162 lines
19 KiB
XML
162 lines
19 KiB
XML
<document id="9538250A97E5BCAE5BF7886A57CD8CAE" ID-CLB-Dataset="34036" ID-DOI="http://dx.doi.org/10.3897/zookeys.619.9951" ID-GBIF-Dataset="dedaf101-3b90-4f1e-adef-5f7ddd95ad4e" ID-PMC="PMC5090162" ID-Pensoft-Pub="1313-2970-619-45" ID-PubMed="27829789" ID-ZooBank="80E9A40681954C1DBB3C0AF732CD478C" ModsDocAuthor="" ModsDocDate="2016" ModsDocID="1313-2970-619-45" ModsDocOrigin="ZooKeys 619" ModsDocTitle="Taxonomic revision of New Guinea diving beetles of the Exocelinadanae group, with the description of ten new species (Coleoptera, Dytiscidae, Copelatinae)" checkinTime="1475024545068" checkinUser="pensoft" docAuthor="Shaverdo, Helena, Sagata, Katayo & Balke, Michael" docDate="2016" docId="FD93AF945D050CB055BFE5DC37ABE63C" docLanguage="en" docName="ZooKeys 619: 45-102" docOrigin="ZooKeys 619" docSource="http://dx.doi.org/10.3897/zookeys.619.9951" docTitle="Exocelina atrata Balfour-Browne 1939" docType="treatment" docVersion="6" lastPageNumber="53" masterDocId="FFE3FFD6FFD3FFBB044E305DFFEA8F13" masterDocTitle="Taxonomic revision of New Guinea diving beetles of the Exocelinadanae group, with the description of ten new species (Coleoptera, Dytiscidae, Copelatinae)" masterLastPageNumber="102" masterPageNumber="45" pageNumber="52" updateTime="1701376044357" updateUser="ExternalLinkService">
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<mods:title id="4EF7DB822535255FF2CCE123D332E96A">Taxonomic revision of New Guinea diving beetles of the Exocelinadanae group, with the description of ten new species (Coleoptera, Dytiscidae, Copelatinae)</mods:title>
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<mods:namePart id="7B6F9A180DA13E9F61DE1E404A2E40E9">Shaverdo, Helena</mods:namePart>
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<mods:namePart id="40DEF37EF4B0FFE12D4D866EBFE65220">Sagata, Katayo</mods:namePart>
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<mods:namePart id="094F7F4795FCA6CF3080EB84C586C5D9">Balke, Michael</mods:namePart>
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<mods:typeOfResource id="70F777785F92E0E87535EE81535DFA25">text</mods:typeOfResource>
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<mods:title id="DA82792933F681B95046D10352D35420">ZooKeys</mods:title>
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<mods:date id="79349FF8C19EAD661338486186054309">2016</mods:date>
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<mods:detail id="261211B2798B31FB1AD30EF8866C19DF" type="volume">
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<mods:number id="76E234038F2082A978E7C5C4AF962EFA">619</mods:number>
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<mods:end id="A8D546C76F67148B444CDD0B57127980">102</mods:end>
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<mods:url id="778EE51D57D1B366184362F98742CBF6">http://dx.doi.org/10.3897/zookeys.619.9951</mods:url>
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<mods:classification id="D62DBD2C529BE4C4E5306031DB2CE4BB">journal article</mods:classification>
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<mods:identifier id="5ED86006B3319BBFF15D0D8744B755D7" type="DOI">http://dx.doi.org/10.3897/zookeys.619.9951</mods:identifier>
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<mods:identifier id="B6CDF927C008F71646A7B610EDDC37BE" type="Pensoft-Pub">1313-2970-619-45</mods:identifier>
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<treatment id="FD93AF945D050CB055BFE5DC37ABE63C" ID-GBIF-Taxon="127890989" LSID="urn:lsid:plazi:treatment:FD93AF945D050CB055BFE5DC37ABE63C" httpUri="http://treatment.plazi.org/id/FD93AF945D050CB055BFE5DC37ABE63C" lastPageId="8" lastPageNumber="53" pageId="7" pageNumber="52">
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<subSubSection id="2DADB8F8CD9921A69D4BF7D4D2CB6E15" pageId="7" pageNumber="52" type="multiple">
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<paragraph id="30955972A846C37B76D5820C9B44639F" pageId="7" pageNumber="52">Taxon classification Animalia Coleoptera Dytiscidae</paragraph>
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</subSubSection>
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<subSubSection id="D470713009C506150653EE7E73453560" pageId="7" pageNumber="52" type="nomenclature">
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<paragraph id="CF4ED5F9D98FA2DD390898139AE29E26" pageId="7" pageNumber="52">
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5.
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<taxonomicName id="CF39E814804A67FACF127009A4B25356" ID-CoL="3DLP2" authority="Balfour-Browne, 1939" authorityName="Balfour-Browne" authorityYear="1939" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina atrata" order="Coleoptera" pageId="7" pageNumber="52" phylum="Arthropoda" rank="species" species="atrata">Exocelina atrata (Balfour-Browne, 1939)</taxonomicName>
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Figs 22, 36
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</paragraph>
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</subSubSection>
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<subSubSection id="CB6FD8CCADABCDBF3BD5204B7247DE66" pageId="7" pageNumber="52" type="reference_group">
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<paragraph id="A6ADDB273508F86129EDF634CEBF01A9" pageId="7" pageNumber="52">
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<taxonomicName id="9F04A7EF237CB5D8D9E045E3E43429C8" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus atratus" order="Coleoptera" pageId="7" pageNumber="52" phylum="Arthropoda" rank="species" species="atratus">Copelatus atratus</taxonomicName>
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J.
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<bibRefCitation id="B770158DDA50429C241BA8FBD88F0809" author="Balfour-Browne, J" journalOrPublisher="Transactions of the Royal entomological Society of London" pageId="35" pageNumber="80" pagination="57 - 88" title="On Copelatus Er. and Leiopterus Steph. (Coleoptera: Dytiscidae) with descriptions of new species." url="10.1111/j.1365-2311.1939.tb00250.x" volume="88" year="1939">Balfour-Browne 1939</bibRefCitation>
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: 66;
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<bibRefCitation id="D10E888E4EC60C17F67DB1B0B5EF4D11" author="Guignot, F" journalOrPublisher="Annales historico-naturales Musei nationalis hungarici (N. S.)" pageId="35" pageNumber="80" pagination="51 - 60" title="Dytiscides recoltes par le Dr. L. Biro en nouvelle Guinee at dans l'ile de Java (Coleoptera)." volume="VII" year="1956">Guignot 1956</bibRefCitation>
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: 55 (catalogue);
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<bibRefCitation id="F4DCA04E5AB289D00DF4FB86E9995C7C" author="Gueorguiev, VB" journalOrPublisher="Izvestiya na Zoologicheskiya Institut (s Muzei) Sofiya" pageId="35" pageNumber="80" pagination="5 - 45" title="Essai de classification des coleopteres Dytiscidae. I. Tribus Copelatini (Colymbetinae)." volume="28" year="1968">
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<normalizedToken id="FCD0CC1DF010348D0E3E98D9AB09BE62" originalValue="Guéorguiev">Gueorguiev</normalizedToken>
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||
1968
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</bibRefCitation>
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: 32 (catalogue);
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<bibRefCitation id="0FBE066D2D8C74D6F342B586CA3DF349" author="Gueorguiev, VB" journalOrPublisher="Frustula entomologica (n. s.)" pageId="35" pageNumber="80" pagination="147 - 166" title="Contibuto alla conoscenza dei Dytiscidae della Nuovo Guinea (Coleoptera)." volume="XV" year="1993">
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<normalizedToken id="E27F36BB0B5548DBDD5A92C282ACD22D" originalValue="Guéorguiev">Gueorguiev</normalizedToken>
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and Rocchi 1993
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</bibRefCitation>
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: 161 (catalogue).
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</paragraph>
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<paragraph id="DABCF4F3799405532BB27E657840AB06" pageId="7" pageNumber="52">
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<taxonomicName id="BAC599585AF4FCBB206643287B0A5F31" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus (Papuadytes) atratus" order="Coleoptera" pageId="7" pageNumber="52" phylum="Arthropoda" rank="species" species="atratus" subGenus="Papuadytes">Copelatus (Papuadytes) atratus</taxonomicName>
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J. Balfour-Browne, 1939:
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<bibRefCitation id="0A751EDE83B3E6634D3CA581F9E2A1D2" author="Balke, M" journalOrPublisher="Annalen des Naturhistorischen Museum Wien" pageId="35" pageNumber="80" pagination="301 - 341" title="Revision of New Guinea Copelatus Erichson, 1832 (Insecta: Coleoptera: Dytiscidae): The running water species, Part I" volume="100 B" year="1998">Balke 1998</bibRefCitation>
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: 326 (notes, diagnosis);
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<bibRefCitation id="AEA613F122C639EA11F415E52CC79069" author="Nilsson, AN" journalOrPublisher="Apollo Books, Stenstrup" pageId="35" pageNumber="80" pagination="1 - 395" title="World catalogue of insects" year="2001">Nilsson 2001</bibRefCitation>
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: 76 (catalogue).
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</paragraph>
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<paragraph id="95420084E0286262297F8A179FF31B54" pageId="7" pageNumber="52">
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<taxonomicName id="A95FEAF767710C277FD33AF70476A1F6" class="Insecta" family="Dytiscidae" genus="Papuadytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Papuadytes atratus" order="Coleoptera" pageId="7" pageNumber="52" phylum="Arthropoda" rank="species" species="atratus">Papuadytes atratus</taxonomicName>
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(J. Balfour-Browne, 1939):
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<bibRefCitation id="78279D06A902666A8E589270C19ED794" author="Nilsson, AN" journalOrPublisher="Koleopterologische Rundschau" pageId="35" pageNumber="80" pagination="55 - 74" title="World Catalogue of Dytiscidae - corrections and additions, 3 (Coleoptera: Dytiscidae)." volume="76" year="2006">Nilsson and Fery 2006</bibRefCitation>
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: 56 (comb. n.).
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</paragraph>
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<paragraph id="64C27A8AD4A232777DCDC2C01FFEBAD8" pageId="7" pageNumber="52">
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<taxonomicName id="A774CB5E352677FB99264272DAE55447" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina atrata" order="Coleoptera" pageId="7" pageNumber="52" phylum="Arthropoda" rank="species" species="atrata">Exocelina atrata</taxonomicName>
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(J. Balfour-Browne, 1939):
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<bibRefCitation id="D944CB87E77CE829C294A8F3EDE91CB1" author="Nilsson, AN" journalOrPublisher="Latissimus" pageId="35" pageNumber="80" pagination="33 - 34" title="Exocelina Broun, 1886, is the valid name of Papuadytes Balke, 1998." volume="23" year="2007">Nilsson 2007</bibRefCitation>
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: 33 (comb. n.).
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</paragraph>
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</subSubSection>
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<subSubSection id="7D94BC3A468B81AC4837D3FD542FE03C" pageId="7" pageNumber="52" type="type locality">
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<paragraph id="A33CEF5839D36EF96AB7F197F1A28449" pageId="7" pageNumber="52">Type locality.</paragraph>
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<paragraph id="A381BB9F7A3E92D551AD2D9A561A3353" pageId="7" pageNumber="52">
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Papua New Guinea: Oro (Northern) Province, Kokoda, approximately
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<geoCoordinate id="0542A79BD09C1F549A90458177BA3873" direction="south" orientation="latitude" precision="925" value="-8.883333">08°53'S</geoCoordinate>
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;
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<geoCoordinate id="4318F0AB0CB25124FFF6A1A854AB8B62" direction="east" orientation="longitude" precision="925" value="147.73334">147°44'E</geoCoordinate>
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, approximately 366 m a.s.l.
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</paragraph>
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</subSubSection>
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<subSubSection id="71D50013145FBA6210ACFECF93F41771" pageId="7" pageNumber="52" type="type material">
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<paragraph id="F292C5E867D03B19F79BD84A8E230D39" pageId="7" pageNumber="52">Type material.</paragraph>
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<paragraph id="A7EB66C1B4F0ED00CC0109FFB5E011EE" pageId="7" pageNumber="52">
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Holotype: male
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<normalizedToken id="8D8145081F9A8E5BA5A7B6B664915DD9" originalValue="“Type”">"Type"</normalizedToken>
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[round, with red bead], "Under stones: river side.", "PAPUA:Kokoda. 1,200ft. viii.1933. L.E.Cheesman. B.M.1933-577.", "
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<taxonomicName id="832C172AC1E6561890CC4A1AA4F06D60" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus atratus" order="Coleoptera" pageId="7" pageNumber="52" phylum="Arthropoda" rank="species" species="atratus">Copelatus atratus</taxonomicName>
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, ♂ Type nov.sp." [hw, blue ink, the word
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<normalizedToken id="76A7F9BB81E6196C9914BA29C443EEFB" originalValue="“type”">"type"</normalizedToken>
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with red ink],
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<normalizedToken id="9F21E816A32F78562BC742BD59D1D3E8" originalValue="“Holotype”">"Holotype"</normalizedToken>
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[red] (BMNH). Paratypes: 1 female
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<normalizedToken id="4F93D52EC4959836300BF59F12525727" originalValue="“Type”">"Type"</normalizedToken>
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[round, with red bead], "Under stones: river side.", "PAPUA:Kokoda. 1,200ft. viii.1933. L.E.Cheesman. B.M.1933-577.", "
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<taxonomicName id="F96A4F9A5838DF5EC8CBD7B616A7A794" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus atratus" order="Coleoptera" pageId="7" pageNumber="52" phylum="Arthropoda" rank="species" species="atratus">Copelatus atratus</taxonomicName>
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, ♀ Type nov.sp." [hw, blue ink, the word
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<normalizedToken id="007CBBDD6E8C5D1B5B626F35185A472F" originalValue="“type”">"type"</normalizedToken>
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with red ink] (BMNH). 4 males, 1 female
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<normalizedToken id="009DEE536B292A3574D4F8FD2CD53285" originalValue="“Co-type”">"Co-type"</normalizedToken>
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[round, with yellow bead], "Under stones: river side.", "PAPUA:Kokoda. 1,200ft. viii.1933. L.E.Cheesman. B.M.1933-577.", "
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<taxonomicName id="D1C4FCE5074E9CE45DBA14AA7A6D7EEB" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus atratus" order="Coleoptera" pageId="7" pageNumber="52" phylum="Arthropoda" rank="species" species="atratus">Copelatus atratus</taxonomicName>
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, ♂ [or ♀] Cotype nov.sp." [hw, blue ink, the word
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<normalizedToken id="B3E7B75D9B80BE72FC032A7595440702" originalValue="“Cotype”">"Cotype"</normalizedToken>
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with red ink] (BMNH). 4 males
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<normalizedToken id="6BD7EF7BAB74B65A77F0FEE689182211" originalValue="“Co-type”">"Co-type"</normalizedToken>
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[round, with yellow bead], "PAPUA:Kokoda. 1,200ft. vi-vii.1933. L.E.Cheesman. B.M.1933-427.", "
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<taxonomicName id="A33761F80E8DD90169474B5B8E85C876" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus atratus" order="Coleoptera" pageId="7" pageNumber="52" phylum="Arthropoda" rank="species" species="atratus">Copelatus atratus</taxonomicName>
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, B-B ♂ Co-type." [hw, black ink] (BMNH).
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</paragraph>
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</subSubSection>
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<subSubSection id="5E7EEE61127D53949AAD94F246B6F05E" pageId="7" pageNumber="52" type="diagnosis">
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<paragraph id="1D4F1A301ACA03EECC2E7B7F843B2C9D" pageId="7" pageNumber="52">Diagnosis.</paragraph>
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<paragraph id="5A9EA602E4B131B813FF07E4A63F0BED" pageId="7" pageNumber="52">Beetle medium-sized, dark brown, with paler, reddish-brown, head and pronotal sides; dorsal surface with fine punctation and evident microreticulation, shiny; pronotum with distinct lateral bead; male antennomeres simple; male protarsomere 4 with large, thick, strongly curved anterolateral hook-like seta; median lobe broad, with almost parallel sides and broadly rounded apex in ventral view and with slightly curved apex, some short distal setae in lateral view; paramere without notch on dorsal side.</paragraph>
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</subSubSection>
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<subSubSection id="EE3A78E0A200B26A8A569E9AFED631E0" lastPageId="8" lastPageNumber="53" pageId="7" pageNumber="52" type="redescription">
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<paragraph id="B2761CCF733631F7B5E6346A7A475506" pageId="7" pageNumber="52">Redescription.</paragraph>
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<paragraph id="BD0AEFF919C7CD15D0489F2E4B72186B" pageId="7" pageNumber="52">Size and shape: Beetle medium-sized (TL-H 4.20-4.25 mm, TL 4.75 mm, MW 2.25 mm), with rather oblong habitus, broadest at elytral middle. Coloration: Head dark brown, with reddish-brown clypeus and vertex; pronotum dark brown on disc and reddish-brown on sides; elytra uniformly dark brown; ventrally pale brown to brown, slightly darker on metacoxal plates; head appendages yellowish-brown to reddish-brown, legs darker distally (Fig. 36).</paragraph>
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<paragraph id="D64C2E45F9EA17423F3CD98BC64E9357" pageId="7" pageNumber="52">Surface sculpture: Head with dense and coarse punctation (spaces between punctures 1-2 times size of punctures); diameter of punctures smaller than diameter of cells of microreticulation. Pronotum with evident, dense punctation, finer, sparser than on head. Elytra with finer, sparser punctation than on pronotum, punctation fine but distinct. Pronotum and elytra with distinct microreticulation, dorsal surface shiny. Head with microreticulation slightly stronger. Metaventrite, metacoxa, and abdominal ventrites distinctly microreticulate, but with cells of microreticulation larger than on dorsal side. Metacoxal plates with longitudinal strioles and transverse wrinkles; abdominal ventrites with strioles. Ventrum with inconspicuous punctation, more evident on metacoxal plates and two last abdominal ventrites.</paragraph>
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<paragraph id="6BB9D8697906E1761FC1DE97157B7C38" pageId="7" pageNumber="52">Structures: Pronotum with distinct lateral bead. Base of prosternum and neck of prosternal process with distinct ridge, smooth anteriorly, without anterolateral extensions. Blade of prosternal process lanceolate, relatively narrow, convex, with distinct bead and few setae laterally; neck and blade of prosternal process evenly jointed. Abdominal ventrite 6 truncate apically.</paragraph>
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<paragraph id="7AD6BD6E05AB568CFFFB0276B7E82BC8" lastPageId="8" lastPageNumber="53" pageId="7" pageNumber="52">
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Male: Antenna simple. Protarsomere 4 with large, thick, strongly curved anterolateral hook-like seta. Protarsomere 5 ventrally with dense anterior band of ca. 70 relatively long, thin setae and posterior row of 15 similar setae (Fig. 22A). Abdominal
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<pageBreakToken id="31622BA55621425D24EFECEEF38DED61" pageId="8" pageNumber="53" start="start">ventrite</pageBreakToken>
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6 with 4-5 lateral striae on each side. Median lobe broad, with almost parallel sides and slightly concave apex in ventral view and with slightly curved apex, some short distal setae situated in small groups under very fine carinas in lateral view; ventral sclerite of medial lobe as long as median lobe or slightly longer (Fig. 22
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<normalizedToken id="F07CEB325418F820A41D080B2C5442A1" originalValue="B–C">B-C</normalizedToken>
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). Paramere without notch, slightly concave on dorsal side, with thin, sparse, inconspicuous proximal setae and thicker, denser, and longer subdistal setae (Fig. 22D).
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</paragraph>
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<paragraph id="3FDA0E14AA909892F9179AE8AA385188" pageId="8" pageNumber="53">Female: Without evident differences in external morphology from males, except for not modified pro- and mesotarsi and abdominal ventrite 6 rounded apically, without striae.</paragraph>
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</subSubSection>
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<subSubSection id="1B597B541FB1902980C50BED3D7177AB" pageId="8" pageNumber="53" type="distribution">
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<paragraph id="ABC1FAE65F22E8A9F15B80946E6E8415" pageId="8" pageNumber="53">Distribution.</paragraph>
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<paragraph id="234405F83A3F2E14C8596EDF671AB710" pageId="8" pageNumber="53">Papua New Guinea: Oro (Northern) Province. The species is known only from its type locality (Fig. 40).</paragraph>
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</subSubSection>
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</treatment>
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</document> |