230 lines
27 KiB
XML
230 lines
27 KiB
XML
<document id="85AC98F9DF09A83C0885DCAF49621022" ID-CLB-Dataset="6498" ID-DOI="10.11646/zootaxa.4966.1.1" ID-GBIF-Dataset="e3a4e21d-e4fd-4d82-a596-de175e91f976" ID-ISSN="1175-5326" ID-Zenodo-Dep="4729328" ID-ZooBank="BD4EE68E-893C-47FC-91CE-418F4A6FE46C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1619760982532" checkinUser="plazi" docAuthor="Jałoszyński, Paweł" docDate="2021" docId="03AF8785A37D276AFF04279E1F4AFF1C" docLanguage="en" docName="zootaxa.4966.1.1.pdf" docOrigin="Zootaxa 4966 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hlavaciellus microtuberculatus Jałoszyński 2021, sp. n." docType="treatment" docVersion="9" lastPageNumber="7" masterDocId="FF96FFFDA379276CFF9323341A13FFE4" masterDocTitle="Six new species of the Oriental genus Hlavaciellus Jałoszyński (Coleoptera Staphylinidae, Scydmaeninae)" masterLastPageNumber="15" masterPageNumber="1" pageNumber="5" updateTime="1698938032643" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="FDB07E1C2643CECA20BE022AD5282CE4">
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<mods:title id="2D84B9B1A8A11585DC0B706F121EA05D">Six new species of the Oriental genus Hlavaciellus Jałoszyński (Coleoptera Staphylinidae, Scydmaeninae)</mods:title>
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<mods:name id="ACB8F5A2E972A430C51656259D65D969" type="personal">
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<mods:roleTerm id="CFA3BAE379D83DC29C7DA2797F99F53C">Author</mods:roleTerm>
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<mods:namePart id="D227CBCF0F5A0EC0F8D29F73A6BC8EBB">Jałoszyński, Paweł</mods:namePart>
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<mods:typeOfResource id="3D75B78082D38A56C75F816EDA43BB69">text</mods:typeOfResource>
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<mods:title id="9BCC7BBBE3B10EA5BA6C61A32C2856B8">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="88E4CFF336F38824C01FFCFC4504DBBB">
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<mods:date id="D2B5BBAC05FDBE15912FC941C5A36989">2021</mods:date>
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<mods:detail id="C5DE51C4BC5E4E4EA224A80DFD695398" type="pubDate">
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<mods:number id="4BA7B5CC37BDB88BB6A72054D0ACFE77">2021-04-30</mods:number>
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<mods:number id="35BE3CF3195E13D8CE0EFEE40EE3E3C6">4966</mods:number>
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<mods:classification id="93203260C26154823CBB30A3DACD495D">journal article</mods:classification>
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<mods:identifier id="01C5C20D0D91CE23A9E230FF3A1D08C6" type="DOI">10.11646/zootaxa.4966.1.1</mods:identifier>
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<treatment id="03AF8785A37D276AFF04279E1F4AFF1C" ID-DOI="http://doi.org/10.5281/zenodo.4791017" ID-GBIF-Taxon="182387822" ID-Zenodo-Dep="4791017" LSID="urn:lsid:plazi:treatment:03AF8785A37D276AFF04279E1F4AFF1C" httpUri="http://treatment.plazi.org/id/03AF8785A37D276AFF04279E1F4AFF1C" lastPageId="6" lastPageNumber="7" pageId="4" pageNumber="5">
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<subSubSection id="C31C6518A37D2768FF04279E184EFB21" box="[151,605,1194,1221]" pageId="4" pageNumber="5" type="nomenclature">
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<paragraph id="8BB93693A37D2768FF04279E184EFB21" blockId="4.[151,605,1194,1257]" box="[151,605,1194,1221]" pageId="4" pageNumber="5">
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<heading id="D0F181FFA37D2768FF04279E184EFB21" bold="true" box="[151,605,1194,1221]" fontSize="11" level="1" pageId="4" pageNumber="5" reason="1">
