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<document id="47F99AC0305B1C152F72A73186A8A348" ID-DOI="10.1080/00222933.2021.1902587" ID-ISSN="1464-5262" ID-Zenodo-Dep="5473916" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630927239686" checkinUser="plazi" docAuthor="Barbosa, Taciano M., Mello-Patiu, Cátia A. &amp; Vasconcelos, Simão D." docDate="2021" docId="03AF87CE3E652B58F119FDCFB6BDFD16" docLanguage="en" docName="JNATHIST.55.5-6.305-340.pdf" docOrigin="Journal of Natural History 55 (5 - 6)" docSource="http://dx.doi.org/10.1080/00222933.2021.1902587" docStyle="DocumentStyle:3C762F9783B768A90048B82ACD135A62.3:JNatHist.2017-.journal_article.0cover" docStyleId="3C762F9783B768A90048B82ACD135A62" docStyleName="JNatHist.2017-.journal_article.0cover" docStyleVersion="3" docTitle="Titanogrypa (Cucullomyia) placida" docType="treatment" docVersion="4" lastPageNumber="334" masterDocId="FF96FFB63E7E2B45F03FFFD3B70CFF92" masterDocTitle="Revision of the New World subgenus Titanogrypa (Cucullomyia) (Diptera: Sarcophagidae), with a reassessment of diagnostic characters" masterLastPageNumber="340" masterPageNumber="305" pageNumber="332" updateTime="1699282807775" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="3B5A243202CF4B49CAABD3AC8661366E">Revision of the New World subgenus Titanogrypa (Cucullomyia) (Diptera: Sarcophagidae), with a reassessment of diagnostic characters</mods:title>
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<mods:namePart id="9FDDE2FDF4C426DDAD6DED52958C366C">Barbosa, Taciano M.</mods:namePart>
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<mods:namePart id="D93ECAA5B670802D73BC95A2892C3834">Mello-Patiu, Cátia A.</mods:namePart>
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<mods:namePart id="0ED2344C6CA582DABDE08F41C897155D">Vasconcelos, Simão D.</mods:namePart>
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<mods:title id="494C85B3D1671A1CFB6F9F115F399C5D">Journal of Natural History</mods:title>
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<mods:part id="0AA2A51F74CC8B07BBCAE45EB1CA77E6">
<mods:date id="31F1D143B6AFCF5BA876D8EFCDC086F6">2021</mods:date>
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<mods:number id="013DAA8DBB443AB01FF2013166FE4F11">2021-06-11</mods:number>
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<mods:detail id="5C09119C6A315251F0BCAEB51190FD49" type="volume">
<mods:number id="DFF872B1C009C26F150D2B1D71F44E75">55</mods:number>
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<mods:number id="169842E0B5A369BA5084E2F593CB2E4A">5 - 6</mods:number>
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<mods:identifier id="8A4BDCAA9AB8AE6642F88F9B598D108F" type="ISSN">1464-5262</mods:identifier>
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<paragraph id="8BB936D83E652B5EF119FDCFB4F2FDA4" blockId="27.[294,1022,540,566]" box="[294,1022,540,566]" pageId="27" pageNumber="332">
<heading id="D0F181B43E652B5EF119FDCFB4F2FDA4" box="[294,1022,540,566]" centered="true" fontSize="11" level="2" pageId="27" pageNumber="332" reason="5">
<taxonomicName id="4C064D5B3E652B5EF119FDCFB481FDA4" ID-CoL="8FGBB" authority="(Aldrich, 1925)" baseAuthorityName="Aldrich" baseAuthorityYear="1925" box="[294,909,540,566]" class="Insecta" family="Sarcophagidae" genus="Titanogrypa" kingdom="Animalia" order="Diptera" pageId="27" pageNumber="332" phylum="Arthropoda" rank="species" species="placida" subGenus="Cucullomyia">
<emphasis id="B972EACA3E652B5EF119FDCFB489FDA4" bold="true" box="[294,901,540,566]" pageId="27" pageNumber="332">
<emphasis id="B972EACA3E652B5EF119FDCFB6B0FDA4" bold="true" box="[294,444,540,566]" italics="true" pageId="27" pageNumber="332">Titanogrypa</emphasis>
(
<emphasis id="B972EACA3E652B5EF1F3FDCFB56EFDA4" bold="true" box="[460,610,540,566]" italics="true" pageId="27" pageNumber="332">Cucullomyia</emphasis>
)
<emphasis id="B972EACA3E652B5EF24CFDCFB5C0FDA4" bold="true" box="[627,716,540,566]" italics="true" pageId="27" pageNumber="332">placida</emphasis>
(
<bibRefCitation id="EF974B293E652B5EF2E3FDCFB489FDA4" author="Aldrich JM" box="[732,901,540,566]" pageId="27" pageNumber="332" pagination="1 - 36" refId="ref17892" refString="Aldrich JM. 1925. New Diptera or two-winged flies in the United States National Museum. Proc US Natl Mus. 66: 1 - 36. doi: 10.5479 / si. 00963801.66 - 2555.1." type="journal article" year="1925">Aldrich, 1925</bibRefCitation>
</emphasis>
)
</taxonomicName>
(
<figureCitation id="133D2A5D3E652B5EF3AAFDCFB4F9FDA4" box="[917,1013,540,566]" captionStart="Figure 6" captionStartId="11.[160,225,1515,1538]" captionTargetBox="[194,1152,150,1482]" captionTargetId="figure-9@11.[163,1153,142,1483]" captionTargetPageId="11" captionText="Figure 6. Titanogrypa (Cucullomyia) placida, male (MNRJ) [lost]. (a) Sternite 5, ventral view. (b) Syntergosternite 7 + 8, epandrium, cercus and surstylus, lateral view. (c) Phallus and associated structures, lateral view. (d) Distiphallus, ventral view. (e) Distiphallus, lateral view. Abbreviations: ar = arm; bp = basiphallus; bs = base; ce = cercus; ea = ejaculatory apodeme; ep = epandrium; ga = postgonal apodeme; hp = hypandrium; ju = juxta; ls = lateral stylus; ms = median stylus; ph = phallapodeme; po = postgonite; pr = pregonite; pp = paraphallus; sur = surstylus; stg 7 + 8 = syntergosternite 7 + 8; ve = vesica." figureDoi="http://doi.org/10.5281/zenodo.5473942" httpUri="https://zenodo.org/record/5473942/files/figure.png" pageId="27" pageNumber="332">Figure 6</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31C65533E652B5EF09FFD82B524FCCF" pageId="27" pageNumber="332" type="reference_group">
<paragraph id="8BB936D83E652B5EF09FFD82B524FCCF" blockId="27.[160,1156,593,861]" pageId="27" pageNumber="332">
<treatmentCitationGroup id="AB1611F63E652B5EF09FFD82B524FCCF" pageId="27" pageNumber="332">
<treatmentCitation id="0AA710C93E652B5EF09FFD82B541FDF8" author="Aldrich JM" box="[160,589,593,618]" page="24" pageId="27" pageNumber="332" year="1925">
<taxonomicName id="4C064D5B3E652B5EF09FFD82B541FDF8" ID-CoL="8945W" authority="Aldrich, 1925: 24" authorityName="Aldrich" authorityPageNumber="24" authorityYear="1925" box="[160,589,593,618]" class="Insecta" family="Sarcophagidae" genus="Sarcophaga" kingdom="Animalia" order="Diptera" pageId="27" pageNumber="332" phylum="Arthropoda" rank="species" species="placida">
<emphasis id="B972EACA3E652B5EF09FFD82B670FDF8" box="[160,380,593,618]" italics="true" pageId="27" pageNumber="332">Sarcophaga placida</emphasis>
<bibRefCitation id="EF974B293E652B5EF1B7FD82B541FDF8" author="Aldrich JM" box="[392,589,593,618]" pageId="27" pageNumber="332" pagination="1 - 36" refId="ref17892" refString="Aldrich JM. 1925. New Diptera or two-winged flies in the United States National Museum. Proc US Natl Mus. 66: 1 - 36. doi: 10.5479 / si. 00963801.66 - 2555.1." type="journal article" year="1925">Aldrich, 1925: 24</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
<typeStatus id="54BD887A3E652B5EF264FD82B59DFDF8" box="[603,657,593,618]" pageId="27" pageNumber="332">Type</typeStatus>
locality:
<collectingCountry id="F31176483E652B5EF2C2FD82B456FDF8" box="[765,858,593,618]" name="Panama" pageId="27" pageNumber="332">Panama</collectingCountry>
, Canal Zone, Ancon. Ref.:
<bibRefCitation id="EF974B293E652B5EF0FFFDA7B646FD1F" author="Dodge RH" box="[192,330,628,653]" pageId="27" pageNumber="332" pagination="421 - 450" refId="ref18607" refString="Dodge RH. 1968. The Sarcophagidae of Barro Colorado Island, Panama (Diptera). Ann Entomol Soc Amer. 61: 421 - 450. doi: 10.1093 / aesa / 61.2.421." type="journal article" year="1968">Dodge 1968</bibRefCitation>
(checklist).
