treatments-xml/data/8C/49/87/8C4987926726FFFFC7B0FCEBFB4AD326.xml

<document id="9AD56BBB0E1F95549BE9481054BFD3AC" ID-DOI="10.11646/zootaxa.3635.5" ID-GBIF-Dataset="6c160e64-940a-41f6-89d4-e7299281bc6e" ID-ISSN="1175-5326" ID-Zenodo-Dep="217427" ID-ZooBank="5E6A3AFD-89BF-441F-A826-A075C932D42D" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460587389638" checkinUser="plazi" docAuthor="Anisyutkin, Leonid N., Anichkin, Alexandr E. &amp; Thinh, Nguyen Van" docDate="2013" docId="8C4987926726FFFFC7B0FCEBFB4AD326" docLanguage="en" docName="zt03635p532.pdf" docOrigin="Zootaxa 3635 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Macrostylopyga bidupi Anisyutkin, Anichkin &amp; Nguyen, sp. nov." docType="treatment" docVersion="6" lastPageNumber="530" masterDocId="7070FFEA6722FFF5C727FF8CFF81D059" masterDocTitle="Macrostylopyga gen. nov., a new genus of cockroaches (Dictyoptera: Blattidae), with descriptions of two new species" masterLastPageNumber="532" masterPageNumber="520" pageNumber="524" updateTime="1698325378222" updateUser="plazi">
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<mods:title id="ED3BAFE79E452276C7C8668BC03514E8">Macrostylopyga gen. nov., a new genus of cockroaches (Dictyoptera: Blattidae), with descriptions of two new species</mods:title>
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<mods:namePart id="09C5FD2AC53B0B36168C0CC8C105D423">Anisyutkin, Leonid N.</mods:namePart>
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<mods:namePart id="1B1A958FD5FD3707026E6773824B142C">Anichkin, Alexandr E.</mods:namePart>
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<mods:namePart id="892E2DD15DD8BDC740B7FD8D172A9143">Thinh, Nguyen Van</mods:namePart>
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<treatment id="8C4987926726FFFFC7B0FCEBFB4AD326" ID-DOI="http://doi.org/10.5281/zenodo.6165482" ID-GBIF-Taxon="119565078" ID-Zenodo-Dep="6165482" LSID="urn:lsid:plazi:treatment:8C4987926726FFFFC7B0FCEBFB4AD326" httpUri="http://treatment.plazi.org/id/8C4987926726FFFFC7B0FCEBFB4AD326" lastPageId="10" lastPageNumber="530" pageId="4" pageNumber="524">
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<heading id="5F1781E86726FFF1C7B0FCEBFC33D3D8" bold="true" box="[151,946,871,897]" fontSize="11" level="1" pageId="4" pageNumber="524" reason="1">
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<taxonomicName id="C3E04D076726FFF1C7B0FCEBFCC9D3D8" ID-CoL="3XBG6" authority="Anisyutkin, Anichkin &amp; Nguyen" authorityName="Anisyutkin, Anichkin &amp; Nguyen" box="[151,840,871,897]" class="Insecta" family="Blattidae" genus="Macrostylopyga" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="524" phylum="Arthropoda" rank="species" species="bidupi" status="sp. nov.">
<emphasis id="3694EA966726FFF1C7B0FCEBFE2AD3D8" bold="true" box="[151,427,871,897]" italics="true" pageId="4" pageNumber="524">Macrostylopyga bidupi</emphasis>
Anisyutkin, Anichkin &amp; Nguyen
</taxonomicName>
, 
<taxonomicNameLabel id="2DA757ED6726FFF1C473FCEBFC33D3D8" box="[852,946,871,897]" pageId="4" pageNumber="524" rank="species">sp. nov.</taxonomicNameLabel>
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<paragraph id="045F36846726FFF1C7B0FC06FDFCD3FB" blockId="4.[151,946,871,930]" box="[151,637,906,930]" pageId="4" pageNumber="524">
(
<figureCitation id="9CDB2A016726FFF1C7B8FC06FEFBD3FB" box="[159,378,906,930]" captionStart="FIGURES 12 – 26" captionStartId="6.[151,264,1797,1820]" captionTargetBox="[253,1329,203,1766]" captionTargetId="figure@6.[239,1348,193,1776]" captionTargetPageId="6" captionText="FIGURES 12 – 26. Macrostylopyga spp., male, holotypes (12 – 17, 23, 24 — M. bidupi sp. nov., 18 – 22 — M. grandis sp. nov., 25, 26 — M. laosana (Anisyutkin, 2010). 12 — facial part of head; 13 — head, thorax and 1 – 7 th abdominal tergites from above; 14, 15 — abdominal apex from above (14) and below (15, hypandrium and genitalia removed); 16 — caudo-medial processes of paraprocts; 17 — hypandrium from below; 18, 22 — hind tarsus from outside (18) and below (22); 19 — arolium and claws; 20, 21, 23 – 26 — metatarsi: right fore (20, 23), right middle (21, 24), left hind (25, 26) from below (20, 21, 23, 24, 26) and outside (25). Dotted area shows membranous parts. Abbreviations: I – X — 1 st – 10 th abdominal tergites respectively, a. s.; ar.; e. r.; i. r.; pr.; par. — see text. Arrows show places of styli attachment. Scale bar (1 mm = e; 2 mm = h, i, k; 3 mm = d, j, l; 5 mm = a – c, f, g): a = 12; b = 13; c = 14; d = 15; e = 16; f = 17; g = 18; h = 19; i = 20, 21; j = 22; k = 23, 24; l = 25, 26." httpUri="https://zenodo.org/record/217429/files/figure.png" pageId="4" pageNumber="524">Figs. 12–17, 23, 24</figureCitation>
, 
<figureCitation id="9CDB2A016726FFF1C6A1FC06FE4FD3FB" box="[390,462,906,930]" captionStart="FIGURES 27 – 38" captionStartId="7.[151,264,1891,1914]" captionTargetBox="[203,1384,204,1860]" captionTargetId="figure@7.[200,1387,193,1870]" captionTargetPageId="7" captionText="FIGURES 27 – 38. Macrostylopyga spp., male genitalia, holotypes (27 – 30 — M. grandis sp. nov., 31 – 34 — M. bidupi sp. nov., 35 – 38 — M. laosana (Anisyutkin, 2010). Left phallomera from above (27, 31, 35), outside (28, 32, 36), inside (29, 33, partly, 37) and below (30, 34, partly, 38). Dotted area shows membranous parts. Figs. 35 – 38 after Anisyutkin, 2010. Abbreviations: a. c. s.; a. p.; c. pr.; d. c. p.; e. c. s.; L 1; L 2; L 3; L 4 C; L 4 C, c. p.; L 4 D; L 4 F; m. l.; r. m. l.; r. o.; s. l.; sla; t. p. — see text. Scale bar 2 mm." httpUri="https://zenodo.org/record/217430/files/figure.png" pageId="4" pageNumber="524">31–34</figureCitation>
, 
<figureCitation id="9CDB2A016726FFF1C6FDFC06FDA2D3FB" box="[474,547,906,930]" captionStart="FIGURES 39 – 49" captionStartId="8.[151,264,1852,1875]" captionTargetBox="[215,1394,216,1820]" captionTargetId="figure@8.[167,1419,193,1847]" captionTargetPageId="8" captionText="FIGURES 39 – 49. Macrostylopyga spp., male genitalia, holotypes (40, 42 – 49) and paratype (39, 41) (39 – 45 — M. grandis sp. nov., 46 – 49 — M. bidupi sp. nov.). 39, 40 — left phallomere from above with sclerite L 1 (39) and from caudal (40); 41 — sclerite L 1 and process sla of left phalomere from inside; 42, 46 — sclerite L 1 from outside (42) and inside (46); 43, 44, 47, 48 — ventral phallomere from side (43, 47) and below (44, 48); 45, 49 — sclerites R 3 from outside. Dotted area shows membranous parts. Abbreviations: L 1; L 2; L 3; L 4 C; L 4 D; m. i.; r. o.; sla; t. p. — see text. Scale bar (2 mm): a = 39; b = 40; c = 41; d = 42; e = 43, 44; f = 45; g = 46; h = 47, 48; i = 49." httpUri="https://zenodo.org/record/217431/files/figure.png" pageId="4" pageNumber="524">46–49</figureCitation>
, 
<figureCitation id="9CDB2A016726FFF1C509FC06FDF4D3FB" box="[558,629,906,930]" captionStart="FIGURES 50 – 56" captionStartId="9.[151,264,1862,1885]" captionTargetBox="[218,1362,203,1830]" captionTargetId="figure@9.[196,1392,193,1841]" captionTargetPageId="9" captionText="FIGURES 50 – 56. Macrostylopyga spp., male genitalia, holotypes (50 – 52 — M. grandis sp. nov., 53 – 55 — M. bidupi sp. nov., 56 — M. laosana (Anisyutkin, 2010). 50, 51, 53, 54 — right phallomere from above (50, 53) and below (51, 54); 52 — sclerite R 1 H; 55 — sclerites R 1 H and R 1 G from dorso-caudal; 56 — outline of sclerite R 1 H processes. Dotted area shows membranous parts. Fig. 56 after Anisyutkin, 2010. Abbreviations: p. s.; R 1 G; R 1 H; R 2; R 3; t. s. — see text. Scale bar (2 mm = a – c, g; 1 mm = d – f): a = 50; b = 51; c = 52; d = 53; e = 54; f = 55; g = 56." httpUri="https://zenodo.org/record/217432/files/figure.png" pageId="4" pageNumber="524">53–55</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="4CFA650F6726FFF1C7B0FC5EFC56D46B" pageId="4" pageNumber="524" type="materials_examined">
<paragraph id="045F36846726FFF1C7B0FC5EFC56D46B" blockId="4.[151,1436,978,2010]" pageId="4" pageNumber="524">
<emphasis id="3694EA966726FFF1C7B0FC5EFE87D3B2" bold="true" box="[151,262,978,1003]" pageId="4" pageNumber="524">Material.</emphasis>
<typeStatus id="DB5B88266726FFF1C62AFC5EFEF4D3B3" box="[269,373,978,1002]" pageId="4" pageNumber="524" type="holotype">Holotype</typeStatus>
male, 
<collectingCountry id="7CF776146726FFF1C6E4FC5EFDA7D3B3" box="[451,550,978,1002]" name="Vietnam" pageId="4" pageNumber="524">Vietnam</collectingCountry>
: Lam Dong Province, Lac Duong District, 
<quantity id="C3189B616726FFF1C328FC5EFBC8D3B3" box="[1039,1097,978,1002]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="4" pageNumber="524" unit="km" value="5.0">5 km</quantity>
NE of Long Lanh Village, Bi Dup–Nui Ba Nature Reserve, 
<geoCoordinate id="61D450436726FFF1C6CEFC7AFDF4D457" box="[489,629,1014,1038]" direction="north" orientation="latitude" pageId="4" pageNumber="524" precision="15" value="12.178889">12°10'44'' N</geoCoordinate>
, 
<geoCoordinate id="61D450436726FFF1C5A6FC7AFC99D457" box="[641,792,1014,1038]" direction="east" orientation="longitude" pageId="4" pageNumber="524" precision="15" value="108.67889">108°40'44'' E</geoCoordinate>
, elevation = 
<quantity id="C3189B616726FFF1C48FFC7AFB81D457" box="[936,1024,1014,1038]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="4" pageNumber="524" unit="m" value="1400.0">1400 m</quantity>
, 
<date id="705E10446726FFF1C32BFC7AFB08D457" box="[1036,1161,1014,1038]" pageId="4" pageNumber="524" value="2009-04">April 2009</date>
, soil trap, Pl. 5–9, probe 23, coll. A. Anichkin (Expedition of the Russia-Vietnam Tropical Centre).
</paragraph>
</subSubSection>
<subSubSection id="4CFA650F6726FFFFC7E0FBB2FB4AD326" lastPageId="10" lastPageNumber="530" pageId="4" pageNumber="524" type="description">
<paragraph id="045F36846726FFF1C7E0FBB2FC9DD783" blockId="4.[151,1436,978,2010]" pageId="4" pageNumber="524">
<emphasis id="3694EA966726FFF1C7E0FBB2FED8D40E" bold="true" box="[199,345,1086,1111]" pageId="4" pageNumber="524">Description.</emphasis>
Male (
<typeStatus id="DB5B88266726FFF1C69EFBB2FDA0D40F" box="[441,545,1086,1110]" pageId="4" pageNumber="524" type="holotype">holotype</typeStatus>
). General coloration reddish black; scapus, medial parts of pro-, meso-, metanotum and abdominal sternites, hypandrium and caudal part of anal plate slightly lighter, reddish; eyes black; anteclypeus, labrum, maxillary and labial palps yellowish brown, apices of labrum and palps pale; thoracic ventral sclerites, femora and, parts of tarsi more bright, reddish brown. Surfaces smooth and lustrous; punctation scarce and weak; facial part of head with weakly expressed transverse wrinkles (
<figureCitation id="9CDB2A016726FFF1C4FDFB42FBAFD4BF" box="[986,1070,1230,1254]" captionStart="FIGURES 12 – 26" captionStartId="6.[151,264,1797,1820]" captionTargetBox="[253,1329,203,1766]" captionTargetId="figure@6.[239,1348,193,1776]" captionTargetPageId="6" captionText="FIGURES 12 – 26. Macrostylopyga spp., male, holotypes (12 – 17, 23, 24 — M. bidupi sp. nov., 18 – 22 — M. grandis sp. nov., 25, 26 — M. laosana (Anisyutkin, 2010). 12 — facial part of head; 13 — head, thorax and 1 – 7 th abdominal tergites from above; 14, 15 — abdominal apex from above (14) and below (15, hypandrium and genitalia removed); 16 — caudo-medial processes of paraprocts; 17 — hypandrium from below; 18, 22 — hind tarsus from outside (18) and below (22); 19 — arolium and claws; 20, 21, 23 – 26 — metatarsi: right fore (20, 23), right middle (21, 24), left hind (25, 26) from below (20, 21, 23, 24, 26) and outside (25). Dotted area shows membranous parts. Abbreviations: I – X — 1 st – 10 th abdominal tergites respectively, a. s.; ar.; e. r.; i. r.; pr.; par. — see text. Arrows show places of styli attachment. Scale bar (1 mm = e; 2 mm = h, i, k; 3 mm = d, j, l; 5 mm = a – c, f, g): a = 12; b = 13; c = 14; d = 15; e = 16; f = 17; g = 18; h = 19; i = 20, 21; j = 22; k = 23, 24; l = 25, 26." httpUri="https://zenodo.org/record/217429/files/figure.png" pageId="4" pageNumber="524">Fig. 12</figureCitation>
). Body comparatively elongated (
<figureCitation id="9CDB2A016726FFF1C7B8FB7EFF75D553" box="[159,244,1266,1290]" captionStart="FIGURES 12 – 26" captionStartId="6.[151,264,1797,1820]" captionTargetBox="[253,1329,203,1766]" captionTargetId="figure@6.[239,1348,193,1776]" captionTargetPageId="6" captionText="FIGURES 12 – 26. Macrostylopyga spp., male, holotypes (12 – 17, 23, 24 — M. bidupi sp. nov., 18 – 22 — M. grandis sp. nov., 25, 26 — M. laosana (Anisyutkin, 2010). 12 — facial part of head; 13 — head, thorax and 1 – 7 th abdominal tergites from above; 14, 15 — abdominal apex from above (14) and below (15, hypandrium and genitalia removed); 16 — caudo-medial processes of paraprocts; 17 — hypandrium from below; 18, 22 — hind tarsus from outside (18) and below (22); 19 — arolium and claws; 20, 21, 23 – 26 — metatarsi: right fore (20, 23), right middle (21, 24), left hind (25, 26) from below (20, 21, 23, 24, 26) and outside (25). Dotted area shows membranous parts. Abbreviations: I – X — 1 st – 10 th abdominal tergites respectively, a. s.; ar.; e. r.; i. r.; pr.; par. — see text. Arrows show places of styli attachment. Scale bar (1 mm = e; 2 mm = h, i, k; 3 mm = d, j, l; 5 mm = a – c, f, g): a = 12; b = 13; c = 14; d = 15; e = 16; f = 17; g = 18; h = 19; i = 20, 21; j = 22; k = 23, 24; l = 25, 26." httpUri="https://zenodo.org/record/217429/files/figure.png" pageId="4" pageNumber="524">Fig. 13</figureCitation>
). Head rounded, longer than wide (
<figureCitation id="9CDB2A016726FFF1C5AAFB7EFD63D553" box="[653,738,1266,1290]" captionStart="FIGURES 12 – 26" captionStartId="6.[151,264,1797,1820]" captionTargetBox="[253,1329,203,1766]" captionTargetId="figure@6.[239,1348,193,1776]" captionTargetPageId="6" captionText="FIGURES 12 – 26. Macrostylopyga spp., male, holotypes (12 – 17, 23, 24 — M. bidupi sp. nov., 18 – 22 — M. grandis sp. nov., 25, 26 — M. laosana (Anisyutkin, 2010). 12 — facial part of head; 13 — head, thorax and 1 – 7 th abdominal tergites from above; 14, 15 — abdominal apex from above (14) and below (15, hypandrium and genitalia removed); 16 — caudo-medial processes of paraprocts; 17 — hypandrium from below; 18, 22 — hind tarsus from outside (18) and below (22); 19 — arolium and claws; 20, 21, 23 – 26 — metatarsi: right fore (20, 23), right middle (21, 24), left hind (25, 26) from below (20, 21, 23, 24, 26) and outside (25). Dotted area shows membranous parts. Abbreviations: I – X — 1 st – 10 th abdominal tergites respectively, a. s.; ar.; e. r.; i. r.; pr.; par. — see text. Arrows show places of styli attachment. Scale bar (1 mm = e; 2 mm = h, i, k; 3 mm = d, j, l; 5 mm = a – c, f, g): a = 12; b = 13; c = 14; d = 15; e = 16; f = 17; g = 18; h = 19; i = 20, 21; j = 22; k = 23, 24; l = 25, 26." httpUri="https://zenodo.org/record/217429/files/figure.png" pageId="4" pageNumber="524">Fig. 12</figureCitation>
); eyes small; ocellar spots absent; distance between eyes 2.3 times eye length; distance between antennal sockets about 1.6 times scape length (
<quantity id="C3189B616726FFF1C36AFA9AFB2BD577" box="[1101,1194,1302,1326]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="4" pageNumber="524" unit="mm" value="1.3">1.3 mm</quantity>
); approximate length ratio of 3rd–5th segments of maxillary palps 1:1:1.2. Pronotum campaniform, slightly wider than long, distinctly narrowed anteriorly; anterior margin truncate, posterior margin weakly projected caudally (
<figureCitation id="9CDB2A016726FFF1C38DFAD2FA81D52F" box="[1194,1280,1374,1398]" captionStart="FIGURES 12 – 26" captionStartId="6.[151,264,1797,1820]" captionTargetBox="[253,1329,203,1766]" captionTargetId="figure@6.[239,1348,193,1776]" captionTargetPageId="6" captionText="FIGURES 12 – 26. Macrostylopyga spp., male, holotypes (12 – 17, 23, 24 — M. bidupi sp. nov., 18 – 22 — M. grandis sp. nov., 25, 26 — M. laosana (Anisyutkin, 2010). 12 — facial part of head; 13 — head, thorax and 1 – 7 th abdominal tergites from above; 14, 15 — abdominal apex from above (14) and below (15, hypandrium and genitalia removed); 16 — caudo-medial processes of paraprocts; 17 — hypandrium from below; 18, 22 — hind tarsus from outside (18) and below (22); 19 — arolium and claws; 20, 21, 23 – 26 — metatarsi: right fore (20, 23), right middle (21, 24), left hind (25, 26) from below (20, 21, 23, 24, 26) and outside (25). Dotted area shows membranous parts. Abbreviations: I – X — 1 st – 10 th abdominal tergites respectively, a. s.; ar.; e. r.; i. r.; pr.; par. — see text. Arrows show places of styli attachment. Scale bar (1 mm = e; 2 mm = h, i, k; 3 mm = d, j, l; 5 mm = a – c, f, g): a = 12; b = 13; c = 14; d = 15; e = 16; f = 17; g = 18; h = 19; i = 20, 21; j = 22; k = 23, 24; l = 25, 26." httpUri="https://zenodo.org/record/217429/files/figure.png" pageId="4" pageNumber="524">Fig. 13</figureCitation>
). Mesonotum and metanotum transverse, dorsally slightly convex, without visible ridges or impressions; posterior margins sinuate (
<figureCitation id="9CDB2A016726FFF1C7D2FA2AFEC9D5E7" box="[245,328,1446,1470]" captionStart="FIGURES 12 – 26" captionStartId="6.[151,264,1797,1820]" captionTargetBox="[253,1329,203,1766]" captionTargetId="figure@6.[239,1348,193,1776]" captionTargetPageId="6" captionText="FIGURES 12 – 26. Macrostylopyga spp., male, holotypes (12 – 17, 23, 24 — M. bidupi sp. nov., 18 – 22 — M. grandis sp. nov., 25, 26 — M. laosana (Anisyutkin, 2010). 12 — facial part of head; 13 — head, thorax and 1 – 7 th abdominal tergites from above; 14, 15 — abdominal apex from above (14) and below (15, hypandrium and genitalia removed); 16 — caudo-medial processes of paraprocts; 17 — hypandrium from below; 18, 22 — hind tarsus from outside (18) and below (22); 19 — arolium and claws; 20, 21, 23 – 26 — metatarsi: right fore (20, 23), right middle (21, 24), left hind (25, 26) from below (20, 21, 23, 24, 26) and outside (25). Dotted area shows membranous parts. Abbreviations: I – X — 1 st – 10 th abdominal tergites respectively, a. s.; ar.; e. r.; i. r.; pr.; par. — see text. Arrows show places of styli attachment. Scale bar (1 mm = e; 2 mm = h, i, k; 3 mm = d, j, l; 5 mm = a – c, f, g): a = 12; b = 13; c = 14; d = 15; e = 16; f = 17; g = 18; h = 19; i = 20, 21; j = 22; k = 23, 24; l = 25, 26." httpUri="https://zenodo.org/record/217429/files/figure.png" pageId="4" pageNumber="524">Fig. 13</figureCitation>
). Thoracal tergites and abdominal tergites II–VII marginated laterally (
<figureCitation id="9CDB2A016726FFF1C345FA2AFB34D5E7" box="[1122,1205,1446,1470]" captionStart="FIGURES 12 – 26" captionStartId="6.[151,264,1797,1820]" captionTargetBox="[253,1329,203,1766]" captionTargetId="figure@6.[239,1348,193,1776]" captionTargetPageId="6" captionText="FIGURES 12 – 26. Macrostylopyga spp., male, holotypes (12 – 17, 23, 24 — M. bidupi sp. nov., 18 – 22 — M. grandis sp. nov., 25, 26 — M. laosana (Anisyutkin, 2010). 12 — facial part of head; 13 — head, thorax and 1 – 7 th abdominal tergites from above; 14, 15 — abdominal apex from above (14) and below (15, hypandrium and genitalia removed); 16 — caudo-medial processes of paraprocts; 17 — hypandrium from below; 18, 22 — hind tarsus from outside (18) and below (22); 19 — arolium and claws; 20, 21, 23 – 26 — metatarsi: right fore (20, 23), right middle (21, 24), left hind (25, 26) from below (20, 21, 23, 24, 26) and outside (25). Dotted area shows membranous parts. Abbreviations: I – X — 1 st – 10 th abdominal tergites respectively, a. s.; ar.; e. r.; i. r.; pr.; par. — see text. Arrows show places of styli attachment. Scale bar (1 mm = e; 2 mm = h, i, k; 3 mm = d, j, l; 5 mm = a – c, f, g): a = 12; b = 13; c = 14; d = 15; e = 16; f = 17; g = 18; h = 19; i = 20, 21; j = 22; k = 23, 24; l = 25, 26." httpUri="https://zenodo.org/record/217429/files/figure.png" pageId="4" pageNumber="524">Fig. 13</figureCitation>
). Anterior margin of fore femur armed as in 
<typeStatus id="DB5B88266726FFF1C694FA47FE62D5BA" box="[435,483,1483,1507]" pageId="4" pageNumber="524">type</typeStatus>
A with 21–22 spines, including 3–4 apical spines. Fore tibiae slightly thickened distally, apical spines not reinforced; mid and hind tibiae not thickened. Tarsi damaged, only right foretarsus (claws damaged) and right metatarsus of middle tarsus present: both metatarsi without euplantulae (
<figureCitation id="9CDB2A016726FFF1C3EEF99EFAD9D673" box="[1225,1368,1554,1578]" captionStart="FIGURES 12 – 26" captionStartId="6.[151,264,1797,1820]" captionTargetBox="[253,1329,203,1766]" captionTargetId="figure@6.[239,1348,193,1776]" captionTargetPageId="6" captionText="FIGURES 12 – 26. Macrostylopyga spp., male, holotypes (12 – 17, 23, 24 — M. bidupi sp. nov., 18 – 22 — M. grandis sp. nov., 25, 26 — M. laosana (Anisyutkin, 2010). 12 — facial part of head; 13 — head, thorax and 1 – 7 th abdominal tergites from above; 14, 15 — abdominal apex from above (14) and below (15, hypandrium and genitalia removed); 16 — caudo-medial processes of paraprocts; 17 — hypandrium from below; 18, 22 — hind tarsus from outside (18) and below (22); 19 — arolium and claws; 20, 21, 23 – 26 — metatarsi: right fore (20, 23), right middle (21, 24), left hind (25, 26) from below (20, 21, 23, 24, 26) and outside (25). Dotted area shows membranous parts. Abbreviations: I – X — 1 st – 10 th abdominal tergites respectively, a. s.; ar.; e. r.; i. r.; pr.; par. — see text. Arrows show places of styli attachment. Scale bar (1 mm = e; 2 mm = h, i, k; 3 mm = d, j, l; 5 mm = a – c, f, g): a = 12; b = 13; c = 14; d = 15; e = 16; f = 17; g = 18; h = 19; i = 20, 21; j = 22; k = 23, 24; l = 25, 26." httpUri="https://zenodo.org/record/217429/files/figure.png" pageId="4" pageNumber="524">Figs. 23, 24</figureCitation>
); fore metatarsus slightly shorter than other segments combined, with 2 rows of spines along lower margin: exterior row with 15 spines (
<figureCitation id="9CDB2A016726FFF1C660F9D6FE18D62B" box="[327,409,1626,1650]" captionStart="FIGURES 12 – 26" captionStartId="6.[151,264,1797,1820]" captionTargetBox="[253,1329,203,1766]" captionTargetId="figure@6.[239,1348,193,1776]" captionTargetPageId="6" captionText="FIGURES 12 – 26. Macrostylopyga spp., male, holotypes (12 – 17, 23, 24 — M. bidupi sp. nov., 18 – 22 — M. grandis sp. nov., 25, 26 — M. laosana (Anisyutkin, 2010). 12 — facial part of head; 13 — head, thorax and 1 – 7 th abdominal tergites from above; 14, 15 — abdominal apex from above (14) and below (15, hypandrium and genitalia removed); 16 — caudo-medial processes of paraprocts; 17 — hypandrium from below; 18, 22 — hind tarsus from outside (18) and below (22); 19 — arolium and claws; 20, 21, 23 – 26 — metatarsi: right fore (20, 23), right middle (21, 24), left hind (25, 26) from below (20, 21, 23, 24, 26) and outside (25). Dotted area shows membranous parts. Abbreviations: I – X — 1 st – 10 th abdominal tergites respectively, a. s.; ar.; e. r.; i. r.; pr.; par. — see text. Arrows show places of styli attachment. Scale bar (1 mm = e; 2 mm = h, i, k; 3 mm = d, j, l; 5 mm = a – c, f, g): a = 12; b = 13; c = 14; d = 15; e = 16; f = 17; g = 18; h = 19; i = 20, 21; j = 22; k = 23, 24; l = 25, 26." httpUri="https://zenodo.org/record/217429/files/figure.png" pageId="4" pageNumber="524">Fig. 23</figureCitation>
