treatments-xml/data/9A/5F/87/9A5F87E8FFC27C4DFF4DAA65ECCDF86A.xml

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<document id="DC4D8951DFA00073319E8713B2DCB48A" ID-DOI="10.11646/zootaxa.5491.1.1" ID-ISSN="1175-5334" ID-Zenodo-Dep="13211849" ID-ZooBank="0612E62C-A53D-42BE-8578-68EC77D34627" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722790779731" checkinUser="plazi" docAuthor="Ahrens, Dirk, Zhao, Ming-Zhi, Pham, Phu Van &amp; Liu, Wan-Gang" docDate="2024" docId="9A5F87E8FFC27C4DFF4DAA65ECCDF86A" docLanguage="en" docName="zootaxa.5491.1.1.pdf" docOrigin="Zootaxa 5491 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5491.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.19:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="19" docTitle="Serica nhiae Ahrens, Zhao, Pham &amp; Liu 2024, new species" docType="treatment" docVersion="2" lastPageNumber="33" masterDocId="6666FF90FFDD7C6DFFDAAD56ED2BFFFF" masterDocTitle="Taxonomic updates on Pachyserica Brenske, 1898 and Serica MacLeay, 1819 reveal 38 new species and new challenges of Sericini systematics regarding DNA barcodes and genus-level diagnostic key characters (Coleoptera: Scarabaeidae: Sericinae)" masterLastPageNumber="89" masterPageNumber="1" pageNumber="32" updateTime="1722792571363" updateUser="ExternalLinkService">
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<mods:title id="4912AD82E625512E06E720E4FCC87719">Taxonomic updates on Pachyserica Brenske, 1898 and Serica MacLeay, 1819 reveal 38 new species and new challenges of Sericini systematics regarding DNA barcodes and genus-level diagnostic key characters (Coleoptera: Scarabaeidae: Sericinae)</mods:title>
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<mods:namePart id="F4A9B3E5BD1A8F9B7DA53510EC42CA5E">Ahrens, Dirk</mods:namePart>
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<mods:affiliation id="6E0B1A0FC68C162AE0341BDFC697F7B2">Zoologisches Forschungsmuseum A. Koenig, Adenauerallee 127, 53113 Bonn, Germany ahrens. dirk _ col @ gmx. de; https: // orcid. org / 0000 - 0003 - 3524 - 7153</mods:affiliation>
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<mods:namePart id="664CCAC04421295265DE6DF446E0ED63">Zhao, Ming-Zhi</mods:namePart>
<mods:nameIdentifier id="F6455BCEC529BBD5C9F995A84C0E6AF6" type="ORCID">0000-0002-3158-6177</mods:nameIdentifier>
<mods:affiliation id="544D9D2E68C2436DA6767342702E5172">College of Plant Protection, South China Agricultural University, 483 Wushan Road, Guangzhou 510642, P. R. China zhaomzhai @ 126. com; https: // orcid. org / 0000 - 0002 - 3158 - 6177</mods:affiliation>
<mods:nameIdentifier id="4D2C99AD39FECA7918009369F26B478C" type="email">zhaomzhai@126.com</mods:nameIdentifier>
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<mods:namePart id="3F1959C861C6012037F5BE4CC32B2240">Pham, Phu Van</mods:namePart>
<mods:nameIdentifier id="B589379CE2C9DD6B58B4B5727613E4C4" type="ORCID">0000-0001-9369-6649</mods:nameIdentifier>
<mods:affiliation id="CCBB424A5532A98B77FCA6081DA06F22">Institute of Ecology and Biological resources, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Cau Giay, Hanoi, Vietnam phupham. iebr @ gmail. com; https: // orcid. org / 0000 - 0001 - 9369 - 6649</mods:affiliation>
<mods:nameIdentifier id="8438A1B077CBCBFC1B6EB0F431F87A79" type="email">phupham.iebr@gmail.com</mods:nameIdentifier>