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<emphasis id="B972EA81A37D2768FF04279E184EFB21" bold="true" box="[151,605,1194,1221]" pageId="4" pageNumber="5">
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<taxonomicName id="4C064D10A37D2768FF04279E1807FB21" ID-CoL="8Y4JQ" authority="Jałoszyński, 2021" authorityName="Jałoszyński" authorityYear="2021" box="[151,532,1194,1221]" class="Insecta" family="Staphylinidae" genus="Hlavaciellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="microtuberculatus" status="sp. nov.">
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<emphasis id="B972EA81A37D2768FF04279E1807FB21" bold="true" box="[151,532,1194,1221]" italics="true" pageId="4" pageNumber="5">Hlavaciellus microtuberculatus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A24157FAA37D2768FD88279F184EFB21" box="[539,605,1195,1221]" pageId="4" pageNumber="5" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C31C6518A37D2768FF0427FB1B50FB0D" box="[151,323,1230,1257]" pageId="4" pageNumber="5" type="description">
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<paragraph id="8BB93693A37D2768FF0427FB1B50FB0D" blockId="4.[151,605,1194,1257]" box="[151,323,1230,1257]" pageId="4" pageNumber="5">
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(
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<figureCitation id="133D2A16A37D2768FF0C27FB1B01FB0D" box="[159,274,1231,1257]" captionStart="FIGURES 9–12" captionStartId="5.[151,264,1428,1452]" captionTargetBox="[191,1431,281,1398]" captionTargetId="figure-62@5.[151,1436,273,1405]" captionTargetPageId="5" captionText="FIGURES 9–12. Hlavaciellus microtuberculatus sp. n., male. Dorsal habitus (9); head in anterior view (10); aedeagus in ventral (11) and lateral (12) views." figureDoi="http://doi.org/10.5281/zenodo.4729334" httpUri="https://zenodo.org/record/4729334/files/figure.png" pageId="4" pageNumber="5">Figs 9–12</figureCitation>
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,
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<figureCitation id="133D2A16A37D2768FE8D27FA1B28FB0D" box="[286,315,1230,1257]" captionStart="FIGURE 25" captionStartId="11.[151,250,1285,1309]" captionTargetBox="[151,1436,237,1261]" captionTargetId="figure-43@11.[151,1436,237,1261]" captionTargetPageId="11" captionText="FIGURE 25. Distribution of nominal species of Hlavaciellus." figureDoi="http://doi.org/10.5281/zenodo.4729342" httpUri="https://zenodo.org/record/4729342/files/figure.png" pageId="4" pageNumber="5">25</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C31C6518A37D2768FF0426231953FA25" pageId="4" pageNumber="5" type="materials_examined">
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<paragraph id="8BB93693A37D2768FF0426231953FA25" blockId="4.[151,1437,1302,2013]" pageId="4" pageNumber="5">
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<emphasis id="B972EA81A37D2768FF0426231B5BFAD4" bold="true" box="[151,328,1302,1329]" pageId="4" pageNumber="5">Type material.</emphasis>
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<materialsCitation id="3B6E3CCEA37D2768FEC12623192DFA9D" ID-GBIF-Occurrence="3118551308" collectionCode="MHNG" collectorName="W. Malaysia & P. Jaloszynski" country="Malaysia" elevation="1400" location="Pahang" pageId="4" pageNumber="5" specimenCount="1" specimenCount-male="1" stateProvince="Pahang" typeStatus="holotype">
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<emphasis id="B972EA81A37D2768FEC1262318F5FAD4" bold="true" box="[338,742,1302,1329]" pageId="4" pageNumber="5">
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<typeStatus id="54BD8831A37D2768FEC126231BD0FAD5" box="[338,451,1303,1329]" pageId="4" pageNumber="5" type="holotype">Holotype</typeStatus>