, 
<emphasis id="3694EA966726FFF1C682F9D0FE46D62A" box="[421,455,1628,1651]" italics="true" pageId="4" pageNumber="524">e.r.</emphasis>
), interior row with 6 larger spines and 2 additional spines—(
<figureCitation id="9CDB2A016726FFF1C353F9D6FB43D62B" box="[1140,1218,1626,1650]" captionStart="FIGURES 12 – 26" captionStartId="6.[151,264,1797,1820]" captionTargetBox="[253,1329,203,1766]" captionTargetId="figure@6.[239,1348,193,1776]" captionTargetPageId="6" captionText="FIGURES 12 – 26. Macrostylopyga spp., male, holotypes (12 – 17, 23, 24 — M. bidupi sp. nov., 18 – 22 — M. grandis sp. nov., 25, 26 — M. laosana (Anisyutkin, 2010). 12 — facial part of head; 13 — head, thorax and 1 – 7 th abdominal tergites from above; 14, 15 — abdominal apex from above (14) and below (15, hypandrium and genitalia removed); 16 — caudo-medial processes of paraprocts; 17 — hypandrium from below; 18, 22 — hind tarsus from outside (18) and below (22); 19 — arolium and claws; 20, 21, 23 – 26 — metatarsi: right fore (20, 23), right middle (21, 24), left hind (25, 26) from below (20, 21, 23, 24, 26) and outside (25). Dotted area shows membranous parts. Abbreviations: I – X — 1 st – 10 th abdominal tergites respectively, a. s.; ar.; e. r.; i. r.; pr.; par. — see text. Arrows show places of styli attachment. Scale bar (1 mm = e; 2 mm = h, i, k; 3 mm = d, j, l; 5 mm = a – c, f, g): a = 12; b = 13; c = 14; d = 15; e = 16; f = 17; g = 18; h = 19; i = 20, 21; j = 22; k = 23, 24; l = 25, 26." httpUri="https://zenodo.org/record/217429/files/figure.png" pageId="4" pageNumber="524">Fig. 23</figureCitation>
, 
<emphasis id="3694EA966726FFF1C3E9F9D0FB6AD62A" box="[1230,1259,1628,1651]" italics="true" pageId="4" pageNumber="524">i.r.</emphasis>
, 
<emphasis id="3694EA966726FFF1C3DEF9D0FA9ED62A" box="[1273,1311,1628,1651]" italics="true" pageId="4" pageNumber="524">a.s.</emphasis>
) located at distal margin of metatarsus (this pair of spines evidently corresponding to pair of spines bordering metatarsal euplantula); second segment with row of 2 spines along lower margin, segments 2?4 with apical euplantulae bordered with pair of spines, arolium vestigial; medial metatarsus (
<figureCitation id="9CDB2A016726FFF1C4BFF94AFC6DD687" box="[920,1004,1734,1758]" captionStart="FIGURES 12 – 26" captionStartId="6.[151,264,1797,1820]" captionTargetBox="[253,1329,203,1766]" captionTargetId="figure@6.[239,1348,193,1776]" captionTargetPageId="6" captionText="FIGURES 12 – 26. Macrostylopyga spp., male, holotypes (12 – 17, 23, 24 — M. bidupi sp. nov., 18 – 22 — M. grandis sp. nov., 25, 26 — M. laosana (Anisyutkin, 2010). 12 — facial part of head; 13 — head, thorax and 1 – 7 th abdominal tergites from above; 14, 15 — abdominal apex from above (14) and below (15, hypandrium and genitalia removed); 16 — caudo-medial processes of paraprocts; 17 — hypandrium from below; 18, 22 — hind tarsus from outside (18) and below (22); 19 — arolium and claws; 20, 21, 23 – 26 — metatarsi: right fore (20, 23), right middle (21, 24), left hind (25, 26) from below (20, 21, 23, 24, 26) and outside (25). Dotted area shows membranous parts. Abbreviations: I – X — 1 st – 10 th abdominal tergites respectively, a. s.; ar.; e. r.; i. r.; pr.; par. — see text. Arrows show places of styli attachment. Scale bar (1 mm = e; 2 mm = h, i, k; 3 mm = d, j, l; 5 mm = a – c, f, g): a = 12; b = 13; c = 14; d = 15; e = 16; f = 17; g = 18; h = 19; i = 20, 21; j = 22; k = 23, 24; l = 25, 26." httpUri="https://zenodo.org/record/217429/files/figure.png" pageId="4" pageNumber="524">Fig. 24</figureCitation>
) with 13 and 8 spines in exterior and interior rows, respectively, and 2 additional spines located at distal margin of metatarsus. Abdominal tergites without visible glandular specializations (
<figureCitation id="9CDB2A016726FFF1C55AF882FC8FD77F" box="[637,782,1806,1830]" captionStart="FIGURES 12 – 26" captionStartId="6.[151,264,1797,1820]" captionTargetBox="[253,1329,203,1766]" captionTargetId="figure@6.[239,1348,193,1776]" captionTargetPageId="6" captionText="FIGURES 12 – 26. Macrostylopyga spp., male, holotypes (12 – 17, 23, 24 — M. bidupi sp. nov., 18 – 22 — M. grandis sp. nov., 25, 26 — M. laosana (Anisyutkin, 2010). 12 — facial part of head; 13 — head, thorax and 1 – 7 th abdominal tergites from above; 14, 15 — abdominal apex from above (14) and below (15, hypandrium and genitalia removed); 16 — caudo-medial processes of paraprocts; 17 — hypandrium from below; 18, 22 — hind tarsus from outside (18) and below (22); 19 — arolium and claws; 20, 21, 23 – 26 — metatarsi: right fore (20, 23), right middle (21, 24), left hind (25, 26) from below (20, 21, 23, 24, 26) and outside (25). Dotted area shows membranous parts. Abbreviations: I – X — 1 st – 10 th abdominal tergites respectively, a. s.; ar.; e. r.; i. r.; pr.; par. — see text. Arrows show places of styli attachment. Scale bar (1 mm = e; 2 mm = h, i, k; 3 mm = d, j, l; 5 mm = a – c, f, g): a = 12; b = 13; c = 14; d = 15; e = 16; f = 17; g = 18; h = 19; i = 20, 21; j = 22; k = 23, 24; l = 25, 26." httpUri="https://zenodo.org/record/217429/files/figure.png" pageId="4" pageNumber="524">Figs. 13, 14</figureCitation>
), posterolateral angles attenuate; tergite VI with caudal margin sinusoidally curved (
<figureCitation id="9CDB2A016726FFF1C6FEF8BEFDADD713" box="[473,556,1842,1866]" captionStart="FIGURES 12 – 26" captionStartId="6.[151,264,1797,1820]" captionTargetBox="[253,1329,203,1766]" captionTargetId="figure@6.[239,1348,193,1776]" captionTargetPageId="6" captionText="FIGURES 12 – 26. Macrostylopyga spp., male, holotypes (12 – 17, 23, 24 — M. bidupi sp. nov., 18 – 22 — M. grandis sp. nov., 25, 26 — M. laosana (Anisyutkin, 2010). 12 — facial part of head; 13 — head, thorax and 1 – 7 th abdominal tergites from above; 14, 15 — abdominal apex from above (14) and below (15, hypandrium and genitalia removed); 16 — caudo-medial processes of paraprocts; 17 — hypandrium from below; 18, 22 — hind tarsus from outside (18) and below (22); 19 — arolium and claws; 20, 21, 23 – 26 — metatarsi: right fore (20, 23), right middle (21, 24), left hind (25, 26) from below (20, 21, 23, 24, 26) and outside (25). Dotted area shows membranous parts. Abbreviations: I – X — 1 st – 10 th abdominal tergites respectively, a. s.; ar.; e. r.; i. r.; pr.; par. — see text. Arrows show places of styli attachment. Scale bar (1 mm = e; 2 mm = h, i, k; 3 mm = d, j, l; 5 mm = a – c, f, g): a = 12; b = 13; c = 14; d = 15; e = 16; f = 17; g = 18; h = 19; i = 20, 21; j = 22; k = 23, 24; l = 25, 26." httpUri="https://zenodo.org/record/217429/files/figure.png" pageId="4" pageNumber="524">Fig. 13</figureCitation>
); tergite VII roundly projected caudally, with small median incision (
<figureCitation id="9CDB2A016726FFF1C21CF8BEFA0ED713" box="[1339,1423,1842,1866]" captionStart="FIGURES 12 – 26" captionStartId="6.[151,264,1797,1820]" captionTargetBox="[253,1329,203,1766]" captionTargetId="figure@6.[239,1348,193,1776]" captionTargetPageId="6" captionText="FIGURES 12 – 26. Macrostylopyga spp., male, holotypes (12 – 17, 23, 24 — M. bidupi sp. nov., 18 – 22 — M. grandis sp. nov., 25, 26 — M. laosana (Anisyutkin, 2010). 12 — facial part of head; 13 — head, thorax and 1 – 7 th abdominal tergites from above; 14, 15 — abdominal apex from above (14) and below (15, hypandrium and genitalia removed); 16 — caudo-medial processes of paraprocts; 17 — hypandrium from below; 18, 22 — hind tarsus from outside (18) and below (22); 19 — arolium and claws; 20, 21, 23 – 26 — metatarsi: right fore (20, 23), right middle (21, 24), left hind (25, 26) from below (20, 21, 23, 24, 26) and outside (25). Dotted area shows membranous parts. Abbreviations: I – X — 1 st – 10 th abdominal tergites respectively, a. s.; ar.; e. r.; i. r.; pr.; par. — see text. Arrows show places of styli attachment. Scale bar (1 mm = e; 2 mm = h, i, k; 3 mm = d, j, l; 5 mm = a – c, f, g): a = 12; b = 13; c = 14; d = 15; e = 16; f = 17; g = 18; h = 19; i = 20, 21; j = 22; k = 23, 24; l = 25, 26." httpUri="https://zenodo.org/record/217429/files/figure.png" pageId="4" pageNumber="524">Fig. 13</figureCitation>
). Anal plate (ultimate tergite X) medially projected, with triangular median incision on caudal margin (
<figureCitation id="9CDB2A016726FFF1C21FF8DAFA0ED737" box="[1336,1423,1878,1902]" captionStart="FIGURES 12 – 26" captionStartId="6.[151,264,1797,1820]" captionTargetBox="[253,1329,203,1766]" captionTargetId="figure@6.[239,1348,193,1776]" captionTargetPageId="6" captionText="FIGURES 12 – 26. Macrostylopyga spp., male, holotypes (12 – 17, 23, 24 — M. bidupi sp. nov., 18 – 22 — M. grandis sp. nov., 25, 26 — M. laosana (Anisyutkin, 2010). 12 — facial part of head; 13 — head, thorax and 1 – 7 th abdominal tergites from above; 14, 15 — abdominal apex from above (14) and below (15, hypandrium and genitalia removed); 16 — caudo-medial processes of paraprocts; 17 — hypandrium from below; 18, 22 — hind tarsus from outside (18) and below (22); 19 — arolium and claws; 20, 21, 23 – 26 — metatarsi: right fore (20, 23), right middle (21, 24), left hind (25, 26) from below (20, 21, 23, 24, 26) and outside (25). Dotted area shows membranous parts. Abbreviations: I – X — 1 st – 10 th abdominal tergites respectively, a. s.; ar.; e. r.; i. r.; pr.; par. — see text. Arrows show places of styli attachment. Scale bar (1 mm = e; 2 mm = h, i, k; 3 mm = d, j, l; 5 mm = a – c, f, g): a = 12; b = 13; c = 14; d = 15; e = 16; f = 17; g = 18; h = 19; i = 20, 21; j = 22; k = 23, 24; l = 25, 26." httpUri="https://zenodo.org/record/217429/files/figure.png" pageId="4" pageNumber="524">Fig. 14</figureCitation>
). Cerci broken off. Paraprocts with hook-like processes on caudo-medial angles (
<figureCitation id="9CDB2A016726FFF1C314F8F6FB12D7CB" box="[1075,1171,1914,1938]" captionStart="FIGURES 12 – 26" captionStartId="6.[151,264,1797,1820]" captionTargetBox="[253,1329,203,1766]" captionTargetId="figure@6.[239,1348,193,1776]" captionTargetPageId="6" captionText="FIGURES 12 – 26. Macrostylopyga spp., male, holotypes (12 – 17, 23, 24 — M. bidupi sp. nov., 18 – 22 — M. grandis sp. nov., 25, 26 — M. laosana (Anisyutkin, 2010). 12 — facial part of head; 13 — head, thorax and 1 – 7 th abdominal tergites from above; 14, 15 — abdominal apex from above (14) and below (15, hypandrium and genitalia removed); 16 — caudo-medial processes of paraprocts; 17 — hypandrium from below; 18, 22 — hind tarsus from outside (18) and below (22); 19 — arolium and claws; 20, 21, 23 – 26 — metatarsi: right fore (20, 23), right middle (21, 24), left hind (25, 26) from below (20, 21, 23, 24, 26) and outside (25). Dotted area shows membranous parts. Abbreviations: I – X — 1 st – 10 th abdominal tergites respectively, a. s.; ar.; e. r.; i. r.; pr.; par. — see text. Arrows show places of styli attachment. Scale bar (1 mm = e; 2 mm = h, i, k; 3 mm = d, j, l; 5 mm = a – c, f, g): a = 12; b = 13; c = 14; d = 15; e = 16; f = 17; g = 18; h = 19; i = 20, 21; j = 22; k = 23, 24; l = 25, 26." httpUri="https://zenodo.org/record/217429/files/figure.png" pageId="4" pageNumber="524">Figs. 15</figureCitation>
, 
<emphasis id="3694EA966726FFF1C384F8F0FB46D7CA" box="[1187,1223,1916,1939]" italics="true" pageId="4" pageNumber="524">p.r.</emphasis>
, 16). Hypandrium nearly symmetrical (
<figureCitation id="9CDB2A016726FFF1C6A4F812FE58D7EF" box="[387,473,1950,1974]" captionStart="FIGURES 12 – 26" captionStartId="6.[151,264,1797,1820]" captionTargetBox="[253,1329,203,1766]" captionTargetId="figure@6.[239,1348,193,1776]" captionTargetPageId="6" captionText="FIGURES 12 – 26. Macrostylopyga spp., male, holotypes (12 – 17, 23, 24 — M. bidupi sp. nov., 18 – 22 — M. grandis sp. nov., 25, 26 — M. laosana (Anisyutkin, 2010). 12 — facial part of head; 13 — head, thorax and 1 – 7 th abdominal tergites from above; 14, 15 — abdominal apex from above (14) and below (15, hypandrium and genitalia removed); 16 — caudo-medial processes of paraprocts; 17 — hypandrium from below; 18, 22 — hind tarsus from outside (18) and below (22); 19 — arolium and claws; 20, 21, 23 – 26 — metatarsi: right fore (20, 23), right middle (21, 24), left hind (25, 26) from below (20, 21, 23, 24, 26) and outside (25). Dotted area shows membranous parts. Abbreviations: I – X — 1 st – 10 th abdominal tergites respectively, a. s.; ar.; e. r.; i. r.; pr.; par. — see text. Arrows show places of styli attachment. Scale bar (1 mm = e; 2 mm = h, i, k; 3 mm = d, j, l; 5 mm = a – c, f, g): a = 12; b = 13; c = 14; d = 15; e = 16; f = 17; g = 18; h = 19; i = 20, 21; j = 22; k = 23, 24; l = 25, 26." httpUri="https://zenodo.org/record/217429/files/figure.png" pageId="4" pageNumber="524">Fig. 17</figureCitation>
), with anterolateral parts (lateral sternal apodemes or apophyses 
<taxonomicName id="C3E04D076726FFF1C3EDF82CFA12D7EF" authority="Klass 1997" authorityName="Klass" authorityYear="1997" box="[1226,1427,1950,1975]" class="Insecta" family="Blattidae" genus="Neostylopyga" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="524" phylum="Arthropoda" rank="species" species="sensu">
<emphasis id="3694EA966726FFF1C3EDF82CFA89D7EE" box="[1226,1288,1952,1975]" italics="true" pageId="4" pageNumber="524">sensu</emphasis>
Klass 1997
</taxonomicName>
) short, posterior part trapezoidal in shape; styli broken off.
</paragraph>
<caption id="509F660C6727FFF0C7B0F8D7FBCFD7B5" httpUri="https://zenodo.org/record/217428/files/figure.png" pageId="5" pageNumber="525" targetBox="[210,1368,212,1821]" targetPageId="5">
<paragraph id="045F36846727FFF0C7B0F8D7FBCFD7B5" blockId="5.[151,1436,1883,2028]" pageId="5" pageNumber="525">
<emphasis id="3694EA966727FFF0C7B0F8D7FEC6D728" bold="true" box="[151,327,1883,1906]" pageId="5" pageNumber="525">FIGURES 1–11.</emphasis>
<taxonomicName id="C3E04D076727FFF0C677F8D0FDC8D72B" box="[336,585,1884,1906]" class="Insecta" family="Blattidae" genus="Macrostylopyga" kingdom="Animalia" order="Blattodea" pageId="5" pageNumber="525" phylum="Arthropoda" rank="species" species="grandis" status="sp. nov.">
<emphasis id="3694EA966727FFF0C677F8D0FDC8D72B" box="[336,585,1884,1906]" italics="true" pageId="5" pageNumber="525">Macrostylopyga grandis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2DA757ED6727FFF0C575F8D0FD24D728" box="[594,677,1884,1905]" pageId="5" pageNumber="525" rank="species">
<emphasis id="3694EA966727FFF0C575F8D0FD24D728" bold="true" box="[594,677,1884,1905]" pageId="5" pageNumber="525">sp. nov.</emphasis>
</taxonomicNameLabel>
(1, 3–7—male, holotype, 2, 8–11–female, paratype). 1, 2—facial part of head; 3—head, thorax and 1–6th abdominal tergites from above; 4–7th abdominal tergite from above; 5, 6, 8, 10—abdominal apex from above (5, 8) and below [6 (hypandrium and genitalia removed), 10]; 7—hypandrium from below; 9—outline of anal plate; 11—right cercus from outside. Dotted area shows membranous parts. Abbreviations: 
<emphasis id="3694EA966727FFF0C364F835FBE9D797" box="[1091,1128,1977,1998]" italics="true" pageId="5" pageNumber="525">I–X</emphasis>
—1st–10th abdominal tergites respectively, 
<emphasis id="3694EA966727FFF0C63BF85AFEC0D7B5" box="[284,321,2006,2028]" italics="true" pageId="5" pageNumber="525">d.e.</emphasis>
; 
<emphasis id="3694EA966727FFF0C669F85BFEEFD7B5" box="[334,366,2007,2028]" italics="true" pageId="5" pageNumber="525">i.o.</emphasis>
; 
<emphasis id="3694EA966727FFF0C65BF85BFE17D7B5" box="[380,406,2007,2028]" italics="true" pageId="5" pageNumber="525">pr.</emphasis>
; 
<emphasis id="3694EA966727FFF0C684F85BFE4AD7B5" box="[419,459,2007,2028]" italics="true" pageId="5" pageNumber="525">par.</emphasis>
—see text. Arrow shows damage of anal plate. Scale bar 5 mm.
</paragraph>
</caption>
<caption id="509F660C6724FFF3C7B0F889FB02D7AB" httpUri="https://zenodo.org/record/217429/files/figure.png" pageId="6" pageNumber="526" targetBox="[253,1329,203,1766]" targetPageId="6">
<paragraph id="045F36846724FFF3C7B0F889FB02D7AB" blockId="6.[151,1436,1797,2034]" pageId="6" pageNumber="526">
<emphasis id="3694EA966724FFF3C7B0F889FED5D743" bold="true" box="[151,340,1797,1820]" pageId="6" pageNumber="526">FIGURES 12–26.</emphasis>
<taxonomicName id="C3E04D076724FFF3C67EF889FE7DD742" box="[345,508,1797,1819]" class="Insecta" family="Blattidae" genus="Macrostylopyga" kingdom="Animalia" order="Blattodea" pageId="6" pageNumber="526" phylum="Arthropoda" rank="genus">
<emphasis id="3694EA966724FFF3C67EF889FE7DD742" box="[345,508,1797,1819]" italics="true" pageId="6" pageNumber="526">Macrostylopyga</emphasis>
</taxonomicName>
spp., male, holotypes (12–17, 23, 24— 
<taxonomicName id="C3E04D076724FFF3C4ABF88AFC6FD742" box="[908,1006,1797,1819]" class="Insecta" family="Blattidae" genus="Macrostylopyga" kingdom="Animalia" order="Blattodea" pageId="6" pageNumber="526" phylum="Arthropoda" rank="species" species="bidupi" status="sp. nov.">
<emphasis id="3694EA966724FFF3C4ABF88AFC6FD742" box="[908,1006,1797,1819]" italics="true" pageId="6" pageNumber="526">M. bidupi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2DA757ED6724FFF3C4D3F88AFBC5D742" box="[1012,1092,1798,1819]" pageId="6" pageNumber="526" rank="species">
<emphasis id="3694EA966724FFF3C4D3F88AFBC5D742" bold="true" box="[1012,1092,1798,1819]" pageId="6" pageNumber="526">sp. nov.</emphasis>
</taxonomicNameLabel>
, 18–22— 
<taxonomicName id="C3E04D076724FFF3C38DF88AFA98D742" box="[1194,1305,1797,1819]" class="Insecta" family="Blattidae" genus="Macrostylopyga" kingdom="Animalia" order="Blattodea" pageId="6" pageNumber="526" phylum="Arthropoda" rank="species" species="grandis" status="sp. nov.">
<emphasis id="3694EA966724FFF3C38DF88AFA98D742" box="[1194,1305,1797,1819]" italics="true" pageId="6" pageNumber="526">M. grandis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2DA757ED6724FFF3C238F88AFAEED742" box="[1311,1391,1798,1819]" pageId="6" pageNumber="526" rank="species">
<emphasis id="3694EA966724FFF3C238F88AFAEED742" bold="true" box="[1311,1391,1798,1819]" pageId="6" pageNumber="526">sp. nov.</emphasis>
</taxonomicNameLabel>
, 25, 26— 
<taxonomicName id="C3E04D076724FFF3C7EEF8A9FD89D763" authority="Anisyutkin, 2010" authorityName="Anisyutkin" authorityYear="2010" box="[201,520,1828,1850]" class="Insecta" family="Blattidae" genus="Macrostylopyga" kingdom="Animalia" order="Blattodea" pageId="6" pageNumber="526" phylum="Arthropoda" rank="species" species="laosana">
<emphasis id="3694EA966724FFF3C7EEF8A9FEBDD763" box="[201,316,1828,1850]" italics="true" pageId="6" pageNumber="526">M. laosana</emphasis>
(Anisyutkin, 2010)
</taxonomicName>
. 12—facial part of head; 13—head, thorax and 1–7th abdominal tergites from above; 14, 15—abdominal apex from above (14) and below (15, hypandrium and genitalia removed); 16—caudo-medial processes of paraprocts; 17—hypandrium from below; 18, 22—hind tarsus from outside (18) and below (22); 19—arolium and claws; 20, 21, 23–26—metatarsi: right fore (20, 23), right middle (21, 24), left hind (25, 26) from below (20, 21, 23, 24, 26) and outside (25). Dotted area shows membranous parts. Abbreviations: 
<emphasis id="3694EA966724FFF3C5D7F82CFC95D7EC" box="[752,788,1952,1973]" italics="true" pageId="6" pageNumber="526">I–X</emphasis>
—1st–10th abdominal tergites respectively, 
<emphasis id="3694EA966724FFF3C3F7F82CFB72D7EC" box="[1232,1267,1952,1973]" italics="true" pageId="6" pageNumber="526">a.s.</emphasis>
; 
<emphasis id="3694EA966724FFF3C227F82CFA9AD7EC" box="[1280,1307,1952,1973]" italics="true" pageId="6" pageNumber="526">ar.</emphasis>
; 
<emphasis id="3694EA966724FFF3C200F82CFAC6D7EC" box="[1319,1351,1952,1973]" italics="true" pageId="6" pageNumber="526">e.r.</emphasis>
; 
<emphasis id="3694EA966724FFF3C274F82CFAEFD7EC" box="[1363,1390,1952,1973]" italics="true" pageId="6" pageNumber="526">i.r.</emphasis>
; 
<emphasis id="3694EA966724FFF3C25CF82CFA17D7EC" box="[1403,1430,1952,1973]" italics="true" pageId="6" pageNumber="526">pr.</emphasis>
; 
<emphasis id="3694EA966724FFF3C7B0F832FF3ED78A" box="[151,191,1982,2003]" italics="true" pageId="6" pageNumber="526">par.</emphasis>
—see text. Arrows show places of styli attachment. Scale bar (1 mm = e; 2 mm = h, i, k; 3 mm = d, j, l; 5 mm = a–c, f, g): a = 12; b = 13; c = 14; d = 15; e = 16; f = 17; g = 18; h = 19; i = 20, 21; j = 22; k = 23, 24; l = 25, 26.
</paragraph>
</caption>
<caption id="509F660C6725FFF2C7B0F8EFFA09D78F" httpUri="https://zenodo.org/record/217430/files/figure.png" pageId="7" pageNumber="527" targetBox="[203,1384,204,1860]" targetPageId="7">
<paragraph id="045F36846725FFF2C7B0F8EFFA09D78F" blockId="7.[151,1437,1891,2006]" pageId="7" pageNumber="527">
<emphasis id="3694EA966725FFF2C7B0F8EFFED4D720" bold="true" box="[151,341,1891,1914]" pageId="7" pageNumber="527">FIGURES 27–38.</emphasis>
<taxonomicName id="C3E04D076725FFF2C67BF8E8FE7ED723" box="[348,511,1892,1914]" class="Insecta" family="Blattidae" genus="Macrostylopyga" kingdom="Animalia" order="Blattodea" pageId="7" pageNumber="527" phylum="Arthropoda" rank="genus">
<emphasis id="3694EA966725FFF2C67BF8E8FE7ED723" box="[348,511,1892,1914]" italics="true" pageId="7" pageNumber="527">Macrostylopyga</emphasis>
</taxonomicName>
spp., male genitalia, holotypes (27–30— 
<taxonomicName id="C3E04D076725FFF2C480F8E9FB99D723" box="[935,1048,1892,1914]" class="Insecta" family="Blattidae" genus="Macrostylopyga" kingdom="Animalia" order="Blattodea" pageId="7" pageNumber="527" phylum="Arthropoda" rank="species" species="grandis" status="sp. nov.">
<emphasis id="3694EA966725FFF2C480F8E9FB99D723" box="[935,1048,1892,1914]" italics="true" pageId="7" pageNumber="527">M. grandis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2DA757ED6725FFF2C338F8E8FBF0D720" box="[1055,1137,1892,1913]" pageId="7" pageNumber="527" rank="species">
<emphasis id="3694EA966725FFF2C338F8E8FBF0D720" bold="true" box="[1055,1137,1892,1913]" pageId="7" pageNumber="527">sp. nov.</emphasis>
</taxonomicNameLabel>
, 31–34— 
<taxonomicName id="C3E04D076725FFF2C3FFF8E9FABDD723" box="[1240,1340,1892,1914]" class="Insecta" family="Blattidae" genus="Macrostylopyga" kingdom="Animalia" order="Blattodea" pageId="7" pageNumber="527" phylum="Arthropoda" rank="species" species="bidupi" status="sp. nov.">
<emphasis id="3694EA966725FFF2C3FFF8E9FABDD723" box="[1240,1340,1892,1914]" italics="true" pageId="7" pageNumber="527">M. bidupi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2DA757ED6725FFF2C264F8E8FA14D720" box="[1347,1429,1892,1913]" pageId="7" pageNumber="527" rank="species">
<emphasis id="3694EA966725FFF2C264F8E8FA14D720" bold="true" box="[1347,1429,1892,1913]" pageId="7" pageNumber="527">sp. nov.</emphasis>
</taxonomicNameLabel>
, 35–38— 
<taxonomicName id="C3E04D076725FFF2C7C8F808FDAED7C0" authority="Anisyutkin, 2010" authorityName="Anisyutkin" authorityYear="2010" box="[239,559,1922,1945]" class="Insecta" family="Blattidae" genus="Macrostylopyga" kingdom="Animalia" order="Blattodea" pageId="7" pageNumber="527" phylum="Arthropoda" rank="species" species="laosana">
<emphasis id="3694EA966725FFF2C7C8F808FEE2D7C0" box="[239,355,1923,1945]" italics="true" pageId="7" pageNumber="527">M. laosana</emphasis>
(Anisyutkin, 2010)
</taxonomicName>
. Left phallomera from above (27, 31, 35), outside (28, 32, 36), inside (29, 33, partly, 37) and below (30, 34, partly, 38). Dotted area shows membranous parts. Figs. 35–38 after Anisyutkin, 2010. Abbreviations: 
<emphasis id="3694EA966725FFF2C7B0F84DFF4DD78F" box="[151,204,1985,2006]" italics="true" pageId="7" pageNumber="527">a.c.s.</emphasis>
; 
<emphasis id="3694EA966725FFF2C7FEF84DFF7ED78F" box="[217,255,1985,2006]" italics="true" pageId="7" pageNumber="527">a.p.</emphasis>
; 
<emphasis id="3694EA966725FFF2C62BF84DFEB8D78F" box="[268,313,1985,2006]" italics="true" pageId="7" pageNumber="527">c.pr.</emphasis>
; 
<emphasis id="3694EA966725FFF2C662F84CFEFCD78F" box="[325,381,1984,2006]" italics="true" pageId="7" pageNumber="527">d.c.p.</emphasis>
; 
<emphasis id="3694EA966725FFF2C6ADF84DFE3FD78F" box="[394,446,1985,2006]" italics="true" pageId="7" pageNumber="527">e.c.s.</emphasis>
; 
<emphasis id="3694EA966725FFF2C6ECF84CFE64D78F" box="[459,485,1984,2006]" italics="true" pageId="7" pageNumber="527">L1</emphasis>
; 
<emphasis id="3694EA966725FFF2C6D4F84CFD8FD78F" box="[499,526,1984,2006]" italics="true" pageId="7" pageNumber="527">L2</emphasis>
; 
<emphasis id="3694EA966725FFF2C53CF84CFDB7D78F" box="[539,566,1984,2006]" italics="true" pageId="7" pageNumber="527">L3</emphasis>
; 
<emphasis id="3694EA966725FFF2C564F84CFDEFD78F" box="[579,622,1984,2006]" italics="true" pageId="7" pageNumber="527">L4C</emphasis>
; 
<emphasis id="3694EA966725FFF2C55BF84CFD58D78F" box="[636,729,1984,2006]" italics="true" pageId="7" pageNumber="527">L4C, c.p.</emphasis>
; 
<emphasis id="3694EA966725FFF2C5C1F84CFC92D78F" box="[742,787,1984,2006]" italics="true" pageId="7" pageNumber="527">L4D</emphasis>
; 
<emphasis id="3694EA966725FFF2C407F84CFCCBD78F" box="[800,842,1984,2006]" italics="true" pageId="7" pageNumber="527">L4F</emphasis>
; 
<emphasis id="3694EA966725FFF2C47FF84CFCFFD78F" box="[856,894,1984,2006]" italics="true" pageId="7" pageNumber="527">m.l.</emphasis>
; 
<emphasis id="3694EA966725FFF2C4ACF84DFC3DD78F" box="[907,956,1984,2006]" italics="true" pageId="7" pageNumber="527">r.m.l.</emphasis>
; 
<emphasis id="3694EA966725FFF2C4EBF84DFC6BD78F" box="[972,1002,1985,2006]" italics="true" pageId="7" pageNumber="527">r.o.</emphasis>
; 
<emphasis id="3694EA966725FFF2C4DDF84CFB96D78F" box="[1018,1047,1984,2006]" italics="true" pageId="7" pageNumber="527">s.l.</emphasis>
; 
<emphasis id="3694EA966725FFF2C303F84CFBC0D78F" box="[1060,1089,1984,2006]" italics="true" pageId="7" pageNumber="527">sla</emphasis>
; 
<emphasis id="3694EA966725FFF2C377F84DFBF1D78F" box="[1104,1136,1985,2006]" italics="true" pageId="7" pageNumber="527">t.p.</emphasis>
—see text. Scale bar 2 mm.
</paragraph>
</caption>
<caption id="509F660C672AFFFDC7B0F8B0FE67D7B2" httpUri="https://zenodo.org/record/217431/files/figure.png" pageId="8" pageNumber="528" targetBox="[215,1394,216,1820]" targetPageId="8">
<paragraph id="045F3684672AFFFDC7B0F8B0FE67D7B2" blockId="8.[151,1437,1852,2027]" pageId="8" pageNumber="528">
<emphasis id="3694EA96672AFFFDC7B0F8B0FED5D708" bold="true" box="[151,340,1852,1875]" pageId="8" pageNumber="528">FIGURES 39–49.</emphasis>
<taxonomicName id="C3E04D07672AFFFDC67BF8B0FE7ED70B" box="[348,511,1852,1874]" class="Insecta" family="Blattidae" genus="Macrostylopyga" kingdom="Animalia" order="Blattodea" pageId="8" pageNumber="528" phylum="Arthropoda" rank="genus">
<emphasis id="3694EA96672AFFFDC67BF8B0FE7ED70B" box="[348,511,1852,1874]" italics="true" pageId="8" pageNumber="528">Macrostylopyga</emphasis>
</taxonomicName>
spp., male genitalia, holotypes (40, 42–49) and paratype (39, 41) (39–45— 
<taxonomicName id="C3E04D07672AFFFDC221F8B1FAF7D70B" box="[1286,1398,1852,1874]" class="Insecta" family="Blattidae" genus="Macrostylopyga" kingdom="Animalia" order="Blattodea" pageId="8" pageNumber="528" phylum="Arthropoda" rank="species" species="grandis" status="sp. nov.">
<emphasis id="3694EA96672AFFFDC221F8B1FAF7D70B" box="[1286,1398,1852,1874]" italics="true" pageId="8" pageNumber="528">M. grandis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2DA757ED672AFFFDC25AF8B1FF42D729" pageId="8" pageNumber="528" rank="species">
<emphasis id="3694EA96672AFFFDC25AF8B1FF42D729" bold="true" pageId="8" pageNumber="528">sp. nov.</emphasis>
</taxonomicNameLabel>
, 46–49— 
<taxonomicName id="C3E04D07672AFFFDC60FF8D0FE0AD728" box="[296,395,1883,1905]" class="Insecta" family="Blattidae" genus="Macrostylopyga" kingdom="Animalia" order="Blattodea" pageId="8" pageNumber="528" phylum="Arthropoda" rank="species" species="bidupi" status="sp. nov.">
<emphasis id="3694EA96672AFFFDC60FF8D0FE0AD728" box="[296,395,1883,1905]" italics="true" pageId="8" pageNumber="528">M. bidupi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2DA757ED672AFFFDC6B6F8D7FE60D729" box="[401,481,1883,1904]" pageId="8" pageNumber="528" rank="species">
<emphasis id="3694EA96672AFFFDC6B6F8D7FE60D729" bold="true" box="[401,481,1883,1904]" pageId="8" pageNumber="528">sp. nov.</emphasis>
</taxonomicNameLabel>
). 39, 40—left phallomere from above with sclerite L1 (39) and from caudal (40); 41—sclerite L1 and process 
<emphasis id="3694EA96672AFFFDC612F8F5FED2D7D6" box="[309,339,1913,1935]" italics="true" pageId="8" pageNumber="528">sla</emphasis>
of left phalomere from inside; 42, 46—sclerite L1 from outside (42) and inside (46); 43, 44, 47, 48—ventral phallomere from side (43, 47) and below (44, 48); 45, 49—sclerites R3 from outside. Dotted area shows membranous parts. Abbreviations: 
<emphasis id="3694EA96672AFFFDC613F83BFECFD794" box="[308,334,1975,1997]" italics="true" pageId="8" pageNumber="528">L1</emphasis>
; 
<emphasis id="3694EA96672AFFFDC67BF83BFEF6D794" box="[348,375,1975,1997]" italics="true" pageId="8" pageNumber="528">L2</emphasis>
; 
<emphasis id="3694EA96672AFFFDC6A3F83BFE1FD794" box="[388,414,1975,1997]" italics="true" pageId="8" pageNumber="528">L3</emphasis>
; 
<emphasis id="3694EA96672AFFFDC68BF83BFE56D794" box="[428,471,1975,1997]" italics="true" pageId="8" pageNumber="528">L4C</emphasis>
; 
<emphasis id="3694EA96672AFFFDC6C3F83BFD90D794" box="[484,529,1975,1997]" italics="true" pageId="8" pageNumber="528">L4D</emphasis>
; 
<emphasis id="3694EA96672AFFFDC538F834FDC4D794" box="[543,581,1976,1997]" italics="true" pageId="8" pageNumber="528">m.i.</emphasis>
; 
<emphasis id="3694EA96672AFFFDC575F834FDF1D794" box="[594,624,1976,1997]" italics="true" pageId="8" pageNumber="528">r.o.</emphasis>
; 
<emphasis id="3694EA96672AFFFDC5A7F83BFD1CD794" box="[640,669,1975,1997]" italics="true" pageId="8" pageNumber="528">sla</emphasis>
; 
<emphasis id="3694EA96672AFFFDC58CF834FD4AD794" box="[683,715,1976,1997]" italics="true" pageId="8" pageNumber="528">t.p.</emphasis>
—see text. Scale bar (2 mm): a = 39; b = 40; c = 41; d = 42; e = 43, 44; f = 45; g = 46; h = 47, 48; i = 49.
</paragraph>
</caption>
<caption id="509F660C672BFFFCC7B0F8CAFD62D7BA" httpUri="https://zenodo.org/record/217432/files/figure.png" pageId="9" pageNumber="529" targetBox="[218,1362,203,1830]" targetPageId="9">
<paragraph id="045F3684672BFFFCC7B0F8CAFD62D7BA" blockId="9.[151,1436,1862,2019]" pageId="9" pageNumber="529">
<emphasis id="3694EA96672BFFFCC7B0F8CAFED4D705" bold="true" box="[151,341,1862,1885]" pageId="9" pageNumber="529">FIGURES 50–56.</emphasis>
<taxonomicName id="C3E04D07672BFFFCC67BF8CBFE7ED704" box="[348,511,1863,1885]" class="Insecta" family="Blattidae" genus="Macrostylopyga" kingdom="Animalia" order="Blattodea" pageId="9" pageNumber="529" phylum="Arthropoda" rank="genus">
<emphasis id="3694EA96672BFFFCC67BF8CBFE7ED704" box="[348,511,1863,1885]" italics="true" pageId="9" pageNumber="529">Macrostylopyga</emphasis>
</taxonomicName>
spp., male genitalia, holotypes (50–52— 
<taxonomicName id="C3E04D07672BFFFCC480F8C4FB99D704" box="[935,1048,1863,1885]" class="Insecta" family="Blattidae" genus="Macrostylopyga" kingdom="Animalia" order="Blattodea" pageId="9" pageNumber="529" phylum="Arthropoda" rank="species" species="grandis" status="sp. nov.">
<emphasis id="3694EA96672BFFFCC480F8C4FB99D704" box="[935,1048,1863,1885]" italics="true" pageId="9" pageNumber="529">M. grandis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2DA757ED672BFFFCC338F8CBFBF0D705" box="[1055,1137,1863,1884]" pageId="9" pageNumber="529" rank="species">
<emphasis id="3694EA96672BFFFCC338F8CBFBF0D705" bold="true" box="[1055,1137,1863,1884]" pageId="9" pageNumber="529">sp. nov.</emphasis>
</taxonomicNameLabel>
, 53–55— 
<taxonomicName id="C3E04D07672BFFFCC3FFF8C4FABDD704" box="[1240,1340,1863,1885]" class="Insecta" family="Blattidae" genus="Macrostylopyga" kingdom="Animalia" order="Blattodea" pageId="9" pageNumber="529" phylum="Arthropoda" rank="species" species="bidupi" status="sp. nov.">
<emphasis id="3694EA96672BFFFCC3FFF8C4FABDD704" box="[1240,1340,1863,1885]" italics="true" pageId="9" pageNumber="529">M. bidupi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2DA757ED672BFFFCC264F8CBFA14D705" box="[1347,1429,1863,1884]" pageId="9" pageNumber="529" rank="species">
<emphasis id="3694EA96672BFFFCC264F8CBFA14D705" bold="true" box="[1347,1429,1863,1884]" pageId="9" pageNumber="529">sp. nov.</emphasis>
</taxonomicNameLabel>
, 56— 
<taxonomicName id="C3E04D07672BFFFCC7EEF8E5FD8AD727" authority="Anisyutkin, 2010" authorityName="Anisyutkin" authorityYear="2010" box="[201,523,1896,1918]" class="Insecta" family="Blattidae" genus="Macrostylopyga" kingdom="Animalia" order="Blattodea" pageId="9" pageNumber="529" phylum="Arthropoda" rank="species" species="laosana">
<emphasis id="3694EA96672BFFFCC7EEF8E5FEBCD727" box="[201,317,1896,1918]" italics="true" pageId="9" pageNumber="529">M. laosana</emphasis>
(Anisyutkin, 2010)
</taxonomicName>
. 50, 51, 53, 54—right phallomere from above (50, 53) and below (51, 54); 52—sclerite R1H; 55—sclerites R1H and R1G from dorso-caudal; 56—outline of sclerite R1H processes. Dotted area shows membranous parts. Fig. 56 after Anisyutkin, 2010. Abbreviations: 
<emphasis id="3694EA96672BFFFCC595F827FD57D799" box="[690,726,1963,1984]" italics="true" pageId="9" pageNumber="529">p.s.</emphasis>
; 
<emphasis id="3694EA96672BFFFCC5C4F826FC93D799" box="[739,786,1962,1984]" italics="true" pageId="9" pageNumber="529">R1G</emphasis>
; 
<emphasis id="3694EA96672BFFFCC438F826FCCFD799" box="[799,846,1962,1984]" italics="true" pageId="9" pageNumber="529">R1H</emphasis>
; 
<emphasis id="3694EA96672BFFFCC47CF826FCF6D799" box="[859,887,1962,1984]" italics="true" pageId="9" pageNumber="529">R2</emphasis>
; 
<emphasis id="3694EA96672BFFFCC4A2F826FC20D799" box="[901,929,1962,1984]" italics="true" pageId="9" pageNumber="529">R3</emphasis>
; 
<emphasis id="3694EA96672BFFFCC488F827FC4DD799" box="[943,972,1963,1984]" italics="true" pageId="9" pageNumber="529">t.s.</emphasis>
—see text. Scale bar (2 mm = a–c, g; 1 mm = d–f): a = 50; b = 51; c = 52; d = 53; e = 54; f = 55; g = 56.
</paragraph>
</caption>
<paragraph id="045F36846728FFFFC7E0FF14FBCED2DD" blockId="10.[151,1436,151,895]" pageId="10" pageNumber="530">
<emphasis id="3694EA966728FFFFC7E0FF14FEF7D0E9" bold="true" box="[199,374,151,176]" pageId="10" pageNumber="530">Male genitalia</emphasis>
. Left phallomere (
<figureCitation id="9CDB2A016728FFFFC574FF1BFD61D0F6" box="[595,736,151,175]" captionStart="FIGURES 27 – 38" captionStartId="7.[151,264,1891,1914]" captionTargetBox="[203,1384,204,1860]" captionTargetId="figure@7.[200,1387,193,1870]" captionTargetPageId="7" captionText="FIGURES 27 – 38. Macrostylopyga spp., male genitalia, holotypes (27 – 30 — M. grandis sp. nov., 31 – 34 — M. bidupi sp. nov., 35 – 38 — M. laosana (Anisyutkin, 2010). Left phallomera from above (27, 31, 35), outside (28, 32, 36), inside (29, 33, partly, 37) and below (30, 34, partly, 38). Dotted area shows membranous parts. Figs. 35 – 38 after Anisyutkin, 2010. Abbreviations: a. c. s.; a. p.; c. pr.; d. c. p.; e. c. s.; L 1; L 2; L 3; L 4 C; L 4 C, c. p.; L 4 D; L 4 F; m. l.; r. m. l.; r. o.; s. l.; sla; t. p. — see text. Scale bar 2 mm." httpUri="https://zenodo.org/record/217430/files/figure.png" pageId="10" pageNumber="530">Figs. 31–34</figureCitation>
, 
<figureCitation id="9CDB2A016728FFFFC5C8FF1BFC8CD0F6" box="[751,781,151,175]" captionStart="FIGURES 39 – 49" captionStartId="8.[151,264,1852,1875]" captionTargetBox="[215,1394,216,1820]" captionTargetId="figure@8.[167,1419,193,1847]" captionTargetPageId="8" captionText="FIGURES 39 – 49. Macrostylopyga spp., male genitalia, holotypes (40, 42 – 49) and paratype (39, 41) (39 – 45 — M. grandis sp. nov., 46 – 49 — M. bidupi sp. nov.). 39, 40 — left phallomere from above with sclerite L 1 (39) and from caudal (40); 41 — sclerite L 1 and process sla of left phalomere from inside; 42, 46 — sclerite L 1 from outside (42) and inside (46); 43, 44, 47, 48 — ventral phallomere from side (43, 47) and below (44, 48); 45, 49 — sclerites R 3 from outside. Dotted area shows membranous parts. Abbreviations: L 1; L 2; L 3; L 4 C; L 4 D; m. i.; r. o.; sla; t. p. — see text. Scale bar (2 mm): a = 39; b = 40; c = 41; d = 42; e = 43, 44; f = 45; g = 46; h = 47, 48; i = 49." httpUri="https://zenodo.org/record/217431/files/figure.png" pageId="10" pageNumber="530">46</figureCitation>
) with sclerite L4C (L2D) large, caudally widened, bent ventrally in cranial part (
<figureCitation id="9CDB2A016728FFFFC688FF30FDB6D08D" box="[431,567,188,212]" captionStart="FIGURES 27 – 38" captionStartId="7.[151,264,1891,1914]" captionTargetBox="[203,1384,204,1860]" captionTargetId="figure@7.[200,1387,193,1870]" captionTargetPageId="7" captionText="FIGURES 27 – 38. Macrostylopyga spp., male genitalia, holotypes (27 – 30 — M. grandis sp. nov., 31 – 34 — M. bidupi sp. nov., 35 – 38 — M. laosana (Anisyutkin, 2010). Left phallomera from above (27, 31, 35), outside (28, 32, 36), inside (29, 33, partly, 37) and below (30, 34, partly, 38). Dotted area shows membranous parts. Figs. 35 – 38 after Anisyutkin, 2010. Abbreviations: a. c. s.; a. p.; c. pr.; d. c. p.; e. c. s.; L 1; L 2; L 3; L 4 C; L 4 C, c. p.; L 4 D; L 4 F; m. l.; r. m. l.; r. o.; s. l.; sla; t. p. — see text. Scale bar 2 mm." httpUri="https://zenodo.org/record/217430/files/figure.png" pageId="10" pageNumber="530">Figs. 32, 33</figureCitation>
, 
<emphasis id="3694EA966728FFFFC564FF31FDEAD08D" box="[579,619,189,212]" italics="true" pageId="10" pageNumber="530">c.p.</emphasis>
); process 
<emphasis id="3694EA966728FFFFC5C7FF30FC81D08D" box="[736,768,188,212]" italics="true" pageId="10" pageNumber="530">sla</emphasis>
simple, directed ventro-medially (
<figureCitation id="9CDB2A016728FFFFC3A1FF30FA8ED08D" box="[1158,1295,188,212]" captionStart="FIGURES 27 – 38" captionStartId="7.[151,264,1891,1914]" captionTargetBox="[203,1384,204,1860]" captionTargetId="figure@7.[200,1387,193,1870]" captionTargetPageId="7" captionText="FIGURES 27 – 38. Macrostylopyga spp., male genitalia, holotypes (27 – 30 — M. grandis sp. nov., 31 – 34 — M. bidupi sp. nov., 35 – 38 — M. laosana (Anisyutkin, 2010). Left phallomera from above (27, 31, 35), outside (28, 32, 36), inside (29, 33, partly, 37) and below (30, 34, partly, 38). Dotted area shows membranous parts. Figs. 35 – 38 after Anisyutkin, 2010. Abbreviations: a. c. s.; a. p.; c. pr.; d. c. p.; e. c. s.; L 1; L 2; L 3; L 4 C; L 4 C, c. p.; L 4 D; L 4 F; m. l.; r. m. l.; r. o.; s. l.; sla; t. p. — see text. Scale bar 2 mm." httpUri="https://zenodo.org/record/217430/files/figure.png" pageId="10" pageNumber="530">Figs. 31, 33</figureCitation>
); sclerite L3 (L3d) as in 
<figureCitation id="9CDB2A016728FFFFC638FF53FE28D0AE" box="[287,425,223,247]" captionStart="FIGURES 27 – 38" captionStartId="7.[151,264,1891,1914]" captionTargetBox="[203,1384,204,1860]" captionTargetId="figure@7.[200,1387,193,1870]" captionTargetPageId="7" captionText="FIGURES 27 – 38. Macrostylopyga spp., male genitalia, holotypes (27 – 30 — M. grandis sp. nov., 31 – 34 — M. bidupi sp. nov., 35 – 38 — M. laosana (Anisyutkin, 2010). Left phallomera from above (27, 31, 35), outside (28, 32, 36), inside (29, 33, partly, 37) and below (30, 34, partly, 38). Dotted area shows membranous parts. Figs. 35 – 38 after Anisyutkin, 2010. Abbreviations: a. c. s.; a. p.; c. pr.; d. c. p.; e. c. s.; L 1; L 2; L 3; L 4 C; L 4 C, c. p.; L 4 D; L 4 F; m. l.; r. m. l.; r. o.; s. l.; sla; t. p. — see text. Scale bar 2 mm." httpUri="https://zenodo.org/record/217430/files/figure.png" pageId="10" pageNumber="530">Figs. 31–34</figureCitation>
; curved sclerite L4F, possibly L4E+L4F, and small sclerite L4D (L 
<date id="705E10446728FFFFC397FF53FB4AD0AE" box="[1200,1227,223,247]" pageId="10" pageNumber="530">3v</date>
) situated on outer side of phallomere (
<figureCitation id="9CDB2A016728FFFFC6A7FE88FE56D145" box="[384,471,260,284]" captionStart="FIGURES 27 – 38" captionStartId="7.[151,264,1891,1914]" captionTargetBox="[203,1384,204,1860]" captionTargetId="figure@7.[200,1387,193,1870]" captionTargetPageId="7" captionText="FIGURES 27 – 38. Macrostylopyga spp., male genitalia, holotypes (27 – 30 — M. grandis sp. nov., 31 – 34 — M. bidupi sp. nov., 35 – 38 — M. laosana (Anisyutkin, 2010). Left phallomera from above (27, 31, 35), outside (28, 32, 36), inside (29, 33, partly, 37) and below (30, 34, partly, 38). Dotted area shows membranous parts. Figs. 35 – 38 after Anisyutkin, 2010. Abbreviations: a. c. s.; a. p.; c. pr.; d. c. p.; e. c. s.; L 1; L 2; L 3; L 4 C; L 4 C, c. p.; L 4 D; L 4 F; m. l.; r. m. l.; r. o.; s. l.; sla; t. p. — see text. Scale bar 2 mm." httpUri="https://zenodo.org/record/217430/files/figure.png" pageId="10" pageNumber="530">Fig. 32</figureCitation>
); large sclerite L2 (L 
<date id="705E10446728FFFFC5E8FE88FD6DD145" box="[719,748,260,284]" pageId="10" pageNumber="530">2v</date>
) occupying ventral side of phallomere, partly extending on inner side (
