136 lines
15 KiB
XML
136 lines
15 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.220.2178" ID-GBIF-Dataset="07cc435f-a623-4a17-b510-a38bf0569844" ID-PMC="PMC3459032" ID-Pensoft-Pub="1313-2970-220-1" ID-PubMed="23077429" ModsDocAuthor="" ModsDocDate="2012" ModsDocID="1313-2970-220-1" ModsDocOrigin="ZooKeys 220" ModsDocTitle="Revision of the Plant Bug Genus Tytthus (Hemiptera, Heteroptera, Miridae, Phylinae)" checkinTime="1451248758882" checkinUser="pensoft" docAuthor="Henry, Thomas J." docDate="2012" docId="57BB8AD7647BF696CC1CA71C9E18D12C" docLanguage="en" docName="ZooKeys 220: 1-114" docOrigin="ZooKeys 220" docSource="http://dx.doi.org/10.3897/zookeys.220.2178" docTitle="Tytthus uniformis Henry, sp. n." docType="treatment" docVersion="4" lastPageNumber="70" masterDocId="FFA2D657D204FFEEFF8DF82F5143FF91" masterDocTitle="Revision of the Plant Bug Genus Tytthus (Hemiptera, Heteroptera, Miridae, Phylinae)" masterLastPageNumber="114" masterPageNumber="1" pageNumber="69" updateTime="1668154436991" updateUser="ExternalLinkService">
|
||
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo>
|
||
<mods:title>Revision of the Plant Bug Genus Tytthus (Hemiptera, Heteroptera, Miridae, Phylinae)</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Henry, Thomas J.</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource>text</mods:typeOfResource>
|
||
<mods:relatedItem type="host">
|
||
<mods:titleInfo>
|
||
<mods:title>ZooKeys</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part>
|
||
<mods:date>2012</mods:date>
|
||
<mods:detail type="volume">
|
||
<mods:number>220</mods:number>
|
||
</mods:detail>
|
||
<mods:extent unit="page">
|
||
<mods:start>1</mods:start>
|
||
<mods:end>114</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:location>
|
||
<mods:url>http://dx.doi.org/10.3897/zookeys.220.2178</mods:url>
|
||
</mods:location>
|
||
<mods:classification>journal article</mods:classification>
|
||
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.220.2178</mods:identifier>
|
||
<mods:identifier type="Pensoft-Pub">1313-2970-220-1</mods:identifier>
|
||
</mods:mods>
|
||
<treatment ID-GBIF-Taxon="152036990" LSID="urn:lsid:zoobank.org:act:682B99D0-90E6-45BE-8565-BC96CDBAA1D9" httpUri="http://treatment.plazi.org/id/57BB8AD7647BF696CC1CA71C9E18D12C" lastPageId="69" lastPageNumber="70" pageId="68" pageNumber="69">
|
||
<subSubSection pageId="68" pageNumber="69" type="nomenclature">
|
||
<paragraph pageId="68" pageNumber="69">
|
||
<taxonomicName LSID="urn:lsid:zoobank.org:act:682B99D0-90E6-45BE-8565-BC96CDBAA1D9" authority="Henry" class="Insecta" family="Miridae" genus="Tytthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tytthus uniformis" order="Hemiptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="uniformis">Tytthus uniformis Henry</taxonomicName>
|
||
<taxonomicNameLabel pageId="68" pageNumber="69">sp. n.</taxonomicNameLabel>
|
||
Figs 45
|
||
<normalizedToken originalValue="–4787–100179–">-4787-100179-</normalizedToken>
|
||
182
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="68" pageNumber="69" type="diagnosis">
|
||
<paragraph pageId="68" pageNumber="69">Diagnosis.</paragraph>
|
||
<paragraph pageId="68" pageNumber="69">This species is distinguished by the contrasting black head and antennal segments I and II and the uniformly pale orange-brown pronotum and legs, and the slightly more smoky orange-brown hemelytra. All males are macropterous and all females examined, except two macropters, are brachpyterous.</paragraph>
|
||
<paragraph pageId="68" pageNumber="69">
|
||
<taxonomicName class="Insecta" family="Miridae" genus="Tytthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tytthus uniformis" order="Hemiptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="uniformis">Tytthus uniformis</taxonomicName>
|
||
is most similar in overall appearance to
|
||
<taxonomicName class="Insecta" family="Miridae" genus="Tytthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tytthus balli" order="Hemiptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="balli">Tytthus balli</taxonomicName>
|
||
and
|
||
<taxonomicName class="Insecta" family="Miridae" genus="Tytthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tytthus insperatus" order="Hemiptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="insperatus">Tytthus insperatus</taxonomicName>
|
||
. It can be distinguished from
|
||
<taxonomicName class="Insecta" family="Miridae" genus="Tytthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tytthus balli" order="Hemiptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="balli">Tytthus balli</taxonomicName>
|
||
by uniformly brownish-orange pronotum and hemelytra;
|
||
<taxonomicName class="Insecta" family="Miridae" genus="Tytthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tytthus balli" order="Hemiptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="balli">Tytthus balli</taxonomicName>
|
||
is usually infuscated on the posterior half of the pronotum, and the inner half of the clavus and apical third of the corium are dark brown. It is distinguished from
|
||
<taxonomicName class="Insecta" family="Miridae" genus="Tytthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tytthus insperatus" order="Hemiptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="insperatus">Tytthus insperatus</taxonomicName>
|
||
by the uniformly pale orange pronotum and legs;
|
||
<taxonomicName class="Insecta" family="Miridae" genus="Tytthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tytthus insperatus" order="Hemiptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="insperatus">Tytthus insperatus</taxonomicName>
|
||
has a dark brown pronotum, each femur has a narrow dorsal and lateral red stripe, and the hind tibiae are fuscous.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="69" lastPageNumber="70" pageId="68" pageNumber="69" type="description">
|
||
<paragraph pageId="68" pageNumber="69">Description.</paragraph>
|
||
<paragraph pageId="68" pageNumber="69">
|
||
<taxonomicName genus="Macropterous" lsidName="Macropterous male" pageId="68" pageNumber="69" rank="species" species="male">Macropterous male</taxonomicName>
|
||
(n= 10, plus holotype in parentheses) (Figs 45, 87, 88): Length to apex of hemelytron 2.45-2.60 mm (2.25 mm); length to base of cuneus 1.70-1.78 mm (1.68 mm); width across hemelytra 0.74-0.77 mm (0.74 mm). Head: Length 0.30-0.32 mm (0.30 mm), width across eyes 0.51-0.53 mm (0.51 mm); interocular width 0.29-0.30 mm (0.27 mm). Labium: Length 0.94-1.01 mm (0.90 mm). Antenna: Segment I, length (0.37 mm (0.34 mm); II, 1.17-1.20 mm (left 0.91 mm, right 1.06 mm); III, 0.67-0.69 mm (0.64 mm); IV, 0.42-0.43 mm (0.40 mm). Pronotum: Length 0.27 mm (0.24 mm), basal width 0.64-0.69 mm (0.67 mm).
|
||
</paragraph>
|
||
<paragraph pageId="68" pageNumber="69">Coloration: Head (Figs 89-91): Semishiny fuscous to black, pale spot on either side of vertex obsolete or indistinct. Labium: Antenna: Uniformly black, segment I very narrowly pale at apex on some specimens. Pronotum: Uniformly pale brownish orange, posterior angles sometimes slightly infuscated; collar pale or whitish; mesoscutum and scutellum uniformly pale brownish orange. Hemelytron: Uniformly pale, translucent, brownish orange; membrane translucent smoky brown. Ostiolar evaporative area (Fig. 92): Brownish orange to orange. Ventral surface: Thoracic segments orange to reddish orange; abdomen in males reddish orange dorsally, paler orange ventrally, with genital capsule becoming fuscous or black; abdomen in females infuscated dorsally, gradually fading to pale orange ventrally. Legs: Uniformly pale brownish orange; claw (Fig. 94).</paragraph>
|
||
<paragraph pageId="69" pageNumber="70">
|
||
<pageBreakToken pageId="69" pageNumber="70" start="start">Structure</pageBreakToken>
|
||
, texture, and vestiture: Head: Wider than long, interocular width subequal to length, impunctate, semishiny; set with a few, scattered, erect, nearly bristlelike dark setae. Labium: Extending beyond metacoxae to base of abbdomen. Pronotum: About 2.5 times as wide at base as long, lateral margins weakly concave, posterior margins moderately flared; set with short, semierect and recumbent setae. Mesoscutum: Broadly exposed. Scutellum: Slightly wider across base than on sides; set with short, semierect and recumbent setae. Hemelytron: Entire in males, cuneus nearly three times as long as wide at base, membrane fully developed, extending well beyond apex of abdomen; set with short, semierect and recumbent setae.