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:
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<collectingCountry id="F3117603A37D2768FE4126221875FAD4" box="[466,614,1302,1328]" name="Malaysia" pageId="4" pageNumber="5">MALAYSIA</collectingCountry>
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(
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<collectingRegion id="49C2F871A37D2768FDEB262218CAFAD4" box="[632,729,1302,1328]" country="Malaysia" name="Pahang" pageId="4" pageNumber="5">Pahang</collectingRegion>
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):
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</emphasis>
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<specimenCount id="9D00FD1AA37D2768FD6326231914FAD5" box="[752,775,1303,1329]" pageId="4" pageNumber="5" type="male">♂</specimenCount>
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, two labels: “
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<collectorName id="26F35345A37D2768FC3426231E2EFAD5" box="[935,1085,1303,1329]" pageId="4" pageNumber="5">W. Malaysia</collectorName>
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: Pahang / Cameron Highlands / trail 9,
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<quantity id="4CFE9B76A37D2768FF6D260F1B5DFAB1" box="[254,334,1339,1365]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="4" pageNumber="5" unit="m" value="1400.0">
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<elevation id="002BD1A0A37D2768FF6D260F1B5DFAB1" box="[254,334,1339,1365]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="4" pageNumber="5" unit="m" value="1400.0">1400m</elevation>
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</quantity>
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, 27.3.93 / Löbl & Calame, #21” [white, printed], “
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<taxonomicName id="4C064D10A37D2768FC0D260F1E73FAB1" authorityName="Jaloszynski" authorityYear="2006" box="[926,1120,1339,1365]" class="Insecta" family="Staphylinidae" genus="Hlavaciellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
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<emphasis id="B972EA81A37D2768FC0D260F1E73FAB1" box="[926,1120,1339,1365]" italics="true" pageId="4" pageNumber="5">HLAVACIELLUS</emphasis>
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</taxonomicName>
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/
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<taxonomicName id="4C064D10A37D2768FBEF260F1F54FAB1" authorityName="Jałoszyński" authorityYear="2021" box="[1148,1351,1339,1365]" class="Insecta" family="Staphylinidae" genus="Hlavaciellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="microtuberculatus">
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<emphasis id="B972EA81A37D2768FBEF260F1F54FAB1" box="[1148,1351,1339,1365]" italics="true" pageId="4" pageNumber="5">microtuberculatus</emphasis>
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</taxonomicName>
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m. /
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<collectorName id="26F35345A37D2768FA1A260F1B08FA9D" pageId="4" pageNumber="5">P. Jałoszyński</collectorName>
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, 2021 / HOLOTYPUS” [red, printed] (
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<collectionCode id="ED17AE56A37D2768FD49266B1925FA9D" box="[730,822,1375,1401]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="4" pageNumber="5" type="Museum">MHNG</collectionCode>
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)
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</materialsCitation>
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||
.
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<materialsCitation id="3B6E3CCEA37D2768FCD9266B1953FA25" ID-GBIF-Occurrence="3119449302" collectionCode="MHNG" collectorName="W. Malaysia" country="Malaysia" elevation="1550" location="Pahang" pageId="4" pageNumber="5" specimenCount="2" specimenCount-female="2" stateProvince="Pahang" typeStatus="paratype">
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||
<emphasis id="B972EA81A37D2768FCD9266B1E3EFA9C" bold="true" box="[842,1069,1374,1401]" pageId="4" pageNumber="5">
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<typeStatus id="54BD8831A37D2768FCD9266B19D1FA9D" box="[842,962,1375,1401]" pageId="4" pageNumber="5" type="paratype">Paratypes</typeStatus>
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(2 exx.):
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</emphasis>
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<specimenCount id="9D00FD1AA37D2768FBA6266B1E73FA9D" box="[1077,1120,1375,1401]" pageId="4" pageNumber="5" type="female">1 ♀</specimenCount>
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, same data as for holotype;
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<specimenCount id="9D00FD1AA37D2768FF0426B71AD0FA79" box="[151,195,1411,1437]" pageId="4" pageNumber="5" type="female">1 ♀</specimenCount>
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, “
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<collectorName id="26F35345A37D2768FF4F26B71B62FA79" box="[220,369,1411,1437]" pageId="4" pageNumber="5">W. Malaysia</collectorName>
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: Pahang # 18b / Cameron Highls,
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||
<quantity id="4CFE9B76A37D2768FD6926B6195AFA79" box="[762,841,1410,1437]" metricMagnitude="3" metricUnit="m" metricValue="1.55" pageId="4" pageNumber="5" unit="m" value="1550.0">
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||
<elevation id="002BD1A0A37D2768FD6926B6195AFA79" box="[762,841,1410,1437]" metricMagnitude="3" metricUnit="m" metricValue="1.55" pageId="4" pageNumber="5" unit="m" value="1550.0">1550m</elevation>
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</quantity>
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/ Gunung Jasar, trail 11 / Löbl & Calame, 24.3.93”; paratypes with yellow “PARATYPUS” labels (
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<collectionCode id="ED17AE56A37D2768FD30269318EEFA25" box="[675,765,1447,1473]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="4" pageNumber="5" type="Museum">MHNG</collectionCode>
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, cPJ).
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</materialsCitation>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C31C6518A37D2768FF5426FE19F3F991" pageId="4" pageNumber="5" type="diagnosis">
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<paragraph id="8BB93693A37D2768FF5426FE19F3F991" blockId="4.[151,1437,1302,2013]" pageId="4" pageNumber="5">
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<emphasis id="B972EA81A37D2768FF5426FE1B51FA00" bold="true" box="[199,322,1482,1508]" pageId="4" pageNumber="5">Diagnosis.</emphasis>
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Body small, not reaching
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<quantity id="4CFE9B76A37D2768FDEB26FF18AFFA01" box="[632,700,1483,1509]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="4" pageNumber="5" unit="mm" value="2.0">2 mm</quantity>
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in length and conspicuously stout, with elytra about as long as broad; head in males with unmodified frons and vertex, but in both sexes with pair of tiny, barely discernible submedian tubercles on vertex; pronotum slightly narrower than elytra; each of antennomeres 3–4 and 7–10 about as long as broad, and 5–6 each indistinctly elongate; aedeagus in ventral view stout, 3 times as long as broad and distinctly narrowing both toward apex and base; parameres with thin subapical setae.
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||
</paragraph>
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</subSubSection>
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||
<subSubSection id="C31C6518A37D2769FF54254A1897F841" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5" type="description">
|
||
<paragraph id="8BB93693A37D2768FF54254A1B95F959" blockId="4.[151,1437,1302,2013]" pageId="4" pageNumber="5">
|
||
<emphasis id="B972EA81A37D2768FF54254A1B4AF97C" bold="true" box="[199,345,1662,1688]" pageId="4" pageNumber="5">Description.</emphasis>
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Body of male (
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||
<figureCitation id="133D2A16A37D2768FD9A254B185EF97D" box="[521,589,1663,1689]" captionStart="FIGURES 9–12" captionStartId="5.[151,264,1428,1452]" captionTargetBox="[191,1431,281,1398]" captionTargetId="figure-62@5.[151,1436,273,1405]" captionTargetPageId="5" captionText="FIGURES 9–12. Hlavaciellus microtuberculatus sp. n., male. Dorsal habitus (9); head in anterior view (10); aedeagus in ventral (11) and lateral (12) views." figureDoi="http://doi.org/10.5281/zenodo.4729334" httpUri="https://zenodo.org/record/4729334/files/figure.png" pageId="4" pageNumber="5">Fig. 9</figureCitation>
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) stout, strongly convex, uniformly dark brown; setae distinctly lighter than cuticle; BL
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||
<quantity id="4CFE9B76A37D2768FE8925971B92F959" box="[282,385,1699,1725]" metricMagnitude="-3" metricUnit="m" metricValue="1.78" pageId="4" pageNumber="5" unit="mm" value="1.78">1.78 mm</quantity>
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||
.