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<mods:namePart id="BF16E4DED9F93CBA0B92D3BB3A5A5E32">Liu, Wan-Gang</mods:namePart>
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<mods:affiliation id="EF48B54BFDFA3340D39D8952035A9379">Zoologisches Forschungsmuseum A. Koenig, Adenauerallee 127, 53113 Bonn, Germany ahrens. dirk _ col @ gmx. de; https: // orcid. org / 0000 - 0003 - 3524 - 7153 &amp; SKLLQG, Institute of Earth and Environment, Chinese Academy of Sciences, Yanxiang Road 97, Yanta District, Xian 710061 P. R. China liuwangang @ ieecas. cn; https: // orcid. org / 0000 - 0003 - 4788 - 7967 &amp; Zoologisches Forschungsmuseum A. Koenig, Adenauerallee 127, 53113 Bonn, Germany ahrens. dirk _ col @ gmx. de; https: // orcid. org / 0000 - 0003 - 3524 - 7153</mods:affiliation>
<mods:nameIdentifier id="5855B780ACB22491847A82DEE20773EE" type="email">ahrens.dirk_col@gmx.de</mods:nameIdentifier>
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<mods:title id="3D212E7875AF93AE7A50CD20534629DF">Zootaxa</mods:title>
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<mods:date id="A7A519C37511799E0C279CB2FA33D7CB">2024</mods:date>
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<treatment id="9A5F87E8FFC27C4DFF4DAA65ECCDF86A" LSID="urn:lsid:plazi:treatment:9A5F87E8FFC27C4DFF4DAA65ECCDF86A" httpUri="http://treatment.plazi.org/id/9A5F87E8FFC27C4DFF4DAA65ECCDF86A" lastPageId="32" lastPageNumber="33" pageId="31" pageNumber="32">
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<heading id="49018192FFC27C72FF4DAA65EE5BF8B3" bold="true" box="[151,880,1842,1869]" fontSize="11" level="2" pageId="31" pageNumber="32" reason="4">
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<taxonomicName id="D5F64D7DFFC27C72FF4DAA65EFF8F8B3" authority="Ahrens, Zhao, Pham &amp; Liu" authorityName="Ahrens, Zhao, Pham &amp; Liu" authorityYear="2024" box="[151,723,1842,1869]" class="Insecta" family="Scarabaeidae" genus="Serica" isSensuLato="true" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="nhiae" status="sp. nov.">
<emphasis id="2082EAECFFC27C72FF4DAA65EDCFF8B2" bold="true" box="[151,228,1843,1869]" italics="true" pageId="31" pageNumber="32">Serica</emphasis>
(s. l.)
<emphasis id="2082EAECFFC27C72FEEAAA64EC5FF8B2" bold="true" box="[304,372,1842,1869]" italics="true" pageId="31" pageNumber="32">nhiae</emphasis>
Ahrens, Zhao, Pham &amp; Liu
</taxonomicName>
,
<taxonomicNameLabel id="3BB15797FFC27C72FD3AAA65EE5BF8B3" box="[736,880,1842,1869]" pageId="31" pageNumber="32" rank="species">new species</taxonomicNameLabel>
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</paragraph>
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<subSubSection id="5AEC6575FFC27C72FF4DAA01EC33F88E" box="[151,280,1879,1905]" pageId="31" pageNumber="32" type="description">
<paragraph id="124936FEFFC27C72FF4DAA01EC33F88E" blockId="31.[151,880,1842,1905]" box="[151,280,1879,1905]" pageId="31" pageNumber="32">
<figureCitation id="8ACD2A7BFFC27C72FF4DAA01EC33F88E" box="[151,280,1879,1905]" captionStart="FIGURE 13" captionStartId="78.[152,255,1773,1797]" captionTargetBox="[160,1426,190,1739]" captionTargetId="figure-21@78.[151,1436,181,1749]" captionTargetPageId="78" captionText="FIGURE 13. AF: Serica nhiae new species (holotype); GK: S. (T.) yexiaohani new species (holotype); A, I: aedeagus, left side lateral view; D, K: aedeagus, right side lateral view; B, J: parameres, dorsal view; C: aedeagus, dorsal view; E, G: habitus, dorsal view; F, H: habitus, lateral view. Scale bar: 0.5 mm. Habitus images not to scale." figureDoi="http://doi.org/10.5281/zenodo.13211891" httpUri="https://zenodo.org/record/13211891/files/figure.png" pageId="31" pageNumber="32">Fig. 13AF</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="5AEC6575FFC27C72FF4DAACEEDD5F805" pageId="31" pageNumber="32" type="materials_examined">
<paragraph id="124936FEFFC27C72FF4DAACEEDD5F805" blockId="31.[151,1436,1944,2042]" pageId="31" pageNumber="32">
<emphasis id="2082EAECFFC27C72FF4DAACEEC97F84D" bold="true" box="[151,444,1944,1970]" pageId="31" pageNumber="32">Type material examined.</emphasis>
<materialsCitation id="A29E3CA3FFC27C72FE18AACEEDD5F805" collectingDate="2011-05-27" collectionCode="MZUF, VNMN" collectorName="L. Bartolozzi &amp; S. Bambi &amp; F. Fabiano &amp; E. Orbach" country="Vietnam" elevation="1900" location="Sa Pa" municipality="Hoang Lien Natl Park" pageId="31" pageNumber="32" specimenCount="1" specimenCount-male="1" stateProvince="Lao Cai" typeStatus="holotype">
<typeStatus id="CD4D885CFFC27C72FE18AACEEF01F84D" box="[450,554,1944,1970]" pageId="31" pageNumber="32" type="holotype">Holotype</typeStatus>
<specimenCount id="04F0FD77FFC27C72FDEAAACFEF6EF84C" box="[560,581,1945,1971]" count="1" pageId="31" pageNumber="32" type="male"></specimenCount>
“N
<collectingCountry id="6AE1766EFFC27C72FDAAAACEEFC3F84D" box="[624,744,1944,1970]" name="Vietnam" pageId="31" pageNumber="32">VIETNAM</collectingCountry>
<collectingRegion id="D032F81CFFC27C72FCD9AACEEEE9F84D" box="[771,962,1944,1970]" country="Vietnam" name="Lao Cai" pageId="31" pageNumber="32">Lao Cai province</collectingRegion>
,
<collectingMunicipality id="F22DAC84FFC27C72FC11AACFE996F84D" box="[971,1213,1944,1971]" pageId="31" pageNumber="32">Hoang Lien Natl Park</collectingMunicipality>
surr.
<location id="17296025FFC27C72FB20AACEE813F84C" LSID="urn:lsid:plazi:treatment:9A5F87E8FFC27C4DFF4DAA65ECCDF86A:17296025FFC27C72FB20AACEE813F84C" box="[1274,1336,1944,1971]" country="Vietnam" municipality="Hoang Lien Natl Park" name="Sa Pa" pageId="31" pageNumber="32" stateProvince="Lao Cai">Sa Pa</location>
pass, at light (~
<quantity id="D50E9B1BFFC27C72FF3DAAEAEC12F829" box="[231,313,1980,2006]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="31" pageNumber="32" unit="m" value="1900.0">
<elevation id="99DBD1CDFFC27C72FF3DAAEAEC12F829" box="[231,313,1980,2006]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="31" pageNumber="32" unit="m" value="1900.0">1900m</elevation>
</quantity>
)
<date id="6648103EFFC27C72FE9DAAEAEC93F829" box="[327,440,1980,2006]" pageId="31" pageNumber="32" value="2011-05-27">
<collectingDate id="760CE9D6FFC27C72FE9DAAEAEC93F829" box="[327,440,1980,2006]" pageId="31" pageNumber="32" value="2011-05-27">27.V.2011</collectingDate>
</date>
/
<collectorName id="BF035328FFC27C72FE16AAEBEF49F829" box="[460,610,1980,2007]" pageId="31" pageNumber="32">L. Bartolozzi</collectorName>
,
<collectorName id="BF035328FFC27C72FDB7AAEAEFFCF829" box="[621,727,1980,2006]" pageId="31" pageNumber="32">S. Bambi</collectorName>
,
<collectorName id="BF035328FFC27C72FD3BAAEBEE72F829" box="[737,857,1980,2007]" pageId="31" pageNumber="32">F. Fabiano</collectorName>
,
<collectorName id="BF035328FFC27C72FCBEAAEBEEFFF829" box="[868,980,1980,2007]" pageId="31" pageNumber="32">E. Orbach</collectorName>
leg. (Num. Maggazzino 2909)” (
<collectionCode id="74E7AE3BFFC27C72FA9FAAEAE8BFF829" box="[1349,1428,1980,2006]" country="Italy" httpUri="http://grbio.org/cool/mvk4-k796" name="Museo Zoologico La Specola, Universita di Firenze" pageId="31" pageNumber="32">MZUF</collectionCode>
/
<collectionCode id="74E7AE3BFFC27C72FF4DAAB6EDD9F805" box="[151,242,2016,2042]" country="Viet Nam" httpUri="http://grbio.org/cool/65ma-2bzq" name="Vietnam National Museum of Nature" pageId="31" pageNumber="32">VNMN</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="5AEC6575FFFD7C4DFF1DADC0EEBAF91E" pageId="32" pageNumber="33" type="description">
<paragraph id="124936FEFFFD7C4DFF1DADC0EC3DFEE2" blockId="32.[151,1437,150,1941]" pageId="32" pageNumber="33">
<emphasis id="2082EAECFFFD7C4DFF1DADC0ECC0FF4E" bold="true" box="[199,491,150,177]" pageId="32" pageNumber="33">
Description of
<typeStatus id="CD4D885CFFFD7C4DFEA4ADC1ECCCFF4E" box="[382,487,151,177]" pageId="32" pageNumber="33" type="holotype">holotype</typeStatus>
.
</emphasis>
Length:
<quantity id="D50E9B1BFFFD7C4DFD8DADC1EFEAFF4E" box="[599,705,151,177]" metricMagnitude="-2" metricUnit="m" metricValue="1.06" pageId="32" pageNumber="33" unit="mm" value="10.6">10.6 mm</quantity>
, length of elytra:
<quantity id="D50E9B1BFFFD7C4DFC4EADC1EEC4FF4E" box="[916,1007,151,177]" metricMagnitude="-3" metricUnit="m" metricValue="8.4" pageId="32" pageNumber="33" unit="mm" value="8.4">8.4 mm</quantity>
, width:
<quantity id="D50E9B1BFFFD7C4DFB8AADC1E987FF4E" box="[1104,1196,151,177]" metricMagnitude="-3" metricUnit="m" metricValue="6.3" pageId="32" pageNumber="33" unit="mm" value="6.3">6.3 mm</quantity>
. Body elongate eggshaped, dark reddish brown, partly with greenish toment, elytra with dark spots, even intervals almost completely dark, antenna yellowish, legs brown; dorsal surface dull and densely covered with short yellow setae being bent posteriorly.
</paragraph>
<paragraph id="124936FEFFFD7C4DFF1DAC71ECA9FD56" blockId="32.[151,1437,150,1941]" pageId="32" pageNumber="33">
Labroclypeus subtrapezoidal, distinctly wider than long, widest at base, lateral margins convex and strongly convergent anteriorly, anterior angles acute; anterior margin deeply and widely emarginate medially; margins weakly reflexed; surface flat and shiny, finely and densely punctate, shortly, finely and densely setose, behind anterior margin glabrous; frontoclypeal suture finely incised, weakly convex; smooth area anterior to eye large and flat, as wide as long; ocular canthus moderately long and slender (1/3 of ocular diameter), with a few fine punctures bearing each a short seta. Frons completely dull and flat, with fine and very dense punctures, evenly covered with short yellow setae being bent backwards and longer erect setae. Eyes small, ratio diameter/ interocular width: 0.49. Antenna with ten antennomeres; antennomeres three to five as wide as long, antennomeres six and seven transverse and short; club with three antennomeres, as long as remaining antennomeres combined and straight.
<collectingRegion id="D032F81CFFFD7C4DFF4DAF3DEDDDFD7A" box="[151,246,619,645]" country="Myanmar" name="Chin" pageId="32" pageNumber="33">Mentum</collectingRegion>
weakly elevated, anteriorly flattened. Labrum transverse, elyptical, produced along middle, moderately emarginate medially.
</paragraph>
<paragraph id="124936FEFFFD7C4DFF1DAFE5EC27FC7E" blockId="32.[151,1437,150,1941]" pageId="32" pageNumber="33">Pronotum subtrapezoidal, widest shortly before base, lateral margins evenly convex and strongly convergent anteriorly; anterior angles distinctly produced and acute; posterior angles strongly rounded; anterior margin medially with a fine marginal line and very weakly produced medially; surface densely and coarsely punctate, with dense short yellow setae bent posteriorly; lateral margins sparsely setose; hypomeron carinate at base, weakly produced ventrally. Scutellum large, triangular, finely and very densely punctate, basal midline smooth, pilosity as in lateral pronotum.</paragraph>
<paragraph id="124936FEFFFD7C4DFF1DAEDDECA7FBCA" blockId="32.[151,1437,150,1941]" pageId="32" pageNumber="33">Elytra oval, widest at middle, striae indistinctly impressed, finely and densely punctate; intervals flat, with fine, very dense punctures; dark spots impunctate or with less dense punctures, with fine, short pilosity as in pronotum, without longer, erect setae; epipleural edge fine, ending at moderately curved external apical angle of elytra, epipleura densely setose, apical margin without a very fine membranous rim composed of minute microtrichomes (magnification 100x).</paragraph>
<paragraph id="124936FEFFFD7C4DFF1DA969E854FB16" blockId="32.[151,1437,150,1941]" pageId="32" pageNumber="33">Ventral surface dull, finely and densely punctate, including metacoxa and abdominal sternites densely shortly setose; abdominal sternites with a transverse row of coarse punctures, each bearing a long seta, otherwise shortly setose as rest of ventral surface; ultimate sternite at middle 1.5 times as long as penultimate one. Mesosternum between mesocoxae half as wide as the slender mesofemur. Ratio of length of metepisternum/ metacoxa: 1/ 1.39. Pygidium strongly convex and dull, finely and densely punctate, smooth midline narrow, with dense short setae.</paragraph>
<paragraph id="124936FEFFFD7C4DFF1DA9A5EFC7F942" blockId="32.[151,1437,150,1941]" pageId="32" pageNumber="33">Legs slender; femora with two longitudinal rows of setae, coarsely and densely punctate between the rows, with robust setae on basal half; metafemur shiny, anterior margin acute, without a continuously serrated line behind anterior edge, posterior margin ventrally serrated in apical half and not widened, dorsal posterior margin completely serrated, in basal half with a few long setae which are half as long as width of metafemur. Metatibia slender and long, widest at apex, ratio of width/ length: 1/ 4.8, dorsally sharply carinate, with two groups of spines, basal group shortly before middle, apical group at three quarters of metatibial length, basally with a few single, robust setae; external face impunctate, with dense superficial longitudinal wrinkles; ventral margin serrated, with two very widely separated robust setae; medial face glabrous and smooth; apex interiorly near tarsal articulation concavely truncate. Tarsomeres ventrally with sparse, minute setae, dorsally smooth. Metatarsomeres ventrally glabrous, laterally and dorsally carinate, with a strongly serrated ridge ventrally; first metatarsomere little shorter than following two tarsomeres combined and almost twice as long as dorsal tibial spur; mesotarsomeres not carinate. Protibia moderately long, bidentate, external margin serrate, with a few small teeth at base; anterior claws asymmetrical, basal tooth of inner claw concavely excavate at apex.</paragraph>
<paragraph id="124936FEFFFD7C4DFF1DAB91EEBAF91E" blockId="32.[151,1437,150,1941]" box="[199,913,1735,1761]" pageId="32" pageNumber="33">
Aedeagus:
<figureCitation id="8ACD2A7BFFFD7C4DFE9EAB91ECE6F91E" box="[324,461,1735,1761]" captionStart="FIGURE 13" captionStartId="78.[152,255,1773,1797]" captionTargetBox="[160,1426,190,1739]" captionTargetId="figure-21@78.[151,1436,181,1749]" captionTargetPageId="78" captionText="FIGURE 13. AF: Serica nhiae new species (holotype); GK: S. (T.) yexiaohani new species (holotype); A, I: aedeagus, left side lateral view; D, K: aedeagus, right side lateral view; B, J: parameres, dorsal view; C: aedeagus, dorsal view; E, G: habitus, dorsal view; F, H: habitus, lateral view. Scale bar: 0.5 mm. Habitus images not to scale." figureDoi="http://doi.org/10.5281/zenodo.13211891" httpUri="https://zenodo.org/record/13211891/files/figure.png" pageId="32" pageNumber="33">Fig. 13AD</figureCitation>
. Habitus:
<figureCitation id="8ACD2A7BFFFD7C4DFDE4AB91EF96F91E" box="[574,701,1735,1761]" captionStart="FIGURE 13" captionStartId="78.[152,255,1773,1797]" captionTargetBox="[160,1426,190,1739]" captionTargetId="figure-21@78.[151,1436,181,1749]" captionTargetPageId="78" captionText="FIGURE 13. AF: Serica nhiae new species (holotype); GK: S. (T.) yexiaohani new species (holotype); A, I: aedeagus, left side lateral view; D, K: aedeagus, right side lateral view; B, J: parameres, dorsal view; C: aedeagus, dorsal view; E, G: habitus, dorsal view; F, H: habitus, lateral view. Scale bar: 0.5 mm. Habitus images not to scale." figureDoi="http://doi.org/10.5281/zenodo.13211891" httpUri="https://zenodo.org/record/13211891/files/figure.png" pageId="32" pageNumber="33">Fig. 13E, F</figureCitation>
. Female unknown.
</paragraph>
</subSubSection>
<subSubSection id="5AEC6575FFFD7C4DFF1DABBCEF95F8B2" pageId="32" pageNumber="33" type="diagnosis">
<paragraph id="124936FEFFFD7C4DFF1DABBCEF95F8B2" blockId="32.[151,1437,150,1941]" pageId="32" pageNumber="33">
<emphasis id="2082EAECFFFD7C4DFF1DABBCEC69F8FB" bold="true" box="[199,322,1770,1796]" pageId="32" pageNumber="33">Diagnosis.</emphasis>
<taxonomicName id="D5F64D7DFFFD7C4DFE91ABBDECF1F8FA" authorityName="Ahrens &amp; Zhao &amp; Pham &amp; Liu" authorityYear="2024" box="[331,474,1771,1797]" class="Insecta" family="Scarabaeidae" genus="Serica" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="nhiae" status="sp. nov.">
<emphasis id="2082EAECFFFD7C4DFE91ABBDECF1F8FA" box="[331,474,1771,1797]" italics="true" pageId="32" pageNumber="33">Serica nhiae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3BB15797FFFD7C4DFE3EABBDEF45F8FA" box="[484,622,1771,1797]" pageId="32" pageNumber="33" rank="species">new species</taxonomicNameLabel>
differs from all other species of the
<taxonomicName id="D5F64D7DFFFD7C4DFBC0ABBDE9BEF8FA" authorityName="Arrow" authorityYear="1946" box="[1050,1173,1771,1797]" class="Insecta" family="Scarabaeidae" genus="Serica" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="26" phylum="Arthropoda" rank="species" species="velutina">
S.
<emphasis id="2082EAECFFFD7C4DFBE1ABBDE9BEF8FA" box="[1083,1173,1771,1797]" italics="true" pageId="32" pageNumber="33">velutina</emphasis>
</taxonomicName>
group by the shape of parameres: both are curved dorsally (lateral view), the right paramere has a large concave basal lobe; the left paramere is at apex evenly but sharply narrowed.
</paragraph>
</subSubSection>
<subSubSection id="5AEC6575FFFD7C4DFF1DAA01ECCDF86A" pageId="32" pageNumber="33" type="etymology">
<paragraph id="124936FEFFFD7C4DFF1DAA01ECCDF86A" blockId="32.[151,1437,150,1941]" pageId="32" pageNumber="33">
<emphasis id="2082EAECFFFD7C4DFF1DAA01EC65F88E" bold="true" box="[199,334,1879,1905]" pageId="32" pageNumber="33">Etymology.</emphasis>
The new species (adjective in genitive case singular) is named after Dr. Nhi Thi Pham in gratitude for her support for Phu Pham.
</paragraph>
</subSubSection>
</treatment>
</document>