<taxonomicName id="4C064D5B3E652B5EF1F9FDA7B5AEFD1F" ID-CoL="5782K" baseAuthorityName="Aldrich" baseAuthorityYear="1925" box="[454,674,628,653]" class="Insecta" family="Sarcophagidae" genus="Cucullomyia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="27" pageNumber="332" phylum="Arthropoda" rank="species" species="placida">
<emphasis id="B972EACA3E652B5EF1F9FDA7B5AEFD1F" box="[454,674,628,653]" italics="true" pageId="27" pageNumber="332">Cucullomyia placida</emphasis>
</taxonomicName>
;
<treatmentCitation id="0AA710C93E652B5EF290FDA7B45BFD1F" author="Lopes HS" box="[687,855,628,653]" page="47" pageId="27" pageNumber="332" year="1969">
<bibRefCitation id="EF974B293E652B5EF290FDA7B45BFD1F" author="Lopes HS" box="[687,855,628,653]" pageId="27" pageNumber="332" refId="ref19006" refString="Lopes HS. 1969. Family Sarcophagidae. In: Papavero N, editor. A catalogue of the Diptera of the Americas south of the United States. Sao Paulo: Departamento de Zoologia, Secretaria de Agricultura; p. 88." type="journal volume" year="1969">Lopes 1969: 47</bibRefCitation>
</treatmentCitation>
(catalogue, new combination);
<treatmentCitation id="0AA710C93E652B5EF132FD45B6D6FD3D" author="Lopes HS" box="[269,474,662,687]" page="79" pageId="27" pageNumber="332" year="1976">
<bibRefCitation id="EF974B293E652B5EF132FD45B6D6FD3D" author="Lopes HS" box="[269,474,662,687]" pageId="27" pageNumber="332" pagination="61 - 87" refId="ref19050" refString="Lopes HS. 1976 a. Some new or little known Neotropical Sarcophagidae (Diptera). Rev Bras Biol. 36: 61 - 87." type="journal article" year="1976">Lopes 1976a: 79</bibRefCitation>
</treatmentCitation>
(redescription);
<treatmentCitation id="0AA710C93E652B5EF295FD45B484FD3D" author="Lopes HS" box="[682,904,662,687]" page="748" pageId="27" pageNumber="332" year="1976">
<bibRefCitation id="EF974B293E652B5EF295FD45B484FD3D" author="Lopes HS" box="[682,904,662,687]" pageId="27" pageNumber="332" pagination="745 - 757" refId="ref19077" refString="Lopes HS. 1976 b. On the genus Cucullomyia Roback (Diptera, Sarcophagidae). Rev Bras Biol. 36: 745 - 757." type="journal article" year="1976">Lopes 1976b: 748</bibRefCitation>
</treatmentCitation>
(key, redescription);
<taxonomicName id="4C064D5B3E652B5EF0FFFD6AB548FD40" ID-CoL="8FGBB" baseAuthorityName="Aldrich" baseAuthorityYear="1925" box="[192,580,697,722]" class="Insecta" family="Sarcophagidae" genus="Titanogrypa" kingdom="Animalia" order="Diptera" pageId="27" pageNumber="332" phylum="Arthropoda" rank="species" species="placida" subGenus="Cucullomyia">
<emphasis id="B972EACA3E652B5EF0FFFD6AB64FFD40" box="[192,323,697,722]" italics="true" pageId="27" pageNumber="332">Titanogrypa</emphasis>
(
<emphasis id="B972EACA3E652B5EF166FD6AB6D3FD40" box="[345,479,697,722]" italics="true" pageId="27" pageNumber="332">Cucullomyia</emphasis>
)
<emphasis id="B972EACA3E652B5EF1CAFD6AB548FD40" box="[501,580,697,722]" italics="true" pageId="27" pageNumber="332">placida</emphasis>
</taxonomicName>
;
<treatmentCitation id="0AA710C93E652B5EF267FD6AB41DFD40" author="Pape T" box="[600,785,697,722]" page="432" pageId="27" pageNumber="332" year="1996">
<bibRefCitation id="EF974B293E652B5EF267FD6AB41DFD40" author="Pape T" box="[600,785,697,722]" pageId="27" pageNumber="332" pagination="1 - 558" refId="ref19466" refString="Pape T. 1996. Catalogue of the Sarcophagidae of the World (Insecta: Diptera). Mem Entomol Int. 8: 1 - 558." type="journal article" year="1996">Pape 1996: 432</bibRefCitation>
</treatmentCitation>
(catalogue, new combination);
<treatmentCitation id="0AA710C93E652B5EF0FFFD0FB6DFFD67" author="Reeves WK &amp; Pape T &amp; Adler PH" box="[192,467,732,757]" page="499" pageId="27" pageNumber="332" year="2000">
<bibRefCitation id="EF974B293E652B5EF0FFFD0FB6DFFD67" author="Reeves WK &amp; Pape T &amp; Adler PH" box="[192,467,732,757]" pageId="27" pageNumber="332" pagination="497 - 500" refId="ref19536" refString="Reeves WK, Pape T, Adler PH. 2000. Biological notes on New World Sarcophagidae (Diptera). Stud Dipterol. 2: 497 - 500." type="journal article" year="2000">Reeves et al. 2000: 499</bibRefCitation>
</treatmentCitation>
(note on biology);
<treatmentCitation id="0AA710C93E652B5EF282FD0FB4C0FD67" author="Giroux M &amp; Pape T &amp; Wheeler TA" box="[701,972,732,757]" page="773" pageId="27" pageNumber="332" year="2010">
<bibRefCitation id="EF974B293E652B5EF282FD0FB4C0FD67" author="Giroux M &amp; Pape T &amp; Wheeler TA" box="[701,972,732,757]" pageId="27" pageNumber="332" pagination="740 - 778" refId="ref18748" refString="Giroux M, Pape T, Wheeler TA. 2010. Towards a phylogeny of the flesh flies (Diptera: Sarcophagidae): morphology and phylogenetic implications of the acrophallus in the subfamily Sarcophaginae. Zool J Linnean Soc. 158: 740 - 778. doi: 10.1111 / j. 1096 - 3642.2009.00561. x." type="journal article" year="2010">Giroux et al. 2010: 773</bibRefCitation>
</treatmentCitation>
(morphological phylogeny);
<treatmentCitation id="0AA710C93E652B5EF16DFD2DB5D9FC85" author="Yepes-Gaurisas D &amp; Sanchez-Rodriguez JD &amp; Mello-Patiu CA &amp; Wolff M" box="[338,725,766,791]" page="1277" pageId="27" pageNumber="332" year="2013">
<bibRefCitation id="EF974B293E652B5EF16DFD2DB5D9FC85" author="Yepes-Gaurisas D &amp; Sanchez-Rodriguez JD &amp; Mello-Patiu CA &amp; Wolff M" box="[338,725,766,791]" pageId="27" pageNumber="332" pagination="1275 - 1287" refId="ref19902" refString="Yepes-Gaurisas D, Sanchez-Rodriguez JD, Mello-Patiu CA, Wolff M. 2013. Synanthropy of Sarcophagidae (Diptera) in La Pintada, Antioquia - Colombia. Int J Trop Biol. 61: 1275 - 1287." type="journal article" year="2013">Yepes-Gaurisas et al. 2013: 1277</bibRefCitation>
</treatmentCitation>
(synantropy);