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||
</paragraph>
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<paragraph id="8BB93693A37D2768FF5425F318CCF871" blockId="4.[151,1437,1302,2013]" pageId="4" pageNumber="5">
|
||
Head (
|
||
<figureCitation id="133D2A16A37D2768FE8125F31B70F905" box="[274,355,1735,1761]" captionStart="FIGURES 9–12" captionStartId="5.[151,264,1428,1452]" captionTargetBox="[191,1431,281,1398]" captionTargetId="figure-62@5.[151,1436,273,1405]" captionTargetPageId="5" captionText="FIGURES 9–12. Hlavaciellus microtuberculatus sp. n., male. Dorsal habitus (9); head in anterior view (10); aedeagus in ventral (11) and lateral (12) views." figureDoi="http://doi.org/10.5281/zenodo.4729334" httpUri="https://zenodo.org/record/4729334/files/figure.png" pageId="4" pageNumber="5">Fig. 10</figureCitation>
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||
) broadest at large, strongly convex and coarsely faceted eyes, HL
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||
<quantity id="4CFE9B76A37D2768FBD925F31EA3F905" box="[1098,1200,1735,1761]" metricMagnitude="-4" metricUnit="m" metricValue="3.3" pageId="4" pageNumber="5" unit="mm" value="0.33">0.33 mm</quantity>
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||
, HW
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||
<quantity id="4CFE9B76A37D2768FB6325F21F44F905" box="[1264,1367,1734,1761]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" pageId="4" pageNumber="5" unit="mm" value="0.45">0.45 mm</quantity>
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||
; frons and vertex unmodified, weakly convex, vertex with pair of tiny submedian tubercles (barely discernible; observations at various angles are necessary to notice tubercles). Punctures on frons and vertex fine and inconspicuous; setae long, moderately dense and suberect. Antennae slender, AnL
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||
<quantity id="4CFE9B76A37D2768FC11240719FBF8A9" box="[898,1000,1843,1869]" metricMagnitude="-3" metricUnit="m" metricValue="1.23" pageId="4" pageNumber="5" unit="mm" value="1.23">1.23 mm</quantity>
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||
, BL/AnL 1.45; scape indistinctly elongate, antennomeres 3–4 and 7–10 each about as long as broad, and 5–6 each indistinctly elongate, 11 distinctly shorter than 9–10 combined, twice as long as broad.
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||
</paragraph>
|
||
<paragraph id="8BB93693A37D2769FF5424AB1F0DFF1D" blockId="4.[151,1437,1302,2013]" lastBlockId="5.[151,1436,151,249]" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5">
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||
Pronotum subrectangular, broadest near anterior third but indistinctly narrowing toward base; PL
|
||
<quantity id="4CFE9B76A37D2768FABC24AA1F8BF85D" box="[1327,1432,1950,1977]" metricMagnitude="-4" metricUnit="m" metricValue="5.8" pageId="4" pageNumber="5" unit="mm" value="0.58">0.58 mm</quantity>
|
||
, PW
|
||
<quantity id="4CFE9B76A37D2768FF5B24F61B3CF839" box="[200,303,1986,2013]" metricMagnitude="-4" metricUnit="m" metricValue="8.5" pageId="4" pageNumber="5" unit="mm" value="0.85">0.85 mm</quantity>
|
||
, PI 0.68; anterior margin weakly rounded; sides with finely crenulate lateral carinae, in posterior 2/3 straight; posterior corners nearly right-angled and with blunt tips; posterior margin shallowly bisinuate; pronotal disc strongly convex, with area at each posterior corner distinctly impressed; antebasal pits slightly closer to posterior than to lateral margins. Punctures on pronotum fine and inconspicuous; setae long, dense, suberect.
