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<document id="88303E4A024941759CA02D4BCB554754" ID-DOI="10.11646/zootaxa.3664.2.1" ID-GBIF-Dataset="a50b71f5-48a6-464f-aea1-f29bbc8ee322" ID-ISSN="1175-5326" ID-Zenodo-Dep="220503" ID-ZooBank="105D178E-6DBF-401D-85BB-CDB041B67E91" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460615203542" checkinUser="plazi" docAuthor="Shi, Hongliang, Sciaky, Riccardo, Liang, Hongbin &amp; Zhou, Hongzhang" docDate="2013" docId="03AA10085649FFFAC1D8FBF1FC8CFB5A" docLanguage="en" docName="zt03664p135.pdf" docOrigin="Zootaxa 3664 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pterostichus (Wraseiellus) kambaiti Andrewes 1947" docType="treatment" docVersion="9" lastPageNumber="120" masterDocId="FF936870565BFFE9C14FFF95FFA9FF9E" masterDocTitle="A new subgenus Wraseiellus of the genus Pterostichus Bonelli (Coleoptera, Carabidae, Pterostichini) and new species descriptions" masterLastPageNumber="135" masterPageNumber="101" pageNumber="119" updateTime="1698330049886" updateUser="plazi">
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<mods:title id="C851557D2417161FF7D21986807AF424">A new subgenus Wraseiellus of the genus Pterostichus Bonelli (Coleoptera, Carabidae, Pterostichini) and new species descriptions</mods:title>
</mods:titleInfo>
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<mods:namePart id="60F6EA82420CFDA1F66FEDAED0F0BC9D">Shi, Hongliang</mods:namePart>
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<mods:namePart id="5A85B79A359C195EF10D239F89160C3B">Sciaky, Riccardo</mods:namePart>
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<mods:namePart id="5A5369FD08A43E9DB462F51400F61552">Liang, Hongbin</mods:namePart>
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<mods:namePart id="69764339A526DC0C0541234FEF806E89">Zhou, Hongzhang</mods:namePart>
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<mods:title id="45DEFD401CF73927A952CA4E42D6A664">Zootaxa</mods:title>
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<mods:part id="76D7F31734DE2FC6CBCF674C660C28F5">
<mods:date id="1D1B52E5FF2F940FD80ADDE737F0CF29">2013</mods:date>
<mods:detail id="94F2B43BC5CFF330603C7090E6ED6F9E" type="volume">
<mods:number id="486B1F7B0E640E9554CDB4F7E592BFBF">3664</mods:number>
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<mods:classification id="9A51B2AFE6CE29BAE54B1DD1DFDE8702">journal article</mods:classification>
<mods:identifier id="730558487A240B972E2C2E6C17BA5101" type="DOI">10.11646/zootaxa.3664.2.1</mods:identifier>
<mods:identifier id="A1DBD2F9BCB63C3411146BAECCCB3246" type="GBIF-Dataset">a50b71f5-48a6-464f-aea1-f29bbc8ee322</mods:identifier>
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<treatment id="03AA10085649FFFAC1D8FBF1FC8CFB5A" ID-DOI="http://doi.org/10.5281/zenodo.6154519" ID-GBIF-Taxon="119568510" ID-Zenodo-Dep="6154519" LSID="urn:lsid:plazi:treatment:03AA10085649FFFAC1D8FBF1FC8CFB5A" httpUri="http://treatment.plazi.org/id/03AA10085649FFFAC1D8FBF1FC8CFB5A" lastPageId="19" lastPageNumber="120" pageId="18" pageNumber="119">
<subSubSection id="C319F2955649FFFBC1D8FBF1FAE3FABF" pageId="18" pageNumber="119" type="nomenclature">
<paragraph id="8BBCA11E5649FFFBC1D8FBF1FC8CFBE3" blockId="18.[151,805,1123,1182]" box="[151,805,1123,1150]" pageId="18" pageNumber="119">
<heading id="D0F416725649FFFBC1D8FBF1FC8CFBE3" bold="true" box="[151,805,1123,1150]" fontSize="11" level="1" pageId="18" pageNumber="119" reason="1">
<taxonomicName id="4C03DA9D5649FFFBC1D8FBF1FC8CFBE3" ID-CoL="4Q4HP" authority="Andrewes, 1947" authorityName="Andrewes" authorityYear="1947" box="[151,805,1123,1150]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="119" phylum="Arthropoda" rank="species" species="kambaiti" subGenus="Wraseiellus">
<emphasis id="B9777D0C5649FFFBC1D8FBF1FC8CFBE3" bold="true" box="[151,805,1123,1150]" pageId="18" pageNumber="119">
<emphasis id="B9777D0C5649FFFBC1D8FBF1FE84FBE0" bold="true" box="[151,301,1124,1150]" italics="true" pageId="18" pageNumber="119">Pterostichus</emphasis>
(
<emphasis id="B9777D0C5649FFFBC072FBF1FE61FBE0" bold="true" box="[317,456,1124,1150]" italics="true" pageId="18" pageNumber="119">Wraseiellus</emphasis>
)
<emphasis id="B9777D0C5649FFFBC095FBF1FDEDFBE0" bold="true" box="[474,580,1124,1150]" italics="true" pageId="18" pageNumber="119">kambaiti</emphasis>
(Andrewes, 1947)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BBCA11E5649FFFBC1D8FB13FD83FB00" blockId="18.[151,805,1123,1182]" box="[151,554,1158,1182]" pageId="18" pageNumber="119">
(
<figureCitation id="1338BD9B5649FFFBC1D1FB13FEEAFB00" box="[158,323,1158,1182]" captionStart="FIGURES 13 – 18" captionStartId="23.[151,264,1749,1772]" captionTargetBox="[158,1427,204,1715]" captionTargetId="figure@23.[151,1436,193,1728]" captionTargetPageId="23" captionText="FIGURES 13 – 18. Habitus of Pterostichus (Wraseiellus) spp.: scale bar = 2.0 mm; 13, Pt. pseudodiversus n. sp., holotype; 14, Pt. pseudodiversus n. sp., a female paratype from Muli; 15, Pt. pseudodiversus n. sp., a male paratype from Deqin; 16, Pt. pseudodiversus n. sp., a male from Weixi; 17, Pt. kambaiti (Andrewes), holotype [photoed by Dr. J. Bergsten]; 18, Pt. kambaiti (Andrewes), holotype of Pterostichus kambaiti Jedlička." httpUri="https://zenodo.org/record/220506/files/figure.png" pageId="18" pageNumber="119">Figures 17, 18</figureCitation>
,
<figureCitation id="1338BD9B5649FFFBC000FB13FE31FB00" box="[335,408,1158,1182]" captionStart="FIGURES 19 – 28" captionStartId="24.[151,264,1308,1331]" captionTargetBox="[166,1426,208,1270]" captionTargetId="figure@24.[151,1436,193,1299]" captionTargetPageId="24" captionText="FIGURES 19 – 28. Labels of name-bearing type specimen: 19, lectotype of Pterostichus andrewesi Jedlička, 1931; 20, holotype of Pterostichus meyeri Jedlička, 1934; 21, holotype of Feronia crebrepunctata Straneo, 1937; 22, holotype of Pterostichus comatus n. sp.; 23, holotype of Pterostichus crassiapex n. sp.; 24, a syntype of Omaseus diversus Fairmaire, 1886; 25, holotype of Pterostichus stictopleurus cangshanensis n. ssp.; 26, holotype of Pterostichus pseudodiversus n. sp.; 27, holotype of Feronia kambaiti Andrewes, 1947 [photoed by Dr. J. Bergsten]; 28, holotype of Pterostichus kambaiti Jedlička, 1965." httpUri="https://zenodo.org/record/220507/files/figure.png" pageId="18" pageNumber="119">27, 28</figureCitation>
,
<figureCitation id="1338BD9B5649FFFBC0ECFB13FE6BFB00" box="[419,450,1158,1182]" captionStart="FIGURES 35 – 40" captionStartId="26.[151,264,1759,1782]" captionTargetBox="[168,1419,197,1735]" captionTargetId="figure@26.[151,1436,193,1738]" captionTargetPageId="26" captionText="FIGURES 35 – 40. Median lobe of male genitalia of Pterostichus (Wraseiellus) spp.: scale bar = 1.0 mm; A, left lateral view of median lobe; B, dorsal view of median lobe; 35, Pt. stictopleurus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 36, Pt. stictopleurus cangshanensis n. ssp., holotype; 37, Pt. pseudodiversus n. sp., holotype; 38, Pt. pseudodiversus n. sp., a paratype from Muli; 39, Pt. pseudodiversus n. sp., a male from Weixi; 40, Pt. kambaiti (Andrewes), holotype of Pterostichus kambaiti Jedlička." httpUri="https://zenodo.org/record/220509/files/figure.png" pageId="18" pageNumber="119">40</figureCitation>
,
<figureCitation id="1338BD9B5649FFFBC082FB13FE42FB00" box="[461,491,1158,1182]" captionStart="FIGURES 67 – 84" captionStartId="29.[151,264,1584,1607]" captionTargetBox="[166,1426,201,1530]" captionTargetId="figure@29.[151,1436,193,1545]" captionTargetPageId="29" captionText="FIGURES 67 – 84. Parameres of male genitalia of Pterostichus (Wraseiellus) spp.: scale bar = 0.5 mm; A, left paramere; B, right paramere: 67, Pt. andrewesi Jedlička, a male from Jinfoshan; 68, Pt. andrewesi Jedlička, a paralectotype from Jinfoshan; 69, Pt. meyeri Jedlička, a male from &quot; Yunnan &quot;; 70, Pt. comatus n. sp., holotype; 71, Pt. crassiapex n. sp., holotype; 72, Pt. stictopleurus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 73, Pt. stictopleurus cangshanensis n. ssp., holotype; 74 – 75, Pt. diversus (Fairmaire), males from Yulongshan; 76, Pt. diversus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 77, Pt. diversus (Fairmaire), a male from &quot; Yunnan &quot;; 78, Pt. kambaiti (Andrewes), holotype of Pterostichus kambaiti Jedlička; 79, Pt. pseudodiversus n. sp., holotype; 80, Pt. pseudodiversus n. sp., a paratype from Muli; 81, Pt. pseudodiversus n. sp., a paratype from Zhongdian; 82, Pt. pseudodiversus n. sp., a paratype from Deqin; 83, Pt. pseudodiversus n. sp.., a paratype from Lanping; 84, Pt. pseudodiversus n. sp., a male from Weixi." httpUri="https://zenodo.org/record/220512/files/figure.png" pageId="18" pageNumber="119">78</figureCitation>
,
<figureCitation id="1338BD9B5649FFFBC0B8FB13FD8BFB00" box="[503,546,1158,1182]" captionStart="FIGURES 98 – 106" captionStartId="32.[151,264,1472,1495]" captionTargetBox="[155,1435,216,1425]" captionTargetId="figure@32.[151,1436,215,1427]" captionTargetPageId="32" captionText="FIGURES 98 – 106. Pronotum hind angle and basal area (right side) of Pterostichus (Wraseiellus) spp.; scale bar = 0.5 mm; 98, Pt. andrewesi Jedlička, a female from Jinfoshan; 99, Pt. meyeri Jedlička, a male from &quot; Yunnan &quot;; 100, Pt. crassiapex n. sp., holotype; 101, Pt. comatus n. sp., holotype; 102, Pt. diversus (Fairmaire), a male from Yulongshan; 103, Pt. pseudodiversus n. sp., holotype; 104, Pt. stictopleurus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 105, Pt. stictopleurus cangshanensis n. ssp., holotype; 106, Pt. kambaiti (Andrewes), holotype of Pterostichus kambaiti Jedlička." httpUri="https://zenodo.org/record/220515/files/figure.png" pageId="18" pageNumber="119">106</figureCitation>
)
</paragraph>
<paragraph id="8BBCA11E5649FFFBC1D8FB5BFA95FB7A" blockId="18.[151,1436,1230,1314]" box="[151,1340,1230,1253]" pageId="18" pageNumber="119">
Andrewes 1947: 22 (Original:
<taxonomicName id="4C03DA9D5649FFFBC082FB45FDB7FB7B" box="[461,542,1232,1253]" class="Insecta" family="Carabidae" genus="Feronia" higherTaxonomySource="CoL" kingdom="Plantae" order="Coleoptera" pageId="18" pageNumber="119" phylum="Arthropoda" rank="genus">
<emphasis id="B9777D0C5649FFFBC082FB45FDB7FB7B" box="[461,542,1232,1253]" italics="true" pageId="18" pageNumber="119">Feronia</emphasis>
</taxonomicName>
;
<typeStatus id="54B81FBC5649FFFBC363FB5BFD2CFB7A" box="[556,645,1230,1252]" pageId="18" pageNumber="119" type="holotype">holotype</typeStatus>
deposited in NHRS); Lorenz 2005: 287 (
<taxonomicName id="4C03DA9D5649FFFBC567FB5AFB0FFB7B" box="[1064,1190,1231,1253]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="119" phylum="Arthropoda" rank="genus">
<emphasis id="B9777D0C5649FFFBC567FB5AFB0FFB7B" box="[1064,1190,1231,1253]" italics="true" pageId="18" pageNumber="119">Pterostichus</emphasis>
</taxonomicName>
(
<emphasis id="B9777D0C5649FFFBC5FBFB45FA8CFB7B" box="[1204,1317,1231,1253]" italics="true" pageId="18" pageNumber="119">incert. sed.</emphasis>
)).
</paragraph>
<paragraph id="8BBCA11E5649FFFBC1D8FB78FAE3FABF" blockId="18.[151,1436,1230,1314]" pageId="18" pageNumber="119">
<emphasis id="B9777D0C5649FFFBC1D8FB78FEACFA9A" bold="true" box="[151,261,1261,1284]" pageId="18" pageNumber="119">Synonym:</emphasis>
<taxonomicName id="4C03DA9D5649FFFBC044FB78FD80FA9D" authority="Jedlicka 1965: 204" authorityName="Jedlicka" authorityPageNumber="204" authorityYear="1965" box="[267,553,1261,1283]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="119" phylum="Arthropoda" rank="species" species="kambaiti">
<emphasis id="B9777D0C5649FFFBC044FB78FECDFA9D" box="[267,356,1261,1283]" italics="true" pageId="18" pageNumber="119">kambaiti</emphasis>
Jedlička 1965: 204
</taxonomicName>
(
<taxonomicName id="4C03DA9D5649FFFBC377FB78FD1FFA9D" box="[568,694,1261,1283]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="119" phylum="Arthropoda" rank="genus">
<emphasis id="B9777D0C5649FFFBC377FB78FD1FFA9D" box="[568,694,1261,1283]" italics="true" pageId="18" pageNumber="119">Pterostichus</emphasis>
</taxonomicName>
(
<emphasis id="B9777D0C5649FFFBC38BFB7BFC9CFA9D" box="[708,821,1261,1283]" italics="true" pageId="18" pageNumber="119">incert. sed.</emphasis>
);
<typeStatus id="54B81FBC5649FFFBC205FB78FC0AFA9D" box="[842,931,1261,1283]" pageId="18" pageNumber="119" type="holotype">holotype</typeStatus>
deposited in NHRS), Junior secondary homonym of
<taxonomicName id="4C03DA9D5649FFFBC1ADFA99FDD0FABC" authority="Andrewes 1947" authorityName="Andrewes" authorityYear="1947" box="[226,633,1291,1314]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="119" phylum="Arthropoda" rank="species" species="kambaiti">
<emphasis id="B9777D0C5649FFFBC1ADFA99FE16FABC" box="[226,447,1292,1314]" italics="true" pageId="18" pageNumber="119">Pterostichus kambaiti</emphasis>
(Andrewes 1947)
</taxonomicName>
; Lorenz 1998: 266 (catalogue; synonymized with
<taxonomicName id="4C03DA9D5649FFFBC53BFA99FA96FABF" authority="Andrewes" authorityName="Andrewes" box="[1140,1343,1291,1314]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="119" phylum="Arthropoda" rank="species" species="kambaiti">
<emphasis id="B9777D0C5649FFFBC53BFA99FB64FABC" box="[1140,1229,1292,1314]" italics="true" pageId="18" pageNumber="119">kambaiti</emphasis>
Andrewes
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C319F2955649FFFBC1D8FADAFC22F966" pageId="18" pageNumber="119" type="materials_examined">
<paragraph id="8BBCA11E5649FFFBC1D8FADAFCC0FAF8" blockId="18.[151,1436,1358,2036]" box="[151,873,1358,1383]" pageId="18" pageNumber="119">
<emphasis id="B9777D0C5649FFFBC1D8FADAFE84FAF9" bold="true" box="[151,301,1358,1383]" pageId="18" pageNumber="119">
<typeStatus id="54B81FBC5649FFFBC1D8FADAFF7BFAF9" box="[151,210,1359,1383]" pageId="18" pageNumber="119">Type</typeStatus>
locality
</emphasis>
.
<collectingCountry id="F314E18E5649FFFBC073FADAFE04FAF9" box="[316,429,1359,1383]" name="Myanmar" pageId="18" pageNumber="119">Myanmar</collectingCountry>
:
<taxonomicName id="4C03DA9D5649FFFBC0F8FADBFD8BFAF8" box="[439,546,1358,1382]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="134" phylum="Arthropoda" rank="species" species="kambaiti">Kambaiti</taxonomicName>
,
<quantity id="4CFB0CFB5649FFFBC361FADBFD2BFAF8" box="[558,642,1358,1382]" metricMagnitude="3" metricUnit="m" metricValue="2.1336" pageId="18" pageNumber="119" unit="ft" value="7000.0">7000 ft</quantity>
. (
<geoCoordinate id="EE37C7D95649FFFBC3DCFADBFD5DFAF8" box="[659,756,1358,1382]" direction="north" orientation="latitude" pageId="18" pageNumber="119" precision="555" value="25.4">N25.40º</geoCoordinate>
,
<geoCoordinate id="EE37C7D95649FFFBC3B0FADBFCF5FAF8" box="[767,860,1358,1382]" direction="east" orientation="longitude" pageId="18" pageNumber="119" precision="555" value="98.13">E98.13º</geoCoordinate>
).
</paragraph>
<paragraph id="8BBCA11E5649FFFBC188FAE6FAEFF9F6" blockId="18.[151,1436,1358,2036]" pageId="18" pageNumber="119">
<emphasis id="B9777D0C5649FFFBC188FAE6FE51FA15" bold="true" box="[199,504,1394,1419]" pageId="18" pageNumber="119">
<typeStatus id="54B81FBC5649FFFBC188FAE6FEADFA15" box="[199,260,1395,1419]" pageId="18" pageNumber="119">Type</typeStatus>
materials examined
</emphasis>
.
<emphasis id="B9777D0C5649FFFBC347FAE6FDDCFA15" bold="true" box="[520,629,1395,1419]" pageId="18" pageNumber="119">
<typeStatus id="54B81FBC5649FFFBC347FAE6FDDCFA15" box="[520,629,1395,1419]" pageId="18" pageNumber="119" type="holotype">Holotype</typeStatus>
</emphasis>
of
<taxonomicName id="4C03DA9D5649FFFBC3D2FAE1FC75FA14" authority="Andrewes" authorityName="Andrewes" box="[669,988,1394,1419]" class="Insecta" family="Carabidae" genus="Feronia" higherTaxonomySource="CoL" kingdom="Plantae" order="Coleoptera" pageId="18" pageNumber="119" phylum="Arthropoda" rank="species" species="kambaiti">
<emphasis id="B9777D0C5649FFFBC3D2FAE1FCCBFA15" box="[669,866,1395,1419]" italics="true" pageId="18" pageNumber="119">Feronia kambaiti</emphasis>
Andrewes
</taxonomicName>
(examined by photo): female (NHRS), body length =
<quantity id="4CFB0CFB5649FFFBC007FA03FE1AFA30" box="[328,435,1430,1454]" metricMagnitude="-2" metricUnit="m" metricValue="1.03" pageId="18" pageNumber="119" unit="mm" value="10.3">10.3 mm</quantity>
, board mounted, &quot;N. E.
<collectingCountry id="F314E18E5649FFFBC394FA02FCE8FA31" box="[731,833,1431,1455]" name="Myanmar" pageId="18" pageNumber="119">BURMA</collectingCountry>
/
<taxonomicName id="4C03DA9D5649FFFBC22DFA03FC64FA30" box="[866,973,1430,1454]" class="Insecta" family="Carabidae" genus="Feronia" higherTaxonomySource="CoL" kingdom="Plantae" order="Coleoptera" pageId="18" pageNumber="134" phylum="Arthropoda" rank="species" species="kambaiti">Kambaiti</taxonomicName>
,
<quantity id="4CFB0CFB5649FFFBC292FA03FB9BFA30" box="[989,1074,1430,1454]" metricMagnitude="3" metricUnit="m" metricValue="2.1336" pageId="18" pageNumber="119" unit="ft" value="7000.0">7000 ft</quantity>
/
<emphasis id="B9777D0C5649FFFBC51EFA03FB6EFA30" box="[1105,1223,1430,1454]" italics="true" pageId="18" pageNumber="119">
<date id="FFBD87DE5649FFFBC51EFA03FB6EFA30" box="[1105,1223,1430,1454]" pageId="18" pageNumber="119" value="1934-04-19">19/4 1934</date>
</emphasis>
/ R. MALAISE&quot;; &quot;
<taxonomicName id="4C03DA9D5649FFFBC1EDFA29FF55FA4D" box="[162,252,1468,1491]" class="Insecta" family="Carabidae" genus="Feronia" higherTaxonomySource="CoL" kingdom="Plantae" order="Coleoptera" pageId="18" pageNumber="119" phylum="Arthropoda" rank="genus">
<emphasis id="B9777D0C5649FFFBC1EDFA29FF55FA4D" box="[162,252,1468,1491]" italics="true" pageId="18" pageNumber="119">Feronia</emphasis>
</taxonomicName>
/
<taxonomicName id="4C03DA9D5649FFFBC05CFA2EFED2FA4D" box="[275,379,1467,1491]" class="Insecta" family="Carabidae" genus="Feronia" higherTaxonomySource="CoL" kingdom="Plantae" order="Coleoptera" pageId="18" pageNumber="134" phylum="Arthropoda" rank="species" species="kambaiti">
<emphasis id="B9777D0C5649FFFBC05CFA2EFED2FA4D" box="[275,379,1467,1491]" italics="true" pageId="18" pageNumber="119">Kambaiti</emphasis>
</taxonomicName>
/
<emphasis id="B9777D0C5649FFFBC0DDFA29FDA6FA4D" box="[402,527,1467,1491]" italics="true" pageId="18" pageNumber="119">
<typeStatus id="54B81FBC5649FFFBC0DDFA29FE61FA4D" box="[402,456,1468,1491]" pageId="18" pageNumber="119">Type</typeStatus>
<collectingCountry id="F314E18E5649FFFBC080FA2EFDA2FA4D" box="[463,523,1467,1491]" name="Andorra" pageId="18" pageNumber="119">Andr</collectingCountry>
.
</emphasis>
/ H.E.Andrewes det.&quot;; &quot;
<typeStatus id="54B81FBC5649FFFBC251FA2FFCC0FA4C" box="[798,873,1466,1490]" pageId="18" pageNumber="119">Typus</typeStatus>
&quot; [red label]; &quot;5534 / E91 +&quot; [blue label]; &quot;NHRS- COLE / 000008537&quot;; &quot;
<taxonomicName id="4C03DA9D5649FFFBC0D1FA74FDAAFA67" box="[414,515,1505,1529]" class="Insecta" family="Carabidae" genus="Feronia" higherTaxonomySource="CoL" kingdom="Plantae" order="Coleoptera" pageId="18" pageNumber="119" phylum="Arthropoda" rank="species" species="kambaiti">kambaiti</taxonomicName>
/
<collectingCountry id="F314E18E5649FFFBC351FA74FDF7FA67" box="[542,606,1505,1529]" name="Andorra" pageId="18" pageNumber="119">Andr</collectingCountry>
.&quot; [box label] [
<figureCitation id="1338BD9B5649FFFBC3B2FA4BFC9BFA72" box="[765,818,1502,1516]" captionStart="FIGURES 13 – 18" captionStartId="23.[151,264,1749,1772]" captionTargetBox="[158,1427,204,1715]" captionTargetId="figure@23.[151,1436,193,1728]" captionTargetPageId="23" captionText="FIGURES 13 – 18. Habitus of Pterostichus (Wraseiellus) spp.: scale bar = 2.0 mm; 13, Pt. pseudodiversus n. sp., holotype; 14, Pt. pseudodiversus n. sp., a female paratype from Muli; 15, Pt. pseudodiversus n. sp., a male paratype from Deqin; 16, Pt. pseudodiversus n. sp., a male from Weixi; 17, Pt. kambaiti (Andrewes), holotype [photoed by Dr. J. Bergsten]; 18, Pt. kambaiti (Andrewes), holotype of Pterostichus kambaiti Jedlička." httpUri="https://zenodo.org/record/220506/files/figure.png" pageId="18" pageNumber="119">Figs. 17</figureCitation>
,
<figureCitation id="1338BD9B5649FFFBC274FA4BFCE5FA72" box="[827,844,1502,1516]" captionStart="FIGURES 19 – 28" captionStartId="24.[151,264,1308,1331]" captionTargetBox="[166,1426,208,1270]" captionTargetId="figure@24.[151,1436,193,1299]" captionTargetPageId="24" captionText="FIGURES 19 – 28. Labels of name-bearing type specimen: 19, lectotype of Pterostichus andrewesi Jedlička, 1931; 20, holotype of Pterostichus meyeri Jedlička, 1934; 21, holotype of Feronia crebrepunctata Straneo, 1937; 22, holotype of Pterostichus comatus n. sp.; 23, holotype of Pterostichus crassiapex n. sp.; 24, a syntype of Omaseus diversus Fairmaire, 1886; 25, holotype of Pterostichus stictopleurus cangshanensis n. ssp.; 26, holotype of Pterostichus pseudodiversus n. sp.; 27, holotype of Feronia kambaiti Andrewes, 1947 [photoed by Dr. J. Bergsten]; 28, holotype of Pterostichus kambaiti Jedlička, 1965." httpUri="https://zenodo.org/record/220507/files/figure.png" pageId="18" pageNumber="119">27</figureCitation>
].
<emphasis id="B9777D0C5649FFFBC22FFA77FC64FA64" bold="true" box="[864,973,1506,1530]" pageId="18" pageNumber="119">
<typeStatus id="54B81FBC5649FFFBC22FFA77FC64FA64" box="[864,973,1506,1530]" pageId="18" pageNumber="119" type="holotype">Holotype</typeStatus>
</emphasis>
of
<taxonomicName id="4C03DA9D5649FFFBC2B8FA74FAF1FA67" authority="Jedlicka" authorityName="Jedlicka" box="[1015,1368,1505,1529]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="119" phylum="Arthropoda" rank="species" species="kambaiti">
<emphasis id="B9777D0C5649FFFBC2B8FA74FB46FA67" box="[1015,1263,1505,1529]" italics="true" pageId="18" pageNumber="119">Pterostichus kambaiti</emphasis>
Jedlička
</taxonomicName>
: male (NHRS), body length =
<quantity id="4CFB0CFB5649FFFBC0E2F990FDA1F983" box="[429,520,1541,1565]" metricMagnitude="-3" metricUnit="m" metricValue="9.9" pageId="18" pageNumber="119" unit="mm" value="9.9">9.9 mm</quantity>
, board mounted, genitalia dissected and glued on mounting board together with specimen, &quot;N. E.
<collectingCountry id="F314E18E5649FFFBC028F9BFFE67F9DC" box="[359,462,1578,1602]" name="Myanmar" pageId="18" pageNumber="119">BURMA</collectingCountry>
/
<taxonomicName id="4C03DA9D5649FFFBC0A4F9BCFDFFF9DF" box="[491,598,1577,1601]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="134" phylum="Arthropoda" rank="species" species="kambaiti">Kambaiti</taxonomicName>
,
<quantity id="4CFB0CFB5649FFFBC329F9BCFD14F9DF" box="[614,701,1577,1601]" metricMagnitude="3" metricUnit="m" metricValue="2.1336" pageId="18" pageNumber="119" unit="ft" value="7000.0">7000 ft</quantity>
. /
<emphasis id="B9777D0C5649FFFBC391F9BCFCFBF9DF" box="[734,850,1577,1601]" italics="true" pageId="18" pageNumber="119">
<date id="FFBD87DE5649FFFBC391F9BCFCFBF9DF" box="[734,850,1577,1601]" pageId="18" pageNumber="119" value="1934-04-19">19/4 1934</date>
</emphasis>
/ R. MALAISE&quot;; &quot;
<emphasis id="B9777D0C5649FFFBC574F9BCFBE0F9DF" box="[1083,1097,1577,1601]" italics="true" pageId="18" pageNumber="119">4</emphasis>
&quot;; &quot;
<typeStatus id="54B81FBC5649FFFBC520F9BCFAA0F9DF" box="[1135,1289,1577,1601]" pageId="18" pageNumber="119" type="holotype">HOLOTYPE</typeStatus>
&quot; [red label]; &quot;
<taxonomicName id="4C03DA9D5649FFFBC1EDF9C5FE87F9F6" box="[162,302,1616,1640]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="119" phylum="Arthropoda" rank="genus">
<emphasis id="B9777D0C5649FFFBC1EDF9C5FE87F9F6" box="[162,302,1616,1640]" italics="true" pageId="18" pageNumber="119">Pterostichus</emphasis>
</taxonomicName>
/
<taxonomicName id="4C03DA9D5649FFFBC00BF9C5FE0FF9F6" box="[324,422,1616,1640]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="119" phylum="Arthropoda" rank="species" species="kambaiti">
<emphasis id="B9777D0C5649FFFBC00BF9C5FE0FF9F6" box="[324,422,1616,1640]" italics="true" pageId="18" pageNumber="119">kambaiti</emphasis>
</taxonomicName>
/
<emphasis id="B9777D0C5649FFFBC0F3F9C4FE58F9F6" box="[444,497,1617,1640]" italics="true" pageId="18" pageNumber="119">sp.n.</emphasis>
/ det. Ing. Jedlička&quot; [pink label]; &quot;5679 / E91 +&quot; [blue label] [
<figureCitation id="1338BD9B5649FFFBC5ECF9D8FB7EF9C5" box="[1187,1239,1613,1627]" captionStart="FIGURES 13 – 18" captionStartId="23.[151,264,1749,1772]" captionTargetBox="[158,1427,204,1715]" captionTargetId="figure@23.[151,1436,193,1728]" captionTargetPageId="23" captionText="FIGURES 13 – 18. Habitus of Pterostichus (Wraseiellus) spp.: scale bar = 2.0 mm; 13, Pt. pseudodiversus n. sp., holotype; 14, Pt. pseudodiversus n. sp., a female paratype from Muli; 15, Pt. pseudodiversus n. sp., a male paratype from Deqin; 16, Pt. pseudodiversus n. sp., a male from Weixi; 17, Pt. kambaiti (Andrewes), holotype [photoed by Dr. J. Bergsten]; 18, Pt. kambaiti (Andrewes), holotype of Pterostichus kambaiti Jedlička." httpUri="https://zenodo.org/record/220506/files/figure.png" pageId="18" pageNumber="119">Figs. 18</figureCitation>
,
<figureCitation id="1338BD9B5649FFFBC591F9D8FB46F9C5" box="[1246,1263,1613,1627]" captionStart="FIGURES 19 – 28" captionStartId="24.[151,264,1308,1331]" captionTargetBox="[166,1426,208,1270]" captionTargetId="figure@24.[151,1436,193,1299]" captionTargetPageId="24" captionText="FIGURES 19 – 28. Labels of name-bearing type specimen: 19, lectotype of Pterostichus andrewesi Jedlička, 1931; 20, holotype of Pterostichus meyeri Jedlička, 1934; 21, holotype of Feronia crebrepunctata Straneo, 1937; 22, holotype of Pterostichus comatus n. sp.; 23, holotype of Pterostichus crassiapex n. sp.; 24, a syntype of Omaseus diversus Fairmaire, 1886; 25, holotype of Pterostichus stictopleurus cangshanensis n. ssp.; 26, holotype of Pterostichus pseudodiversus n. sp.; 27, holotype of Feronia kambaiti Andrewes, 1947 [photoed by Dr. J. Bergsten]; 28, holotype of Pterostichus kambaiti Jedlička, 1965." httpUri="https://zenodo.org/record/220507/files/figure.png" pageId="18" pageNumber="119">28</figureCitation>
,
<figureCitation id="1338BD9B5649FFFBC5BAF9D8FAAFF9C5" box="[1269,1286,1613,1627]" captionStart="FIGURES 35 – 40" captionStartId="26.[151,264,1759,1782]" captionTargetBox="[168,1419,197,1735]" captionTargetId="figure@26.[151,1436,193,1738]" captionTargetPageId="26" captionText="FIGURES 35 – 40. Median lobe of male genitalia of Pterostichus (Wraseiellus) spp.: scale bar = 1.0 mm; A, left lateral view of median lobe; B, dorsal view of median lobe; 35, Pt. stictopleurus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 36, Pt. stictopleurus cangshanensis n. ssp., holotype; 37, Pt. pseudodiversus n. sp., holotype; 38, Pt. pseudodiversus n. sp., a paratype from Muli; 39, Pt. pseudodiversus n. sp., a male from Weixi; 40, Pt. kambaiti (Andrewes), holotype of Pterostichus kambaiti Jedlička." httpUri="https://zenodo.org/record/220509/files/figure.png" pageId="18" pageNumber="119">40</figureCitation>
,
<figureCitation id="1338BD9B5649FFFBC443F9D8FAB4F9C5" box="[1292,1309,1613,1627]" captionStart="FIGURES 67 – 84" captionStartId="29.[151,264,1584,1607]" captionTargetBox="[166,1426,201,1530]" captionTargetId="figure@29.[151,1436,193,1545]" captionTargetPageId="29" captionText="FIGURES 67 – 84. Parameres of male genitalia of Pterostichus (Wraseiellus) spp.: scale bar = 0.5 mm; A, left paramere; B, right paramere: 67, Pt. andrewesi Jedlička, a male from Jinfoshan; 68, Pt. andrewesi Jedlička, a paralectotype from Jinfoshan; 69, Pt. meyeri Jedlička, a male from &quot; Yunnan &quot;; 70, Pt. comatus n. sp., holotype; 71, Pt. crassiapex n. sp., holotype; 72, Pt. stictopleurus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 73, Pt. stictopleurus cangshanensis n. ssp., holotype; 74 – 75, Pt. diversus (Fairmaire), males from Yulongshan; 76, Pt. diversus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 77, Pt. diversus (Fairmaire), a male from &quot; Yunnan &quot;; 78, Pt. kambaiti (Andrewes), holotype of Pterostichus kambaiti Jedlička; 79, Pt. pseudodiversus n. sp., holotype; 80, Pt. pseudodiversus n. sp., a paratype from Muli; 81, Pt. pseudodiversus n. sp., a paratype from Zhongdian; 82, Pt. pseudodiversus n. sp., a paratype from Deqin; 83, Pt. pseudodiversus n. sp.., a paratype from Lanping; 84, Pt. pseudodiversus n. sp., a male from Weixi." httpUri="https://zenodo.org/record/220512/files/figure.png" pageId="18" pageNumber="119">78</figureCitation>
,
<figureCitation id="1338BD9B5649FFFBC46CF9D8FA92F9C5" box="[1315,1339,1613,1627]" captionStart="FIGURES 98 – 106" captionStartId="32.[151,264,1472,1495]" captionTargetBox="[155,1435,216,1425]" captionTargetId="figure@32.[151,1436,215,1427]" captionTargetPageId="32" captionText="FIGURES 98 – 106. Pronotum hind angle and basal area (right side) of Pterostichus (Wraseiellus) spp.; scale bar = 0.5 mm; 98, Pt. andrewesi Jedlička, a female from Jinfoshan; 99, Pt. meyeri Jedlička, a male from &quot; Yunnan &quot;; 100, Pt. crassiapex n. sp., holotype; 101, Pt. comatus n. sp., holotype; 102, Pt. diversus (Fairmaire), a male from Yulongshan; 103, Pt. pseudodiversus n. sp., holotype; 104, Pt. stictopleurus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 105, Pt. stictopleurus cangshanensis n. ssp., holotype; 106, Pt. kambaiti (Andrewes), holotype of Pterostichus kambaiti Jedlička." httpUri="https://zenodo.org/record/220515/files/figure.png" pageId="18" pageNumber="119">106</figureCitation>
].
</paragraph>
<paragraph id="8BBCA11E5649FFFBC188F9E0FC22F966" blockId="18.[151,1436,1358,2036]" pageId="18" pageNumber="119">
<emphasis id="B9777D0C5649FFFBC188F9E0FE28F913" bold="true" box="[199,385,1653,1677]" pageId="18" pageNumber="119">
Notes on
<typeStatus id="54B81FBC5649FFFBC00CF9E0FE28F913" box="[323,385,1653,1677]" pageId="18" pageNumber="119">types</typeStatus>
</emphasis>
.
<taxonomicName id="4C03DA9D5649FFFBC0D8F9E1FD47F913" authority="Andrewes" authorityName="Andrewes" box="[407,750,1652,1677]" class="Insecta" family="Carabidae" genus="Feronia" higherTaxonomySource="CoL" kingdom="Plantae" order="Coleoptera" pageId="18" pageNumber="119" phylum="Arthropoda" rank="species" species="kambaiti">
<emphasis id="B9777D0C5649FFFBC0D8F9E1FD47F913" bold="true" box="[407,750,1652,1677]" pageId="18" pageNumber="119">
<emphasis id="B9777D0C5649FFFBC0D8F9E1FDC1F913" bold="true" box="[407,616,1652,1677]" italics="true" pageId="18" pageNumber="119">Feronia kambaiti</emphasis>
Andrewes
</emphasis>
</taxonomicName>
: The original description indicated that this species was published on two females, and in the preface noted that all the &quot;
<typeStatus id="54B81FBC5649FFFBC2D5F90DFC73F92E" box="[922,986,1688,1712]" pageId="18" pageNumber="119">types</typeStatus>
&quot; were in Stockholm Museum and the &quot;
<typeStatus id="54B81FBC5649FFFBC1D0F929FEB9F94A" box="[159,272,1724,1748]" pageId="18" pageNumber="119" type="paratype">paratypes</typeStatus>
&quot; were in the collection of Andrewes and British Museum. So, the one we examined by photo with the help of Dr. J. Bergsten is the
<typeStatus id="54B81FBC5649FFFBC091F975FDE9F966" box="[478,576,1760,1784]" pageId="18" pageNumber="119" type="holotype">holotype</typeStatus>
of this species (
<figureCitation id="1338BD9B5649FFFBC3BAF975FCFBF966" box="[757,850,1760,1784]" captionStart="FIGURES 13 – 18" captionStartId="23.[151,264,1749,1772]" captionTargetBox="[158,1427,204,1715]" captionTargetId="figure@23.[151,1436,193,1728]" captionTargetPageId="23" captionText="FIGURES 13 – 18. Habitus of Pterostichus (Wraseiellus) spp.: scale bar = 2.0 mm; 13, Pt. pseudodiversus n. sp., holotype; 14, Pt. pseudodiversus n. sp., a female paratype from Muli; 15, Pt. pseudodiversus n. sp., a male paratype from Deqin; 16, Pt. pseudodiversus n. sp., a male from Weixi; 17, Pt. kambaiti (Andrewes), holotype [photoed by Dr. J. Bergsten]; 18, Pt. kambaiti (Andrewes), holotype of Pterostichus kambaiti Jedlička." httpUri="https://zenodo.org/record/220506/files/figure.png" pageId="18" pageNumber="119">Figs. 17</figureCitation>
,
<figureCitation id="1338BD9B5649FFFBC211F975FCD7F966" box="[862,894,1760,1784]" captionStart="FIGURES 19 – 28" captionStartId="24.[151,264,1308,1331]" captionTargetBox="[166,1426,208,1270]" captionTargetId="figure@24.[151,1436,193,1299]" captionTargetPageId="24" captionText="FIGURES 19 – 28. Labels of name-bearing type specimen: 19, lectotype of Pterostichus andrewesi Jedlička, 1931; 20, holotype of Pterostichus meyeri Jedlička, 1934; 21, holotype of Feronia crebrepunctata Straneo, 1937; 22, holotype of Pterostichus comatus n. sp.; 23, holotype of Pterostichus crassiapex n. sp.; 24, a syntype of Omaseus diversus Fairmaire, 1886; 25, holotype of Pterostichus stictopleurus cangshanensis n. ssp.; 26, holotype of Pterostichus pseudodiversus n. sp.; 27, holotype of Feronia kambaiti Andrewes, 1947 [photoed by Dr. J. Bergsten]; 28, holotype of Pterostichus kambaiti Jedlička, 1965." httpUri="https://zenodo.org/record/220507/files/figure.png" pageId="18" pageNumber="119">27</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C319F2955649FFFBC188F891FAD5F8DE" pageId="18" pageNumber="119" type="description">
<paragraph id="8BBCA11E5649FFFBC188F891FAD5F8DE" blockId="18.[151,1436,1358,2036]" pageId="18" pageNumber="119">
<taxonomicName id="4C03DA9D5649FFFBC188F891FDAAF883" authority="Jedli" authorityName="Jedli" box="[199,515,1796,1821]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="119" phylum="Arthropoda" rank="species" species="kambaiti">
<emphasis id="B9777D0C5649FFFBC188F891FDAAF883" bold="true" box="[199,515,1796,1821]" pageId="18" pageNumber="119">
<emphasis id="B9777D0C5649FFFBC188F891FE6BF883" bold="true" box="[199,450,1796,1821]" italics="true" pageId="18" pageNumber="119">Pterostichus kambaiti</emphasis>
Jedli
</emphasis>
</taxonomicName>
č
<emphasis id="B9777D0C5649FFFBC340F890FD85F883" bold="true" box="[527,556,1797,1821]" pageId="18" pageNumber="119">ka</emphasis>
: The original description clearly indicated that this species was described on a single male. Therefore, the one we borrowed from NHRS is certainly the
<typeStatus id="54B81FBC5649FFFBC280F8BDFB98F8DE" box="[975,1073,1832,1856]" pageId="18" pageNumber="119" type="holotype">holotype</typeStatus>
of this species (
<figureCitation id="1338BD9B5649FFFBC5A9F8BDFAEDF8DE" box="[1254,1348,1832,1856]" captionStart="FIGURES 13 – 18" captionStartId="23.[151,264,1749,1772]" captionTargetBox="[158,1427,204,1715]" captionTargetId="figure@23.[151,1436,193,1728]" captionTargetPageId="23" captionText="FIGURES 13 – 18. Habitus of Pterostichus (Wraseiellus) spp.: scale bar = 2.0 mm; 13, Pt. pseudodiversus n. sp., holotype; 14, Pt. pseudodiversus n. sp., a female paratype from Muli; 15, Pt. pseudodiversus n. sp., a male paratype from Deqin; 16, Pt. pseudodiversus n. sp., a male from Weixi; 17, Pt. kambaiti (Andrewes), holotype [photoed by Dr. J. Bergsten]; 18, Pt. kambaiti (Andrewes), holotype of Pterostichus kambaiti Jedlička." httpUri="https://zenodo.org/record/220506/files/figure.png" pageId="18" pageNumber="119">Figs. 18</figureCitation>
,
<figureCitation id="1338BD9B5649FFFBC400F8BDFAC6F8DE" box="[1359,1391,1832,1856]" captionStart="FIGURES 19 – 28" captionStartId="24.[151,264,1308,1331]" captionTargetBox="[166,1426,208,1270]" captionTargetId="figure@24.[151,1436,193,1299]" captionTargetPageId="24" captionText="FIGURES 19 – 28. Labels of name-bearing type specimen: 19, lectotype of Pterostichus andrewesi Jedlička, 1931; 20, holotype of Pterostichus meyeri Jedlička, 1934; 21, holotype of Feronia crebrepunctata Straneo, 1937; 22, holotype of Pterostichus comatus n. sp.; 23, holotype of Pterostichus crassiapex n. sp.; 24, a syntype of Omaseus diversus Fairmaire, 1886; 25, holotype of Pterostichus stictopleurus cangshanensis n. ssp.; 26, holotype of Pterostichus pseudodiversus n. sp.; 27, holotype of Feronia kambaiti Andrewes, 1947 [photoed by Dr. J. Bergsten]; 28, holotype of Pterostichus kambaiti Jedlička, 1965." httpUri="https://zenodo.org/record/220507/files/figure.png" pageId="18" pageNumber="119">28</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C319F2955649FFFAC188F8D9FBD1FEDE" lastPageId="19" lastPageNumber="120" pageId="18" pageNumber="119" type="diagnosis">
<paragraph id="8BBCA11E5649FFFBC188F8D9FC80F86A" blockId="18.[151,1436,1358,2036]" pageId="18" pageNumber="119">
<emphasis id="B9777D0C5649FFFBC188F8D9FEE8F8FB" bold="true" box="[199,321,1868,1893]" pageId="18" pageNumber="119">Diagnosis.</emphasis>
In the external characters, this species is most similar to
<emphasis id="B9777D0C5649FFFBC281F8D8FB6FF8FA" box="[974,1222,1868,1892]" italics="true" pageId="18" pageNumber="119">
<taxonomicName id="4C03DA9D5649FFFBC281F8D8FB2EF8FA" box="[974,1159,1868,1892]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="119" phylum="Arthropoda" rank="species" sensu="stricto" species="stictopleurus">Pt. stictopleurus</taxonomicName>
<taxonomicNameLabel id="A244C0775649FFFBC5C0F8D8FB6FF8FA" box="[1167,1222,1869,1892]" pageId="18" pageNumber="119" sensu="stricto">s.str.</taxonomicNameLabel>
</emphasis>
These two species can be distinguished from other members of this subgenus by the combination of the following characters: (1) 5th tarsomere glabrous beneath; (2) males with isodiametric microsculpture on elytra; (3) pronotum with area between outer basal foveal groove and lateral margin slightly convex, not forming a distinct ridge; (4) lateral margins straight before hind angle, hind angles obtuse, not pointed.
</paragraph>
<paragraph id="8BBCA11E5648FFFAC188FF0CFBD1FEDE" blockId="19.[151,1436,151,1221]" pageId="19" pageNumber="120">
<taxonomicName id="4C03DA9D5648FFFAC188FF0CFEF8FF2E" box="[199,337,152,176]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="120" phylum="Arthropoda" rank="species" species="kambaiti">
<emphasis id="B9777D0C5648FFFAC188FF0CFEF8FF2E" box="[199,337,152,176]" italics="true" pageId="19" pageNumber="120">Pt. kambaiti</emphasis>
</taxonomicName>
is different from
<emphasis id="B9777D0C5648FFFAC354FF0CFCBBFF2E" box="[539,786,152,176]" italics="true" pageId="19" pageNumber="120">
<taxonomicName id="4C03DA9D5648FFFAC354FF0CFD7DFF2E" box="[539,724,152,176]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="120" phylum="Arthropoda" rank="species" sensu="stricto" species="stictopleurus">Pt. stictopleurus</taxonomicName>
<taxonomicNameLabel id="A244C0775648FFFAC393FF0CFCBBFF2E" box="[732,786,153,176]" pageId="19" pageNumber="120" sensu="stricto">s.str.</taxonomicNameLabel>
</emphasis>
in: (1) scutellar striae located between 1st and 2nd striae (scutellar striae located between 1st stria and elytral suture in
<emphasis id="B9777D0C5648FFFAC206FF28FB97FF4A" box="[841,1086,188,212]" italics="true" pageId="19" pageNumber="120">
<taxonomicName id="4C03DA9D5648FFFAC206FF28FBA8FF4A" box="[841,1025,188,212]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="120" phylum="Arthropoda" rank="species" sensu="stricto" species="stictopleurus">Pt. stictopleurus</taxonomicName>
<taxonomicNameLabel id="A244C0775648FFFAC548FF28FB97FF4A" box="[1031,1086,189,212]" pageId="19" pageNumber="120" sensu="stricto">s.str.</taxonomicNameLabel>
</emphasis>
); (2) metaepisterna impunctate (distinctly punctate in
<emphasis id="B9777D0C5648FFFAC0D2FF74FD33FF66" box="[413,666,224,248]" italics="true" pageId="19" pageNumber="120">
<taxonomicName id="4C03DA9D5648FFFAC0D2FF74FDF0FF66" box="[413,601,224,248]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="120" phylum="Arthropoda" rank="species" sensu="stricto" species="stictopleurus">Pt. stictopleurus</taxonomicName>
<taxonomicNameLabel id="A244C0775648FFFAC32CFF74FD33FF66" box="[611,666,225,248]" pageId="19" pageNumber="120" sensu="stricto">s.str.</taxonomicNameLabel>
</emphasis>
); (3) body size smaller,
<quantity id="4CFB0CFB5648FFFAC2F1FF4AFBFEFF69" box="[958,1111,223,247]" metricMagnitude="-2" metricUnit="m" metricValue="1.01" metricValueMax="1.03" metricValueMin="0.99" pageId="19" pageNumber="120" unit="mm" value="10.1" valueMax="10.3" valueMin="9.9">9.9–10.3 mm</quantity>
(usually much larger in
<emphasis id="B9777D0C5648FFFAC433FF74FEC0FE82" italics="true" pageId="19" pageNumber="120">
<taxonomicName id="4C03DA9D5648FFFAC433FF74FE81FE82" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="120" phylum="Arthropoda" rank="species" sensu="stricto" species="stictopleurus">Pt. stictopleurus</taxonomicName>
<taxonomicNameLabel id="A244C0775648FFFAC07DFE90FEC0FE82" box="[306,361,261,284]" pageId="19" pageNumber="120" sensu="stricto">s.str.</taxonomicNameLabel>
</emphasis>
,
<quantity id="4CFB0CFB5648FFFAC034FE91FD8CFE82" box="[379,549,260,284]" metricMagnitude="-2" metricUnit="m" metricValue="1.1949999999999998" metricValueMax="1.31" metricValueMin="1.08" pageId="19" pageNumber="120" unit="mm" value="11.95" valueMax="13.1" valueMin="10.8">10.8–13.1 mm</quantity>
); (4) right paramere much wider than in
<emphasis id="B9777D0C5648FFFAC545FE90FAAEFE82" box="[1034,1287,260,284]" italics="true" pageId="19" pageNumber="120">
<taxonomicName id="4C03DA9D5648FFFAC545FE90FB6CFE82" box="[1034,1221,260,284]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="120" phylum="Arthropoda" rank="species" sensu="stricto" species="stictopleurus">Pt. stictopleurus</taxonomicName>
<taxonomicNameLabel id="A244C0775648FFFAC59FFE90FAAEFE82" box="[1232,1287,261,284]" pageId="19" pageNumber="120" sensu="stricto">s.str.</taxonomicNameLabel>
</emphasis>
; (5) apex of median lobe of male genitalia more bent to the dorsal side than in.
<emphasis id="B9777D0C5648FFFAC2CCFEBCFBD1FEDE" box="[899,1144,296,320]" italics="true" pageId="19" pageNumber="120">
<taxonomicName id="4C03DA9D5648FFFAC2CCFEBCFB93FEDE" box="[899,1082,296,320]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="120" phylum="Arthropoda" rank="species" sensu="stricto" species="stictopleurus">Pt. stictopleurus</taxonomicName>
<taxonomicNameLabel id="A244C0775648FFFAC50EFEBCFBD1FEDE" box="[1089,1144,297,320]" pageId="19" pageNumber="120" sensu="stricto">s.str.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C319F2955648FFFAC188FEDEFC89FB91" pageId="19" pageNumber="120" type="description">
<paragraph id="8BBCA11E5648FFFAC188FEDEFC89FB91" blockId="19.[151,1436,151,1221]" pageId="19" pageNumber="120">
<emphasis id="B9777D0C5648FFFAC188FEDEFEF8FEFA" bold="true" box="[199,337,331,356]" pageId="19" pageNumber="120">Description</emphasis>
. Body length
<quantity id="4CFB0CFB5648FFFAC0B9FED9FD39FEFA" box="[502,656,332,356]" metricMagnitude="-2" metricUnit="m" metricValue="1.01" metricValueMax="1.03" metricValueMin="0.99" pageId="19" pageNumber="120" unit="mm" value="10.1" valueMax="10.3" valueMin="9.9">9.9–10.3 mm</quantity>
; dorsal side black, elytra somewhat lighter, shining in males. Males with elytral microsculpture isodiametric.
<emphasis id="B9777D0C5648FFFAC33FFEE5FD19FE16" bold="true" box="[624,688,368,392]" pageId="19" pageNumber="120">Head</emphasis>
. Frons and vertex impunctate, a few punctures present in frontal furrows; antennae reaching elytra basal eighth; 3rd antennomere glabrous except apical setae.
<emphasis id="B9777D0C5648FFFAC5F6FE01FA9BFE32" bold="true" box="[1209,1330,404,428]" pageId="19" pageNumber="120">Pronotum</emphasis>
rounded, widest at about middle, PW/PL = 1.30; lateral margins fully rounded near the middle and then gradually curved posteriorly, almost straight before hind angles; hind angles obtuse, not pointed; one mid-lateral seta present at about anterior fourth; basal fovea shallow, faintly defined; inner groove of basal fovea weak, subparallel to median line, gradually disappeared posteriorly, ended at a distance from the posterior margin of pronotum; outer groove of basal fovea about half length as inner one, somewhat more distinct than the inner one, reaching the basal margin of pronotum; ridge between outer groove and lateral margin slightly convex; area between inner and outer basal foveal grooves hardly convex; some faint punctures present in basal fovea area and the area between the two basal foveae.
<emphasis id="B9777D0C5648FFFAC1BCFD21FE96FD52" bold="true" box="[243,319,692,716]" pageId="19" pageNumber="120">Elytra</emphasis>
with base slightly depressed between 3rd and 5th intervals; elytral shoulders slightly narrowed, humeral teeth small; intervals slightly convex; striae distinct, without punctures; scutellar striae nearly complete, located between 1st and 2nd striae; umbilical pore series on 9th interval sparse in the middle, composed of 15 pores (5, 2, 8).
<emphasis id="B9777D0C5648FFFAC043FCB5FE0BFCA6" bold="true" box="[268,418,799,824]" pageId="19" pageNumber="120">Ventral side</emphasis>
. Proepisterna and metaepisterna impunctate; terminal sternum of male without special structure.
<emphasis id="B9777D0C5648FFFAC046FCD1FEE8FCC2" bold="true" box="[265,321,836,860]" pageId="19" pageNumber="120">Legs</emphasis>
. 5th tarsomere without seta beneath.
<emphasis id="B9777D0C5648FFFAC3BDFCD1FC36FCC2" bold="true" box="[754,927,835,860]" pageId="19" pageNumber="120">Male genitalia</emphasis>
. Median lobe of male genitalia bent at a 90 degrees angle at about basal two-fifth; ventral margin almost straight in the middle, apical part slightly bent to the ventral side and then the apical lamella gradually bent to the dorsal side (
<figureCitation id="1338BD9B5648FFFAC2ADFC19FB9EFC3A" box="[994,1079,908,932]" captionStart="FIGURES 35 – 40" captionStartId="26.[151,264,1759,1782]" captionTargetBox="[168,1419,197,1735]" captionTargetId="figure@26.[151,1436,193,1738]" captionTargetPageId="26" captionText="FIGURES 35 – 40. Median lobe of male genitalia of Pterostichus (Wraseiellus) spp.: scale bar = 1.0 mm; A, left lateral view of median lobe; B, dorsal view of median lobe; 35, Pt. stictopleurus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 36, Pt. stictopleurus cangshanensis n. ssp., holotype; 37, Pt. pseudodiversus n. sp., holotype; 38, Pt. pseudodiversus n. sp., a paratype from Muli; 39, Pt. pseudodiversus n. sp., a male from Weixi; 40, Pt. kambaiti (Andrewes), holotype of Pterostichus kambaiti Jedlička." httpUri="https://zenodo.org/record/220509/files/figure.png" pageId="19" pageNumber="120">Fig. 40</figureCitation>
A); apical orifice large, slightly turned to the left side, apical lamella twisted to the left (
<figureCitation id="1338BD9B5648FFFAC252FC3AFCDAFC59" box="[797,883,943,967]" captionStart="FIGURES 35 – 40" captionStartId="26.[151,264,1759,1782]" captionTargetBox="[168,1419,197,1735]" captionTargetId="figure@26.[151,1436,193,1738]" captionTargetPageId="26" captionText="FIGURES 35 – 40. Median lobe of male genitalia of Pterostichus (Wraseiellus) spp.: scale bar = 1.0 mm; A, left lateral view of median lobe; B, dorsal view of median lobe; 35, Pt. stictopleurus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 36, Pt. stictopleurus cangshanensis n. ssp., holotype; 37, Pt. pseudodiversus n. sp., holotype; 38, Pt. pseudodiversus n. sp., a paratype from Muli; 39, Pt. pseudodiversus n. sp., a male from Weixi; 40, Pt. kambaiti (Andrewes), holotype of Pterostichus kambaiti Jedlička." httpUri="https://zenodo.org/record/220509/files/figure.png" pageId="19" pageNumber="120">Fig. 40</figureCitation>
B); apical lamella laminar, about 1.5 times as its basal width, apex rounded. Right paramere short, apical part triangular and curved, narrowed apically, apex moderately wide (
<figureCitation id="1338BD9B5648FFFAC02DFC62FE1CFB91" box="[354,437,1015,1039]" captionStart="FIGURES 67 – 84" captionStartId="29.[151,264,1584,1607]" captionTargetBox="[166,1426,201,1530]" captionTargetId="figure@29.[151,1436,193,1545]" captionTargetPageId="29" captionText="FIGURES 67 – 84. Parameres of male genitalia of Pterostichus (Wraseiellus) spp.: scale bar = 0.5 mm; A, left paramere; B, right paramere: 67, Pt. andrewesi Jedlička, a male from Jinfoshan; 68, Pt. andrewesi Jedlička, a paralectotype from Jinfoshan; 69, Pt. meyeri Jedlička, a male from &quot; Yunnan &quot;; 70, Pt. comatus n. sp., holotype; 71, Pt. crassiapex n. sp., holotype; 72, Pt. stictopleurus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 73, Pt. stictopleurus cangshanensis n. ssp., holotype; 74 – 75, Pt. diversus (Fairmaire), males from Yulongshan; 76, Pt. diversus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 77, Pt. diversus (Fairmaire), a male from &quot; Yunnan &quot;; 78, Pt. kambaiti (Andrewes), holotype of Pterostichus kambaiti Jedlička; 79, Pt. pseudodiversus n. sp., holotype; 80, Pt. pseudodiversus n. sp., a paratype from Muli; 81, Pt. pseudodiversus n. sp., a paratype from Zhongdian; 82, Pt. pseudodiversus n. sp., a paratype from Deqin; 83, Pt. pseudodiversus n. sp.., a paratype from Lanping; 84, Pt. pseudodiversus n. sp., a male from Weixi." httpUri="https://zenodo.org/record/220512/files/figure.png" pageId="19" pageNumber="120">Fig. 78</figureCitation>
B). Female genitalia not studied.
</paragraph>
</subSubSection>
<subSubSection id="C319F2955648FFFAC188FB8EFC8CFB5A" pageId="19" pageNumber="120" type="distribution">
<paragraph id="8BBCA11E5648FFFAC188FB8EFBF2FBAA" blockId="19.[151,1436,151,1221]" box="[199,1115,1051,1077]" pageId="19" pageNumber="120">
<emphasis id="B9777D0C5648FFFAC188FB8EFEF0FBAA" bold="true" box="[199,345,1051,1076]" pageId="19" pageNumber="120">Distribution</emphasis>
. Only known from the
<typeStatus id="54B81FBC5648FFFAC32FFB88FD39FBAB" box="[608,656,1053,1077]" pageId="19" pageNumber="120">type</typeStatus>
locality,
<taxonomicName id="4C03DA9D5648FFFAC3B6FB89FCCDFBAA" box="[761,868,1052,1076]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="134" phylum="Arthropoda" rank="species" species="kambaiti">Kambaiti</taxonomicName>
,
<collectingCountry id="F314E18E5648FFFAC23FFB88FC74FBAB" box="[880,989,1053,1077]" name="Myanmar" pageId="19" pageNumber="120">Myanmar</collectingCountry>
(
<figureCitation id="1338BD9B5648FFFAC2A3FB89FBE7FBAA" box="[1004,1102,1052,1076]" captionStart="FIGURES 110 – 111" captionStartId="33.[151,264,1885,1908]" captionTargetBox="[224,1312,207,1827]" captionTargetId="figure@33.[224,1343,193,1844]" captionTargetPageId="33" captionText="FIGURES 110 – 111. Distribution maps for species of subgenus Pterostichus (Wraseiellus): - Pt. andrewesi Jedlička; « Pt. meyeri Jedlička; ˑ Pt. crassiapex n. sp.; ¿ Pt. comatus n. sp.; ● Pt. diversus (Fairmaire); Ο Pt. pseudodiversus n. sp.; ˔ Pt. stictopleurus stictopleurus (Fairmaire); r Pt. stictopleurus cangshanensis n. ssp.; ․ Pt. kambaiti (Andrewes). 111. shows an enlarged scale of the area in figure 110 marked by the dotted line square." httpUri="https://zenodo.org/record/220516/files/figure.png" pageId="19" pageNumber="120">Fig. 110</figureCitation>
).
</paragraph>
<paragraph id="8BBCA11E5648FFFAC188FBD5FC8CFB5A" blockId="19.[151,1436,151,1221]" pageId="19" pageNumber="120">
<emphasis id="B9777D0C5648FFFAC188FBD5FE0DFBC6" bold="true" box="[199,420,1088,1112]" pageId="19" pageNumber="120">Notes on synonym</emphasis>
. From the collecting data label, the
<typeStatus id="54B81FBC5648FFFAC20CFBAAFC19FBC9" box="[835,944,1087,1111]" pageId="19" pageNumber="120" type="holotype">holotypes</typeStatus>
of
<taxonomicName id="4C03DA9D5648FFFAC297FBD4FAA1FBC9" authority="Jedlicka 1965" authorityName="Jedlicka" authorityYear="1965" box="[984,1288,1087,1112]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="120" phylum="Arthropoda" rank="species" species="kambaiti">
<emphasis id="B9777D0C5648FFFAC297FBD4FBCAFBC6" box="[984,1123,1088,1112]" italics="true" pageId="19" pageNumber="120">Pt. kambaiti</emphasis>
Jedlička 1965
</taxonomicName>
and
<taxonomicName id="4C03DA9D5648FFFAC40DFBD4FE1DFBE2" authority="Andrewes 1947" authorityName="Andrewes" authorityYear="1947" class="Insecta" family="Carabidae" genus="Feronia" higherTaxonomySource="CoL" kingdom="Plantae" order="Coleoptera" pageId="19" pageNumber="120" phylum="Arthropoda" rank="species" species="kambaiti">
<emphasis id="B9777D0C5648FFFAC40DFBD4FF50FBE2" italics="true" pageId="19" pageNumber="120">Feronia kambaiti</emphasis>
Andrewes 1947
</taxonomicName>
were collected at the same day in the locality of
<taxonomicName id="4C03DA9D5648FFFAC2A6FBF1FBF8FBE2" box="[1001,1105,1124,1148]" class="Insecta" family="Carabidae" genus="Feronia" higherTaxonomySource="CoL" kingdom="Plantae" order="Coleoptera" pageId="19" pageNumber="134" phylum="Arthropoda" rank="species" species="kambaiti">Kambaiti</taxonomicName>
by Malaise. We did not find any significant difference between the two
<typeStatus id="54B81FBC5648FFFAC331FB12FD46FB01" box="[638,751,1159,1183]" pageId="19" pageNumber="120" type="holotype">holotypes</typeStatus>
. So, there is no doubt that
<taxonomicName id="4C03DA9D5648FFFAC553FB1CFAE1FB01" authority="Jedlicka 1965" authorityName="Jedlicka" authorityYear="1965" box="[1052,1352,1159,1184]" class="Insecta" family="Carabidae" genus="Feronia" higherTaxonomySource="CoL" kingdom="Plantae" order="Coleoptera" pageId="19" pageNumber="120" phylum="Arthropoda" rank="species" species="kambaiti">
<emphasis id="B9777D0C5648FFFAC553FB1CFB0CFB3E" box="[1052,1189,1160,1184]" italics="true" pageId="19" pageNumber="120">Pt. kambaiti</emphasis>
Jedlička 1965
</taxonomicName>
is both synonym and homonym of
<taxonomicName id="4C03DA9D5648FFFAC084FB38FC88FB5A" authority="Andrewes 1947" authorityName="Andrewes" authorityYear="1947" box="[459,801,1196,1220]" class="Insecta" family="Carabidae" genus="Feronia" higherTaxonomySource="CoL" kingdom="Plantae" order="Coleoptera" pageId="19" pageNumber="120" phylum="Arthropoda" rank="species" species="kambaiti">
<emphasis id="B9777D0C5648FFFAC084FB38FDFDFB5A" box="[459,596,1196,1220]" italics="true" pageId="19" pageNumber="120">Pt. kambaiti</emphasis>
(Andrewes 1947)
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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