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<document id="3A8B4FB43F2C8929DA5BBE76309B40C9" ID-DOI="10.5281/zenodo.157530" ID-GBIF-Dataset="030814c6-571c-4244-8332-5f865a334c1a" ID-ISSN="1175­5326" ID-Zenodo-Dep="157530" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461022234774" checkinUser="plazi" docAuthor="Philips, T. Keith &amp; Foster, Douglas E." docDate="2004" docId="5A5C87D0FFC9FFC3BC390CABFDA4F883" docLanguage="en" docName="zt00577.pdf" docOrigin="Zootaxa 577" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Cryptopeniculus nigrosetus Philips, new species" docType="treatment" docVersion="9" lastPageNumber="10" masterDocId="A665FFA8FFCDFFCABD31095FFFF9FFEE" masterDocTitle="Cryptopeniculus nigrosetus n. g., n. sp. (Coleoptera: Ptinidae) from the Namaqualand region of South Africa" masterLastPageNumber="11" masterPageNumber="1" pageNumber="5" updateTime="1698570451452" updateUser="plazi">
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<mods:title id="048011A34CE17FC74EFD98E8C2E2B642">Cryptopeniculus nigrosetus n. g., n. sp. (Coleoptera: Ptinidae) from the Namaqualand region of South Africa</mods:title>
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<mods:namePart id="4E713EEA4D2FE1D6A7DC658377364E8E">Philips, T. Keith</mods:namePart>
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<mods:namePart id="BFEDFD9707A6F57A160CDBEED3BA3453">Foster, Douglas E.</mods:namePart>
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<mods:date id="2010923AC6DC1E773055FBDB5CA3AAF8">2004</mods:date>
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<treatment id="5A5C87D0FFC9FFC3BC390CABFDA4F883" ID-DOI="http://doi.org/10.5281/zenodo.6270794" ID-GBIF-Taxon="119605534" ID-Zenodo-Dep="6270794" LSID="urn:lsid:plazi:treatment:5A5C87D0FFC9FFC3BC390CABFDA4F883" httpUri="http://treatment.plazi.org/id/5A5C87D0FFC9FFC3BC390CABFDA4F883" lastPageId="9" lastPageNumber="10" pageId="4" pageNumber="5">
<subSubSection id="9AEF654DFFC9FFCEBC390CABFC11F9E0" box="[264,1000,1524,1550]" pageId="4" pageNumber="5" type="nomenclature">
<paragraph id="D24A36C6FFC9FFCEBC390CABFC11F9E0" blockId="4.[264,1000,1524,1550]" box="[264,1000,1524,1550]" pageId="4" pageNumber="5">
<heading id="890281AAFFC9FFCEBC390CABFC11F9E0" bold="true" box="[264,1000,1524,1550]" fontSize="11" level="1" pageId="4" pageNumber="5" reason="1">
<emphasis id="E081EAD4FFC9FFCEBC390CABFC11F9E0" bold="true" box="[264,1000,1524,1550]" pageId="4" pageNumber="5">
<taxonomicName id="15F54D45FFC9FFCEBC390CABFD4AF9E0" authority="Philips" authorityName="Philips" box="[264,691,1524,1550]" class="Insecta" family="Ptinidae" genus="Cryptopeniculus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="nigrosetus" status="sp. nov.">
<emphasis id="E081EAD4FFC9FFCEBC390CABFDAAF9E0" bold="true" box="[264,595,1524,1550]" italics="true" pageId="4" pageNumber="5">Cryptopeniculus nigrosetus</emphasis>
Philips
</taxonomicName>
,
<taxonomicNameLabel id="FBB257AFFFC9FFCEBFF10CABFCA9F9E0" box="[704,848,1524,1550]" pageId="4" pageNumber="5" rank="species">new species</taxonomicNameLabel>
(
<figureCitation id="4ACE2A43FFC9FFCEBE6E0CABFC26F9E0" box="[863,991,1524,1550]" captionStart-0="FIGURES 1 3" captionStart-1="FIGURES 9 11" captionStart-2="FIGURES 12 15" captionStart-3="FIGURE 16" captionStartId-0="2.[264,383,1699,1723]" captionStartId-1="7.[264,383,1820,1844]" captionStartId-2="8.[264,382,1484,1508]" captionStartId-3="9.[264,367,1547,1571]" captionTargetBox-0="[264,1323,880,1681]" captionTargetBox-1="[470,1108,642,1799]" captionTargetBox-2="[271,1317,553,1457]" captionTargetBox-3="[346,1261,315,1528]" captionTargetId-0="figure@2.[264,1323,880,1681]" captionTargetId-1="figure@7.[425,1139,616,1824]" captionTargetId-2="figure@8.[271,1328,549,1457]" captionTargetId-3="figure@9.[310,1262,315,1529]" captionTargetPageId-0="2" captionTargetPageId-1="7" captionTargetPageId-2="8" captionTargetPageId-3="9" captionText-0="FIGURES 1 3. Cryptopeniculus nigrosetus external views. 1, dorsal habitus; 2, lateral habitus; 3, dorsal view of the pronotum." captionText-1="FIGURES 9 11. Cryptopeniculus nigroseta genitalia. 9, male genitalia, ventral view; 10, male genitalia, lateral view; 11, female genitalia, dorsal view." captionText-2="FIGURES 12 15. Cryptopeniculus nigroseta mouthparts. 12, maxillary palp; 13, labium, ventral view; 14, mentum, ventral view; 15, mandible, dorsal view." captionText-3="FIGURE 16. Currently known distribution of Cryptopeniculus nigrosetus in western South Africa near the coast as indicated by the circles." httpUri-0="https://zenodo.org/record/169099/files/figure.png" httpUri-1="https://zenodo.org/record/157532/files/figure.png" httpUri-2="https://zenodo.org/record/157533/files/figure.png" httpUri-3="https://zenodo.org/record/169100/files/figure.png" pageId="4" pageNumber="5">Figs. 116</figureCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="9AEF654DFFC9FFCEBC390F60FD22F89F" pageId="4" pageNumber="5" type="diagnosis">
<paragraph id="D24A36C6FFC9FFCEBC390F60FD22F89F" blockId="4.[264,1321,1599,1905]" pageId="4" pageNumber="5">
DIAGNOSIS. This species can be recognized by the two elongate pronotal crests consisting of flocculant setae covered by combs of darker elongate but thick setae with their tips converging dorsally (
<figureCitation id="4ACE2A43FFC9FFCEBF330FD0FD95F947" box="[514,620,1679,1705]" captionStart="FIGURES 1 3" captionStartId="2.[264,383,1699,1723]" captionTargetBox="[264,1323,880,1681]" captionTargetId="figure@2.[264,1323,880,1681]" captionTargetPageId="2" captionText="FIGURES 1 3. Cryptopeniculus nigrosetus external views. 1, dorsal habitus; 2, lateral habitus; 3, dorsal view of the pronotum." httpUri="https://zenodo.org/record/169099/files/figure.png" pageId="4" pageNumber="5">Figs 1, 3</figureCitation>
). The crests are separated most widely anteriorly and converge posteriorly, becoming fused near the pronotal base. Additionally, the smooth, globular shape of the elytra (
<figureCitation id="4ACE2A43FFC9FFCEBF150F80FD91F917" box="[548,616,1759,1785]" captionStart="FIGURES 1 3" captionStartId="2.[264,383,1699,1723]" captionTargetBox="[264,1323,880,1681]" captionTargetId="figure@2.[264,1323,880,1681]" captionTargetPageId="2" captionText="FIGURES 1 3. Cryptopeniculus nigrosetus external views. 1, dorsal habitus; 2, lateral habitus; 3, dorsal view of the pronotum." httpUri="https://zenodo.org/record/169099/files/figure.png" pageId="4" pageNumber="5">Fig. 1</figureCitation>
), the constriction and short longitudinal ridges at the elytral bases (Fig. 6), the ratio of ventrite lengths (Fig. 8), and the pronounced antennal fossae (Fig. 4) extending laterally from the antennal insertions should help differentiate this species from the other known African taxa.
</paragraph>
</subSubSection>
<subSubSection id="9AEF654DFFC8FFCCBC090843FEADFC60" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6" type="description">
<paragraph id="D24A36C6FFC8FFCCBC090843FCA0FCD0" blockId="5.[264,1323,284,1873]" lastBlockId="6.[264,1323,284,1870]" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6">
DESCRIPTION.
<typeStatus id="0D4E8864FFC8FFCFBF340843FD8FFED8" box="[517,630,284,310]" pageId="5" pageNumber="6" type="holotype">Holotype</typeStatus>
. Sex not determined. Length from anterior edge of pronotum to apex of elytra
<quantity id="150D9B23FFC8FFCFBF20081BFD86FEB0" box="[529,639,324,350]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="5" pageNumber="6" unit="mm" value="2.1">2.10 mm</quantity>
. Color light to dark reddish black (
<figureCitation id="4ACE2A43FFC8FFCFB92F081BFB70FEB0" box="[1054,1161,324,350]" captionStart="FIGURES 1 3" captionStartId="2.[264,383,1699,1723]" captionTargetBox="[264,1323,880,1681]" captionTargetId="figure@2.[264,1323,880,1681]" captionTargetPageId="2" captionText="FIGURES 1 3. Cryptopeniculus nigrosetus external views. 1, dorsal habitus; 2, lateral habitus; 3, dorsal view of the pronotum." httpUri="https://zenodo.org/record/169099/files/figure.png" pageId="5" pageNumber="6">Figs 12</figureCitation>
).
<emphasis id="E081EAD4FFC8FFCFB99B081BFB13FEB3" box="[1194,1258,324,349]" italics="true" pageId="5" pageNumber="6">Head</emphasis>
(Fig. 4): surface covered with scattered usually appressed short stout setae, those on frons above antennal fossae more erect, vertex surface adjacent to pronotum rough, frons distinctly smoother; antennal fossae separated by a ridge about 2/3 the maximum scape width, laterally with deep groove; clypeus crescent shaped with a pronounced posterior margin forming a ridge; labrum slightly emarginate, anterior margin with row of dense setae; labial and maxillary palps fusiform (
<figureCitation id="4ACE2A43FFC8FFCFBF780B6BFD2EFDA0" box="[585,727,564,590]" captionStart="FIGURES 12 15" captionStartId="8.[264,382,1484,1508]" captionTargetBox="[271,1317,553,1457]" captionTargetId="figure@8.[271,1328,549,1457]" captionTargetPageId="8" captionText="FIGURES 12 15. Cryptopeniculus nigroseta mouthparts. 12, maxillary palp; 13, labium, ventral view; 14, mentum, ventral view; 15, mandible, dorsal view." httpUri="https://zenodo.org/record/157533/files/figure.png" pageId="5" pageNumber="6">Figs 1213</figureCitation>
), mentum equilateral triangular shape (
<figureCitation id="4ACE2A43FFC8FFCFB98C0B6BFAE3FDA0" box="[1213,1306,564,590]" captionStart="FIGURES 12 15" captionStartId="8.[264,382,1484,1508]" captionTargetBox="[271,1317,553,1457]" captionTargetId="figure@8.[271,1328,549,1457]" captionTargetPageId="8" captionText="FIGURES 12 15. Cryptopeniculus nigroseta mouthparts. 12, maxillary palp; 13, labium, ventral view; 14, mentum, ventral view; 15, mandible, dorsal view." httpUri="https://zenodo.org/record/157533/files/figure.png" pageId="5" pageNumber="6">Fig. 14</figureCitation>
), mandible inner surface with indistinct tooth near middle (
<figureCitation id="4ACE2A43FFC8FFCFBE9C0B03FBFDFD98" box="[941,1028,604,630]" captionStart="FIGURES 12 15" captionStartId="8.[264,382,1484,1508]" captionTargetBox="[271,1317,553,1457]" captionTargetId="figure@8.[271,1328,549,1457]" captionTargetPageId="8" captionText="FIGURES 12 15. Cryptopeniculus nigroseta mouthparts. 12, maxillary palp; 13, labium, ventral view; 14, mentum, ventral view; 15, mandible, dorsal view." httpUri="https://zenodo.org/record/157533/files/figure.png" pageId="5" pageNumber="6">Fig. 15</figureCitation>
); eyes small and slightly projecting especially posteriorly, maximum length about as long as fourth antennomere, approximately 12 ommatidia at maximum length, 9 ommatidia at maximum width; antennae after scape gradually expanding in diameter to apex, second through 10 th antennomeres subequal, ultimate about 50% longer than penultimate.
<emphasis id="E081EAD4FFC8FFCFBEDF0BA0FB9BFCF6" box="[1006,1122,767,792]" italics="true" pageId="5" pageNumber="6">Pronotum</emphasis>
(
<figureCitation id="4ACE2A43FFC8FFCFB9430BA0FB27FCF7" box="[1138,1246,767,793]" captionStart="FIGURES 1 3" captionStartId="2.[264,383,1699,1723]" captionTargetBox="[264,1323,880,1681]" captionTargetId="figure@2.[264,1323,880,1681]" captionTargetPageId="2" captionText="FIGURES 1 3. Cryptopeniculus nigrosetus external views. 1, dorsal habitus; 2, lateral habitus; 3, dorsal view of the pronotum." httpUri="https://zenodo.org/record/169099/files/figure.png" pageId="5" pageNumber="6">Figs 13</figureCitation>
): setal clumps forming two crests in very broad “V” shape arising at anterior 1/5 at edge of abrupt chitinous triangular shaped ridge on either side of middle, lateral edges curving gradually posteriorly converging through to posterior 1/3 with setal crests continuing to posterior margin; crests formed of tightly spaced, juxtaposed, overlapping, long, brown to nearly black narrow setae, setal origins visible as projections around perimeter of crests, angled towards middle of crest, apical portion of setae strongly curved, tips converging at crest peak; basally setae angled anterio­obliquely medially; beneath elongate setae a layer of paler colored flocculant setae in shape of broad “U”; anterior edge lined with erect setae similar to those forming external covering of crests, setae continuing onto lateral edge but not restricted, clump of setae laterally also forming loose tuft with setal tips converging dorsally to height below crest; latero­ventrally pronotum covered with white or light colored closely appressed setae obscuring surface, setae extending along lateral posterior edge; perimeter of pronotum with large deep punctures approximately in a row, others more scattered, setae arising from conical projections producing distinctly uneven surface, medially a distinct narrow longitudinal groove between crests with setal inserts on either side.
<emphasis id="E081EAD4FFC8FFCFBC630C20FE62FA76" box="[338,411,1407,1432]" italics="true" pageId="5" pageNumber="6">Elytra</emphasis>
(
<figureCitation id="4ACE2A43FFC8FFCFBC9F0C20FE03FA77" box="[430,506,1407,1433]" captionStart="FIGURES 1 3" captionStartId="2.[264,383,1699,1723]" captionTargetBox="[264,1323,880,1681]" captionTargetId="figure@2.[264,1323,880,1681]" captionTargetPageId="2" captionText="FIGURES 1 3. Cryptopeniculus nigrosetus external views. 1, dorsal habitus; 2, lateral habitus; 3, dorsal view of the pronotum." httpUri="https://zenodo.org/record/169099/files/figure.png" pageId="5" pageNumber="6">Figs 1</figureCitation>
, 6): Surface glabrous except for very small widely spaced puncture rows, more visible laterally; punctures sometimes with small, short, usually appressed setae, sometimes more numerous near elytral apices, occasionally at base between grooves with thicker, erect, white appressed setae; laterally with single row of small brown recumbent overlapping setae extending from near base to about midpoint, a second row below of similarly located and spaced, appressed white setae more obliquely aligned, a third row extending along epipleura to about midpoint of white, closely appressed, very dense, obliquely aligned setae; near apex along suture with oblique parallel fine wrinkles.
<emphasis id="E081EAD4FFC8FFCFB9E70FC8FEA7F936" italics="true" pageId="5" pageNumber="6">Ve nt r al surface</emphasis>
(Figs. 5, 78): Prothoracic process elongate, parallel sided, truncate at apex, extending posterior of procoxae into mesosternal groove, surface obscured with recumbent light or white colored setae, scattered recumbent setae; meso­ and metasternum with moderately dense appressed white setae, large, scattered, setose round punctures with posteriorly recumbent white setae; ventrites with some appressed white setae mainly around the perimeter although some extending medially along ventrite sutures, surface with scattered setose punctures smaller than those on metasternum and sometimes slightly less rounded, recumbent setae similar to those on metasternum; also with very fine punctures and fine regular reticular texture mainly along apices and fifth ventrite except at middle; ratio of ventrite lengths at middle starting anteriorly 16:18:14:13:26.
<emphasis id="E081EAD4FFCBFFCCBED708BBFBE7FE13" box="[998,1054,484,509]" italics="true" pageId="6" pageNumber="7">Legs</emphasis>
(
<figureCitation id="4ACE2A43FFCBFFCCB91F08BBFB63FE10" box="[1070,1178,484,510]" captionStart="FIGURES 1 3" captionStartId="2.[264,383,1699,1723]" captionTargetBox="[264,1323,880,1681]" captionTargetId="figure@2.[264,1323,880,1681]" captionTargetPageId="2" captionText="FIGURES 1 3. Cryptopeniculus nigrosetus external views. 1, dorsal habitus; 2, lateral habitus; 3, dorsal view of the pronotum." httpUri="https://zenodo.org/record/169099/files/figure.png" pageId="6" pageNumber="7">Figs 12</figureCitation>
): With scattered white recumbent setae, apices of femora with stouter brown setae, femora approximately parallel sided, expanding to club about middle on pro­ and mesolegs, at apical 3/5 on metalegs; first metatarsomere slightly longer than fifth, second through fourth shorter and equal in length, covered with scattered white and brown recumbent setae.
<emphasis id="E081EAD4FFCBFFCCB9850BDBFADAFD73" box="[1204,1315,644,669]" italics="true" pageId="6" pageNumber="7">Genitalia</emphasis>
: Male parameres (
<figureCitation id="4ACE2A43FFCBFFCCBCD10BF3FDA4FD28" box="[480,605,684,710]" captionStart="FIGURES 9 11" captionStartId="7.[264,383,1820,1844]" captionTargetBox="[470,1108,642,1799]" captionTargetId="figure@7.[425,1139,616,1824]" captionTargetPageId="7" captionText="FIGURES 9 11. Cryptopeniculus nigroseta genitalia. 9, male genitalia, ventral view; 10, male genitalia, lateral view; 11, female genitalia, dorsal view." httpUri="https://zenodo.org/record/157532/files/figure.png" pageId="6" pageNumber="7">Figs 910</figureCitation>
) gradually decreasing in width towards apex, tips curved inwards, blunt, apical half with scattered setae; median lobe gradually narrowed to apex. Female genitalic coxites elongate (
<figureCitation id="4ACE2A43FFCBFFCCBFAC0BA3FD0AFCF8" box="[669,755,764,790]" captionStart="FIGURES 9 11" captionStartId="7.[264,383,1820,1844]" captionTargetBox="[470,1108,642,1799]" captionTargetId="figure@7.[425,1139,616,1824]" captionTargetPageId="7" captionText="FIGURES 9 11. Cryptopeniculus nigroseta genitalia. 9, male genitalia, ventral view; 10, male genitalia, lateral view; 11, female genitalia, dorsal view." httpUri="https://zenodo.org/record/157532/files/figure.png" pageId="6" pageNumber="7">Fig. 11</figureCitation>
); stylus apex broad, slightly excavated; baculum elongate; spermatheca unsclerotized, not apparent.
</paragraph>
<paragraph id="D24A36C6FFCBFFCCBC090A13FEADFC60" blockId="6.[264,1323,284,1870]" pageId="6" pageNumber="7">
<emphasis id="E081EAD4FFCBFFCCBC090A13FDE5FC8B" box="[312,540,844,869]" italics="true" pageId="6" pageNumber="7">Sexual dimorphism</emphasis>
. No consistent external differences between the sexes have been found.
</paragraph>
</subSubSection>
<subSubSection id="9AEF654DFFCBFFCDBC090AC3FC5CFDA0" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7" type="materials_examined">
<paragraph id="D24A36C6FFCBFFCCBC090AC3FE0BFC58" blockId="6.[264,1323,284,1870]" box="[312,498,924,950]" pageId="6" pageNumber="7">
<typeStatus id="0D4E8864FFCBFFCCBC090AC3FE78FC58" box="[312,385,924,950]" pageId="6" pageNumber="7">
<collectionCode id="B4E4AE03FFCBFFCCBC090AC3FE78FC58" box="[312,385,924,950]" pageId="6" pageNumber="7">TYPE</collectionCode>
</typeStatus>
SERIES.
</paragraph>
<paragraph id="D24A36C6FFCBFFCCBC090A9BFAD2F8A0" blockId="6.[264,1323,284,1870]" pageId="6" pageNumber="7">
Data on separate labels on the pins is indicated by the “/” and is as follows:
<emphasis id="E081EAD4FFCBFFCCB9840A9BFAD3FC33" box="[1205,1322,964,989]" italics="true" pageId="6" pageNumber="7">
<typeStatus id="0D4E8864FFCBFFCCB9840A9BFADAFC33" box="[1205,1315,964,989]" pageId="6" pageNumber="7" type="holotype">Holotype</typeStatus>
:
</emphasis>
S. Afr. SW Cape, Seweputs coast,
<geoCoordinate id="B7C15001FFCBFFCCBF960AB3FCF8FBE8" box="[679,769,1004,1030]" direction="south" orientation="latitude" pageId="6" pageNumber="7" precision="555" value="-31.39">31.39 S</geoCoordinate>
<geoCoordinate id="B7C15001FFCBFFCCBE130AB3FC87FBE8" box="[802,894,1004,1030]" direction="east" orientation="longitude" pageId="6" pageNumber="7" precision="555" value="18.17">18.17 E</geoCoordinate>
/
<date id="A64B1006FFCBFFCCBEA10AB3FBF3FBE8" box="[912,1034,1004,1030]" pageId="6" pageNumber="7" value="1981-08-23">23.8.1981</date>
: E­Y:1836, groundtraps, 64 days, leg. Endrödy­Younga/ groundtrap with meat bait (
<collectionCode id="B4E4AE03FFCBFFCCBE870D4BFBE8FBC0" box="[950,1041,1044,1070]" pageId="6" pageNumber="7">TVMC</collectionCode>
).
<emphasis id="E081EAD4FFCBFFCCB91A0D4BFB52FBC3" box="[1067,1195,1044,1069]" italics="true" pageId="6" pageNumber="7">
<typeStatus id="0D4E8864FFCBFFCCB91A0D4BFB5CFBC3" box="[1067,1189,1044,1069]" pageId="6" pageNumber="7" type="paratype">Paratypes</typeStatus>
:
</emphasis>
Same data as
<typeStatus id="0D4E8864FFCBFFCCBC190D63FE6CFBB8" box="[296,405,1084,1110]" pageId="6" pageNumber="7" type="holotype">Holotype</typeStatus>
(2). Same data as
<typeStatus id="0D4E8864FFCBFFCCBF460D63FD1DFBB8" box="[631,740,1084,1110]" pageId="6" pageNumber="7" type="holotype">Holotype</typeStatus>
except banana bait (7); faeces bait (4); meat bait (3); ferm. banana bait (2). Same data as
<typeStatus id="0D4E8864FFCBFFCCBFD40D3BFCABFB90" box="[741,850,1124,1150]" pageId="6" pageNumber="7" type="holotype">Holotype</typeStatus>
except Seweputs Farm
<geoCoordinate id="B7C15001FFCBFFCCB9400D3BFB33FB90" box="[1137,1226,1124,1150]" direction="south" orientation="latitude" pageId="6" pageNumber="7" precision="555" value="-31.39">31.39 S</geoCoordinate>
<geoCoordinate id="B7C15001FFCBFFCCB9D90D3BFEE3FB48" direction="east" orientation="longitude" pageId="6" pageNumber="7" precision="555" value="18.22">18.22 E</geoCoordinate>
/E­Y:1835/ faeces bait (1); ferm. banana bait (5); meat bait (2); E­Y:1838/ ferm. banana bait (2). All the following are S. Afr., S. W. Cape: Soutpan,
<quantity id="150D9B23FFCBFFCCBEE40DEBFBD8FB20" box="[981,1057,1204,1230]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="6" pageNumber="7" unit="km" value="20.0">20 km</quantity>
E.
<geoCoordinate id="B7C15001FFCBFFCCB97A0DEBFB5FFB20" box="[1099,1190,1204,1230]" direction="south" orientation="latitude" pageId="6" pageNumber="7" precision="555" value="-31.12">31.12 S</geoCoordinate>
<geoCoordinate id="B7C15001FFCBFFCCB9F70DEBFADBFB20" box="[1222,1314,1204,1230]" direction="east" orientation="longitude" pageId="6" pageNumber="7" precision="555" value="18.06">18.06 E</geoCoordinate>
/
<date id="A64B1006FFCBFFCCBC390D83FE79FB18" box="[264,384,1244,1270]" pageId="6" pageNumber="7" value="1985-09-13">13.9.1985</date>
, E­Y:2240, ground and vegetation, leg. Endrödy­Younga (16); Nortier farm
<geoCoordinate id="B7C15001FFCBFFCCBC390C5BFE9DFAF0" box="[264,356,1284,1310]" direction="south" orientation="latitude" pageId="6" pageNumber="7" precision="555" value="-32.03">32.03 S</geoCoordinate>
<geoCoordinate id="B7C15001FFCBFFCCBCB90C5BFE1CFAF0" box="[392,485,1284,1310]" direction="east" orientation="longitude" pageId="6" pageNumber="7" precision="555" value="18.19">18.19 E</geoCoordinate>
/
<date id="A64B1006FFCBFFCCBCC90C5BFD8AFAF0" box="[504,627,1284,1310]" pageId="6" pageNumber="7" value="1981-08-25">25.8.1981</date>
,E­Y:1845, groundtraps, leg Endrödy­Younga/ groundtraps with faeces bait (2); meat bait (2); Titiesbaai,
<geoCoordinate id="B7C15001FFCBFFCCBE080C73FC6CFAA8" box="[825,917,1324,1350]" direction="south" orientation="latitude" pageId="6" pageNumber="7" precision="555" value="-31.1">31.10 S</geoCoordinate>
<geoCoordinate id="B7C15001FFCBFFCCBE880C73FBEEFAA8" box="[953,1047,1324,1350]" direction="east" orientation="longitude" pageId="6" pageNumber="7" precision="555" value="17.46">17.46 E</geoCoordinate>
/
<date id="A64B1006FFCBFFCCB91A0C73FB5AFAA8" box="[1067,1187,1324,1350]" pageId="6" pageNumber="7" value="1985-12-09">12.9.1985</date>
, E­Y:2239, veget. coastal dunes, leg. Endrödy­Younga (3); Papendorp dunes,
<geoCoordinate id="B7C15001FFCBFFCCB9080C0BFB61FA80" box="[1081,1176,1364,1390]" direction="south" orientation="latitude" pageId="6" pageNumber="7" precision="555" value="-31.38">31.38 S</geoCoordinate>
<geoCoordinate id="B7C15001FFCBFFCCB9F00C0BFAD8FA80" box="[1217,1313,1364,1390]" direction="east" orientation="longitude" pageId="6" pageNumber="7" precision="555" value="18.12">18.12 E</geoCoordinate>
/
<date id="A64B1006FFCBFFCCBC390C23FE79FA78" box="[264,384,1404,1430]" pageId="6" pageNumber="7" value="1981-08-22">22.8.1981</date>
, E­Y:1830, groundtraps, 64 days, leg. Endrödy­Younga/ groundtraps with meat bait (1), faeces bait (1), ferm. banana bait (1); Lamberts Bay, N,
<geoCoordinate id="B7C15001FFCBFFCCB90A0CFBFB60FA50" box="[1083,1177,1444,1470]" direction="south" orientation="latitude" pageId="6" pageNumber="7" precision="555" value="-32.04">32.04 S</geoCoordinate>
<geoCoordinate id="B7C15001FFCBFFCCB9F00CFBFAD8FA50" box="[1217,1313,1444,1470]" direction="east" orientation="longitude" pageId="6" pageNumber="7" precision="555" value="18.19">18.19 E</geoCoordinate>
/
<date id="A64B1006FFCBFFCCBC390C93FE79FA08" box="[264,384,1484,1510]" pageId="6" pageNumber="7" value="1981-08-25">25.8.1981</date>
, E­Y:1844, groundtraps, leg. Endrödy­Younga/ groundtraps with faeces bait (1); Koekenaap dunes,
<geoCoordinate id="B7C15001FFCBFFCCBCC40CABFDAFF9E0" box="[501,598,1524,1550]" direction="south" orientation="latitude" pageId="6" pageNumber="7" precision="555" value="-31.32">31.32 S</geoCoordinate>
<geoCoordinate id="B7C15001FFCBFFCCBFB30CABFD1DF9E0" box="[642,740,1524,1550]" direction="east" orientation="longitude" pageId="6" pageNumber="7" precision="555" value="18.14">18.14 E</geoCoordinate>
/
<date id="A64B1006FFCBFFCCBFCD0CABFC8DF9E0" box="[764,884,1524,1550]" pageId="6" pageNumber="7" value="1994-09-22">22.9.1994</date>
, E­Y:3030, ground and hummocks Endrödy &amp; Bellamy (2); Dermbergsdraaifarm,
<geoCoordinate id="B7C15001FFCBFFCCBE0C0F43FC60F9D8" box="[829,921,1564,1590]" direction="south" orientation="latitude" pageId="6" pageNumber="7" precision="555" value="-30.47">30.47 S</geoCoordinate>
<geoCoordinate id="B7C15001FFCBFFCCBE8D0F43FBE0F9D8" box="[956,1049,1564,1590]" direction="east" orientation="longitude" pageId="6" pageNumber="7" precision="555" value="17.43">17.43 E</geoCoordinate>
/
<date id="A64B1006FFCBFFCCB91D0F43FB5AF9D8" box="[1068,1187,1564,1590]" pageId="6" pageNumber="7" value="1994-09-18">18.9.1994</date>
, E­Y:3014, groundtraps, 7 days, Endrödy &amp; Bellamy/ groundtraps with meat bait (3), banana bait (3), faeces bait (1). All the following are S. Afr., Namaqualand: Klein Kogelfontein,
<geoCoordinate id="B7C15001FFCBFFCCB98B0F33FAEDF968" box="[1210,1300,1644,1670]" direction="south" orientation="latitude" pageId="6" pageNumber="7" precision="555" value="-31.1">31.10 S</geoCoordinate>
<geoCoordinate id="B7C15001FFCBFFCCBC390FCBFE9FF940" box="[264,358,1684,1710]" direction="east" orientation="longitude" pageId="6" pageNumber="7" precision="555" value="17.5">17.50 E</geoCoordinate>
/
<date id="A64B1006FFCBFFCCBC4A0FCBFE09F940" box="[379,496,1684,1710]" pageId="6" pageNumber="7" value="1979-08-27">27.8.1979</date>
/E­Y:1606, groundtraps, 62 days, leg. Endrödy­Younga/ groundtrap with banana bait (1); groundtrap with meat bait (1); Rooidam farm,
<geoCoordinate id="B7C15001FFCBFFCCB9750FE3FB59F938" box="[1092,1184,1724,1750]" direction="south" orientation="latitude" pageId="6" pageNumber="7" precision="555" value="-31.02">31.02 S</geoCoordinate>
<geoCoordinate id="B7C15001FFCBFFCCB9F50FE3FAD8F938" box="[1220,1313,1724,1750]" direction="east" orientation="longitude" pageId="6" pageNumber="7" precision="555" value="17.46">17.46 E</geoCoordinate>
/
<date id="A64B1006FFCBFFCCBC390FBBFE84F910" box="[264,381,1764,1790]" pageId="6" pageNumber="7" value="1994-09-20">20.9.1994</date>
/E­Y:3026, ground &amp; at light, leg. Endrödy­Younga (2); Katdoringvlei,
<geoCoordinate id="B7C15001FFCBFFCCB9E10FBBFAD3F910" box="[1232,1322,1764,1790]" direction="south" orientation="latitude" pageId="6" pageNumber="7" precision="555" value="-31.07">31.07 S</geoCoordinate>
<geoCoordinate id="B7C15001FFCBFFCCBC110E53FE85F8C8" box="[288,380,1804,1830]" direction="east" orientation="longitude" pageId="6" pageNumber="7" precision="555" value="17.52">17.52 E</geoCoordinate>
/
<date id="A64B1006FFCBFFCCBCBF0E53FDEAF8C8" box="[398,531,1804,1830]" pageId="6" pageNumber="7" value="1979-10-28">28.10.1979</date>
/E­Y:1664, singled on sand, day, leg. Endrödy­Younga (2); Island Point,
<quantity id="150D9B23FFCBFFCCBC6A0E6BFE67F8A0" box="[347,414,1844,1870]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="6" pageNumber="7" unit="km" value="4.0">4 km</quantity>
S,
<geoCoordinate id="B7C15001FFCBFFCCBCE00E6BFDCBF8A0" box="[465,562,1844,1870]" direction="south" orientation="latitude" pageId="6" pageNumber="7" precision="555" value="-30.56">30.56 S</geoCoordinate>
<geoCoordinate id="B7C15001FFCBFFCCBF6C0E6BFD47F8A0" box="[605,702,1844,1870]" direction="east" orientation="longitude" pageId="6" pageNumber="7" precision="555" value="17.38">17.38 E</geoCoordinate>
/
<date id="A64B1006FFCBFFCCBFE40E6BFCB2F8A0" box="[725,843,1844,1870]" pageId="6" pageNumber="7" value="1979-08-25">25.8.1979</date>
/E­Y:1595, groundtraps, 63 days, leg.