|
||
</paragraph>
|
||
<paragraph pageId="69" pageNumber="70">Male genitalia (Fig. 93): Left paramere (Fig. 179): Mitt-shaped; right arm long, slender, apically crenulate; left arm short, apically acute. Right paramere (Fig. 180): Oval. Endosoma (Fig. 181): Strongly C-shaped, apex bluntly pointed. Phallotheca (Fig. 182) slender, apically pointed.</paragraph>
|
||
<paragraph pageId="69" pageNumber="70">Macropterous female (n= 2) (Fig. 46): Length to apex of hemelytron 2.65-2.75 mm, length to base of cuneus 1.88-2.00 mm, width across hemelytra 0.91-1.08 mm. Head: Length 0.32-0.34 mm, width across eyes 0.54 mm; interocular width 0.32 mm. Labium: Length 0.96-0.98 mm, extending to bases of hind coxae. Antenna: Segment I, length 0.34-0.35 mm, II 0.99-1.07 mm; III 0.74-0.77 mm; IV 0.50-0.51 mm. Pronotum: Length 0.30 mm, basal width 0.75 mm.</paragraph>
|
||
<paragraph pageId="69" pageNumber="70">Brachypterous female (n= 8) (Figs 47, 95-100): Length to apex of abdomen 1.90-2.23, length to base of cuneus 1.63-1.75 mm, width across hemelytra 0.90-0.93 mm. Head: Length 0.34-0.35 mm, width across eyes 0.54-0.56 mm; interocular width 0.34 mm. Labium: Length 0.96-0.98 mm, extending to second or third visible abdominal segment. Antenna: Segment I, length 0.30-0.32 mm; II, 0.94-1.04 mm; III, 0.40-0.42 mm; IV, 0.40-0.45 mm. Pronotum: Length 0.26-0.27 mm, basal width 0.62-0.67 mm.</paragraph>
|
||
<paragraph pageId="69" pageNumber="70">The hemelytra are abbreviated in nearly all females examined. The cuneal fracture is visible but the cuneus is greatly shortened, with only the narrow basal area of membrane along abbreviated cuneus present.</paragraph>
|
||
<caption pageId="69" pageNumber="70">
|
||
<paragraph pageId="69" pageNumber="70">
|
||
Figures 179-186. Male genitalia 179-182
|
||
<taxonomicName class="Insecta" family="Miridae" genus="Tytthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tytthus uniformis" order="Hemiptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="uniformis">Tytthus uniformis</taxonomicName>
|
||
179 left paramere 180 right paramere 181 endosoma 182 phallotheca 183-186
|
||
<taxonomicName class="Insecta" family="Miridae" genus="Tytthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tytthus vagus" order="Hemiptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="vagus">Tytthus vagus</taxonomicName>
|
||
183 left paramere 184 right paramere 185 endosoma 186 phallotheca.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection pageId="69" pageNumber="70" type="etymology">
|
||
<paragraph pageId="69" pageNumber="70">Etymology.</paragraph>
|
||
<paragraph pageId="69" pageNumber="70">Named for the uniformly pale brownish-orange pronotum, hemelytra, and legs.</paragraph>
|
||
<paragraph pageId="69" pageNumber="70">
|
||
Host. Big sacaton,
|
||
<taxonomicName class="Liliopsida" family="Poaceae" genus="Sporobolus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Sporobolus wrightii" order="Poales" pageId="69" pageNumber="70" phylum="Tracheophyta" rank="species" species="wrightii">Sporobolus wrightii</taxonomicName>
|
||
Munro ex Scribn. [
|
||
<taxonomicName genus="Poaceae" lsidName="Poaceae" pageId="69" pageNumber="70" rank="genus">Poaceae</taxonomicName>
|
||
].