<treatmentCitation id="0AA710C93E652B5EF3BCFD2DB38FFC85" author="Mello-Patiu CA" box="[899,1155,766,791]" page="895" pageId="27" pageNumber="332" year="2016">
<bibRefCitation id="EF974B293E652B5EF3BCFD2DB38FFC85" author="Mello-Patiu CA" box="[899,1155,766,791]" pageId="27" pageNumber="332" pagination="884 - 903" refId="ref19176" refString="Mello-Patiu CA. 2016. Family Sarcophagidae. Zootaxa. 4122: 884 - 903. doi: 10.11646 / zootaxa. 4122.1.75." type="journal article" year="2016">Mello-Patiu 2016: 895</bibRefCitation>
</treatmentCitation>
(catalogue);
<treatmentCitation id="0AA710C93E652B5EF174FCF2B598FCA8" author="Valverde-Castro C &amp; Buenaventura E &amp; Sanchez-Rodriguez JD &amp; Wolff M" box="[331,660,801,826]" page="4" pageId="27" pageNumber="332" year="2017">
<bibRefCitation id="EF974B293E652B5EF174FCF2B598FCA8" author="Valverde-Castro C &amp; Buenaventura E &amp; Sanchez-Rodriguez JD &amp; Wolff M" box="[331,660,801,826]" pageId="27" pageNumber="332" pagination="1 - 11" refId="ref19800" refString="Valverde-Castro C, Buenaventura E, Sanchez-Rodriguez JD, Wolff M. 2017. Flesh flies (Diptera: Sarcophagidae: Sarcophaginae) from the Colombian Guajira biogeographic province, an approach to their ecology and distribution. Zoologia. 34: 1 - 11. doi: 10.3897 / zoologia. 34. e 12277." type="journal article" year="2017">Valverde-Castro et al. 2017: 4</bibRefCitation>
</treatmentCitation>
(species list),
<treatmentCitation id="0AA710C93E652B5EF30EFCF2B7E5FCCF" author="Buenaventura E &amp; Pape T" page="819" pageId="27" pageNumber="332" year="2018">
<bibRefCitation id="EF974B293E652B5EF30EFCF2B7E5FCCF" author="Buenaventura E &amp; Pape T" pageId="27" pageNumber="332" pagination="808 - 906" refId="ref18282" refString="Buenaventura E, Pape T. 2018. Phylogeny, evolution and male terminalia functionality of Sarcophaginae (Diptera: Sarcophagidae). Zool J Linnean Soc. 183: 808 - 906. doi: 10.1093 / zoolinnean / zlx 070." type="journal article" year="2018">Buenaventura and Pape 2018: 819</bibRefCitation>
</treatmentCitation>
(morphological phylogeny).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C31C65533E652B5EF09FFC5BB5E2FBE0" pageId="27" pageNumber="332" type="materials_examined">
<paragraph id="8BB936D83E652B5EF09FFC5BB5E2FBE0" blockId="27.[160,1157,904,1138]" pageId="27" pageNumber="332">
<emphasis id="B972EACA3E652B5EF09FFC5BB500FC30" bold="true" box="[160,524,904,930]" italics="true" pageId="27" pageNumber="332">Examined material examined.</emphasis>
<materialsCitation id="3B6E3C853E652B5EF223FC5AB453FC30" box="[540,863,905,930]" country="Mexico" location="Oaxaca" pageId="27" pageNumber="332" specimenCount="1" specimenCount-male="1" stateProvince="Oaxaca">
<collectingCountry id="F31176483E652B5EF223FC5AB57AFC30" box="[540,630,905,930]" name="Mexico" pageId="27" pageNumber="332">MEXICO</collectingCountry>
<specimenCount id="9D00FD513E652B5EF2A3FC5AB5C6FC33" box="[668,714,905,930]" pageId="27" pageNumber="332" type="male">
1
<emphasis id="B972EACA3E652B5EF289FC59B5C6FC33" box="[694,714,906,929]" italics="true" pageId="27" pageNumber="332"></emphasis>
</specimenCount>
:
<collectingRegion id="49C2F83A3E652B5EF2E3FC5AB43CFC30" box="[732,816,905,930]" country="Mexico" name="Oaxaca" pageId="27" pageNumber="332">Oaxaca</collectingRegion>
,
<date id="FFB810183E652B5EF37FFC5AB453FC30" box="[832,863,905,930]" pageId="27" pageNumber="332" value="1951-06-30">30</date>
</materialsCitation>
<materialsCitation id="3B6E3C853E652B5EF360FC5AB479FC30" box="[863,885,905,930]" country="Mexico" isTailEnumeration="true" location="Oaxaca" pageId="27" pageNumber="332" specimenCount="1" stateProvince="Oaxaca">.vi</materialsCitation>
.1951, W
<materialsCitation id="3B6E3C853E652B5EF3E3FC5AB4FEFC30" box="[988,1010,905,930]" country="Mexico" isTailEnumeration="true" location="Oaxaca" pageId="27" pageNumber="332" specimenCount="1" stateProvince="Oaxaca">.G</materialsCitation>
. Downs leg
<materialsCitation id="3B6E3C853E652B5EF4BFFC5AB634FC7A" collectionCode="MNRJ" collectorName="W. G. Downs" country="Mexico" location="Vera Cruz" municipality="Xalapa" pageId="27" pageNumber="332" specimenCount="1" specimenCount-male="1">
. [donated and compared with
<typeStatus id="54BD887A3E652B5EF23EFC7FB53FFC57" box="[513,563,940,965]" pageId="27" pageNumber="332">type</typeStatus>
series by
<collectorName id="26F3530E3E652B5EF290FC7FB431FC57" box="[687,829,940,965]" pageId="27" pageNumber="332">W.G. Downs</collectorName>
] (
<collectionCode id="ED17AE1D3E652B5EF36CFC7FB49AFC57" box="[851,918,940,965]" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="27" pageNumber="332">MNRJ</collectionCode>
)
<emphasis id="B972EACA3E652B5EF398FC7FB4E8FC57" bold="true" box="[935,996,940,965]" pageId="27" pageNumber="332">[lost]</emphasis>
;
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1
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:
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,
<location id="8ED960033E652B5EF09FFC1CB603FC7A" LSID="urn:lsid:plazi:treatment:03AF87CE3E652B58F119FDCFB6BDFD16:8ED960033E652B5EF09FFC1CB603FC7A" box="[160,271,975,1000]" country="Mexico" municipality="Xalapa" name="Vera Cruz" pageId="27" pageNumber="332">Vera Cruz</location>
,
<date id="FFB810183E652B5EF125FC1CB634FC7A" box="[282,312,975,1000]" pageId="27" pageNumber="332" value="1950-09-25">25</date>
</materialsCitation>
<materialsCitation id="3B6E3C853E652B5EF107FC1CB641FC7A" box="[312,333,975,1000]" collectionCode="MNRJ" collectorName="W. G. Downs" country="Mexico" isTailEnumeration="true" location="Vera Cruz" municipality="Xalapa" pageId="27" pageNumber="332" specimenCount="1">.ix</materialsCitation>
.1950, W
<materialsCitation id="3B6E3C853E652B5EF18EFC1CB6CBFC7A" box="[433,455,975,1000]" collectionCode="MNRJ" collectorName="W. G. Downs" country="Mexico" isTailEnumeration="true" location="Vera Cruz" municipality="Xalapa" pageId="27" pageNumber="332" specimenCount="1">.G</materialsCitation>