<figureCitation id="9CDB2A016728FFFFC63FFEABFE25D166" box="[280,420,295,319]" captionStart="FIGURES 27 – 38" captionStartId="7.[151,264,1891,1914]" captionTargetBox="[203,1384,204,1860]" captionTargetId="figure@7.[200,1387,193,1870]" captionTargetPageId="7" captionText="FIGURES 27 – 38. Macrostylopyga spp., male genitalia, holotypes (27 – 30 — M. grandis sp. nov., 31 – 34 — M. bidupi sp. nov., 35 – 38 — M. laosana (Anisyutkin, 2010). Left phallomera from above (27, 31, 35), outside (28, 32, 36), inside (29, 33, partly, 37) and below (30, 34, partly, 38). Dotted area shows membranous parts. Figs. 35 – 38 after Anisyutkin, 2010. Abbreviations: a. c. s.; a. p.; c. pr.; d. c. p.; e. c. s.; L 1; L 2; L 3; L 4 C; L 4 C, c. p.; L 4 D; L 4 F; m. l.; r. m. l.; r. o.; s. l.; sla; t. p. — see text. Scale bar 2 mm." httpUri="https://zenodo.org/record/217430/files/figure.png" pageId="10" pageNumber="530">Figs. 33, 34</figureCitation>
), with forked spiniform curved process caudolaterally (
<figureCitation id="9CDB2A016728FFFFC338FEABFB2AD166" box="[1055,1195,295,319]" captionStart="FIGURES 27 – 38" captionStartId="7.[151,264,1891,1914]" captionTargetBox="[203,1384,204,1860]" captionTargetId="figure@7.[200,1387,193,1870]" captionTargetPageId="7" captionText="FIGURES 27 – 38. Macrostylopyga spp., male genitalia, holotypes (27 – 30 — M. grandis sp. nov., 31 – 34 — M. bidupi sp. nov., 35 – 38 — M. laosana (Anisyutkin, 2010). Left phallomera from above (27, 31, 35), outside (28, 32, 36), inside (29, 33, partly, 37) and below (30, 34, partly, 38). Dotted area shows membranous parts. Figs. 35 – 38 after Anisyutkin, 2010. Abbreviations: a. c. s.; a. p.; c. pr.; d. c. p.; e. c. s.; L 1; L 2; L 3; L 4 C; L 4 C, c. p.; L 4 D; L 4 F; m. l.; r. m. l.; r. o.; s. l.; sla; t. p. — see text. Scale bar 2 mm." httpUri="https://zenodo.org/record/217430/files/figure.png" pageId="10" pageNumber="530">Figs. 32, 34</figureCitation>
, 
<emphasis id="3694EA966728FFFFC39EFEA5FB68D119" box="[1209,1257,297,320]" italics="true" pageId="10" pageNumber="530">c.pr.</emphasis>
), caudally with membranous lobe; &quot;additional elongated curved sclerite&quot;, probably separated part of L2, situated on inner side of phallomere dorsally to L2 (
<figureCitation id="9CDB2A016728FFFFC6E0FEE3FD98D1DE" box="[455,537,367,391]" captionStart="FIGURES 27 – 38" captionStartId="7.[151,264,1891,1914]" captionTargetBox="[203,1384,204,1860]" captionTargetId="figure@7.[200,1387,193,1870]" captionTargetPageId="7" captionText="FIGURES 27 – 38. Macrostylopyga spp., male genitalia, holotypes (27 – 30 — M. grandis sp. nov., 31 – 34 — M. bidupi sp. nov., 35 – 38 — M. laosana (Anisyutkin, 2010). Left phallomera from above (27, 31, 35), outside (28, 32, 36), inside (29, 33, partly, 37) and below (30, 34, partly, 38). Dotted area shows membranous parts. Figs. 35 – 38 after Anisyutkin, 2010. Abbreviations: a. c. s.; a. p.; c. pr.; d. c. p.; e. c. s.; L 1; L 2; L 3; L 4 C; L 4 C, c. p.; L 4 D; L 4 F; m. l.; r. m. l.; r. o.; s. l.; sla; t. p. — see text. Scale bar 2 mm." httpUri="https://zenodo.org/record/217430/files/figure.png" pageId="10" pageNumber="530">Fig. 33</figureCitation>
, 
<emphasis id="3694EA966728FFFFC502FEFDFDDCD1D1" box="[549,605,369,392]" italics="true" pageId="10" pageNumber="530">e.c.s.</emphasis>
), accompanied with moderately sclerotized lobe, covered with sclerotized tubercles in medial part (
<figureCitation id="9CDB2A016728FFFFC68BFE18FDB2D1F5" box="[428,563,404,428]" captionStart="FIGURES 27 – 38" captionStartId="7.[151,264,1891,1914]" captionTargetBox="[203,1384,204,1860]" captionTargetId="figure@7.[200,1387,193,1870]" captionTargetPageId="7" captionText="FIGURES 27 – 38. Macrostylopyga spp., male genitalia, holotypes (27 – 30 — M. grandis sp. nov., 31 – 34 — M. bidupi sp. nov., 35 – 38 — M. laosana (Anisyutkin, 2010). Left phallomera from above (27, 31, 35), outside (28, 32, 36), inside (29, 33, partly, 37) and below (30, 34, partly, 38). Dotted area shows membranous parts. Figs. 35 – 38 after Anisyutkin, 2010. Abbreviations: a. c. s.; a. p.; c. pr.; d. c. p.; e. c. s.; L 1; L 2; L 3; L 4 C; L 4 C, c. p.; L 4 D; L 4 F; m. l.; r. m. l.; r. o.; s. l.; sla; t. p. — see text. Scale bar 2 mm." httpUri="https://zenodo.org/record/217430/files/figure.png" pageId="10" pageNumber="530">Figs. 31, 33</figureCitation>
, 
<emphasis id="3694EA966728FFFFC519FE18FDDFD1F5" box="[574,606,404,428]" italics="true" pageId="10" pageNumber="530">s.l.</emphasis>
) and protruded membranous lobe caudally (
<figureCitation id="9CDB2A016728FFFFC36AFE18FB7FD1F5" box="[1101,1278,404,428]" captionStart="FIGURES 27 – 38" captionStartId="7.[151,264,1891,1914]" captionTargetBox="[203,1384,204,1860]" captionTargetId="figure@7.[200,1387,193,1870]" captionTargetPageId="7" captionText="FIGURES 27 – 38. Macrostylopyga spp., male genitalia, holotypes (27 – 30 — M. grandis sp. nov., 31 – 34 — M. bidupi sp. nov., 35 – 38 — M. laosana (Anisyutkin, 2010). Left phallomera from above (27, 31, 35), outside (28, 32, 36), inside (29, 33, partly, 37) and below (30, 34, partly, 38). Dotted area shows membranous parts. Figs. 35 – 38 after Anisyutkin, 2010. Abbreviations: a. c. s.; a. p.; c. pr.; d. c. p.; e. c. s.; L 1; L 2; L 3; L 4 C; L 4 C, c. p.; L 4 D; L 4 F; m. l.; r. m. l.; r. o.; s. l.; sla; t. p. — see text. Scale bar 2 mm." httpUri="https://zenodo.org/record/217430/files/figure.png" pageId="10" pageNumber="530">Figs. 31, 33, 34</figureCitation>
, 
<emphasis id="3694EA966728FFFFC22EFE18FAB2D1F5" box="[1289,1331,404,428]" italics="true" pageId="10" pageNumber="530">m.l.</emphasis>
); sclerite L1 simple, large, plate-like, tripartite caudally (
<figureCitation id="9CDB2A016728FFFFC5E3FE3BFC9CD196" box="[708,797,439,463]" captionStart="FIGURES 39 – 49" captionStartId="8.[151,264,1852,1875]" captionTargetBox="[215,1394,216,1820]" captionTargetId="figure@8.[167,1419,193,1847]" captionTargetPageId="8" captionText="FIGURES 39 – 49. Macrostylopyga spp., male genitalia, holotypes (40, 42 – 49) and paratype (39, 41) (39 – 45 — M. grandis sp. nov., 46 – 49 — M. bidupi sp. nov.). 39, 40 — left phallomere from above with sclerite L 1 (39) and from caudal (40); 41 — sclerite L 1 and process sla of left phalomere from inside; 42, 46 — sclerite L 1 from outside (42) and inside (46); 43, 44, 47, 48 — ventral phallomere from side (43, 47) and below (44, 48); 45, 49 — sclerites R 3 from outside. Dotted area shows membranous parts. Abbreviations: L 1; L 2; L 3; L 4 C; L 4 D; m. i.; r. o.; sla; t. p. — see text. Scale bar (2 mm): a = 39; b = 40; c = 41; d = 42; e = 43, 44; f = 45; g = 46; h = 47, 48; i = 49." httpUri="https://zenodo.org/record/217431/files/figure.png" pageId="10" pageNumber="530">Fig. 46</figureCitation>
). Ventral phallomere L4G (VP) elongated and slightly curved (
<figureCitation id="9CDB2A016728FFFFC7DDFE50FE12D1AD" box="[250,403,476,500]" captionStart="FIGURES 39 – 49" captionStartId="8.[151,264,1852,1875]" captionTargetBox="[215,1394,216,1820]" captionTargetId="figure@8.[167,1419,193,1847]" captionTargetPageId="8" captionText="FIGURES 39 – 49. Macrostylopyga spp., male genitalia, holotypes (40, 42 – 49) and paratype (39, 41) (39 – 45 — M. grandis sp. nov., 46 – 49 — M. bidupi sp. nov.). 39, 40 — left phallomere from above with sclerite L 1 (39) and from caudal (40); 41 — sclerite L 1 and process sla of left phalomere from inside; 42, 46 — sclerite L 1 from outside (42) and inside (46); 43, 44, 47, 48 — ventral phallomere from side (43, 47) and below (44, 48); 45, 49 — sclerites R 3 from outside. Dotted area shows membranous parts. Abbreviations: L 1; L 2; L 3; L 4 C; L 4 D; m. i.; r. o.; sla; t. p. — see text. Scale bar (2 mm): a = 39; b = 40; c = 41; d = 42; e = 43, 44; f = 45; g = 46; h = 47, 48; i = 49." httpUri="https://zenodo.org/record/217431/files/figure.png" pageId="10" pageNumber="530">Figs. 47, 48</figureCitation>
). Right phallomere complex in shape (
<figureCitation id="9CDB2A016728FFFFC454FE50FB85D1AD" box="[883,1028,476,500]" captionStart="FIGURES 50 – 56" captionStartId="9.[151,264,1862,1885]" captionTargetBox="[218,1362,203,1830]" captionTargetId="figure@9.[196,1392,193,1841]" captionTargetPageId="9" captionText="FIGURES 50 – 56. Macrostylopyga spp., male genitalia, holotypes (50 – 52 — M. grandis sp. nov., 53 – 55 — M. bidupi sp. nov., 56 — M. laosana (Anisyutkin, 2010). 50, 51, 53, 54 — right phallomere from above (50, 53) and below (51, 54); 52 — sclerite R 1 H; 55 — sclerites R 1 H and R 1 G from dorso-caudal; 56 — outline of sclerite R 1 H processes. Dotted area shows membranous parts. Fig. 56 after Anisyutkin, 2010. Abbreviations: p. s.; R 1 G; R 1 H; R 2; R 3; t. s. — see text. Scale bar (2 mm = a – c, g; 1 mm = d – f): a = 50; b = 51; c = 52; d = 53; e = 54; f = 55; g = 56." httpUri="https://zenodo.org/record/217432/files/figure.png" pageId="10" pageNumber="530">Figs. 53–55</figureCitation>
); basal sclerite (R2) rounded and dorsoventrally compressed, with transverse groove; sclerite R1H with single long, slender process directed caudomedio-ventrally (
<figureCitation id="9CDB2A016728FFFFC67EFDA8FE60D265" box="[345,481,548,572]" captionStart="FIGURES 50 – 56" captionStartId="9.[151,264,1862,1885]" captionTargetBox="[218,1362,203,1830]" captionTargetId="figure@9.[196,1392,193,1841]" captionTargetPageId="9" captionText="FIGURES 50 – 56. Macrostylopyga spp., male genitalia, holotypes (50 – 52 — M. grandis sp. nov., 53 – 55 — M. bidupi sp. nov., 56 — M. laosana (Anisyutkin, 2010). 50, 51, 53, 54 — right phallomere from above (50, 53) and below (51, 54); 52 — sclerite R 1 H; 55 — sclerites R 1 H and R 1 G from dorso-caudal; 56 — outline of sclerite R 1 H processes. Dotted area shows membranous parts. Fig. 56 after Anisyutkin, 2010. Abbreviations: p. s.; R 1 G; R 1 H; R 2; R 3; t. s. — see text. Scale bar (2 mm = a – c, g; 1 mm = d – f): a = 50; b = 51; c = 52; d = 53; e = 54; f = 55; g = 56." httpUri="https://zenodo.org/record/217432/files/figure.png" pageId="10" pageNumber="530">Figs. 53–55</figureCitation>
), its dorsal part forming transverse plate-like subsclerite (
<figureCitation id="9CDB2A016728FFFFC345FDA8FB68D265" box="[1122,1257,548,572]" captionStart="FIGURES 50 – 56" captionStartId="9.[151,264,1862,1885]" captionTargetBox="[218,1362,203,1830]" captionTargetId="figure@9.[196,1392,193,1841]" captionTargetPageId="9" captionText="FIGURES 50 – 56. Macrostylopyga spp., male genitalia, holotypes (50 – 52 — M. grandis sp. nov., 53 – 55 — M. bidupi sp. nov., 56 — M. laosana (Anisyutkin, 2010). 50, 51, 53, 54 — right phallomere from above (50, 53) and below (51, 54); 52 — sclerite R 1 H; 55 — sclerites R 1 H and R 1 G from dorso-caudal; 56 — outline of sclerite R 1 H processes. Dotted area shows membranous parts. Fig. 56 after Anisyutkin, 2010. Abbreviations: p. s.; R 1 G; R 1 H; R 2; R 3; t. s. — see text. Scale bar (2 mm = a – c, g; 1 mm = d – f): a = 50; b = 51; c = 52; d = 53; e = 54; f = 55; g = 56." httpUri="https://zenodo.org/record/217432/files/figure.png" pageId="10" pageNumber="530">Figs. 53, 55</figureCitation>
, 
<emphasis id="3694EA966728FFFFC3D3FDA9FA9AD265" box="[1268,1307,549,572]" italics="true" pageId="10" pageNumber="530">p.s.</emphasis>
) with long, sclerotized tubular structure, evidently of glandular nature (
<figureCitation id="9CDB2A016728FFFFC46FFDCBFC1CD206" box="[840,925,583,607]" captionStart="FIGURES 50 – 56" captionStartId="9.[151,264,1862,1885]" captionTargetBox="[218,1362,203,1830]" captionTargetId="figure@9.[196,1392,193,1841]" captionTargetPageId="9" captionText="FIGURES 50 – 56. Macrostylopyga spp., male genitalia, holotypes (50 – 52 — M. grandis sp. nov., 53 – 55 — M. bidupi sp. nov., 56 — M. laosana (Anisyutkin, 2010). 50, 51, 53, 54 — right phallomere from above (50, 53) and below (51, 54); 52 — sclerite R 1 H; 55 — sclerites R 1 H and R 1 G from dorso-caudal; 56 — outline of sclerite R 1 H processes. Dotted area shows membranous parts. Fig. 56 after Anisyutkin, 2010. Abbreviations: p. s.; R 1 G; R 1 H; R 2; R 3; t. s. — see text. Scale bar (2 mm = a – c, g; 1 mm = d – f): a = 50; b = 51; c = 52; d = 53; e = 54; f = 55; g = 56." httpUri="https://zenodo.org/record/217432/files/figure.png" pageId="10" pageNumber="530">Fig. 53</figureCitation>
, 
<emphasis id="3694EA966728FFFFC48BFDC5FC4CD239" box="[940,973,585,608]" italics="true" pageId="10" pageNumber="530">t.s.</emphasis>
); R1H fused with swollen sclerite R1G (
<figureCitation id="9CDB2A016728FFFFC7B8FDE0FEA8D2DD" box="[159,297,620,644]" captionStart="FIGURES 50 – 56" captionStartId="9.[151,264,1862,1885]" captionTargetBox="[218,1362,203,1830]" captionTargetId="figure@9.[196,1392,193,1841]" captionTargetPageId="9" captionText="FIGURES 50 – 56. Macrostylopyga spp., male genitalia, holotypes (50 – 52 — M. grandis sp. nov., 53 – 55 — M. bidupi sp. nov., 56 — M. laosana (Anisyutkin, 2010). 50, 51, 53, 54 — right phallomere from above (50, 53) and below (51, 54); 52 — sclerite R 1 H; 55 — sclerites R 1 H and R 1 G from dorso-caudal; 56 — outline of sclerite R 1 H processes. Dotted area shows membranous parts. Fig. 56 after Anisyutkin, 2010. Abbreviations: p. s.; R 1 G; R 1 H; R 2; R 3; t. s. — see text. Scale bar (2 mm = a – c, g; 1 mm = d – f): a = 50; b = 51; c = 52; d = 53; e = 54; f = 55; g = 56." httpUri="https://zenodo.org/record/217432/files/figure.png" pageId="10" pageNumber="530">Figs. 53–55</figureCitation>
); plate-like curved sclerite R3 situated craniolaterally (
<figureCitation id="9CDB2A016728FFFFC4A8FDE0FC6CD2DD" box="[911,1005,620,644]" captionStart="FIGURES 39 – 49" captionStartId="8.[151,264,1852,1875]" captionTargetBox="[215,1394,216,1820]" captionTargetId="figure@8.[167,1419,193,1847]" captionTargetPageId="8" captionText="FIGURES 39 – 49. Macrostylopyga spp., male genitalia, holotypes (40, 42 – 49) and paratype (39, 41) (39 – 45 — M. grandis sp. nov., 46 – 49 — M. bidupi sp. nov.). 39, 40 — left phallomere from above with sclerite L 1 (39) and from caudal (40); 41 — sclerite L 1 and process sla of left phalomere from inside; 42, 46 — sclerite L 1 from outside (42) and inside (46); 43, 44, 47, 48 — ventral phallomere from side (43, 47) and below (44, 48); 45, 49 — sclerites R 3 from outside. Dotted area shows membranous parts. Abbreviations: L 1; L 2; L 3; L 4 C; L 4 D; m. i.; r. o.; sla; t. p. — see text. Scale bar (2 mm): a = 39; b = 40; c = 41; d = 42; e = 43, 44; f = 45; g = 46; h = 47, 48; i = 49." httpUri="https://zenodo.org/record/217431/files/figure.png" pageId="10" pageNumber="530">Figs. 49</figureCitation>
, 
<figureCitation id="9CDB2A016728FFFFC4DFFDE0FBC3D2DD" box="[1016,1090,620,644]" captionStart="FIGURES 50 – 56" captionStartId="9.[151,264,1862,1885]" captionTargetBox="[218,1362,203,1830]" captionTargetId="figure@9.[196,1392,193,1841]" captionTargetPageId="9" captionText="FIGURES 50 – 56. Macrostylopyga spp., male genitalia, holotypes (50 – 52 — M. grandis sp. nov., 53 – 55 — M. bidupi sp. nov., 56 — M. laosana (Anisyutkin, 2010). 50, 51, 53, 54 — right phallomere from above (50, 53) and below (51, 54); 52 — sclerite R 1 H; 55 — sclerites R 1 H and R 1 G from dorso-caudal; 56 — outline of sclerite R 1 H processes. Dotted area shows membranous parts. Fig. 56 after Anisyutkin, 2010. Abbreviations: p. s.; R 1 G; R 1 H; R 2; R 3; t. s. — see text. Scale bar (2 mm = a – c, g; 1 mm = d – f): a = 50; b = 51; c = 52; d = 53; e = 54; f = 55; g = 56." httpUri="https://zenodo.org/record/217432/files/figure.png" pageId="10" pageNumber="530">53, 54</figureCitation>
).
</paragraph>
<paragraph id="045F36846728FFFFC7E0FD03FE11D2FE" blockId="10.[151,1436,151,895]" box="[199,400,655,679]" pageId="10" pageNumber="530">Female unknown.</paragraph>
<paragraph id="045F36846728FFFFC7E0FD39FBADD2B6" blockId="10.[151,1436,151,895]" pageId="10" pageNumber="530">Measurements (mm). Head length 6.5, head width 5.2; pronotum length 8.5, pronotum width 9.6; mesonotum length 5.5, mesonotum width 10.5; metanotum length 5.3, metanotum width 10.8.</paragraph>
<paragraph id="045F36846728FFFFC7E0FD77FB4AD326" blockId="10.[151,1436,151,895]" pageId="10" pageNumber="530">
<emphasis id="3694EA966728FFFFC7E0FD77FEE3D34D" bold="true" box="[199,354,763,788]" pageId="10" pageNumber="530">Comparison.</emphasis>
The new species can readily be distinguished from other species of the genus (
<taxonomicName id="C3E04D076728FFFFC3D3FD71FAF3D34D" box="[1268,1394,764,788]" class="Insecta" family="Blattidae" genus="Macrostylopyga" kingdom="Animalia" order="Blattodea" pageId="10" pageNumber="530" phylum="Arthropoda" rank="species" species="grandis" status="sp. nov.">
<emphasis id="3694EA966728FFFFC3D3FD71FAF3D34D" box="[1268,1394,764,788]" italics="true" pageId="10" pageNumber="530">M. grandis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2DA757ED6728FFFFC25DFD70FF49D361" pageId="10" pageNumber="530" rank="species">
<emphasis id="3694EA966728FFFFC25DFD70FF49D361" bold="true" pageId="10" pageNumber="530">sp. nov.</emphasis>
</taxonomicNameLabel>
and 
<taxonomicName id="C3E04D076728FFFFC7DAFCADFEFDD361" box="[253,380,800,824]" class="Insecta" family="Blattidae" genus="Macrostylopyga" kingdom="Animalia" order="Blattodea" pageId="10" pageNumber="530" phylum="Arthropoda" rank="species" species="laosana">
<emphasis id="3694EA966728FFFFC7DAFCADFEFDD361" box="[253,380,800,824]" italics="true" pageId="10" pageNumber="530">M. laosana</emphasis>
</taxonomicName>
) by the structure of the paraprocts (the presence of hook-like caudomedial processes, 
<figureCitation id="9CDB2A016728FFFFC21CFC93FF34D305" captionStart="FIGURES 12 – 26" captionStartId="6.[151,264,1797,1820]" captionTargetBox="[253,1329,203,1766]" captionTargetId="figure@6.[239,1348,193,1776]" captionTargetPageId="6" captionText="FIGURES 12 – 26. Macrostylopyga spp., male, holotypes (12 – 17, 23, 24 — M. bidupi sp. nov., 18 – 22 — M. grandis sp. nov., 25, 26 — M. laosana (Anisyutkin, 2010). 12 — facial part of head; 13 — head, thorax and 1 – 7 th abdominal tergites from above; 14, 15 — abdominal apex from above (14) and below (15, hypandrium and genitalia removed); 16 — caudo-medial processes of paraprocts; 17 — hypandrium from below; 18, 22 — hind tarsus from outside (18) and below (22); 19 — arolium and claws; 20, 21, 23 – 26 — metatarsi: right fore (20, 23), right middle (21, 24), left hind (25, 26) from below (20, 21, 23, 24, 26) and outside (25). Dotted area shows membranous parts. Abbreviations: I – X — 1 st – 10 th abdominal tergites respectively, a. s.; ar.; e. r.; i. r.; pr.; par. — see text. Arrows show places of styli attachment. Scale bar (1 mm = e; 2 mm = h, i, k; 3 mm = d, j, l; 5 mm = a – c, f, g): a = 12; b = 13; c = 14; d = 15; e = 16; f = 17; g = 18; h = 19; i = 20, 21; j = 22; k = 23, 24; l = 25, 26." httpUri="https://zenodo.org/record/217429/files/figure.png" pageId="10" pageNumber="530">Figs. 15, 16</figureCitation>
), and that of the male genitalia: shape of the sclerites L4C, L4F, L2, L1 (compare 
<figureCitation id="9CDB2A016728FFFFC372FCC8FB5BD305" box="[1109,1242,836,860]" captionStart="FIGURES 27 – 38" captionStartId="7.[151,264,1891,1914]" captionTargetBox="[203,1384,204,1860]" captionTargetId="figure@7.[200,1387,193,1870]" captionTargetPageId="7" captionText="FIGURES 27 – 38. Macrostylopyga spp., male genitalia, holotypes (27 – 30 — M. grandis sp. nov., 31 – 34 — M. bidupi sp. nov., 35 – 38 — M. laosana (Anisyutkin, 2010). Left phallomera from above (27, 31, 35), outside (28, 32, 36), inside (29, 33, partly, 37) and below (30, 34, partly, 38). Dotted area shows membranous parts. Figs. 35 – 38 after Anisyutkin, 2010. Abbreviations: a. c. s.; a. p.; c. pr.; d. c. p.; e. c. s.; L 1; L 2; L 3; L 4 C; L 4 C, c. p.; L 4 D; L 4 F; m. l.; r. m. l.; r. o.; s. l.; sla; t. p. — see text. Scale bar 2 mm." httpUri="https://zenodo.org/record/217430/files/figure.png" pageId="10" pageNumber="530">Figs. 31–34</figureCitation>
and 
<figureCitation id="9CDB2A016728FFFFC237FCC8FF5FD326" captionStart="FIGURES 27 – 38" captionStartId="7.[151,264,1891,1914]" captionTargetBox="[203,1384,204,1860]" captionTargetId="figure@7.[200,1387,193,1870]" captionTargetPageId="7" captionText="FIGURES 27 – 38. Macrostylopyga spp., male genitalia, holotypes (27 – 30 — M. grandis sp. nov., 31 – 34 — M. bidupi sp. nov., 35 – 38 — M. laosana (Anisyutkin, 2010). Left phallomera from above (27, 31, 35), outside (28, 32, 36), inside (29, 33, partly, 37) and below (30, 34, partly, 38). Dotted area shows membranous parts. Figs. 35 – 38 after Anisyutkin, 2010. Abbreviations: a. c. s.; a. p.; c. pr.; d. c. p.; e. c. s.; L 1; L 2; L 3; L 4 C; L 4 C, c. p.; L 4 D; L 4 F; m. l.; r. m. l.; r. o.; s. l.; sla; t. p. — see text. Scale bar 2 mm." httpUri="https://zenodo.org/record/217430/files/figure.png" pageId="10" pageNumber="530">Figs. 27–30, 35–38</figureCitation>
) and, especially, in the shape of the process R1H (compare 
<figureCitation id="9CDB2A016728FFFFC45CFCEBFC7ED326" box="[891,1023,871,895]" captionStart="FIGURES 50 – 56" captionStartId="9.[151,264,1862,1885]" captionTargetBox="[218,1362,203,1830]" captionTargetId="figure@9.[196,1392,193,1841]" captionTargetPageId="9" captionText="FIGURES 50 – 56. Macrostylopyga spp., male genitalia, holotypes (50 – 52 — M. grandis sp. nov., 53 – 55 — M. bidupi sp. nov., 56 — M. laosana (Anisyutkin, 2010). 50, 51, 53, 54 — right phallomere from above (50, 53) and below (51, 54); 52 — sclerite R 1 H; 55 — sclerites R 1 H and R 1 G from dorso-caudal; 56 — outline of sclerite R 1 H processes. Dotted area shows membranous parts. Fig. 56 after Anisyutkin, 2010. Abbreviations: p. s.; R 1 G; R 1 H; R 2; R 3; t. s. — see text. Scale bar (2 mm = a – c, g; 1 mm = d – f): a = 50; b = 51; c = 52; d = 53; e = 54; f = 55; g = 56." httpUri="https://zenodo.org/record/217432/files/figure.png" pageId="10" pageNumber="530">Figs. 53–55</figureCitation>
and 
<figureCitation id="9CDB2A016728FFFFC311FCEBFB3FD326" box="[1078,1214,871,895]" captionStart="FIGURES 50 – 56" captionStartId="9.[151,264,1862,1885]" captionTargetBox="[218,1362,203,1830]" captionTargetId="figure@9.[196,1392,193,1841]" captionTargetPageId="9" captionText="FIGURES 50 – 56. Macrostylopyga spp., male genitalia, holotypes (50 – 52 — M. grandis sp. nov., 53 – 55 — M. bidupi sp. nov., 56 — M. laosana (Anisyutkin, 2010). 50, 51, 53, 54 — right phallomere from above (50, 53) and below (51, 54); 52 — sclerite R 1 H; 55 — sclerites R 1 H and R 1 G from dorso-caudal; 56 — outline of sclerite R 1 H processes. Dotted area shows membranous parts. Fig. 56 after Anisyutkin, 2010. Abbreviations: p. s.; R 1 G; R 1 H; R 2; R 3; t. s. — see text. Scale bar (2 mm = a – c, g; 1 mm = d – f): a = 50; b = 51; c = 52; d = 53; e = 54; f = 55; g = 56." httpUri="https://zenodo.org/record/217432/files/figure.png" pageId="10" pageNumber="530">Figs. 52, 56</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>