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="69" pageNumber="70" type="distribution">
|
||
<paragraph pageId="69" pageNumber="70">Distribution.</paragraph>
|
||
<paragraph pageId="69" pageNumber="70">United States: Arizona and New Mexico.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="69" pageNumber="70" type="type material">
|
||
<paragraph pageId="69" pageNumber="70">Type material.</paragraph>
|
||
<paragraph pageId="69" pageNumber="70">
|
||
Holotype ♂ (00162002) (USNM): UNITED STATES:New Mexico:Hidalgo Co.: Rt 338, 26.3 km S of Animas,
|
||
<geoCoordinate direction="north" orientation="latitude" precision="1" value="31.71636">31.71636°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate direction="west" orientation="longitude" precision="1" value="-108.82239">108.82239°W</geoCoordinate>
|
||
, 1457 m, 15 May 2004, A. G. Wheeler, Jr.,
|
||
<taxonomicName class="Liliopsida" family="Poaceae" genus="Sporobolus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Sporobolus wrightii" order="Poales" pageId="69" pageNumber="70" phylum="Tracheophyta" rank="species" species="wrightii">Sporobolus wrightii</taxonomicName>
|
||
(
|
||
<taxonomicName genus="Poaceae" lsidName="Poaceae" pageId="69" pageNumber="70" rank="genus">Poaceae</taxonomicName>
|
||
). Paratypes:UNITED STATES:Arizona:Cochise Co.: Huachuca Mountains, 5354 Ash Canyon Road, 0.5 mi W of Hwy 92,
|
||
<geoCoordinate direction="north" orientation="latitude" precision="1" value="31.38194">31.38194°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate direction="west" orientation="longitude" precision="1" value="-110.22444">110.22444°W</geoCoordinate>
|
||
, 1554 m, 17 Aug 1992, N. McFarland, 1 ♀ (00162047) (USNM). Santa Cruz Co.: Audubon Research Ranch, SE of Elgin, 12 - 14 May 2004, A. G. Wheeler, Jr.,
|
||
<taxonomicName class="Liliopsida" family="Poaceae" genus="Sporobolus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Sporobolus wrightii" order="Poales" pageId="69" pageNumber="70" phylum="Tracheophyta" rank="species" species="wrightii">Sporobolus wrightii</taxonomicName>
|
||
(
|
||
<taxonomicName genus="Poaceae" lsidName="Poaceae" pageId="69" pageNumber="70" rank="genus">Poaceae</taxonomicName>
|
||
), 9 ♂♂ (00162021, 00162033 - 00162039, 00162046), 17 ♀♀ (00162022 - 00162032, 00162040 - 00162045) (USNM). New Mexico:Hidalgo Co.: Rt 338, 26.3 km S of Animas,
|
||
<geoCoordinate direction="north" orientation="latitude" precision="1" value="31.71636">31.71636°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate direction="west" orientation="longitude" precision="1" value="-108.82239">108.82239°W</geoCoordinate>
|
||
, 1457 m, 15 May 2004, A. G. Wheeler, Jr.,
|
||
<taxonomicName class="Liliopsida" family="Poaceae" genus="Sporobolus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Sporobolus wrightii" order="Poales" pageId="69" pageNumber="70" phylum="Tracheophyta" rank="species" species="wrightii">Sporobolus wrightii</taxonomicName>
|
||
(
|
||
<taxonomicName genus="Poaceae" lsidName="Poaceae" pageId="69" pageNumber="70" rank="genus">Poaceae</taxonomicName>
|
||
), 4 ♂♂ (00162033 - 00162036) (USNM).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |