258 lines
29 KiB
XML
258 lines
29 KiB
XML
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<mods:title id="B2C6B612F7F15BAA7F679824EF2769B6">Updated catalogue and taxonomic notes on the Old-World scorpion genus Buthus Leach, 1815 (Scorpiones, Buthidae)</mods:title>
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<mods:namePart id="9C64813326B1F6520F177C4A2554E569">Sousa, Pedro</mods:namePart>
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<mods:namePart id="3F983CC9BDCD9E224113CBE132F5D7C0">Arnedo, Miquel A.</mods:namePart>
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<treatment id="8042C2C0F1A7D0303F3D7852F4E858F4" ID-GBIF-Taxon="132648862" LSID="urn:lsid:plazi:treatment:8042C2C0F1A7D0303F3D7852F4E858F4" httpUri="http://treatment.plazi.org/id/8042C2C0F1A7D0303F3D7852F4E858F4" lastPageNumber="34" pageId="19" pageNumber="34">
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<paragraph id="B7E187D588A1DE48BC2BE51EC4CC1F87" pageId="19" pageNumber="34">
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43.
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<taxonomicName id="AE4B2B5880BB78FBF97F64188164E76A" ID-CoL="NYZY" authority="C. L. Koch, 1839" authorityName="C. L. Koch" authorityYear="1839" class="Arachnida" family="Buthidae" genus="Buthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Buthus paris" order="Scorpiones" pageId="19" pageNumber="34" phylum="Arthropoda" rank="species" species="paris">Buthus paris (C. L. Koch, 1839)</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="2BDA0CC6FD37B3AE9131EEFEE9EB3D68" pageId="19" pageNumber="34" type="reference_group">
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<paragraph id="72DDDCDF03FF9EF70BFBD6A7533729A8" pageId="19" pageNumber="34">
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=
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<taxonomicName id="B10479F3E6D34299F5CCE1CEDF6C2182" class="Arachnida" family="Buthidae" genus="Androctonus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Androctonus clytoneus" order="Scorpiones" pageId="19" pageNumber="34" phylum="Arthropoda" rank="species" species="clytoneus">Androctonus clytoneus</taxonomicName>
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: C. L.
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<bibRefCitation id="A61A708948BC4EF65A67C1B29971622A" author="Koch, CL" journalOrPublisher="Zeh'schen Buchhandlung, Nuernberg" pageId="29" pageNumber="44" title="Die Arachniden. Getreu nach der Natur abgebildet und beschrieben. 5. C. H." url="http://www.biodiversitylibrary.org/item/110884" year="1839 a">Koch 1839a</bibRefCitation>
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: 70-72, pl. CLXIII, fig. 384 (synonymized by
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<bibRefCitation id="33EC8EDBD5FA6BA037E19C8EAE00E3D3" author="Vachon, M" journalOrPublisher="Archives de l'Institut Pasteur d'Algerie" pageId="35" pageNumber="50" pagination="334 - 396" title="Etudes sur les Scorpions. III (suite). Description des Scorpions du Nord de l'Afrique." volume="27" year="1949 c">Vachon 1949c</bibRefCitation>
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: 380-381). Types lost; Africa.
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</paragraph>
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<paragraph id="C36FAD470722DDC68D23FCFEBFF9968A" pageId="19" pageNumber="34">
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<taxonomicName id="A41FCEF8AE84FB6A79D52A8F92E91C6C" class="Arachnida" family="Buthidae" genus="Androctonus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Androctonus paris" order="Scorpiones" pageId="19" pageNumber="34" phylum="Arthropoda" rank="species" species="paris">Androctonus paris</taxonomicName>
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: C. L.
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<bibRefCitation id="11DB0B22499F5C1456E46D7F669DC75E" author="Koch, CL" journalOrPublisher="Zeh'schen Buchhandlung, Nuernberg" pageId="29" pageNumber="44" title="Die Arachniden. Getreu nach der Natur abgebildet und beschrieben. 5. C. H." url="http://www.biodiversitylibrary.org/item/110884" year="1839 a">Koch 1839a</bibRefCitation>
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: 25-28, pl. CLI, fig. 352; C. L.
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<bibRefCitation id="56E8D8C045A757CF9C9208D49ABA66F4" author="Koch, CL" journalOrPublisher="Zeh'schen Buchhandlung, Nuernberg" pageId="29" pageNumber="44" url="http://www.biodiversitylibrary.org/bibliography/43744" year="1850">Koch 1850</bibRefCitation>
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: 90.
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</paragraph>
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<paragraph id="3861B8E0BD8F0A8F7FCFBEF8DE80E25F" pageId="19" pageNumber="34">
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<taxonomicName id="C93FCAF14F60A2CCFCA32806144CC564" class="Arachnida" family="Buthidae" genus="Androctonus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Androctonus clytonicus" order="Scorpiones" pageId="19" pageNumber="34" phylum="Arthropoda" rank="species" species="clytonicus">Androctonus clytonicus</taxonomicName>
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(ISS):
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<bibRefCitation id="858012EF4084FBEEE394A6ADDAB3EF5E" pageId="19" pageNumber="34">Gervais 1844a</bibRefCitation>
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: 43.
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</paragraph>
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<paragraph id="724D3CA23CBEB7D80D2CE97D9665FF76" pageId="19" pageNumber="34">
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<taxonomicName id="FFD0D032AA61CF904DE16DF9D23D0ED8" class="Arachnida" family="Buthidae" genus="Androctonus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Androctonus clytoneus" order="Scorpiones" pageId="19" pageNumber="34" phylum="Arthropoda" rank="species" species="clytoneus">Androctonus clytoneus</taxonomicName>
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: C. L.
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<bibRefCitation id="3EAD4FC425C1D8A9E3BE5968740E201C" author="Koch, CL" journalOrPublisher="Zeh'schen Buchhandlung, Nuernberg" pageId="29" pageNumber="44" url="http://www.biodiversitylibrary.org/bibliography/43744" year="1850">Koch 1850</bibRefCitation>
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: 90.
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</paragraph>
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<paragraph id="03B4234C37779C91F8471DC8B4224633" pageId="19" pageNumber="34">
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<taxonomicName id="984F55CE6A0737F7FE58D7D47FA9C869" class="Arachnida" family="Buthidae" genus="Buthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Buthus occitanus" order="Scorpiones" pageId="19" pageNumber="34" phylum="Arthropoda" rank="subspecies" species="occitanus" subspecies="paris">Buthus occitanus paris</taxonomicName>
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:
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<bibRefCitation id="41373D36AC51B6958B2128FA989C60D8" author="Birula, AA" journalOrPublisher="Bulletin de l'Academie Imperiale des Sciences de St-Petersbourg [Izviestiia Imperatorskoi akademii nauk]" pageId="23" pageNumber="38" pagination="105 - 113" title="Bemerkungen ueber einige neue oder wenig bekaunte Skorpionnenformen Nord-Afrikas." volume="19" year="1903">Birula 1903</bibRefCitation>
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: 107;
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<bibRefCitation id="29814361F1C1992648D152452E484D65" author="Birula, AA" journalOrPublisher="Horae Societatis Entomologicae Rossicae" pageId="23" pageNumber="38" pagination="115 - 192" title="Ueber Scorpio maurus Linne und seine Unterarten." url="https://www.ntnu.no/ub/scorpion-files/Birula_1910.pdf" volume="39" year="1910">Birula 1910</bibRefCitation>
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: 118, 155;
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<bibRefCitation id="156188A7527CBA57FBC299751C2C1D51" pageId="19" pageNumber="34">Giltay 1929</bibRefCitation>
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: 196;
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<bibRefCitation id="584FF35914E3A2FFF21CDDB10B205C9B" author="Werner, F" journalOrPublisher="Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, Wien" pageId="37" pageNumber="52" pagination="285 - 306" title="Ergebnisse einer zoologischen Forschungsreise nach Marokko. VI. Skorpione." url="http://www.zobodat.at/publikation_articles.php?id=180394" volume="141" year="1932">Werner 1932</bibRefCitation>
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: 300-305;
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<bibRefCitation id="07FE433341451D09288752ECF748FFCB" author="Vachon, M" journalOrPublisher="Archives de l'Institut Pasteur d'Algerie" pageId="35" pageNumber="50" pagination="334 - 396" title="Etudes sur les Scorpions. III (suite). Description des Scorpions du Nord de l'Afrique." volume="27" year="1949 c">Vachon 1949c</bibRefCitation>
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: 380-388, fig. 356, 400, 445-455;
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<bibRefCitation id="881AC480D26103CA0A7878F20C40944A" pageId="19" pageNumber="34">Vachon 1951b</bibRefCitation>
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: 621;
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<bibRefCitation id="BF69654D4183129F246230FA2D747377" author="Vachon, M" journalOrPublisher="Institut Pasteur d'Algerie, Alger" pageId="35" pageNumber="50" title="Etudes sur les scorpions. (published 1948 - 1951 in Archives de l'Institut Pasteur d'Algerie, 1948, 26: 25 - 90, 162 - 208, 288 - 316, 441 - 481; 1949, 27: 66 - 100, 134 - 169, 281 - 288, 334 - 396; 1950, 28: 152 - 216, 382 - 413; 1951, 29: 46 - 104)." url="https://www.ntnu.no/ub/scorpion-files/vachon.php" year="1952 a">Vachon 1952a</bibRefCitation>
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: 308-316, fig. 356, 400 445-455;
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<bibRefCitation id="174EB59C9F50CAC108D29ABB7B97318D" author="Malhomme, J" journalOrPublisher="Bulletin de la Societe des Sciences Naturelles et Physiques du Maroc" pageId="32" pageNumber="47" pagination="25 - 30" title="Apercu sur la distribution des scorpions de la region de Marrakech." volume="34" year="1954">Malhomme 1954</bibRefCitation>
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: 29;
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<bibRefCitation id="A6B99B9CAA2BC96EE965A206683CC13F" author="Arroyo, H" journalOrPublisher="Archives de l'Institut Pasteur d'Algerie" pageId="23" pageNumber="38" pagination="186 - 189" title="Etude sur la variete oranaise du scorpion Buthusoccitanus (Amoreux, 1789)." volume="39" year="1961">Arroyo 1961</bibRefCitation>
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||
: 186-189; Le Corroller 1967: 63;
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<normalizedToken id="DFCD1E21FD02A04CE4A2AAF0DF4EE384" originalValue="Peréz">Perez</normalizedToken>
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1974: 23; Levy and Amitai 1980: 16;
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<bibRefCitation id="296AC4E293452747A4AB65E4289B0524" author="El-Hennawy, HK" journalOrPublisher="Serket" pageId="25" pageNumber="40" pagination="95 - 153" title="A catalogue of the scorpions described from the Arab countries (1758 - 1990) (Arachnida: Scorpionida)." volume="2" year="1992">El-Hennawy 1992</bibRefCitation>
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: 98, 121;
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<bibRefCitation id="A4AA1DAF6351B9FD49E6E2359873C7DC" author="Kovarik, F" journalOrPublisher="Klapalekiana" pageId="29" pageNumber="44" pagination="19 - 21" title="First report of Orthochirusinnesi (Scorpionida: Buthidae) from Morocco." volume="31" year="1995">
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<normalizedToken id="49524FB05AAFA0D1D4F09C9A7C4DFA59" originalValue="Kovařík">Kovarik</normalizedToken>
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1995
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</bibRefCitation>
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: 20;
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<bibRefCitation id="DA5ED72B2FD7513953B584DADE0E5405" pageId="19" pageNumber="34">Gantenbein et al. 1998</bibRefCitation>
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a: 51;
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<bibRefCitation id="EB7DC5B2922E47D31A9D975A607624E7" author="Kovarik, F" journalOrPublisher="Madagaskar, Jihlava" pageId="29" pageNumber="44" title="Stiri." year="1998">
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<normalizedToken id="A5AFC82277D1A821E4E2803DE2199E0C" originalValue="Kovařík">Kovarik</normalizedToken>
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1998
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</bibRefCitation>
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: 106;
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<bibRefCitation id="CAF12020111E0240407AA613E0BD68EF" pageId="19" pageNumber="34">Fet and Lowe 2000</bibRefCitation>
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: 96;
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<bibRefCitation id="61D6E91E6E1D334F15CD4DEAC7734EE9" author="Touloun, O" journalOrPublisher="Journal of Venomous Animals and Toxins" pageId="35" pageNumber="50" pagination="1 - 9" title="Epidemiological survey of scorpion envenomation in southwestern Morocco." url="https://doi.org/10.1590/S0104-79302001000200006" volume="7" year="2001">Touloun et al. 2001</bibRefCitation>
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: 2;
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<bibRefCitation id="86208B74073A002339F91980F74156FC" author="Gantenbein, B" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="26" pageNumber="41" pagination="119 - 130" title="The phylogeographic importance of the Strait of Gibraltar as a gene flow barrier in terrestrial arthropods: a case study with the scorpion Buthusoccitanus as model organism." url="https://doi.org/10.1016/S1055-7903(03)00031-9" volume="28" year="2003">
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Gantenbein and
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<normalizedToken id="2602AA504AB6EAC0C9BB5A6EF922B162" originalValue="Largiadèr">Largiader</normalizedToken>
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2003
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||
</bibRefCitation>
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||
: 120, 122;
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<bibRefCitation id="24A37267BEA25C75F75D56EDB5A3206F" pageId="19" pageNumber="34">Touloun 2012</bibRefCitation>
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: 35, 104, 108, fig.5B.
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</paragraph>
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<paragraph id="3F82A70F18D138181C4F0EFD7AC66D30" pageId="19" pageNumber="34">
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<taxonomicName id="A1FCBBB7826656C2520286F1B991541C" class="Arachnida" family="Buthidae" genus="Buthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Buthus (Buthus) occitanus" order="Scorpiones" pageId="19" pageNumber="34" phylum="Arthropoda" rank="subspecies" species="occitanus" subGenus="Buthus" subspecies="paris">Buthus (Buthus) occitanus paris</taxonomicName>
|
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:
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<bibRefCitation id="9FBE79FE1794E9D8671FB4A3015C10CC" author="Birula, AA" journalOrPublisher="Horae Societatis Entomologicae Rossicae" pageId="23" pageNumber="38" pagination="115 - 192" title="Ueber Scorpio maurus Linne und seine Unterarten." url="https://www.ntnu.no/ub/scorpion-files/Birula_1910.pdf" volume="39" year="1910">Birula 1910</bibRefCitation>
|
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. 145, 155;
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<bibRefCitation id="78443547DE666FD5411C492EC26B3128" author="Birula, AA" journalOrPublisher="Zapiski Kavkazskogo Muzeya, Seriia A [Memoires du Musee du Caucase, Se ́ rie A]" pageId="23" pageNumber="38" pagination="1 - 253" title="[authored as Byalynitskii-Birulya, A. A.] ArachnoideaArthrogastra Caucasica. Pars I. Scorpiones." volume="5" year="1917 a">Birula 1917a</bibRefCitation>
|
||
: 223.
|
||
</paragraph>
|
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<paragraph id="012ACAFD6BF2D73451F6D1096A50114A" pageId="19" pageNumber="34">
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<taxonomicName id="A8EEDC5D7E4464D59E95FC389D1C67F2" class="Arachnida" family="Buthidae" genus="Buthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Buthus paris" order="Scorpiones" pageId="19" pageNumber="34" phylum="Arthropoda" rank="species" species="paris">Buthus paris</taxonomicName>
|
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:
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<bibRefCitation id="BCC4FC13047B2B4079A02FF8E20C3715" author="Lourenco, WR" journalOrPublisher="Revue Suisse de Zoologie" pageId="30" pageNumber="45" pagination="875 - 912" title="Complements a la faune de scorpions (Arachnida) de l'Afrique du Nord, avec des considerations sur le genre Buthus Leach, 1815." volume="110" year="2003">
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<normalizedToken id="3802EEEB70F9C82346B02960C6B5F602" originalValue="Lourenço">Lourenco</normalizedToken>
|
||
2003
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||
</bibRefCitation>
|
||
: 896-897, fig. 52-56;
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<bibRefCitation id="7C087AE2CC75A6A1DB1A18CD8F06C54D" author="Kovarik, F" journalOrPublisher="Euscorpius" pageId="29" pageNumber="44" pagination="1 - 16" title="Review of Tunisian Species of the Genus Buthus with Descriptions of Two New Species and a Discussion of Ehrenberg's Types (Scorpiones: Buthidae)." url="http://www.science.marshall.edu/fet/euscorpius/p2006_34.pdf" volume="34" year="2006">
|
||
<normalizedToken id="7905C9B36B2B8AD267CF30747DA5D4F8" originalValue="Kovařík">Kovarik</normalizedToken>
|
||
2006
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</bibRefCitation>
|
||
: 2, 6, 8, 15, fig. 10-11;
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<bibRefCitation id="2AA7946201B5F3C14BF6310D3598F062" author="Lourenco, WR" journalOrPublisher="Entomologische Mitteilungen aus dem Zoologischen Museum Hamburg" pageId="31" pageNumber="46" pagination="63 - 68" title="A new species of Buthus Leach, 1815 from Algeria (Scorpiones, Buthidae)." volume="16" year="2013">
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<normalizedToken id="BFAF5F962033BA494B22343B266411C2" originalValue="Lourenço">Lourenco</normalizedToken>
|
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2013
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||
</bibRefCitation>
|
||
: 65-66; Rossi, Tropea and
|
||
<normalizedToken id="39086B52812519CEF8FC377FDFA7A931" originalValue="Yağmur">Yagmur</normalizedToken>
|
||
2013: 3, 5, 7;
|
||
<bibRefCitation id="EB0D3AACA88CCEBFBF5C7189E3144DAE" author="Aboumaad, B" journalOrPublisher="Bulletin de la Societe de pathologie exotique" pageId="22" pageNumber="37" pagination="1 - 9" title="L'envenimation scorpionique au Maroc: scorpions du genre Androctonus, Buthus et Hottentota." url="https://doi.org/10.1007/s13149-014-0332-7" volume="107" year="2014">
|
||
<normalizedToken id="254BE0DF86A49F407874C31B7421E208" originalValue="Aboumaâd">Aboumaad</normalizedToken>
|
||
et al. 2014
|
||
</bibRefCitation>
|
||
: 6;
|
||
<bibRefCitation id="E0EB2D868198A43CC57E7FBFD0A50DB8" author="Touloun, O" journalOrPublisher="Serket" pageId="35" pageNumber="50" pagination="72 - 79" title="Inventory and distribution of scorpion fauna in eastern Morocco (Arachnida, Scorpiones)." volume="14" year="2014">Touloun et al. 2014</bibRefCitation>
|
||
: 77-78;
|
||
<bibRefCitation id="C33B8E02D1D8556748BF9FDDB58A62C2" author="Lourenco, WR" journalOrPublisher="Revista Iberica de Aracnologia" pageId="31" pageNumber="46" pagination="13 - 17" title="One more new species of Buthus Leach; 1815 from Algeria (Scorpiones: Buthidae)." volume="28" year="2016">
|
||
<normalizedToken id="084E65E18158FA4DED55F16A473CEEDD" originalValue="Lourenço">Lourenco</normalizedToken>
|
||
and Sadine 2016
|
||
</bibRefCitation>
|
||
: 14-15.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="54C2D92693664F4419B36A437286DF16" pageId="19" pageNumber="34" type="type material">
|
||
<paragraph id="23DF128D7FCA4E41CB122D91A7664419" pageId="19" pageNumber="34">Type material.</paragraph>
|
||
<paragraph id="32CE2818A4FE670C9E7E262F03B47B1F" pageId="19" pageNumber="34">
|
||
Holotype lost according to
|
||
<bibRefCitation id="6944D799760D2DA58340F464638AB10A" pageId="19" pageNumber="34">Fet and Lowe (2000)</bibRefCitation>
|
||
, Algeria.
|
||
<bibRefCitation id="F07C9F8D885B13C167D7F9E9098CCDA5" author="Vachon, M" journalOrPublisher="Archives de l'Institut Pasteur d'Algerie" pageId="35" pageNumber="50" pagination="334 - 396" title="Etudes sur les Scorpions. III (suite). Description des Scorpions du Nord de l'Afrique." volume="27" year="1949 c">Vachon (1949c</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="DA9683CB6E8D57675F77880386FFBFC9" author="Vachon, M" journalOrPublisher="Institut Pasteur d'Algerie, Alger" pageId="35" pageNumber="50" title="Etudes sur les scorpions. (published 1948 - 1951 in Archives de l'Institut Pasteur d'Algerie, 1948, 26: 25 - 90, 162 - 208, 288 - 316, 441 - 481; 1949, 27: 66 - 100, 134 - 169, 281 - 288, 334 - 396; 1950, 28: 152 - 216, 382 - 413; 1951, 29: 46 - 104)." url="https://www.ntnu.no/ub/scorpion-files/vachon.php" year="1952 a">1952a</bibRefCitation>
|
||
) wrote that the types came from Alger without further explanation.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B66DAF4762181AD55C68071E23125741" pageId="19" pageNumber="34" type="distribution">
|
||
<paragraph id="F0C33BCEAC757EAC804680F5D237A577" pageId="19" pageNumber="34">Distribution.</paragraph>
|
||
<paragraph id="23D6A8CA00F3D756432E36300EB61DE1" pageId="19" pageNumber="34">the species is currently distributed across Algeria, Morocco and Tunisia.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B65BCA0A383456BDF2C22892234FEF34" pageId="19" pageNumber="34" type="remarks">
|
||
<paragraph id="6ADD378E1E4AC33D4A1DC2A7B9134E5B" pageId="19" pageNumber="34">Remarks.</paragraph>
|
||
<paragraph id="C0788754447B14D923A405F4B951AC32" pageId="19" pageNumber="34">
|
||
Because of the description of new species from Algeria, and the lack of both type specimen and locality (beyond the country), a neotype for
|
||
<taxonomicName id="4603DA2F07A311C085BEE8AAD4B20F02" lsidName="B. paris" pageId="19" pageNumber="34" rank="species" species="paris">B. paris</taxonomicName>
|
||
is necessary to stabilize the taxonomy of Algerian
|
||
<taxonomicName id="AF8875AAF9CF63DE139527DB2763DA7E" class="Arachnida" family="Buthidae" genus="Buthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Buthus" order="Scorpiones" pageId="19" pageNumber="34" phylum="Arthropoda" rank="genus">Buthus</taxonomicName>
|
||
, which may challenge the status of some newly described species. This is further complicated by recent diagnoses of
|
||
<taxonomicName id="913CDCC0A6A8E76DB48CF1DBBD7A9D29" lsidName="B. paris" pageId="19" pageNumber="34" rank="species" species="paris">B. paris</taxonomicName>
|
||
(at least in part:
|
||
<bibRefCitation id="9327C4BC7D14CFAA444C0D04D868F168" author="Lourenco, WR" journalOrPublisher="Revue Suisse de Zoologie" pageId="30" pageNumber="45" pagination="875 - 912" title="Complements a la faune de scorpions (Arachnida) de l'Afrique du Nord, avec des considerations sur le genre Buthus Leach, 1815." volume="110" year="2003">
|
||
<normalizedToken id="74BB3BBD94770A5412AC2F236934BCD9" originalValue="Lourenço">Lourenco</normalizedToken>
|
||
2003
|
||
</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="3AF3EBDFDFF16DD6B1EBCBC0CC9D2977" author="Kovarik, F" journalOrPublisher="Euscorpius" pageId="29" pageNumber="44" pagination="1 - 16" title="Review of Tunisian Species of the Genus Buthus with Descriptions of Two New Species and a Discussion of Ehrenberg's Types (Scorpiones: Buthidae)." url="http://www.science.marshall.edu/fet/euscorpius/p2006_34.pdf" volume="34" year="2006">
|
||
<normalizedToken id="4DF2CBFB341E4252CD4CC54A7DBC9363" originalValue="Kovařík">Kovarik</normalizedToken>
|
||
2006
|
||
</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="CB1AB5BD35EFEF61B69879249CE88F60" author="Lourenco, WR" journalOrPublisher="Revista Iberica de Aracnologia" pageId="31" pageNumber="46" pagination="13 - 17" title="One more new species of Buthus Leach; 1815 from Algeria (Scorpiones: Buthidae)." volume="28" year="2016">
|
||
<normalizedToken id="ECA4757B0EBA7EC9616075468E1D8D0E" originalValue="Lourenço">Lourenco</normalizedToken>
|
||
and Sadine 2016
|
||
</bibRefCitation>
|
||
) that differ from those offered by
|
||
<bibRefCitation id="83B2B84369FFFD3AF7B85E2D56313DC8" author="Vachon, M" journalOrPublisher="Institut Pasteur d'Algerie, Alger" pageId="35" pageNumber="50" title="Etudes sur les scorpions. (published 1948 - 1951 in Archives de l'Institut Pasteur d'Algerie, 1948, 26: 25 - 90, 162 - 208, 288 - 316, 441 - 481; 1949, 27: 66 - 100, 134 - 169, 281 - 288, 334 - 396; 1950, 28: 152 - 216, 382 - 413; 1951, 29: 46 - 104)." url="https://www.ntnu.no/ub/scorpion-files/vachon.php" year="1952 a">Vachon (1952a)</bibRefCitation>
|
||
. The differences between the different diagnoses include the number of rows in the movable finger, the aspect ratio of the first metasomal segment, the body chaetotaxie, the aculeus to vesicle length and the type of sexual dimorphism of the pedipalp chelae.
|
||
<bibRefCitation id="A0C00554756DF7063CB92341DED3486A" author="Vachon, M" journalOrPublisher="Institut Pasteur d'Algerie, Alger" pageId="35" pageNumber="50" title="Etudes sur les scorpions. (published 1948 - 1951 in Archives de l'Institut Pasteur d'Algerie, 1948, 26: 25 - 90, 162 - 208, 288 - 316, 441 - 481; 1949, 27: 66 - 100, 134 - 169, 281 - 288, 334 - 396; 1950, 28: 152 - 216, 382 - 413; 1951, 29: 46 - 104)." url="https://www.ntnu.no/ub/scorpion-files/vachon.php" year="1952 a">Vachon (1952a)</bibRefCitation>
|
||
studied a large number of specimens from the entire Maghreb region, unrivalled by any subsequent study, which leads us to consider
|
||
<normalizedToken id="81FE7380BA6B537851ED9E7B4AD5A004" originalValue="Vachon’s">Vachon's</normalizedToken>
|
||
description as the "gold standard". Vachon himself stated that most of the specimens used in his redescriptions were stored at the MNHN. If this material is ever located, it should have priority in the designation of a neotype. Although
|
||
<bibRefCitation id="777061CF3D98290FF4696A996B84E98A" author="Vachon, M" journalOrPublisher="Institut Pasteur d'Algerie, Alger" pageId="35" pageNumber="50" title="Etudes sur les scorpions. (published 1948 - 1951 in Archives de l'Institut Pasteur d'Algerie, 1948, 26: 25 - 90, 162 - 208, 288 - 316, 441 - 481; 1949, 27: 66 - 100, 134 - 169, 281 - 288, 334 - 396; 1950, 28: 152 - 216, 382 - 413; 1951, 29: 46 - 104)." url="https://www.ntnu.no/ub/scorpion-files/vachon.php" year="1952 a">Vachon (1952a)</bibRefCitation>
|
||
did not formally describe any varieties within
|
||
<taxonomicName id="93B38D9E94519F7BE3181830476B3129" lsidName="B. paris" pageId="19" pageNumber="34" rank="species" species="paris">B. paris</taxonomicName>
|
||
, he split the specimens that compose the species into three regions: 1) The typical region (from Algiers to northern Tunisia); 2) Specimens from the Oujda region (Morocco); 3) Those from northern Morocco and the Middle Atlas flanks. Interestingly the split of
|
||
<taxonomicName id="93A76B12F23B9F19CE2F2D611B88EA2F" lsidName="B. paris" pageId="19" pageNumber="34" rank="species" species="paris">B. paris</taxonomicName>
|
||
into these three regions corresponds well with the distribution of the genetic variability in the cox1 gene (
|
||
<bibRefCitation id="36B258AACC2BE88223535258BB45FD10" author="Gantenbein, B" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="26" pageNumber="41" pagination="119 - 130" title="The phylogeographic importance of the Strait of Gibraltar as a gene flow barrier in terrestrial arthropods: a case study with the scorpion Buthusoccitanus as model organism." url="https://doi.org/10.1016/S1055-7903(03)00031-9" volume="28" year="2003">
|
||
Gantenbein and
|
||
<normalizedToken id="D5A813320904EFCBDC302C8A977C2E87" originalValue="Largiadèr">Largiader</normalizedToken>
|
||
2003
|
||
</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="9C0DD7183DDCD36566F06F42D9EE5B6D" author="Sousa, P" journalOrPublisher="Journal of Zoology" pageId="34" pageNumber="49" pagination="66 - 75" title="Phylogeographic patterns of Buthus scorpions (Scorpiones: Buthidae) in the Maghreb and South-Western Europe based on CO 1 mtDNA sequences." url="https://doi.org/10.1111/j.1469-7998.2012.00925.x" volume="288" year="2012">Sousa et al. 2012</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="D4A7FE0B780D70E97F4F8CD4545BAF65" author="Pedroso, D" journalOrPublisher="African Invertebrates" pageId="33" pageNumber="48" pagination="298 - 308" title="Phylogeography of Buthus Leach, 1815 (Scorpiones: Buthidae): a multigene molecular approach reveals a further complex evolutionary history in the Maghreb." url="https://doi.org/10.3377/004.048.0216" volume="48" year="2013">Pedroso et al. 2013</bibRefCitation>
|
||
), placing
|
||
<taxonomicName id="DD68295DFFC89FF91AC760A2DF967398" lsidName="B. paris" pageId="19" pageNumber="34" rank="species" species="paris">B. paris</taxonomicName>
|
||
in two different groups: in
|
||
<taxonomicName id="13555E9CDDED124CAD5F1C57E01CFA3E" genus="Buthoidea" lsidName="Buthoidea occitanus" pageId="19" pageNumber="34" rank="species" species="occitanus">occitanus</taxonomicName>
|
||
which include all
|
||
<taxonomicName id="C8471D4AE96FA908F18F9C357778AE1A" lsidName="B. paris" pageId="19" pageNumber="34" rank="species" species="paris">B. paris</taxonomicName>
|
||
specimens from regions two and three, and
|
||
<taxonomicName id="8AB779C2DE80396415DC06962C3ED16D" genus="Buthoidea" lsidName="Buthoidea tunetanus" pageId="19" pageNumber="34" rank="species" species="tunetanus">tunetanus</taxonomicName>
|
||
, which include
|
||
<taxonomicName id="14DA3525E924790CC46CD192F0560298" lsidName="B. paris" pageId="19" pageNumber="34" rank="species" species="paris">B. paris</taxonomicName>
|
||
specimens from the typical region one, along the split of the two cox1 groups in the middle of Algeria (Fig. 7). If confirmed, this will mean that
|
||
<taxonomicName id="CF6420D53E9B4C9259CD957B2668BE44" lsidName="B. paris" pageId="19" pageNumber="34" rank="species" species="paris">B. paris</taxonomicName>
|
||
does not occur in Morocco. It is unclear if the variety from the third region above might correspond to
|
||
<taxonomicName id="67CF374DC67BE4D0D73FD717E38978C2" lsidName="B. confluens" pageId="19" pageNumber="34" rank="species" species="confluens">B. confluens</taxonomicName>
|
||
<normalizedToken id="C173E53BD8573FFA0CF3849DEBEC71A6" originalValue="Lourenço">Lourenco</normalizedToken>
|
||
, Touloun & Boumezzough, 2012, although these authors (page 22) refrained from suggesting this possibility because they could not find any of the material used by Vachon, purportedly to be in the MNHN, to describe this variety.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |