treatments-xml/data/03/A8/21/03A82104FF8BFF91FF0AF9F194267ABB.xml
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<document id="60929FA37C335651E02E629B3BFB42F7" ID-CLB-Dataset="5593" ID-DOI="10.11646/zootaxa.3797.1.13" ID-GBIF-Dataset="f4cb3f8a-03a4-4173-acef-e13274ef48ad" ID-ISSN="1175-5326" ID-Zenodo-Dep="4915425" ID-ZooBank="058AE196-A5DE-480D-BE32-ED4E81DC2ABD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1623199967774" checkinUser="felipe" docAuthor="Lombardo, Francesco, Stiewe, Martin B. D., Ippolito, Salvatrice &amp; Marletta, Alessandro" docDate="2014" docId="03A82104FF8BFF91FF0AF9F194267ABB" docLanguage="en" docName="zt03797p193.pdf.imf" docOrigin="Zootaxa 3797 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Otomantis minima Lombardo &amp; Stiewe &amp; Ippolito &amp; Marletta 2014, sp. n." docType="treatment" docVersion="8" lastPageNumber="185" masterDocId="FF91597CFF84FF81FF9DFFB1902F7E7E" masterDocTitle="A taxonomic revision of Otomantis Bolivar, 1890 (Mantodea: Hymenopodidae, Acromantinae) with description of five new species" masterLastPageNumber="193" masterPageNumber="169" pageNumber="184" updateTime="1699007449450" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="0639577D5B73195E83F57FCE8AD66664">A taxonomic revision of Otomantis Bolivar, 1890 (Mantodea: Hymenopodidae, Acromantinae) with description of five new species</mods:title>
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<mods:namePart id="E96CDE987F76331879F10A31ACECF83D">Lombardo, Francesco</mods:namePart>
<mods:affiliation id="1FB9D3F78183BCB489F1EEE37F94BD01">University of Catania, Department of Biological, Geological and Environmental Sciences, Section of Animal Biology, Via Androne 81, Catania 95124, Italy. Email: lombafra @ unict. it</mods:affiliation>
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<mods:namePart id="CDC50DD6B5F76D3F25B7AD20926659E3">Stiewe, Martin B. D.</mods:namePart>
<mods:affiliation id="6300418B84A1BD9E49391D5316728CD0">The Natural History Museum, London, Scientific Associate, Department of Entomology, Cromwell Road, SW 7 5 BD, London, UK. Email: m. stiewe @ nhm. ac. uk</mods:affiliation>
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<mods:namePart id="EA4655D55F5339E404B9CE889CFF6E95">Ippolito, Salvatrice</mods:namePart>
<mods:affiliation id="258B757CBCAFEFA89738882EBFCF21AA">University of Catania, Department of Biological, Geological and Environmental Sciences, Section of Animal Biology, Via Androne 81, Catania 95124, Italy. Email: lombafra @ unict. it</mods:affiliation>
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<mods:namePart id="FAF5F01F3E8985729B2DB98693F82CB8">Marletta, Alessandro</mods:namePart>
<mods:affiliation id="0A8367C3A1F62AF778CD8E8BA1D058E8">University of Catania, Department of Biological, Geological and Environmental Sciences, Section of Animal Biology, Via Androne 81, Catania 95124, Italy. Email: lombafra @ unict. it</mods:affiliation>
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<mods:number id="864E0192BB617BE9A40EF608BFA6F66E">3797</mods:number>
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<subSubSection id="C31BC399FF8BFF8EFF0AF9F192377824" box="[151,536,1600,1626]" pageId="15" pageNumber="184" type="nomenclature">