</paragraph>
<paragraph id="D24A36C6FFCAFFCDBC390843FC5CFDA0" blockId="7.[264,1323,284,590]" pageId="7" pageNumber="8">
Endrödy­Younga/ groundtraps with ferm. banana bait (1); Hoekbaai,
<quantity id="150D9B23FFCAFFCDB9000843FB94FED8" box="[1073,1133,284,310]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="7" pageNumber="8" unit="km" value="2.0">2 km</quantity>
ENE,
<geoCoordinate id="B7C15001FFCAFFCDB98D0843FAEDFED8" box="[1212,1300,284,310]" direction="south" orientation="latitude" pageId="7" pageNumber="8" precision="555" value="-31.11">31.11 S</geoCoordinate>
<geoCoordinate id="B7C15001FFCAFFCDBC39081BFE9FFEB0" box="[264,358,324,350]" direction="east" orientation="longitude" pageId="7" pageNumber="8" precision="555" value="17.47">17.47 E</geoCoordinate>
/
<date id="A64B1006FFCAFFCDBC4A081BFE09FEB0" box="[379,496,324,350]" pageId="7" pageNumber="8" value="1979-08-27">27.8.1979</date>
/E­Y:1610, groundtraps, 62 days, leg. Endrödy­Younga/ groundtrap with faeces bait.
<typeStatus id="0D4E8864FFCAFFCDBCE70833FDBAFE68" box="[470,579,364,390]" pageId="7" pageNumber="8" type="holotype">Holotype</typeStatus>
and
<typeStatus id="0D4E8864FFCAFFCDBF4A0833FD12FE68" box="[635,747,364,390]" pageId="7" pageNumber="8" type="paratype">paratypes</typeStatus>
are deposited in the Museum of Natural History, Northern Flagship Institution (formerly the Transvaal Museum). Additional
<typeStatus id="0D4E8864FFCAFFCDB9BC08CBFB04FE40" box="[1165,1277,404,430]" pageId="7" pageNumber="8" type="paratype">paratypes</typeStatus>
are deposited in the following collections: Canadian Museum of Nature, Gatineau, Quebec (
<collectionCode id="B4E4AE03FFCAFFCDBC2108BBFE95FE10" box="[272,364,484,510]" pageId="7" pageNumber="8">CMNC</collectionCode>
), Natural History Museum, London (
<collectionCode id="B4E4AE03FFCAFFCDBE1F08BBFC75FE10" box="[814,908,484,510]" httpUri="http://grbio.org/cool/4819-0jtd" name="Natural History Museum, Rangoon" pageId="7" pageNumber="8">NHMC</collectionCode>
),
<collectingCountry id="AAE27656FFCAFFCDBE9208BBFBBAFE10" box="[931,1091,484,510]" name="United States of America" pageId="7" pageNumber="8">United States</collectingCountry>
National Museum, Washington D.C. (
<collectionCode id="B4E4AE03FFCAFFCDBCD50B53FDC6FDC8" LSID="urn:lsid:biocol.org:col:34871" box="[484,575,524,550]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="7" pageNumber="8">USNM</collectionCode>
), Muséum national d'Histoire naturelle, Paris (
<collectionCode id="B4E4AE03FFCAFFCDB9570B53FB3FFDC8" LSID="urn:lsid:biocol.org:col:34988" box="[1126,1222,524,550]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="7" pageNumber="8">MNHN</collectionCode>
), Xavier Bellés collection and the T.K. Philips collection (
<collectionCode id="B4E4AE03FFCAFFCDBE760B6BFC60FDA0" box="[839,921,564,590]" pageId="7" pageNumber="8">TKPC</collectionCode>
).
</paragraph>
</subSubSection>
<caption id="868A664EFFCAFFCDBC390E43FCA4F8B8" httpUri="https://zenodo.org/record/157532/files/figure.png" pageId="7" pageNumber="8" targetBox="[470,1108,642,1799]" targetPageId="7">
<paragraph id="D24A36C6FFCAFFCDBC390E43FCA4F8B8" blockId="7.[264,1323,1820,1878]" pageId="7" pageNumber="8">
<emphasis id="E081EAD4FFCAFFCDBC390E43FE3DF8DA" bold="true" box="[264,452,1820,1844]" pageId="7" pageNumber="8">FIGURES 911.</emphasis>
<taxonomicName id="15F54D45FFCAFFCDBCFE0E42FD17F8DA" box="[463,750,1821,1844]" class="Insecta" family="Ptinidae" genus="Cryptopeniculus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="nigroseta">
<emphasis id="E081EAD4FFCAFFCDBCFE0E42FD17F8DA" box="[463,750,1821,1844]" italics="true" pageId="7" pageNumber="8">Cryptopeniculus nigroseta</emphasis>
</taxonomicName>
genitalia. 9, male genitalia, ventral view; 10, male genitalia, lateral view; 11, female genitalia, dorsal view.
</paragraph>
</caption>
<subSubSection id="9AEF654DFFC5FFC2BC090843FDE7FE38" pageId="8" pageNumber="9" type="description">
<paragraph id="D24A36C6FFC5FFC2BC090843FDE7FE38" blockId="8.[264,1322,284,470]" pageId="8" pageNumber="9">
VARIATION. Length from anterior edge of pronotum to apex of elytra averages 1.97
<quantity id="150D9B23FFC5FFC2BC7D081BFE4EFEB0" box="[332,439,324,350]" metricMagnitude="-4" metricUnit="m" metricValue="1.6" pageId="8" pageNumber="9" unit="mm" value="0.16">0.16 mm</quantity>
(n = 25). Specimens from the most northern site have more pronounced carinae along the lateral margins of the interantennal space. They are also slightly darker, have two widely spaced long erect setal rows on the dorsal surface of the elytra, and the legs are slightly stouter.
</paragraph>
</subSubSection>
<caption id="868A664EFFC5FFC2BC390C93FC7AF9E8" httpUri="https://zenodo.org/record/157533/files/figure.png" pageId="8" pageNumber="9" targetBox="[271,1317,553,1457]" targetPageId="8">
<paragraph id="D24A36C6FFC5FFC2BC390C93FC7AF9E8" blockId="8.[264,1322,1484,1542]" pageId="8" pageNumber="9">
<emphasis id="E081EAD4FFC5FFC2BC390C93FE29FA0A" bold="true" box="[264,464,1484,1508]" pageId="8" pageNumber="9">FIGURES 1215.</emphasis>
<taxonomicName id="15F54D45FFC5FFC2BCE80C92FD0FFA0A" box="[473,758,1485,1508]" class="Insecta" family="Ptinidae" genus="Cryptopeniculus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="nigroseta">
<emphasis id="E081EAD4FFC5FFC2BCE80C92FD0FFA0A" box="[473,758,1485,1508]" italics="true" pageId="8" pageNumber="9">Cryptopeniculus nigroseta</emphasis>
</taxonomicName>
mouthparts. 12, maxillary palp; 13, labium, ventral view; 14, mentum, ventral view; 15, mandible, dorsal view.
</paragraph>
</caption>
<subSubSection id="9AEF654DFFC5FFC2BC090FD5FD0FF922" pageId="8" pageNumber="9" type="etymology">
<paragraph id="D24A36C6FFC5FFC2BC090FD5FD0FF922" blockId="8.[264,1325,1674,1860]" pageId="8" pageNumber="9">ETYMOLOGY. Based on the Latin adjective for black and noun for hair in reference to the coarse black setae on the pronotum.</paragraph>
</subSubSection>
<subSubSection id="9AEF654DFFC5FFC2BC090F85FBF3F8AA" pageId="8" pageNumber="9" type="distribution">
<paragraph id="D24A36C6FFC5FFC2BC090F85FBF3F8AA" blockId="8.[264,1325,1674,1860]" pageId="8" pageNumber="9">
DISTRIBUTION. This monotypic genus ranges from about
<quantity id="150D9B23FFC5FFC2B9300F85FBB2F91A" box="[1025,1099,1754,1780]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="9" unit="km" value="10.0">10 km</quantity>
north of Lamberts Bay in the Western Cape Province northwards through to near Nariep (approximately
<quantity id="150D9B23FFC5FFC2B83E0E5DFED4F8AA" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="8" pageNumber="9" unit="km" value="30.0">30 km</quantity>
southwest of Garies) in the Northern Cape Province (
<figureCitation id="4ACE2A43FFC5FFC2BE940E75FC04F8AA" box="[933,1021,1834,1860]" captionStart="FIGURE 16" captionStartId="9.[264,367,1547,1571]" captionTargetBox="[346,1261,315,1528]" captionTargetId="figure@9.[310,1262,315,1529]" captionTargetPageId="9" captionText="FIGURE 16. Currently known distribution of Cryptopeniculus nigrosetus in western South Africa near the coast as indicated by the circles." httpUri="https://zenodo.org/record/169100/files/figure.png" pageId="8" pageNumber="9">Fig. 16</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="868A664EFFC4FFC3BC390F54FD45F9AA" httpUri="https://zenodo.org/record/169100/files/figure.png" pageId="9" pageNumber="10" targetBox="[346,1261,315,1528]" targetPageId="9">
<paragraph id="D24A36C6FFC4FFC3BC390F54FD45F9AA" blockId="9.[264,1322,1547,1604]" pageId="9" pageNumber="10">
<emphasis id="E081EAD4FFC4FFC3BC390F54FE6EF9CD" bold="true" box="[264,407,1547,1571]" pageId="9" pageNumber="10">FIGURE 16.</emphasis>
Currently known distribution of
<taxonomicName id="15F54D45FFC4FFC3BE310F54FBDCF9CC" box="[768,1061,1547,1570]" class="Insecta" family="Ptinidae" genus="Cryptopeniculus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="nigrosetus">
<emphasis id="E081EAD4FFC4FFC3BE310F54FBDCF9CC" box="[768,1061,1547,1570]" italics="true" pageId="9" pageNumber="10">Cryptopeniculus nigrosetus</emphasis>
</taxonomicName>
in western South Africa near the coast as indicated by the circles.
</paragraph>
</caption>
<subSubSection id="9AEF654DFFC4FFC3BC090FD4FDA4F883" pageId="9" pageNumber="10" type="biology_ecology">
<paragraph id="D24A36C6FFC4FFC3BC090FD4FDA4F883" blockId="9.[264,1322,1675,1901]" pageId="9" pageNumber="10">BIOLOGY. This species may be associated with coastal hummocks, vegetated dunes and other sandy habitats where it is undoubtedly scavenging on organic matter, most likely dung or other accumulated detritus. It has been captured with pitfalls usually set for 60 days and baited with faeces, fermenting bananas, and meat. Like most ptinines in these habitats, individuals are opportunistic and will feed on and attempt to breed in a wide variety of potential food sources.</paragraph>
</subSubSection>
</treatment>
</document>