. Downs leg
<materialsCitation id="3B6E3C853E652B5EF273FC1CB69DFB98" collectionCode="MNRJ" collectorName="W. G. Downs" country="Mexico" pageId="27" pageNumber="332" specimenCount="1">
. [donated and compared with
<typeStatus id="54BD887A3E652B5EF393FC1CB4D2FC7A" box="[940,990,975,1000]" pageId="27" pageNumber="332">type</typeStatus>
series by
<collectorName id="26F3530E3E652B5EF46FFC1CB7FCFB98" pageId="27" pageNumber="332">W.G. Downs</collectorName>
] (
<collectionCode id="ED17AE1D3E652B5EF13CFC22B64AFB98" box="[259,326,1009,1034]" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="27" pageNumber="332">MNRJ</collectionCode>
)
<emphasis id="B972EACA3E652B5EF16BFC22B69DFB98" bold="true" box="[340,401,1009,1034]" pageId="27" pageNumber="332">[lost]</emphasis>
</materialsCitation>
;
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<collectingCountry id="F31176483E652B5EF1A0FC22B525FB98" box="[415,553,1009,1034]" name="Costa Rica" pageId="27" pageNumber="332">COSTA RICA</collectingCountry>
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1
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:
<collectingRegion id="49C2F83A3E652B5EF241FC22B49BFB98" box="[638,919,1009,1034]" country="Costa Rica" name="Guanacaste" pageId="27" pageNumber="332">Província de Guanacaste</collectingRegion>
,
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,
<date id="FFB810183E652B5EF0DCFBC7B7E0FBBF" box="[227,236,1044,1069]" pageId="27" pageNumber="332" value="1983-02">ii</date>
</materialsCitation>
.1983, D
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. Janzen &amp; W. Hall Wacks leg
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. (
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)
<emphasis id="B972EACA3E652B5EF309FBC7B47FFBBF" bold="true" box="[822,883,1044,1069]" pageId="27" pageNumber="332">[lost]</emphasis>
;
<specimenCount id="9D00FD513E652B5EF3BAFBC7B4BFFBB9" box="[901,947,1044,1069]" pageId="27" pageNumber="332" type="male">
1
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:
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<collectingRegion id="49C2F83A3E652B5EF09FFBE4B629FBC2" box="[160,293,1079,1104]" country="Costa Rica" name="Guanacaste" pageId="27" pageNumber="332">Guanacaste</collectingRegion>
,
<date id="FFB810183E652B5EF104FBE4B68DFBC2" box="[315,385,1079,1104]" pageId="27" pageNumber="332" value="2010-08-14" valueMax="2010-08-18" valueMin="2010-08-14">1418</date>
</materialsCitation>
<materialsCitation id="3B6E3C853E652B5EF1BEFBE4B6BBFBC2" box="[385,439,1079,1104]" collectionCode="MNRJ" country="Costa Rica" isTailEnumeration="true" location="Parque Nacional" pageId="27" pageNumber="332" specimenCount="1" stateProvince="Guanacaste">. viii</materialsCitation>
.2010, S
<materialsCitation id="3B6E3C853E652B5EF223FBE4B5A8FBC2" box="[540,676,1079,1104]" collectionCode="MNRJ" collectorName="Nihei" country="Costa Rica" isTailEnumeration="true" location="Parque Nacional" pageId="27" pageNumber="332" specimenCount="1" stateProvince="Guanacaste">
.
<collectorName id="26F3530E3E652B5EF20DFBE4B560FBC2" box="[562,620,1079,1104]" pageId="27" pageNumber="332">Nihei</collectorName>
leg
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<materialsCitation id="3B6E3C853E652B5EF29BFBE4B647FBE0" collectionCode="MNRJ" country="Costa Rica" location="Parque Nacional" pageId="27" pageNumber="332" specimenCount="1" specimenCount-male="1" stateProvince="Guanacaste">
. (
<collectionCode id="ED17AE1D3E652B5EF2FFFBE4B40FFBC2" box="[704,771,1079,1104]" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="27" pageNumber="332">MNRJ</collectionCode>
)
<emphasis id="B972EACA3E652B5EF322FBE4B456FBC2" bold="true" box="[797,858,1079,1104]" pageId="27" pageNumber="332">[lost]</emphasis>
;
<specimenCount id="9D00FD513E652B5EF34DFBE4B4ABFBDC" box="[882,935,1079,1104]" pageId="27" pageNumber="332" type="male">
1
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</specimenCount>
:
<location id="8ED960033E652B5EF3FFFBE4B38FFBC2" LSID="urn:lsid:plazi:treatment:03AF87CE3E652B58F119FDCFB6BDFD16:8ED960033E652B5EF3FFFBE4B38FFBC2" box="[960,1155,1079,1104]" country="Costa Rica" name="Parque Nacional" pageId="27" pageNumber="332" stateProvince="Guanacaste">Parque Nacional</location>
<collectingRegion id="49C2F83A3E652B5EF09FFB8AB629FBE0" box="[160,293,1113,1138]" country="Costa Rica" name="Guanacaste" pageId="27" pageNumber="332">Guanacaste</collectingRegion>
,
<date id="FFB810183E652B5EF110FB8AB647FBE0" box="[303,331,1113,1138]" pageId="27" pageNumber="332" value="2010-08-15">15</date>
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<materialsCitation id="3B6E3C853E652B5EF174FB8AB661FBE0" box="[331,365,1113,1138]" collectionCode="MNRJ" country="Costa Rica" isTailEnumeration="true" location="Parque Nacional" pageId="27" pageNumber="332" specimenCount="1" stateProvince="Guanacaste">.viii</materialsCitation>
.2010, J
<materialsCitation id="3B6E3C853E652B5EF1FAFB8AB547FBE0" box="[453,587,1113,1138]" collectionCode="MNRJ" collectorName="Gibson" country="Costa Rica" isTailEnumeration="true" location="Parque Nacional" pageId="27" pageNumber="332" specimenCount="1" stateProvince="Guanacaste">
.
<collectorName id="26F3530E3E652B5EF1EFFB8AB513FBE0" box="[464,543,1113,1138]" pageId="27" pageNumber="332">Gibson</collectorName>
leg
</materialsCitation>
. (
<collectionCode id="ED17AE1D3E652B5EF264FB8AB592FBE0" box="[603,670,1113,1138]" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="27" pageNumber="332">MNRJ</collectionCode>
)
<emphasis id="B972EACA3E652B5EF294FB8AB5E4FBE0" bold="true" box="[683,744,1113,1138]" pageId="27" pageNumber="332">[lost]</emphasis>
.
</paragraph>
</subSubSection>
<subSubSection id="C31C65533E652B5EF09FFB4DB64FFAF7" pageId="27" pageNumber="332" type="diagnosis">
<paragraph id="8BB936D83E652B5EF09FFB4DB64FFAF7" blockId="27.[160,1156,1182,1381]" pageId="27" pageNumber="332">
<emphasis id="B972EACA3E652B5EF09FFB4DB616FB2A" bold="true" box="[160,282,1182,1208]" italics="true" pageId="27" pageNumber="332">Diagnosis.</emphasis>
Scutellum with a lateral patch of whitish hair-like setulae; vesica longer than wide, without projection at the base, with a narrow and more sclerotised sheath along the distal margin, facing outwards in ventral view (
<figureCitation id="133D2A5D3E652B5EF299FB37B40CFB6F" box="[678,768,1252,1277]" captionStart="Figure 6" captionStartId="11.[160,225,1515,1538]" captionTargetBox="[194,1152,150,1482]" captionTargetId="figure-9@11.[163,1153,142,1483]" captionTargetPageId="11" captionText="Figure 6. Titanogrypa (Cucullomyia) placida, male (MNRJ) [lost]. (a) Sternite 5, ventral view. (b) Syntergosternite 7 + 8, epandrium, cercus and surstylus, lateral view. (c) Phallus and associated structures, lateral view. (d) Distiphallus, ventral view. (e) Distiphallus, lateral view. Abbreviations: ar = arm; bp = basiphallus; bs = base; ce = cercus; ea = ejaculatory apodeme; ep = epandrium; ga = postgonal apodeme; hp = hypandrium; ju = juxta; ls = lateral stylus; ms = median stylus; ph = phallapodeme; po = postgonite; pr = pregonite; pp = paraphallus; sur = surstylus; stg 7 + 8 = syntergosternite 7 + 8; ve = vesica." figureDoi="http://doi.org/10.5281/zenodo.5473942" httpUri="https://zenodo.org/record/5473942/files/figure.png" pageId="27" pageNumber="332">Figure 6</figureCitation>
(e)); juxta composed of broad lobe, as long as paraphallus, and with internal sclerotised band (
<figureCitation id="133D2A5D3E652B5EF371FAD4B4A1FAB2" box="[846,941,1287,1312]" captionStart="Figure 6" captionStartId="11.[160,225,1515,1538]" captionTargetBox="[194,1152,150,1482]" captionTargetId="figure-9@11.[163,1153,142,1483]" captionTargetPageId="11" captionText="Figure 6. Titanogrypa (Cucullomyia) placida, male (MNRJ) [lost]. (a) Sternite 5, ventral view. (b) Syntergosternite 7 + 8, epandrium, cercus and surstylus, lateral view. (c) Phallus and associated structures, lateral view. (d) Distiphallus, ventral view. (e) Distiphallus, lateral view. Abbreviations: ar = arm; bp = basiphallus; bs = base; ce = cercus; ea = ejaculatory apodeme; ep = epandrium; ga = postgonal apodeme; hp = hypandrium; ju = juxta; ls = lateral stylus; ms = median stylus; ph = phallapodeme; po = postgonite; pr = pregonite; pp = paraphallus; sur = surstylus; stg 7 + 8 = syntergosternite 7 + 8; ve = vesica." figureDoi="http://doi.org/10.5281/zenodo.5473942" httpUri="https://zenodo.org/record/5473942/files/figure.png" pageId="27" pageNumber="332">Figure 6</figureCitation>
(ce)); median and lateral styli very long, surpassing the juxta in lateral view, both with spines at the apex (
<figureCitation id="133D2A5D3E652B5EF097FA9FB609FAF7" box="[168,261,1356,1381]" captionStart="Figure 6" captionStartId="11.[160,225,1515,1538]" captionTargetBox="[194,1152,150,1482]" captionTargetId="figure-9@11.[163,1153,142,1483]" captionTargetPageId="11" captionText="Figure 6. Titanogrypa (Cucullomyia) placida, male (MNRJ) [lost]. (a) Sternite 5, ventral view. (b) Syntergosternite 7 + 8, epandrium, cercus and surstylus, lateral view. (c) Phallus and associated structures, lateral view. (d) Distiphallus, ventral view. (e) Distiphallus, lateral view. Abbreviations: ar = arm; bp = basiphallus; bs = base; ce = cercus; ea = ejaculatory apodeme; ep = epandrium; ga = postgonal apodeme; hp = hypandrium; ju = juxta; ls = lateral stylus; ms = median stylus; ph = phallapodeme; po = postgonite; pr = pregonite; pp = paraphallus; sur = surstylus; stg 7 + 8 = syntergosternite 7 + 8; ve = vesica." figureDoi="http://doi.org/10.5281/zenodo.5473942" httpUri="https://zenodo.org/record/5473942/files/figure.png" pageId="27" pageNumber="332">Figure 6</figureCitation>
(d,e)).
</paragraph>
</subSubSection>
<subSubSection id="C31C65533E652B58F09FFA42B409FEA9" lastPageId="29" lastPageNumber="334" pageId="27" pageNumber="332" type="description">
<paragraph id="8BB936D83E652B5EF09FFA42B422FA39" blockId="27.[160,814,1425,1451]" box="[160,814,1425,1451]" pageId="27" pageNumber="332">
<emphasis id="B972EACA3E652B5EF09FFA42B64AFA39" bold="true" box="[160,326,1425,1451]" italics="true" pageId="27" pageNumber="332">Redescription.</emphasis>
MALE (n = 5). Body length = 11.012.0 mm.
</paragraph>
<paragraph id="8BB936D83E652B59F09FFA05B5DDFE08" blockId="27.[160,1156,1494,1728]" lastBlockId="28.[160,1156,142,410]" lastPageId="28" lastPageNumber="333" pageId="27" pageNumber="332">
<emphasis id="B972EACA3E652B5EF09FFA05B7E9FA62" bold="true" box="[160,229,1494,1520]" italics="true" pageId="27" pageNumber="332">Head.</emphasis>
Parafacial and fronto-orbital plates with golden pruinosity; fronto-orbital plate with sparse setulae; parafacial plate with row of setulae close to eye and measuring about 0.48 of distance between the vibrissae; frons about 0.33 head width at level of ocellar triangle; frontal vitta entirely blackish; rows of frontal setae parallel except anteriormost 2 slightly divergent, 78 well-developed frontal setae reaching level of base of pedicel, the uppermost one shorter than the others; reclinate orbital seta present, proclinate orbital setae absent; outer vertical setae 0.5x inner vertical setae and divergent; ocellar setae as developed as upper frontals; ocellar triangle dark brown; postocellar and paravertical setae present; postocular area with golden pruinosity; gena with golden pruinosity at the top and silver at the bottom, black setae in anterior part and whitish setae close to occiput; postgena with silvery pruinosity and numerous whitish setae; face with silvery pruinosity; facial ridge black with silvery pruinosity, with short setulae in lower half; antenna dark brown, total length 0.77 of distance from insertion to vibrissal level, first flagellomere brown with grey pruinosity and approximately 2.0x longer than pedicel; arista long plumose on basal 2/3; palpus blackish.
</paragraph>
<paragraph id="8BB936D83E622B59F09FFE15B47AFBC2" blockId="28.[160,1156,454,1104]" pageId="28" pageNumber="333">
<emphasis id="B972EACA3E622B59F09FFE15B7F6FE72" bold="true" box="[160,250,454,480]" italics="true" pageId="28" pageNumber="333">Thorax.</emphasis>
Black with silvery-grey pruinosity, postpronotum and notopleuron with slightly yellowish grey pruinosity; chaetotaxy: acrostichals 1 2 + 1, dorsocentrals 4 5 poorly differentiated + 4 (posteriormost 2 well developed), intra-alars 2 + 2, supra-alars 2 + 3, postpronotals 3, notopleurals 4; katepisternals 3, almost in a straight line; postalar wall setulose; postalar callus with 2 setae; scutellum with a patch of whitish hair-like setulae on the lateral margin and reaching ventral part, pairs of basal, lateral and subapical setae (lateral pair shorter), a pair of preapical discal setae, and no apical setae; meral setae 78; proepisternum bare; prosternum setulose.
<emphasis id="B972EACA3E622B59F2B9FD6AB5B2FD40" box="[646,702,697,722]" italics="true" pageId="28" pageNumber="333">Wing</emphasis>
. Hyaline, with dark brown veins; tegula dark brown; basicosta yellowish; vein R1 bare; vein R4+5 with setulae dorsally on 1/2 of distance to crossvein r-m; cell r4+5 open at wing margin; third costal sector bare ventrally.
<emphasis id="B972EACA3E622B59F09FFCF2B7DCFCA8" box="[160,208,801,826]" italics="true" pageId="28" pageNumber="333">Legs</emphasis>
. Blackish-brown with silvery pruinosity, pulvilli yellowish-brown; mid femur with a row of 3 4 median anterior setae, a row of anteroventral setae, 2 preapical posterior setae, 2 median setae and a ctenidium of 10 spiniform setae in posteroventral margin; mid tibia with 1 median anterodorsal, 1 basal and 1 median posterior setae, and 1 median posterodorsal seta; hind femur with a row of anterodorsal setae, a row of anterior setae in the proximal half, 1 apical dorsal and 1 apical posterodorsal seta, and rows of anteroventral and posteroventral setae; hind tibia with 1 median anterior seta, 1 basal, 1 median and 1 apical seta in the same position on the anterodorsal and posterodorsal sufarces, and 1 strong ventral seta; hind coxa and trochanter with normal setae.