<paragraph id="8BBE9012FF8BFF8EFF0AF9F192377824" blockId="15.[151,536,1600,1660]" box="[151,536,1600,1626]" pageId="15" pageNumber="184">
<heading id="D0F6277EFF8BFF8EFF0AF9F192377824" box="[151,536,1600,1626]" level="2" pageId="15" pageNumber="184">
<taxonomicName id="4C01EB91FF8BFF8EFF0AF9F191E07824" ID-CoL="4B7HV" authority="Stiewe" authorityName="Lombardo &amp; Stiewe &amp; Ippolito &amp; Marletta" authorityYear="2014" box="[151,463,1600,1626]" class="Insecta" family="Hymenopodidae" genus="Otomantis" kingdom="Animalia" order="Mantodea" pageId="15" pageNumber="184" phylum="Arthropoda" rank="species" species="minima" status="sp. nov.">
<emphasis id="B9754C00FF8BFF8EFF0AF9F191567824" bold="true" box="[151,377,1600,1626]" italics="true" pageId="15" pageNumber="184">Otomantis minima</emphasis>
Stiewe
</taxonomicName>
<emphasis id="B9754C00FF8BFF8EFE4BF9F192377824" bold="true" box="[470,536,1600,1626]" pageId="15" pageNumber="184">
<taxonomicNameLabel id="A246F17BFF8BFF8EFE4BF9F192377824" box="[470,536,1600,1626]" pageId="15" pageNumber="184" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31BC399FF8BFF8EFF0AF9D2947678BA" pageId="15" pageNumber="184" type="description">
<paragraph id="8BBE9012FF8BFF8EFF0AF9D2919C7802" blockId="15.[151,536,1600,1660]" box="[151,435,1635,1660]" pageId="15" pageNumber="184">
(
<figureCitation id="133A8C97FF8BFF8EFF02F9D290D77802" box="[159,248,1635,1660]" captionStart="FIGURE 2" captionStartId="4.[151,250,1506,1528]" captionTargetBox="[151,1436,411,1482]" captionTargetId="figure-290@4.[151,1436,411,1482]" captionText="FIGURE 2. Head of: A (♂), G (♀), O. scutigera; B (♂), H (♀), O. aurita; C (♂), I (♀), O. rendalli; D (♂), L (♀), O. casaica; E (♂), M (♀), O. bolivari sp. n.; F (♂), O. minima sp. n., Scale = 1mm." figureDoi="http://doi.org/10.5281/zenodo.5083266" httpUri="https://zenodo.org/record/5083266/files/figure.png" pageId="15" pageNumber="184">Figs 2F</figureCitation>
; 3F; 5E; 7C; 13)
</paragraph>
<paragraph id="8BBE9012FF8BFF8EFF0AF91A947678BA" blockId="15.[151,1437,1707,1984]" box="[151,1113,1707,1732]" pageId="15" pageNumber="184">
<emphasis id="B9754C00FF8BFF8EFF0AF91A916A78BA" bold="true" box="[151,325,1707,1732]" inLineHeading="true" pageId="15" pageNumber="184" reason="1">Type material.</emphasis>
<typeStatus id="54BA2EB0FF8BFF8EFECEF91A919478BA" box="[339,443,1707,1732]" pageId="15" pageNumber="184" type="holotype">Holotype</typeStatus>
<specimenCount id="9D075B9BFF8BFF8EFE5FF91A91C978BA" box="[450,486,1707,1732]" pageId="15" pageNumber="184" type="male">1♂</specimenCount>
, AFRICA-S: Stiewe genital slide Nr. OT.SA.03 (NHM)
</paragraph>
</subSubSection>
<subSubSection id="C31BC399FF8BFF8EFF5AF97E92BD79BE" pageId="15" pageNumber="184" type="diagnosis">
<paragraph id="8BBE9012FF8BFF8EFF5AF97E92BD79BE" blockId="15.[151,1437,1707,1984]" pageId="15" pageNumber="184">
<emphasis id="B9754C00FF8BFF8EFF5AF97E916E7896" bold="true" box="[199,321,1743,1768]" inLineHeading="true" pageId="15" pageNumber="184" reason="1">Diagnosis.</emphasis>
This new species
<taxonomicName id="4C01EB91FF8BFF8EFD8AF96192BF7899" authority="Lombardo &amp; Stiewe &amp; Ippolito &amp; Marletta, 2014" authorityName="Lombardo &amp; Stiewe &amp; Ippolito &amp; Marletta" authorityYear="2014" box="[535,656,1744,1767]" class="Insecta" family="Hymenopodidae" genus="Otomantis" kingdom="Animalia" order="Mantodea" pageId="15" pageNumber="184" phylum="Arthropoda" rank="species" species="minima" status="sp. nov.">
<emphasis id="B9754C00FF8BFF8EFD8AF96192BF7899" box="[535,656,1744,1767]" italics="true" pageId="15" pageNumber="184">O. minima</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A246F17BFF8BFF8EFD04F96192F77899" box="[665,728,1744,1767]" pageId="15" pageNumber="184" rank="species">sp. n.</taxonomicNameLabel>
represents the smallest member of the genus. The copulatory apparatus of this new species resembles that of
<taxonomicName id="4C01EB91FF8BFF8EFD51F94293667972" box="[716,841,1779,1804]" class="Insecta" family="Hymenopodidae" genus="Otomantis" kingdom="Animalia" order="Mantodea" pageId="15" pageNumber="184" phylum="Arthropoda" rank="species" species="rendalli">
<emphasis id="B9754C00FF8BFF8EFD51F94293667972" box="[716,841,1779,1804]" italics="true" pageId="15" pageNumber="184">O. rendalli</emphasis>
</taxonomicName>
in regards of the shape of the ventral phallomere. However, there are some striking differences between these two species. The ventral phallomere is strongly extended distally in
<taxonomicName id="4C01EB91FF8BFF8EFEE7F88A91DF792D" authority="Lombardo &amp; Stiewe &amp; Ippolito &amp; Marletta, 2014" authorityName="Lombardo &amp; Stiewe &amp; Ippolito &amp; Marletta" authorityYear="2014" box="[378,496,1851,1875]" class="Insecta" family="Hymenopodidae" genus="Otomantis" kingdom="Animalia" order="Mantodea" pageId="15" pageNumber="184" phylum="Arthropoda" rank="species" species="minima" status="sp. nov.">
<emphasis id="B9754C00FF8BFF8EFEE7F88A91DF792D" box="[378,496,1851,1875]" italics="true" pageId="15" pageNumber="184">O. minima</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A246F17BFF8BFF8EFE6AF88A921A792C" box="[503,565,1851,1874]" pageId="15" pageNumber="184" rank="species">sp. n.</taxonomicNameLabel>
, which is not the case in
<taxonomicName id="4C01EB91FF8BFF8EFCCDF88A93E5792A" box="[848,970,1851,1876]" class="Insecta" family="Hymenopodidae" genus="Otomantis" kingdom="Animalia" order="Mantodea" pageId="15" pageNumber="184" phylum="Arthropoda" rank="species" species="rendalli">
<emphasis id="B9754C00FF8BFF8EFCCDF88A93E5792A" box="[848,970,1851,1876]" italics="true" pageId="15" pageNumber="184">O. rendalli</emphasis>
</taxonomicName>
, the lateral process of the left phallomere is conspicuously long and developed in
<taxonomicName id="4C01EB91FF8BFF8EFDC8F8D192E47909" authority="Lombardo &amp; Stiewe &amp; Ippolito &amp; Marletta, 2014" authorityName="Lombardo &amp; Stiewe &amp; Ippolito &amp; Marletta" authorityYear="2014" box="[597,715,1888,1911]" class="Insecta" family="Hymenopodidae" genus="Otomantis" kingdom="Animalia" order="Mantodea" pageId="15" pageNumber="184" phylum="Arthropoda" rank="species" species="minima" status="sp. nov.">
<emphasis id="B9754C00FF8BFF8EFDC8F8D192E47909" box="[597,715,1888,1911]" italics="true" pageId="15" pageNumber="184">O. minima</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A246F17BFF8BFF8EFD4CF8D193217909" box="[721,782,1888,1911]" pageId="15" pageNumber="184" rank="species">sp. n.</taxonomicNameLabel>
, and its pronotum is much shorter than in rendalli. Finally, the mesofemora of
<taxonomicName id="4C01EB91FF8BFF8EFEE9F83291C179E2" box="[372,494,1923,1948]" class="Insecta" family="Hymenopodidae" genus="Otomantis" kingdom="Animalia" order="Mantodea" pageId="15" pageNumber="184" phylum="Arthropoda" rank="species" species="rendalli">
<emphasis id="B9754C00FF8BFF8EFEE9F83291C179E2" box="[372,494,1923,1948]" italics="true" pageId="15" pageNumber="184">O. rendalli</emphasis>
</taxonomicName>
has a well-developed distal lobe, whereas the same is only indicated in
<taxonomicName id="4C01EB91FF8BFF8EFAB9F83295B379E5" authority="Lombardo &amp; Stiewe &amp; Ippolito &amp; Marletta, 2014" authorityName="Lombardo &amp; Stiewe &amp; Ippolito &amp; Marletta" authorityYear="2014" box="[1316,1436,1923,1947]" class="Insecta" family="Hymenopodidae" genus="Otomantis" kingdom="Animalia" order="Mantodea" pageId="15" pageNumber="184" phylum="Arthropoda" rank="species" species="minima" status="sp. nov.">
<emphasis id="B9754C00FF8BFF8EFAB9F83295B379E5" box="[1316,1436,1923,1947]" italics="true" pageId="15" pageNumber="184">O. minima</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A246F17BFF8BFF8EFF0AF81990FC79C1" box="[151,211,1960,1983]" pageId="15" pageNumber="184" rank="species">sp. n.</taxonomicNameLabel>
The female is unknown for the species.
</paragraph>
</subSubSection>
<subSection id="E28E89F9FF94FF91FF0AFF2694267ABB" pageId="16" pageNumber="185" type="multiple">
<subSubSection id="C31BC399FF94FF91FF0AFF2691AF7F8B" pageId="16" pageNumber="185" type="description">
<paragraph id="8BBE9012FF94FF91FF0AFF2691467ECE" blockId="16.[151,1437,151,1112]" box="[151,361,151,176]" pageId="16" pageNumber="185">
<emphasis id="B9754C00FF94FF91FF0AFF2691467ECE" bold="true" box="[151,361,151,176]" inLineHeading="true" pageId="16" pageNumber="185" reason="1">Description male.</emphasis>
</paragraph>
<paragraph id="8BBE9012FF94FF91FF0AFF0D90F27E86" blockId="16.[151,1437,151,1112]" pageId="16" pageNumber="185">General coloration: Body lutescent, without dark spots. Head dark brown, with some blackish spots. Antennae ochre.</paragraph>
<paragraph id="8BBE9012FF94FF91FF5AFEB595517F63" blockId="16.[151,1437,151,1112]" box="[199,1406,260,285]" pageId="16" pageNumber="185">Measurements (mm): TL12.7; HW 4.1; PL 3.8; SDW 2.2: MPW 1.25; CL 3.5; FL 4.6; MFW 2.0; WL 14.7.</paragraph>
<paragraph id="8BBE9012FF94FF91FF5AFE9692A37FF6" blockId="16.[151,1437,151,1112]" pageId="16" pageNumber="185">
Head (
<figureCitation id="133A8C97FF94FF91FE8BFE96915E7F3E" box="[278,369,295,320]" captionStart="FIGURE 2" captionStartId="4.[151,250,1506,1528]" captionTargetBox="[151,1436,411,1482]" captionTargetId="figure-290@4.[151,1436,411,1482]" captionText="FIGURE 2. Head of: A (♂), G (♀), O. scutigera; B (♂), H (♀), O. aurita; C (♂), I (♀), O. rendalli; D (♂), L (♀), O. casaica; E (♂), M (♀), O. bolivari sp. n.; F (♂), O. minima sp. n., Scale = 1mm." figureDoi="http://doi.org/10.5281/zenodo.5083266" httpUri="https://zenodo.org/record/5083266/files/figure.png" pageId="16" pageNumber="185">Fig. 2F</figureCitation>
): Transverse, vertex nearly straight, vertical process of vertex small triangular, juxta-ocular tubercles conical; eyes kidney-shaped with a distinct short apical spine; frontal shield transversal about 2.5 times wider than high, apex arched and acuminate.
</paragraph>
<paragraph id="8BBE9012FF94FF91FF0AFE2591AF7F8B" blockId="16.[151,1437,151,1112]" pageId="16" pageNumber="185">
Pronotum (
<figureCitation id="133A8C97FF94FF91FE8AFE2591407FD3" box="[279,367,404,429]" captionStart="FIGURE 3" captionStartId="5.[151,250,1201,1223]" captionTargetBox="[151,1436,327,1187]" captionTargetId="figure-396@5.[151,1436,327,1187]" captionTargetPageId="5" captionText="FIGURE 3. Male pronotum of: A (dorsal view), G (lateral view), O. scutigera; B (dorsal view), I (lateral view), O. aurita; C (dorsal view), O. rendalli; D (dorsal view), H (lateral view), O. casaica; E (dorsal view), L (lateral view), O. bolivari sp. n.; F (dorsal view) O. minima sp. n., Scale = 1mm." figureDoi="http://doi.org/10.5281/zenodo.4915431" httpUri="https://zenodo.org/record/4915431/files/figure.png" pageId="16" pageNumber="185">Fig. 3F</figureCitation>
): Relatively small, 1.7 times longer than wide; prozone narrow and elongated, disc with two indicated bumps; metazone with a longitudinal median carina; lateral margins of supracoxal dilation with some small dark tubercles.
</paragraph>
</subSubSection>
<subSubSection id="C31BC399FF94FF91FF5AFE4E91697C8E" pageId="16" pageNumber="185" type="materials_examined">
<paragraph id="8BBE9012FF94FF91FF5AFE4E92F07CFD" blockId="16.[151,1437,151,1112]" pageId="16" pageNumber="185">
Forelegs: Coxae with anterior margin with some very small teeth; internal apical lobes divergent. Femora flattened (
<figureCitation id="133A8C97FF94FF91FE91FD9591487C43" box="[268,359,548,573]" captionStart="FIGURE 5" captionStartId="7.[151,250,1460,1482]" captionTargetBox="[151,1436,807,1435]" captionTargetId="figure-413@7.[151,1436,807,1435]" captionTargetPageId="7" captionText="FIGURE 5. Inner surface of fore femur (males): A, O. scutigera; B, O. aurita; C, O. rendalli; D, O. casaica; E, O. minima sp. n.; F, O. bolivari sp. n., Scale = 1mm." figureDoi="http://doi.org/10.5281/zenodo.4915437" httpUri="https://zenodo.org/record/4915437/files/figure.png" pageId="16" pageNumber="185">Fig. 5E</figureCitation>
), 2.3 times longer than wide, dorsal margin homogeneously curved with some small spines between external spines, well-developed genicular spine. Spination formula F=4ES/11IS/4DS and
<collectionCode id="ED1008D7FF94FF91FB7BFDF694D77C1E" box="[1254,1272,583,608]" name="Tavera, Department of Geology and Geophysics" pageId="16" pageNumber="185">T</collectionCode>
=10ES/9IS, all spines of femora and of tibiae ochre with dark apex.
</paragraph>
<paragraph id="8BBE9012FF94FF91FF0AFD2191697C8E" blockId="16.[151,1437,151,1112]" pageId="16" pageNumber="185">
Meso and metathoracic legs: Mesofemora with very small distal lobe which is indicated but undeveloped; metafemora with small distal lobe, bigger than the same on the mesofemora; metatibiae and meta-basitarsi missing in the
<typeStatus id="54BA2EB0FF94FF91FF40FD66916C7C8E" box="[221,323,727,752]" pageId="16" pageNumber="185" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="C31BC399FF94FF91FF5AFD4D91107A26" pageId="16" pageNumber="185" type="description">
<paragraph id="8BBE9012FF94FF91FF5AFD4D94387D46" blockId="16.[151,1437,151,1112]" pageId="16" pageNumber="185">Wings: Exceeding the apex of abdomen; Metathoracic wings about 3.5 times longer than maximum width; costal field opaque, narrow; discoidal area hyaline. Metathoracic wings hyaline.</paragraph>
<paragraph id="8BBE9012FF94FF91FF5AFCF593E97DFE" blockId="16.[151,1437,151,1112]" pageId="16" pageNumber="185">Abdomen: Cylindrical, supra-anal plate triangular in shape with rounded apex, cerci cylindrical; subgenital plate longer than wide, distal edge deeply emarginated between the styli.</paragraph>
<paragraph id="8BBE9012FF94FF91FF5AFC3D91107A26" blockId="16.[151,1437,151,1112]" pageId="16" pageNumber="185">
External genitalia (
<figureCitation id="133A8C97FF94FF91FE03FC3D91D77DDB" box="[414,504,908,933]" captionStart="FIGURE 7" captionStartId="10.[151,250,1090,1112]" captionTargetBox="[151,1435,267,1066]" captionTargetId="figure-374@10.[151,1436,267,1066]" captionTargetPageId="10" captionText="FIGURE 7. External genitalia of (all phallomeres in ventral view): A, O. rendalli (holotype,); B, O. aurita (holotype, ventral view); C, O. minima sp. n. (holotype, left phallomere ventral view, remaining phallomeres dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.4915447" httpUri="https://zenodo.org/record/4915447/files/figure.png" pageId="16" pageNumber="185">Fig. 7C</figureCitation>
): Ventral phallomere longer than wide, distal margin rounded, strongly protruding, lateral with one big thorn-like distal process (dp) and a second anterior sclerotized one which is more elongated. Left phallomere with short dorsal lamina (dl) and sclerotized in its anterior part; ventral lamina (vl) wide and sclerotized, the anterior process is well developed; both laminas form a long conspicuous lateral process (lp); phalloid apophysis (af) enlarged, sclerotized, bilobed and the surface fine-grained. Right phallomere inconspicuous.
</paragraph>
</subSubSection>
<subSubSection id="C31BC399FF94FF91FF5AFB36924F7ADE" box="[199,608,1159,1184]" pageId="16" pageNumber="185" type="distribution">
<paragraph id="8BBE9012FF94FF91FF5AFB36924F7ADE" blockId="16.[199,1033,1159,1221]" box="[199,608,1159,1184]" pageId="16" pageNumber="185">
<emphasis id="B9754C00FF94FF91FF5AFB36914F7ADE" bold="true" box="[199,352,1159,1184]" inLineHeading="true" pageId="16" pageNumber="185" reason="1">Distribution.</emphasis>
<collectingCountry id="F316D082FF94FF91FEFAFB3691D97ADE" box="[359,502,1159,1184]" name="South Africa" pageId="16" pageNumber="185">South Africa</collectingCountry>
(
<figureCitation id="133A8C97FF94FF91FD9AFB36927C7ADE" box="[519,595,1159,1184]" captionStart="FIGURE 13" captionStartId="19.[151,250,1883,1905]" captionTargetBox="[168,1419,628,1830]" captionTargetId="figure-276@19.[168,1419,628,1859]" captionTargetPageId="19" captionText="FIGURE 13. Geographic distribution of Otomantis spp. in Southern Africa and Madagascar: black circle= O casaica; black square= O. bolivari sp. n.; black triangle= O. rendalli; black diamond= O. scutigera; black shape star= O. aurita; open triangle= O. capirica; open square= O. centralis sp. n.; open circle= O. gracilis sp. n.; open diamond= O. trimacula sp. n. (note: question mark refers to the distribution of O. minima sp. n., known only from “South Africa”)." figureDoi="http://doi.org/10.5281/zenodo.4915465" httpUri="https://zenodo.org/record/4915465/files/figure.png" pageId="16" pageNumber="185">Fig.13</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C31BC399FF94FF91FF5AFB1D94267ABB" box="[199,1033,1196,1221]" pageId="16" pageNumber="185" type="etymology">
<paragraph id="8BBE9012FF94FF91FF5AFB1D94267ABB" blockId="16.[199,1033,1159,1221]" box="[199,1033,1196,1221]" pageId="16" pageNumber="185">
<emphasis id="B9754C00FF94FF91FF5AFB1D91617ABB" bold="true" box="[199,334,1196,1221]" inLineHeading="true" pageId="16" pageNumber="185" reason="1">Etymology.</emphasis>
Named in reference to the very small size of this new species.
</paragraph>
</subSubSection>
</subSection>
</treatment>
</document>