collectingCountry id="F31176628E4BFFDCFE20F817FD8385BD" box="[510,608,2000,2024]" name="Uruguay" pageId="4" pageNumber="283">Uruguay</collectingCountry>
” (typeset on faded green paper), b) “H. &amp; A. HOWDEN / COLLECTION /
<emphasis id="B972EAE08E4AFFDDFF49FF5EFF4C82E5" box="[151,175,153,176]" italics="true" pageId="5" pageNumber="284">ex</emphasis>
.
<collectorName id="26F353248E4AFFDDFF63FF5FFED882E5" box="[189,315,152,176]" pageId="5" pageNumber="284">A.Martínez</collectorName>
coll.” (typeset with black border).
<collectorName id="26F353248E4AFFDDFD63FF5FFD0882E5" box="[701,747,152,176]" pageId="5" pageNumber="284">One</collectorName>
male
<typeStatus id="54BD88508E4AFFDDFCEEFF5FFC6D82E5" box="[816,910,152,176]" pageId="5" pageNumber="284" type="paratype">paratype</typeStatus>
at
<collectionCode id="ED17AE378E4AFFDDFC6FFF5FFBE782FA" box="[945,1028,152,175]" pageId="5" pageNumber="284">BMNH</collectionCode>
labeled a) “18234” (handwritten), b) “
<collectorName id="26F353248E4AFFDDFF7FFF7BFEE38280" box="[161,256,188,213]" pageId="5" pageNumber="284">La Plata</collectorName>
/
<collectorName id="26F353248E4AFFDDFEC4FF7AFE638280" box="[282,384,188,213]" pageId="5" pageNumber="284">M Video</collectorName>
” (handwritten), c) “
<collectorName id="26F353248E4AFFDDFDBFFF7BFD2A8280" box="[609,713,188,213]" pageId="5" pageNumber="284">Fry Coll.</collectorName>
/ 1905-100.” (typeset).
<collectorName id="26F353248E4AFFDDFC06FF7BFBE58281" box="[984,1030,188,212]" pageId="5" pageNumber="284">One</collectorName>
male
<typeStatus id="54BD88508E4AFFDDFB90FF7AFB4F8280" box="[1102,1196,189,213]" pageId="5" pageNumber="284" type="paratype">paratype</typeStatus>
at
<collectionCode id="ED17AE378E4AFFDDFB0CFF7AFAC68281" box="[1234,1317,189,212]" pageId="5" pageNumber="284">BMNH</collectionCode>
labeled a) “Monte- / video” (typeset), b) “
<collectingDate id="EFFCE9DA8E4AFFDDFE33FF27FDB082AD" box="[493,595,224,248]" pageId="5" pageNumber="284" value="1909" valueMax="1974">
<date id="FFB810328E4AFFDDFE33FF27FDB082AD" box="[493,595,224,248]" pageId="5" pageNumber="284" value="1909" valueMax="1974">190974</date>
</collectingDate>
</materialsCitation>
.
<materialsCitation id="3B6E3CAF8E4AFFDDFD89FF27FD1583A0" ID-GBIF-Occurrence="1913888854" collectingDate="1933-09-24" collectingDateMax="1934-09-12" collectingDateMin="1933-09-24" collectionCode="BMNH, MNHN, MUSEUM, PARIS, USNM" collectorName="Coll. Leon &amp; One &amp; Cerro &amp; Apterops &amp; Chand. &amp; Collection &amp; Two &amp; All" country="Brazil" location="Montevideo" pageId="5" pageNumber="284" specimenCount="6" stateProvince="Montevideo" typeStatus="paratype">
” (typeset), c) “21.” (handwritten). One female
<typeStatus id="54BD88508E4AFFDDFB87FF27FB5482AD" box="[1113,1207,224,248]" pageId="5" pageNumber="284" type="paratype">paratype</typeStatus>
at
<collectionCode id="ED17AE378E4AFFDDFB06FF27FAC882A2" box="[1240,1323,224,247]" pageId="5" pageNumber="284">BMNH</collectionCode>
labeled a) “
<collectingCountry id="F31176628E4AFFDDFF7FFEC3FF0B8348" box="[161,232,260,285]" name="Brazil" pageId="5" pageNumber="284">Brazil</collectingCountry>
” (handwritten), b) “2032.” (handwritten), c) “67-45” (handwritten), d) “inflate Reiche / Brasilia” (handwritten), e) “So named / in Reiches / Collection. / C.W.” (typeset). One female
<typeStatus id="54BD88508E4AFFDDFB91FEEFFB4E8315" box="[1103,1197,296,320]" pageId="5" pageNumber="284" type="paratype">paratype</typeStatus>
at
<collectionCode id="ED17AE378E4AFFDDFB0CFEEFFAC4836A" box="[1234,1319,296,319]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="5" pageNumber="284">MNHN</collectionCode>
labeled a) “Plata / Dolle” (handwritten), b) “
<collectionCode id="ED17AE378E4AFFDDFDCCFE8BFD018331" box="[530,738,332,357]" pageId="5" pageNumber="284">MUSÉUM PARIS</collectionCode>
/ 1906 /
<collectorName id="26F353248E4AFFDDFC92FE8BFC208331" box="[844,963,332,357]" pageId="5" pageNumber="284">Coll. Léon</collectorName>
FAIRMAIRE” (handwritten).
<collectorName id="26F353248E4AFFDDFAC5FE8BFAAA8331" box="[1307,1353,332,356]" pageId="5" pageNumber="284">One</collectorName>
female
<typeStatus id="54BD88508E4AFFDDFF49FEB7FF1683DD" box="[151,245,368,392]" pageId="5" pageNumber="284" type="paratype">paratype</typeStatus>
at
<collectionCode id="ED17AE378E4AFFDDFECBFEB7FE8683DD" box="[277,357,368,392]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="5" pageNumber="284">USNM</collectionCode>
labeled a) “
<collectingRegion id="49C2F8108E4AFFDDFE38FEA8FD8E83DD" box="[486,621,367,392]" country="Uruguay" name="Montevideo" pageId="5" pageNumber="284">Montevideo</collectingRegion>
/
<collectorName id="26F353248E4AFFDDFD5FFEB7FD5C83DD" box="[641,703,368,392]" pageId="5" pageNumber="284">Cerro</collectorName>
/
<collectingDate id="EFFCE9DA8E4AFFDDFD0AFEB7FCA083DD" box="[724,835,368,392]" pageId="5" pageNumber="284" value="1933-09-24">
<date id="FFB810328E4AFFDDFD0AFEB7FCA083DD" box="[724,835,368,392]" pageId="5" pageNumber="284" value="1933-09-24">24. IX. 33</date>
</collectingDate>
” (handwritten), b) “
<collectorName id="26F353248E4AFFDDFBC3FEB7FB6783DD" box="[1053,1156,368,392]" pageId="5" pageNumber="284">Apterops</collectorName>
/ translucidus / (
<collectorName id="26F353248E4AFFDDFAE6FEA8FA6B83DD" box="[1336,1416,367,392]" pageId="5" pageNumber="284">Chand.</collectorName>
i. litt.)” (handwritten), c) “OLCartwright /
<collectorName id="26F353248E4AFFDDFDBBFE53FD3483F8" box="[613,727,404,429]" pageId="5" pageNumber="284">Collection</collectorName>
/ 1959” (typeset).
<collectorName id="26F353248E4AFFDDFC6CFE52FC0183F9" box="[946,994,405,428]" pageId="5" pageNumber="284">Two</collectorName>
female
<typeStatus id="54BD88508E4AFFDDFB9CFE52FB4883F8" box="[1090,1195,405,429]" pageId="5" pageNumber="284" type="paratype">paratypes</typeStatus>
at
<collectionCode id="ED17AE378E4AFFDDFB0DFE53FAC083F9" box="[1235,1315,404,428]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="5" pageNumber="284">USNM</collectionCode>
labeled a) “
<collectingRegion id="49C2F8108E4AFFDDFF7EFE70FEC48385" box="[160,295,439,464]" country="Uruguay" name="Montevideo" pageId="5" pageNumber="284">Montevideo</collectingRegion>
/
<collectorName id="26F353248E4AFFDDFE9FFE7FFE9C8385" box="[321,383,440,464]" pageId="5" pageNumber="284">Cerro</collectorName>
/
<collectingDate id="EFFCE9DA8E4AFFDDFE47FE7FFDE38385" box="[409,512,440,464]" pageId="5" pageNumber="284" value="1934-09-12">
<date id="FFB810328E4AFFDDFE47FE7FFDE38385" box="[409,512,440,464]" pageId="5" pageNumber="284" value="1934-09-12">12 IX 34</date>
</collectingDate>
” (handwritten).
<collectorName id="26F353248E4AFFDDFD66FE70FD388385" box="[696,731,439,464]" pageId="5" pageNumber="284">All</collectorName>
<typeStatus id="54BD88508E4AFFDDFD3AFE7FFCAE8385" box="[740,845,440,464]" pageId="5" pageNumber="284" type="paratype">paratypes</typeStatus>
listed above also bear a yellow
<typeStatus id="54BD88508E4AFFDDFB64FE7FFAFB8385" box="[1210,1304,440,464]" pageId="5" pageNumber="284" type="paratype">paratype</typeStatus>
label and a database label with the database number and a barcode.
</materialsCitation>
</paragraph>
</subSubSection>
<caption id="DF79667A8E4AFFDDFF49F975FD5A84B3" httpUri="https://zenodo.org/record/1439678/files/figure.png" pageId="5" pageNumber="284" startId="5.[151,264,1714,1736]" targetBox="[335,1263,553,1700]" targetPageId="5">
<paragraph id="8BB936F28E4AFFDDFF49F975FD5A84B3" blockId="5.[151,1436,1714,1766]" pageId="5" pageNumber="284">
<emphasis id="B972EAE08E4AFFDDFF49F975FEA9849D" bold="true" box="[151,330,1714,1736]" pageId="5" pageNumber="284">FIGURES 610.</emphasis>
<taxonomicName id="4C064D718E4AFFDDFE8DF975FD1F849D" authority="Smith &amp; Evans" authorityName="Smith &amp; Evans" authorityYear="2018" box="[339,764,1714,1736]" class="Insecta" family="Scarabaeidae" genus="Apteroathlia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="284" phylum="Arthropoda" rank="species" species="translucida" status="sp. nov.">
<emphasis id="B972EAE08E4AFFDDFE8DF975FE31849D" box="[339,466,1714,1736]" italics="true" pageId="5" pageNumber="284">Apteroathlia</emphasis>
<emphasis id="B972EAE08E4AFFDDFE02F975FDAC849D" box="[476,591,1714,1736]" italics="true" pageId="5" pageNumber="284">translucida</emphasis>
Smith &amp; Evans
</taxonomicName>
,
<taxonomicNameLabel id="A241579B8E4AFFDDFCD4F974FC69849D" box="[778,906,1714,1736]" pageId="5" pageNumber="284" rank="species">new species</taxonomicNameLabel>
, holotype. 6, dorsal habitus; 7, oblique habitus; 8, ventral habitus; 9, head and pronotum; 10, parameres.
</paragraph>
</caption>
<subSubSection id="C31C65798E4AFFDEFF19F8C9FE6783AD" lastPageId="6" lastPageNumber="285" pageId="5" pageNumber="284" type="description">
<paragraph id="8BB936F28E4AFFDEFF19F8C9FE9D83FA" blockId="5.[151,1436,1806,2011]" lastBlockId="6.[151,1437,151,938]" lastPageId="6" lastPageNumber="285" pageId="5" pageNumber="284">
<emphasis id="B972EAE08E4AFFDDFF19F8C9FE008572" bold="true" box="[199,483,1806,1831]" pageId="5" pageNumber="284">
Description of
<typeStatus id="54BD88508E4AFFDDFEA2F8C9FE008572" box="[380,483,1806,1831]" pageId="5" pageNumber="284" type="holotype">holotype</typeStatus>
</emphasis>
(
<figureCitation id="133D2A778E4AFFDDFE2BF8C9FD908572" box="[501,627,1806,1831]" captionStart="FIGURES 610" captionStartId="5.[151,264,1714,1736]" captionTargetBox="[335,1263,553,1700]" captionTargetId="figure@5.[335,1263,553,1700]" captionTargetPageId="5" captionText="FIGURES 610. Apteroathlia translucida Smith &amp; Evans, new species, holotype. 6, dorsal habitus; 7, oblique habitus; 8, ventral habitus; 9, head and pronotum; 10, parameres." httpUri="https://zenodo.org/record/1439678/files/figure.png" pageId="5" pageNumber="284">Figs. 610</figureCitation>
). Male. Length
<quantity id="4CFE9B178E4AFFDDFCF2F8C9FC648573" box="[812,903,1806,1831]" metricMagnitude="-3" metricUnit="m" metricValue="6.6" pageId="5" pageNumber="284" unit="mm" value="6.6">6.6 mm</quantity>
, width
<quantity id="4CFE9B178E4AFFDDFC01F8C8FBD98573" box="[991,1082,1807,1831]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="5" pageNumber="284" unit="mm" value="3.1">3.1 mm</quantity>
. Dorsal and ventral color light brown, elytra translucent. Dorsal surface glabrous.
<emphasis id="B972EAE08E4AFFDDFD0BF8F5FCF1851F" box="[725,786,1842,1866]" italics="true" pageId="5" pageNumber="284">Head</emphasis>
: surface moderately punctate. Labrum with apex reflexed backward and convexly parabolic medially; underside concave medially. Clypeus with apex straight, abutted to labrum. Frontoclypeal suture visible. Eyes round, slightly protruding laterally. Antennae with 8 antennomeres; antennal club with 3 antennomeres, club distinctly shorter in length to antennomeres 15 combined, club surface thickly setose. Maxillae robust, triangular, basal width slightly less than width of mentum; maxillary palpus with 4 palpomeres, length approximately ¾ that of antennae. Mentum prominent, slightly longer than wide, apex straight, surface slightly convex; labial palpus reduced.
<emphasis id="B972EAE08E49FFDEFD65FF7AFCC98281" box="[699,810,189,212]" italics="true" pageId="6" pageNumber="285">Pronotum</emphasis>
: surface moderately to densely punctate; apical angle acute, basal angle squared. Pronotum widest medially with smooth margins.
<emphasis id="B972EAE08E49FFDEFC28FF27FBDF82AD" box="[1014,1084,224,248]" italics="true" pageId="6" pageNumber="285">Elytra</emphasis>
: surface moderately to densely punctate, striae punctate but obscured by other surface punctures.
<emphasis id="B972EAE08E49FFDEFCA9FEC2FC588349" box="[887,955,261,284]" italics="true" pageId="6" pageNumber="285">Wings</emphasis>
: apterous.
<emphasis id="B972EAE08E49FFDEFBE8FEC2FB988349" box="[1078,1147,261,284]" italics="true" pageId="6" pageNumber="285">Venter</emphasis>
: metasternum with dense punctures. Pygidium flat; surface moderately to sparsely punctate with microsculpturing.
<emphasis id="B972EAE08E49FFDEFB76FEEEFB3E8315" box="[1192,1245,297,320]" italics="true" pageId="6" pageNumber="285">Legs</emphasis>
: protibia with 3 apical teeth on outer margin of apical half; 3rd tooth weak. Claws symmetrical with medial tooth. Protibial spur present, metatibial spurs approximately equal in length.
<emphasis id="B972EAE08E49FFDEFCC7FEB4FC6083DE" box="[793,899,371,395]" italics="true" pageId="6" pageNumber="285">Genitalia</emphasis>
: parameres long, slender, tapering into needlelike apices (
<figureCitation id="133D2A778E49FFDEFEC3FE51FE9383FA" box="[285,368,406,431]" captionStart="FIGURES 610" captionStartId="5.[151,264,1714,1736]" captionTargetBox="[335,1263,553,1700]" captionTargetId="figure@5.[335,1263,553,1700]" captionTargetPageId="5" captionText="FIGURES 610. Apteroathlia translucida Smith &amp; Evans, new species, holotype. 6, dorsal habitus; 7, oblique habitus; 8, ventral habitus; 9, head and pronotum; 10, parameres." httpUri="https://zenodo.org/record/1439678/files/figure.png" pageId="6" pageNumber="285">Fig. 10</figureCitation>
).
</paragraph>
<paragraph id="8BB936F28E49FFDEFF19FE7DFE6783AD" blockId="6.[151,1437,151,938]" pageId="6" pageNumber="285">
<emphasis id="B972EAE08E49FFDEFF19FE7DFEDF8386" bold="true" box="[199,316,442,467]" pageId="6" pageNumber="285">Variation.</emphasis>
Length
<quantity id="4CFE9B178E49FFDEFE79FE7DFDD48387" box="[423,567,442,467]" metricMagnitude="-3" metricUnit="m" metricValue="6.8" metricValueMax="7.7" metricValueMin="5.9" pageId="6" pageNumber="285" unit="mm" value="6.8" valueMax="7.7" valueMin="5.9">5.97.7 mm</quantity>
. Dorsal color tan to brown. Females with slightly more inflated, convex abdominal segments.
</paragraph>
</subSubSection>
<subSubSection id="C31C65798E49FFDEFF19FDC5FDB38036" pageId="6" pageNumber="285" type="etymology">
<paragraph id="8BB936F28E49FFDEFF19FDC5FDB38036" blockId="6.[151,1437,151,938]" pageId="6" pageNumber="285">
<emphasis id="B972EAE08E49FFDEFF19FDC5FEAD804E" bold="true" box="[199,334,514,539]" pageId="6" pageNumber="285">Etymology.</emphasis>
This name is in reference to the translucent elytra and an homage to “
<emphasis id="B972EAE08E49FFDEFB51FDC4FA6B804E" box="[1167,1416,515,539]" italics="true" pageId="6" pageNumber="285">Apterops translucidus</emphasis>
”, which was used on older determination labels for some specimens but was never published. It should be treated as an adjective in the nominative singular.
</paragraph>
</subSubSection>
<subSubSection id="C31C65798E49FFDEFF19FDA9FCD2809A" pageId="6" pageNumber="285" type="distribution">
<paragraph id="8BB936F28E49FFDEFF19FDA9FC9180FE" blockId="6.[151,1437,151,938]" pageId="6" pageNumber="285">
<emphasis id="B972EAE08E49FFDEFF19FDA9FEBA80D2" bold="true" box="[199,345,622,647]" pageId="6" pageNumber="285">Distribution</emphasis>
(
<figureCitation id="133D2A778E49FFDEFEB3FDA9FE2680D2" box="[365,453,622,647]" captionStart="FIGURE 66" captionStartId="21.[151,250,1961,1983]" captionTargetBox="[153,1433,499,1939]" captionTargetId="figure@21.[153,1433,499,1939]" captionTargetPageId="21" captionText="FIGURE 66. Distribution map of Apteroathlia nox Smith &amp; Evans, new species (black circles) and Apteroathlia translucida Smith &amp; Evans, new species (red circles)." httpUri="https://zenodo.org/record/1439706/files/figure.png" pageId="6" pageNumber="285">Fig. 66</figureCitation>
).
<collectingCountry id="F31176628E49FFDEFE02FDA8FD8680D2" box="[476,613,623,647]" name="Uruguay" pageId="6" pageNumber="285">URUGUAY</collectingCountry>
(14):
<collectingRegion id="49C2F8108E49FFDEFD6DFDA9FCD980D2" box="[691,826,622,647]" country="Uruguay" name="Montevideo" pageId="6" pageNumber="285">Montevideo</collectingRegion>
(8): Cerro,
<collectingRegion id="49C2F8108E49FFDEFC15FDA9FBB580D2" box="[971,1110,622,647]" country="Uruguay" name="Montevideo" pageId="6" pageNumber="285">Montevideo</collectingRegion>
; no data (6).
<collectingCountry id="F31176628E49FFDEFB27FDA8FA7F80D2" box="[1273,1436,623,647]" name="Argentina" pageId="6" pageNumber="285">ARGENTINA</collectingCountry>
(14):
<collectingRegion id="49C2F8108E49FFDEFF0AFD54FE8F80FE" box="[212,364,658,683]" country="Argentina" name="Ciudad Autonoma de Buenos Aires" pageId="6" pageNumber="285">Buenos Aires</collectingRegion>
(14): Azul, La Plata. NO DATA (1): “
<collectingCountry id="F31176628E49FFDEFCC6FD55FC8080FE" box="[792,867,658,683]" name="Brazil" pageId="6" pageNumber="285">Brazil</collectingCountry>
”.
</paragraph>
<paragraph id="8BB936F28E49FFDEFF19FD71FCD2809A" blockId="6.[151,1437,151,938]" box="[199,817,694,720]" pageId="6" pageNumber="285">
<emphasis id="B972EAE08E49FFDEFF19FD71FE9E809A" bold="true" box="[199,381,694,719]" pageId="6" pageNumber="285">Temporal data.</emphasis>
May (13), September (4), October (2).
</paragraph>
</subSubSection>
<subSubSection id="C31C65798E49FFDEFF19FD1DFE0081FC" pageId="6" pageNumber="285" type="discussion">
<paragraph id="8BB936F28E49FFDEFF19FD1DFE0081FC" blockId="6.[151,1437,151,938]" pageId="6" pageNumber="285">
<emphasis id="B972EAE08E49FFDEFF19FD1DFED780A6" bold="true" box="[199,308,730,755]" pageId="6" pageNumber="285">Remarks</emphasis>
.
<taxonomicName id="4C064D718E49FFDEFE9DFD1CFDBB80A6" authority="Smith &amp; Evans, 2018" authorityName="Smith &amp; Evans" authorityYear="2018" box="[323,600,731,755]" class="Insecta" family="Scarabaeidae" genus="Apteroathlia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="285" phylum="Arthropoda" rank="species" species="translucida" status="sp. nov.">
<emphasis id="B972EAE08E49FFDEFE9DFD1CFDBB80A6" box="[323,600,731,755]" italics="true" pageId="6" pageNumber="285">Apteroathlia translucida</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A241579B8E49FFDEFDBFFD1CFD0A80A6" box="[609,745,730,755]" pageId="6" pageNumber="285" rank="species">new species</taxonomicNameLabel>
is mainly found around the gulf shores of Río de la Plata in
<collectingCountry id="F31176628E49FFDEFF49FCC5FF1A814F" box="[151,249,770,794]" name="Uruguay" pageId="6" pageNumber="285">Uruguay</collectingCountry>
and
<collectingRegion id="49C2F8108E49FFDEFEEDFCC5FDD8814F" box="[307,571,769,794]" country="Argentina" name="Buenos Aires" pageId="6" pageNumber="285">Buenos Aires Province</collectingRegion>
,
<collectingCountry id="F31176628E49FFDEFD96FCC6FD58814F" box="[584,699,769,794]" name="Argentina" pageId="6" pageNumber="285">Argentina</collectingCountry>
. One 19th Century specimen is labeled “
<collectingCountry id="F31176628E49FFDEFB5BFCC6FB2C814F" box="[1157,1231,769,794]" name="Brazil" pageId="6" pageNumber="285">Brazil</collectingCountry>
”, but we consider this record doubtful and in need of further verification. The specimens collected in May were probably collected fresh while those obtained in September and October appear to have been found dead and dried. The conservation status of this species should be investigated since most of the specimens were collected from coastal habitats in highly populated urban areas.
</paragraph>
</subSubSection>
</treatment>
</document>