|
||
</paragraph>
|
||
<caption id="DF79661BA37C2769FF0426A01BC1FA34" ID-DOI="http://doi.org/10.5281/zenodo.4729334" ID-Zenodo-Dep="4729334" httpUri="https://zenodo.org/record/4729334/files/figure.png" pageId="5" pageNumber="6" startId="5.[151,264,1428,1452]" targetBox="[191,1431,281,1398]" targetPageId="5">
|
||
<paragraph id="8BB93693A37C2769FF0426A01BC1FA34" blockId="5.[151,1436,1428,1488]" pageId="5" pageNumber="6">
|
||
<emphasis id="B972EA81A37C2769FF0426A01B5BFA48" bold="true" box="[151,328,1428,1452]" pageId="5" pageNumber="6">FIGURES 9–12.</emphasis>
|
||
<taxonomicName id="4C064D10A37C2769FEDD26A0189FFA48" authority="Jałoszyński, 2021" authorityName="Jałoszyński" authorityYear="2021" box="[334,652,1428,1452]" class="Insecta" family="Staphylinidae" genus="Hlavaciellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="microtuberculatus" status="sp. nov.">
|
||
<emphasis id="B972EA81A37C2769FEDD26A0189FFA48" box="[334,652,1428,1452]" italics="true" pageId="5" pageNumber="6">Hlavaciellus microtuberculatus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B972EA81A37C2769FD0026A018DFFA48" bold="true" box="[659,716,1428,1452]" pageId="5" pageNumber="6">
|
||
<taxonomicNameLabel id="A24157FAA37C2769FD0026A018DFFA48" box="[659,716,1428,1452]" pageId="5" pageNumber="6" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, male. Dorsal habitus (9); head in anterior view (10); aedeagus in ventral (11) and lateral (12) views.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BB93693A37C2769FF5426CA1B9DF961" blockId="5.[151,1437,1534,2029]" pageId="5" pageNumber="6">
|
||
Elytra together oval, broadest distinctly in front of middle; EL
|
||
<quantity id="4CFE9B76A37C2769FC0426CB19ECF9FD" box="[919,1023,1535,1561]" metricMagnitude="-4" metricUnit="m" metricValue="8.8" pageId="5" pageNumber="6" unit="mm" value="0.88">0.88 mm</quantity>
|
||
, EW
|
||
<quantity id="4CFE9B76A37C2769FBD326CB1EBBF9FD" box="[1088,1192,1535,1561]" metricMagnitude="-4" metricUnit="m" metricValue="9.3" pageId="5" pageNumber="6" unit="mm" value="0.93">0.93 mm</quantity>
|
||
, EI 0.95; subhumeral lines sharply marked, each developed as step-wise border between elevated humeral region and median area, slightly exceeding 0.3 EL; sides of elytra strongly rounded; elytral apices separately rounded. Punctures and setae similar to those on pronotum.
|
||
</paragraph>
|
||
<paragraph id="8BB93693A37C2769FF5425BB1840F94C" blockId="5.[151,1437,1534,2029]" box="[199,595,1678,1705]" pageId="5" pageNumber="6">Legs long and slender, unmodified.</paragraph>
|
||
<paragraph id="8BB93693A37C2769FF5425861F39F8DD" blockId="5.[151,1437,1534,2029]" pageId="5" pageNumber="6">
|
||
Aedeagus (
|
||
<figureCitation id="133D2A16A37C2769FED525861BD5F929" box="[326,454,1714,1741]" captionStart="FIGURES 9–12" captionStartId="5.[151,264,1428,1452]" captionTargetBox="[191,1431,281,1398]" captionTargetId="figure-62@5.[151,1436,273,1405]" captionTargetPageId="5" captionText="FIGURES 9–12. Hlavaciellus microtuberculatus sp. n., male. Dorsal habitus (9); head in anterior view (10); aedeagus in ventral (11) and lateral (12) views." figureDoi="http://doi.org/10.5281/zenodo.4729334" httpUri="https://zenodo.org/record/4729334/files/figure.png" pageId="5" pageNumber="6">Figs 11–12</figureCitation>
|
||
) elongate but not very slender; AeL
|
||
<quantity id="4CFE9B76A37C2769FCF8258719C7F929" box="[875,980,1715,1741]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="6" unit="mm" value="0.3">0.30 mm</quantity>
|
||
; median lobe in ventral view 3 times as long as broad, broadest near sub-basal third and with broadly rounded sides; endophallic structures distinct, in ventral view appearing as single elongate subtriangular element, but in lateral view two separate tubular structures can be seen, proximal one bent at sharp angle near base of median lobe; parameres with thin subapical setae.