</paragraph>
<paragraph id="8BB936D83E622B59F09FFBA8B318FAD0" blockId="28.[160,1155,1147,1346]" pageId="28" pageNumber="333">
<emphasis id="B972EACA3E622B59F09FFBA8B615FB07" bold="true" box="[160,281,1147,1173]" italics="true" pageId="28" pageNumber="333">Abdomen.</emphasis>
Dark brown with yellowish grey pruinosity on T1 + 2 T3; T4 T5 reddish brown with yellowish pruinosity, more intense on T5; T1 + 2 T4 with lateral marginal setae; T3 T5 with a set of long setae with wavy apex on ventral surface, more numerous on T5; T4 with a pair of median marginal setae; T5 with a complete row of marginal setae (
<emphasis id="B972EACA3E622B59F3C6FB37B31EFB6F" box="[1017,1042,1252,1277]" italics="true" pageId="28" pageNumber="333">ca</emphasis>
. 12 14); ST2 4 rectangular with long setae in distal half; ST5 deeply cleft with short base, long and narrow arms, longer setae at base and spiniform setae in distal half (
<figureCitation id="133D2A5D3E622B59F3ADFAFAB4E7FAD0" box="[914,1003,1321,1346]" captionStart="Figure 6" captionStartId="11.[160,225,1515,1538]" captionTargetBox="[194,1152,150,1482]" captionTargetId="figure-9@11.[163,1153,142,1483]" captionTargetPageId="11" captionText="Figure 6. Titanogrypa (Cucullomyia) placida, male (MNRJ) [lost]. (a) Sternite 5, ventral view. (b) Syntergosternite 7 + 8, epandrium, cercus and surstylus, lateral view. (c) Phallus and associated structures, lateral view. (d) Distiphallus, ventral view. (e) Distiphallus, lateral view. Abbreviations: ar = arm; bp = basiphallus; bs = base; ce = cercus; ea = ejaculatory apodeme; ep = epandrium; ga = postgonal apodeme; hp = hypandrium; ju = juxta; ls = lateral stylus; ms = median stylus; ph = phallapodeme; po = postgonite; pr = pregonite; pp = paraphallus; sur = surstylus; stg 7 + 8 = syntergosternite 7 + 8; ve = vesica." figureDoi="http://doi.org/10.5281/zenodo.5473942" httpUri="https://zenodo.org/record/5473942/files/figure.png" pageId="28" pageNumber="333">Figure 6</figureCitation>
(a)).
</paragraph>
<paragraph id="8BB936D83E622B58F09FFABDB55CFF7F" blockId="28.[160,1156,1390,1728]" lastBlockId="29.[160,1155,142,237]" lastPageId="29" lastPageNumber="334" pageId="28" pageNumber="333">
<emphasis id="B972EACA3E622B59F09FFABDB62AFA1A" bold="true" box="[160,294,1390,1416]" italics="true" pageId="28" pageNumber="333">
<taxonomicName id="4C064D5B3E622B59F09FFABDB62DFA1A" box="[160,289,1390,1416]" class="Magnoliopsida" family="Combretaceae" genus="Terminalia" kingdom="Plantae" order="Myrtales" pageId="28" pageNumber="333" phylum="Tracheophyta" rank="genus">Terminalia</taxonomicName>
.
</emphasis>
Reddish brown; syntergosternite 7 + 8 with yellowish pruinosity and sparse setulae; epandrium with yellowish pruinosity, a pair of developed dorsal seta, and long and sparse setulae; surstylus short, clavate but with a short projection in the posterior margin and some apical setae (
<figureCitation id="133D2A5D3E622B59F1C4FA04B55BFA62" box="[507,599,1495,1520]" captionStart="Figure 6" captionStartId="11.[160,225,1515,1538]" captionTargetBox="[194,1152,150,1482]" captionTargetId="figure-9@11.[163,1153,142,1483]" captionTargetPageId="11" captionText="Figure 6. Titanogrypa (Cucullomyia) placida, male (MNRJ) [lost]. (a) Sternite 5, ventral view. (b) Syntergosternite 7 + 8, epandrium, cercus and surstylus, lateral view. (c) Phallus and associated structures, lateral view. (d) Distiphallus, ventral view. (e) Distiphallus, lateral view. Abbreviations: ar = arm; bp = basiphallus; bs = base; ce = cercus; ea = ejaculatory apodeme; ep = epandrium; ga = postgonal apodeme; hp = hypandrium; ju = juxta; ls = lateral stylus; ms = median stylus; ph = phallapodeme; po = postgonite; pr = pregonite; pp = paraphallus; sur = surstylus; stg 7 + 8 = syntergosternite 7 + 8; ve = vesica." figureDoi="http://doi.org/10.5281/zenodo.5473942" httpUri="https://zenodo.org/record/5473942/files/figure.png" pageId="28" pageNumber="333">Figure 6</figureCitation>
(b)); cercal prongs narrow and parallel in posterior view, with distal end slightly enlarged and rounded in profile (
<figureCitation id="133D2A5D3E622B59F36EFA29B4A1F981" box="[849,941,1530,1555]" captionStart="Figure 6" captionStartId="11.[160,225,1515,1538]" captionTargetBox="[194,1152,150,1482]" captionTargetId="figure-9@11.[163,1153,142,1483]" captionTargetPageId="11" captionText="Figure 6. Titanogrypa (Cucullomyia) placida, male (MNRJ) [lost]. (a) Sternite 5, ventral view. (b) Syntergosternite 7 + 8, epandrium, cercus and surstylus, lateral view. (c) Phallus and associated structures, lateral view. (d) Distiphallus, ventral view. (e) Distiphallus, lateral view. Abbreviations: ar = arm; bp = basiphallus; bs = base; ce = cercus; ea = ejaculatory apodeme; ep = epandrium; ga = postgonal apodeme; hp = hypandrium; ju = juxta; ls = lateral stylus; ms = median stylus; ph = phallapodeme; po = postgonite; pr = pregonite; pp = paraphallus; sur = surstylus; stg 7 + 8 = syntergosternite 7 + 8; ve = vesica." figureDoi="http://doi.org/10.5281/zenodo.5473942" httpUri="https://zenodo.org/record/5473942/files/figure.png" pageId="28" pageNumber="333">Figure 6</figureCitation>
(b)); pregonite long with spatulate apex and a median depression on anterior margin (
<figureCitation id="133D2A5D3E622B59F3BFF9CEB4D7F9A4" box="[896,987,1565,1590]" captionStart="Figure 6" captionStartId="11.[160,225,1515,1538]" captionTargetBox="[194,1152,150,1482]" captionTargetId="figure-9@11.[163,1153,142,1483]" captionTargetPageId="11" captionText="Figure 6. Titanogrypa (Cucullomyia) placida, male (MNRJ) [lost]. (a) Sternite 5, ventral view. (b) Syntergosternite 7 + 8, epandrium, cercus and surstylus, lateral view. (c) Phallus and associated structures, lateral view. (d) Distiphallus, ventral view. (e) Distiphallus, lateral view. Abbreviations: ar = arm; bp = basiphallus; bs = base; ce = cercus; ea = ejaculatory apodeme; ep = epandrium; ga = postgonal apodeme; hp = hypandrium; ju = juxta; ls = lateral stylus; ms = median stylus; ph = phallapodeme; po = postgonite; pr = pregonite; pp = paraphallus; sur = surstylus; stg 7 + 8 = syntergosternite 7 + 8; ve = vesica." figureDoi="http://doi.org/10.5281/zenodo.5473942" httpUri="https://zenodo.org/record/5473942/files/figure.png" pageId="28" pageNumber="333">Figure 6</figureCitation>
(c)); postgonite about 0.5x the pregonite, with a long median setae and small setulae (
<figureCitation id="133D2A5D3E622B59F3C5F9ECB357F9CA" box="[1018,1115,1599,1624]" captionStart="Figure 6" captionStartId="11.[160,225,1515,1538]" captionTargetBox="[194,1152,150,1482]" captionTargetId="figure-9@11.[163,1153,142,1483]" captionTargetPageId="11" captionText="Figure 6. Titanogrypa (Cucullomyia) placida, male (MNRJ) [lost]. (a) Sternite 5, ventral view. (b) Syntergosternite 7 + 8, epandrium, cercus and surstylus, lateral view. (c) Phallus and associated structures, lateral view. (d) Distiphallus, ventral view. (e) Distiphallus, lateral view. Abbreviations: ar = arm; bp = basiphallus; bs = base; ce = cercus; ea = ejaculatory apodeme; ep = epandrium; ga = postgonal apodeme; hp = hypandrium; ju = juxta; ls = lateral stylus; ms = median stylus; ph = phallapodeme; po = postgonite; pr = pregonite; pp = paraphallus; sur = surstylus; stg 7 + 8 = syntergosternite 7 + 8; ve = vesica." figureDoi="http://doi.org/10.5281/zenodo.5473942" httpUri="https://zenodo.org/record/5473942/files/figure.png" pageId="28" pageNumber="333">Figure 6</figureCitation>
(c)); basiphallus about 3x paraphallus length, paraphallus with 2 dorsal keels (
<figureCitation id="133D2A5D3E622B59F3C3F9B1B357F9E9" box="[1020,1115,1634,1659]" captionStart="Figure 6" captionStartId="11.[160,225,1515,1538]" captionTargetBox="[194,1152,150,1482]" captionTargetId="figure-9@11.[163,1153,142,1483]" captionTargetPageId="11" captionText="Figure 6. Titanogrypa (Cucullomyia) placida, male (MNRJ) [lost]. (a) Sternite 5, ventral view. (b) Syntergosternite 7 + 8, epandrium, cercus and surstylus, lateral view. (c) Phallus and associated structures, lateral view. (d) Distiphallus, ventral view. (e) Distiphallus, lateral view. Abbreviations: ar = arm; bp = basiphallus; bs = base; ce = cercus; ea = ejaculatory apodeme; ep = epandrium; ga = postgonal apodeme; hp = hypandrium; ju = juxta; ls = lateral stylus; ms = median stylus; ph = phallapodeme; po = postgonite; pr = pregonite; pp = paraphallus; sur = surstylus; stg 7 + 8 = syntergosternite 7 + 8; ve = vesica." figureDoi="http://doi.org/10.5281/zenodo.5473942" httpUri="https://zenodo.org/record/5473942/files/figure.png" pageId="28" pageNumber="333">Figure 6</figureCitation>
(c)); vesica longer than wide, without projection at the base, with a narrow and more sclerotised sheath along the distal margin, facing outwards in ventral view (
<figureCitation id="133D2A5D3E622B59F3C2F974B356F952" box="[1021,1114,1703,1728]" captionStart="Figure 6" captionStartId="11.[160,225,1515,1538]" captionTargetBox="[194,1152,150,1482]" captionTargetId="figure-9@11.[163,1153,142,1483]" captionTargetPageId="11" captionText="Figure 6. Titanogrypa (Cucullomyia) placida, male (MNRJ) [lost]. (a) Sternite 5, ventral view. (b) Syntergosternite 7 + 8, epandrium, cercus and surstylus, lateral view. (c) Phallus and associated structures, lateral view. (d) Distiphallus, ventral view. (e) Distiphallus, lateral view. Abbreviations: ar = arm; bp = basiphallus; bs = base; ce = cercus; ea = ejaculatory apodeme; ep = epandrium; ga = postgonal apodeme; hp = hypandrium; ju = juxta; ls = lateral stylus; ms = median stylus; ph = phallapodeme; po = postgonite; pr = pregonite; pp = paraphallus; sur = surstylus; stg 7 + 8 = syntergosternite 7 + 8; ve = vesica." figureDoi="http://doi.org/10.5281/zenodo.5473942" httpUri="https://zenodo.org/record/5473942/files/figure.png" pageId="28" pageNumber="333">Figure 6</figureCitation>
(e)); juxta composed of broad lobe, as long as paraphallus, and with internal sclerotised band (
<figureCitation id="133D2A5D3E632B58F097FF62B608FF58" box="[168,260,177,202]" captionStart="Figure 6" captionStartId="11.[160,225,1515,1538]" captionTargetBox="[194,1152,150,1482]" captionTargetId="figure-9@11.[163,1153,142,1483]" captionTargetPageId="11" captionText="Figure 6. Titanogrypa (Cucullomyia) placida, male (MNRJ) [lost]. (a) Sternite 5, ventral view. (b) Syntergosternite 7 + 8, epandrium, cercus and surstylus, lateral view. (c) Phallus and associated structures, lateral view. (d) Distiphallus, ventral view. (e) Distiphallus, lateral view. Abbreviations: ar = arm; bp = basiphallus; bs = base; ce = cercus; ea = ejaculatory apodeme; ep = epandrium; ga = postgonal apodeme; hp = hypandrium; ju = juxta; ls = lateral stylus; ms = median stylus; ph = phallapodeme; po = postgonite; pr = pregonite; pp = paraphallus; sur = surstylus; stg 7 + 8 = syntergosternite 7 + 8; ve = vesica." figureDoi="http://doi.org/10.5281/zenodo.5473942" httpUri="https://zenodo.org/record/5473942/files/figure.png" pageId="29" pageNumber="334">Figure 6</figureCitation>
(c,d)); median and lateral styli very long, surpassing the juxta in lateral view, both with spines at the apex (
<figureCitation id="133D2A5D3E632B58F189FF07B51FFF7F" box="[438,531,212,237]" captionStart="Figure 6" captionStartId="11.[160,225,1515,1538]" captionTargetBox="[194,1152,150,1482]" captionTargetId="figure-9@11.[163,1153,142,1483]" captionTargetPageId="11" captionText="Figure 6. Titanogrypa (Cucullomyia) placida, male (MNRJ) [lost]. (a) Sternite 5, ventral view. (b) Syntergosternite 7 + 8, epandrium, cercus and surstylus, lateral view. (c) Phallus and associated structures, lateral view. (d) Distiphallus, ventral view. (e) Distiphallus, lateral view. Abbreviations: ar = arm; bp = basiphallus; bs = base; ce = cercus; ea = ejaculatory apodeme; ep = epandrium; ga = postgonal apodeme; hp = hypandrium; ju = juxta; ls = lateral stylus; ms = median stylus; ph = phallapodeme; po = postgonite; pr = pregonite; pp = paraphallus; sur = surstylus; stg 7 + 8 = syntergosternite 7 + 8; ve = vesica." figureDoi="http://doi.org/10.5281/zenodo.5473942" httpUri="https://zenodo.org/record/5473942/files/figure.png" pageId="29" pageNumber="334">Figure 6</figureCitation>
(d,e)).
</paragraph>
<paragraph id="8BB936D83E632B58F09FFEF2B409FEA9" blockId="29.[160,773,289,315]" box="[160,773,289,315]" pageId="29" pageNumber="334">
<emphasis id="B972EACA3E632B58F09FFEF2B7F1FEA9" bold="true" box="[160,253,289,315]" italics="true" pageId="29" pageNumber="334">Female.</emphasis>
Redescribed and illustrated in
<bibRefCitation id="EF974B293E632B58F25CFEF1B40DFEA9" author="Lopes HS" box="[611,769,290,315]" pageId="29" pageNumber="334" pagination="61 - 87" refId="ref19050" refString="Lopes HS. 1976 a. Some new or little known Neotropical Sarcophagidae (Diptera). Rev Bras Biol. 36: 61 - 87." type="journal article" year="1976">Lopes (1976a)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C31C65533E632B58F09FFEBCB514FE39" pageId="29" pageNumber="334" type="distribution">
<paragraph id="8BB936D83E632B58F09FFEBCB514FE39" blockId="29.[160,1155,367,427]" pageId="29" pageNumber="334">
<emphasis id="B972EACA3E632B58F09FFEBCB586FE1B" bold="true" box="[160,650,367,393]" pageId="29" pageNumber="334">
Distribution
<emphasis id="B972EACA3E632B58F100FEBCB586FE1B" bold="true" box="[319,650,367,393]" italics="true" pageId="29" pageNumber="334">(Nearctic and Neotropical).</emphasis>
</emphasis>
<collectingCountry id="F31176483E632B58F2A6FEA3B5E5FE1B" box="[665,745,368,393]" name="Mexico" pageId="29" pageNumber="334">Mexico</collectingCountry>
(
<collectingRegion id="49C2F83A3E632B58F2C3FEA3B45DFE1B" box="[764,849,368,393]" country="Mexico" name="Sonora" pageId="29" pageNumber="334">Sonora</collectingRegion>
),
<collectingCountry id="F31176483E632B58F358FEA3B4D9FE1B" box="[871,981,368,393]" name="Colombia" pageId="29" pageNumber="334">Colômbia</collectingCountry>
,
<collectingCountry id="F31176483E632B58F3DAFEA3B36CFE1B" box="[997,1120,368,393]" name="Costa Rica" pageId="29" pageNumber="334">Costa Rica</collectingCountry>
,
<collectingCountry id="F31176483E632B58F44FFEA3B60FFE39" name="El Salvador" pageId="29" pageNumber="334">El Salvador</collectingCountry>
,
<collectingCountry id="F31176483E632B58F132FE41B672FE39" box="[269,382,402,427]" name="Nicaragua" pageId="29" pageNumber="334">Nicarágua</collectingCountry>
and
<collectingCountry id="F31176483E632B58F187FE41B519FE39" box="[440,533,402,427]" name="Panama" pageId="29" pageNumber="334">Panamá</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C31C65533E632B58F09FFE33B6BDFD16" pageId="29" pageNumber="334" type="discussion">
<paragraph id="8BB936D83E632B58F09FFE33B6BDFD16" blockId="29.[160,1156,480,644]" pageId="29" pageNumber="334">
<emphasis id="B972EACA3E632B58F09FFE33B629FE68" bold="true" box="[160,293,480,506]" italics="true" pageId="29" pageNumber="334">Comments.</emphasis>
<taxonomicName id="4C064D5B3E632B58F10AFE32B6B4FE68" authorityName=", Pape" authorityYear="1996" box="[309,440,481,506]" class="Insecta" family="Sarcophagidae" genus="Titanogrypa" kingdom="Animalia" order="Diptera" pageId="29" pageNumber="334" phylum="Arthropoda" rank="genus">
<emphasis id="B972EACA3E632B58F10AFE32B6B4FE68" box="[309,440,481,506]" italics="true" pageId="29" pageNumber="334">Titanogrypa</emphasis>
</taxonomicName>
<taxonomicName id="4C064D5B3E632B58F1FEFE32B530FE68" baseAuthorityName="Aldrich" baseAuthorityYear="1925" box="[449,572,481,506]" class="Insecta" family="Sarcophagidae" genus="Titanogrypa" kingdom="Animalia" order="Diptera" pageId="29" pageNumber="334" phylum="Arthropoda" rank="species" species="placida" subGenus="Cucullomyia">
(
<emphasis id="B972EACA3E632B58F1F7FE32B6DBFE68" box="[456,471,481,506]" italics="true" pageId="29" pageNumber="334">C</emphasis>
.)
<emphasis id="B972EACA3E632B58F1D2FE32B530FE68" box="[493,572,481,506]" italics="true" pageId="29" pageNumber="334">placida</emphasis>
</taxonomicName>
closely resembles
<emphasis id="B972EACA3E632B58F32DFE32B413FE68" box="[786,799,481,506]" italics="true" pageId="29" pageNumber="334">T</emphasis>
.
<taxonomicName id="4C064D5B3E632B58F312FE32B4E4FE68" baseAuthorityName="Lopes" baseAuthorityYear="1976" box="[813,1000,481,506]" class="Insecta" family="Sarcophagidae" genus="Titanogrypa" kingdom="Animalia" order="Diptera" pageId="29" pageNumber="334" phylum="Arthropoda" rank="species" species="albuquerquei" subGenus="Cucullomyia">
(
<emphasis id="B972EACA3E632B58F30BFE32B44FFE68" box="[820,835,481,506]" italics="true" pageId="29" pageNumber="334">C</emphasis>
.)
<emphasis id="B972EACA3E632B58F366FE32B4E4FE68" box="[857,1000,481,506]" italics="true" pageId="29" pageNumber="334">albuquerquei</emphasis>
</taxonomicName>
as both have vesica long and ST 5 with narrow arms (
<figureCitation id="133D2A5D3E632B58F258FDD0B5C3FD8E" box="[615,719,515,540]" captionStart="Figure 2" captionStartId="7.[160,225,1612,1635]" captionTargetBox="[164,1152,143,1579]" captionTargetId="figure-9@7.[163,1153,142,1580]" captionTargetPageId="7" captionText="Figure 2. Titanogrypa (Cucullomyia) albuquerquei, holotype male (MNRJ) [lost]. (a) Sternite 5, ventral view. (b) Epandrium, cercus and surstylus, lateral view. (c) Phallus and associated structures, lateral view. (d) Distiphallus, ventral view. (e) Distiphallus, lateral view. Abbreviations: ar = arm; bp = basiphallus; bs = base; ce = cercus; ea = ejaculatory apodeme; ep = epandrium; ga = postgonal apodeme; hp = hypandrium;ju = juxta; ls = lateral stylus; ms = median stylus; ph = phallapodeme;po = postgonite; pr = pregonite; pp = paraphallus; sur = surstylus; stg 7 + 8 = syntergosternite 7 + 8; ve = vesica." figureDoi="http://doi.org/10.5281/zenodo.5473924" httpUri="https://zenodo.org/record/5473924/files/figure.png" pageId="29" pageNumber="334">Figures 2</figureCitation>
(a,e), 6(a,e)), but this species is remarkably different from other congeneric species, mainly in having a long vesica without a membranous projection, juxta with broad and long lobes, and styli very long and exposed (
<figureCitation id="133D2A5D3E632B58F131FDB8B678FD16" box="[270,372,619,644]" captionStart="Figure 6" captionStartId="11.[160,225,1515,1538]" captionTargetBox="[194,1152,150,1482]" captionTargetId="figure-9@11.[163,1153,142,1483]" captionTargetPageId="11" captionText="Figure 6. Titanogrypa (Cucullomyia) placida, male (MNRJ) [lost]. (a) Sternite 5, ventral view. (b) Syntergosternite 7 + 8, epandrium, cercus and surstylus, lateral view. (c) Phallus and associated structures, lateral view. (d) Distiphallus, ventral view. (e) Distiphallus, lateral view. Abbreviations: ar = arm; bp = basiphallus; bs = base; ce = cercus; ea = ejaculatory apodeme; ep = epandrium; ga = postgonal apodeme; hp = hypandrium; ju = juxta; ls = lateral stylus; ms = median stylus; ph = phallapodeme; po = postgonite; pr = pregonite; pp = paraphallus; sur = surstylus; stg 7 + 8 = syntergosternite 7 + 8; ve = vesica." figureDoi="http://doi.org/10.5281/zenodo.5473942" httpUri="https://zenodo.org/record/5473942/files/figure.png" pageId="29" pageNumber="334">Figures 6</figureCitation>
(ce))
</paragraph>
</subSubSection>
</treatment>
</document>