|
||
</paragraph>
|
||
<paragraph id="8BB93693A37C2769FF5424761897F841" blockId="5.[151,1437,1534,2029]" pageId="5" pageNumber="6">
|
||
Female. Externally similar to male but with clearly shorter antennae in relation to body. BL
|
||
<quantity id="4CFE9B76A37C2769FB5424761F7EF8B9" box="[1223,1389,1858,1885]" metricMagnitude="-3" metricUnit="m" metricValue="1.815" metricValueMax="1.85" metricValueMin="1.78" pageId="5" pageNumber="6" unit="mm" value="1.815" valueMax="1.85" valueMin="1.78">1.78–1.85 mm</quantity>
|
||
; HL
|
||
<quantity id="4CFE9B76A37C2769FF0424531B2EF865" box="[151,317,1895,1921]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="5" pageNumber="6" unit="mm" value="0.35" valueMax="0.4" valueMin="0.3">0.30–0.40 mm</quantity>
|
||
, HW
|
||
<quantity id="4CFE9B76A37C2769FE1224521834F865" box="[385,551,1894,1921]" metricMagnitude="-4" metricUnit="m" metricValue="4.6499999999999995" metricValueMax="4.8" metricValueMin="4.5" pageId="5" pageNumber="6" unit="mm" value="0.46499999999999997" valueMax="0.48" valueMin="0.45">0.45–0.48 mm</quantity>
|
||
, BL/AnL 1.82–1.95; PL
|
||
<quantity id="4CFE9B76A37C2769FCD1245219FBF865" box="[834,1000,1894,1921]" metricMagnitude="-4" metricUnit="m" metricValue="5.6499999999999995" metricValueMax="5.8" metricValueMin="5.5" pageId="5" pageNumber="6" unit="mm" value="0.565" valueMax="0.58" valueMin="0.55">0.55–0.58 mm</quantity>
|
||
, PW
|
||
<quantity id="4CFE9B76A37C2769FBB424521EDEF865" box="[1063,1229,1894,1921]" metricMagnitude="-4" metricUnit="m" metricValue="8.75" metricValueMax="9.0" metricValueMin="8.5" pageId="5" pageNumber="6" unit="mm" value="0.875" valueMax="0.9" valueMin="0.85">0.85–0.90 mm</quantity>
|
||
, PI 0.64–0.65; EL
|
||
<quantity id="4CFE9B76A37C2769FF0424BF1AEEF841" box="[151,253,1931,1957]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="5" pageNumber="6" unit="mm" value="0.9">0.90 mm</quantity>
|
||
, EW
|
||
<quantity id="4CFE9B76A37C2769FEA824BF1BF3F841" box="[315,480,1931,1957]" metricMagnitude="-4" metricUnit="m" metricValue="9.15" metricValueMax="9.3" metricValueMin="9.0" pageId="5" pageNumber="6" unit="mm" value="0.915" valueMax="0.93" valueMin="0.9">0.90–0.93 mm</quantity>
|
||
, EI 0.97–1.00.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31C6518A37C2769FF54249A18A8F82D" box="[199,699,1966,1993]" pageId="5" pageNumber="6" type="distribution">
|
||
<paragraph id="8BB93693A37C2769FF54249A18A8F82D" blockId="5.[151,1437,1534,2029]" box="[199,699,1966,1993]" pageId="5" pageNumber="6">
|
||
<emphasis id="B972EA81A37C2769FF54249A1B4AF82C" bold="true" box="[199,345,1966,1992]" pageId="5" pageNumber="6">Distribution</emphasis>
|
||
. Peninsular
|
||
<collectingCountry id="F3117603A37C2769FE76249A185FF82C" box="[485,588,1966,1992]" name="Malaysia" pageId="5" pageNumber="6">Malaysia</collectingCountry>
|
||
(
|
||
<figureCitation id="133D2A16A37C2769FDC8249A18BCF82D" box="[603,687,1966,1993]" captionStart="FIGURE 25" captionStartId="11.[151,250,1285,1309]" captionTargetBox="[151,1436,237,1261]" captionTargetId="figure-43@11.[151,1436,237,1261]" captionTargetPageId="11" captionText="FIGURE 25. Distribution of nominal species of Hlavaciellus." figureDoi="http://doi.org/10.5281/zenodo.4729342" httpUri="https://zenodo.org/record/4729342/files/figure.png" pageId="5" pageNumber="6">Fig. 25</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31C6518A37C2769FF5424E71F67F809" box="[199,1396,2002,2029]" pageId="5" pageNumber="6" type="etymology">
|
||
<paragraph id="8BB93693A37C2769FF5424E71F67F809" blockId="5.[151,1437,1534,2029]" box="[199,1396,2002,2029]" pageId="5" pageNumber="6">
|
||
<emphasis id="B972EA81A37C2769FF5424E71B5DF809" bold="true" box="[199,334,2003,2029]" pageId="5" pageNumber="6">Etymology.</emphasis>
|
||
The adjective
|
||
<taxonomicName id="4C064D10A37C2769FE6024E618ADF808" authorityName="Jałoszyński" authorityYear="2021" box="[499,702,2002,2028]" class="Insecta" family="Staphylinidae" genus="Hlavaciellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="microtuberculatus">
|
||
<emphasis id="B972EA81A37C2769FE6024E618ADF808" box="[499,702,2002,2028]" italics="true" pageId="5" pageNumber="6">microtuberculatus</emphasis>
|
||
</taxonomicName>
|
||
refers to the pair of tiny tubercles on the vertex in both sexes.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31C6518A37F276AFF5423A21F4AFF1C" pageId="6" pageNumber="7" type="discussion">
|
||
<paragraph id="8BB93693A37F276AFF5423A21F4AFF1C" blockId="6.[151,1437,150,249]" pageId="6" pageNumber="7">
|
||
<emphasis id="B972EA81A37F276AFF5423A21B28FF54" bold="true" box="[199,315,150,176]" pageId="6" pageNumber="7">Remarks.</emphasis>
|
||
<taxonomicName id="4C064D10A37F276AFED123A218B2FF54" authorityName="Jałoszyński" authorityYear="2021" box="[322,673,150,176]" class="Insecta" family="Staphylinidae" genus="Hlavaciellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="microtuberculatus">
|
||
<emphasis id="B972EA81A37F276AFED123A218B2FF54" box="[322,673,150,176]" italics="true" pageId="6" pageNumber="7">Hlavaciellus microtuberculatus</emphasis>
|
||
</taxonomicName>
|
||
belongs to small-bodied species, clearly below
|
||
<quantity id="4CFE9B76A37F276AFB2A23A21EE9FF55" box="[1209,1274,150,177]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="6" pageNumber="7" unit="mm" value="2.0">2 mm</quantity>
|
||
in length. The most conspicuous feature of this otherwise unremarkable species is a very stout body, with elytra about as long as broad, or even indistinctly broader than long; in all remaining species the elytra are at least slightly elongate.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |