253 lines
34 KiB
XML
253 lines
34 KiB
XML
<document id="6683D19CD50653E49EB9B764E26BD4F6" ID-CLB-Dataset="27401" ID-DOI="10.11646/zootaxa.4577.1.13" ID-GBIF-Dataset="84173e4f-fbfc-43a8-bf19-1124b12af112" ID-ISSN="1175-5326" ID-Zenodo-Dep="2628566" ID-ZooBank="00C85EF8-68EA-4183-80F3-EE5DE9F872FF" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1554387052592" checkinUser="plazi" docAuthor="Kim, Junggon, Kim, Wongun, Park, Jihwan & Jung, Sunghoon" docDate="2019" docId="3449F157FFBA116117A0FB8FFBFDAD48" docLanguage="en" docName="zootaxa.4577.1.13.pdf" docOrigin="Zootaxa 4577 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pachycoleus japonicum, Comb. n." docType="treatment" docVersion="9" lastPageNumber="197" masterDocId="C870892FFFBB11631737FFFBFF9FAF53" masterDocTitle="First record of family Dipsocoridae (Hemiptera: Heteroptera: Dipsocoromorpha) from the Korean Peninsula and a new generic combination for Cryptostemma japonicum Miyamoto" masterLastPageNumber="200" masterPageNumber="195" pageNumber="196" updateTime="1698718421455" updateUser="ExternalLinkService">
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<mods:title id="5B5D561B7138AAA19A957BBE427D1D50">First record of family Dipsocoridae (Hemiptera: Heteroptera: Dipsocoromorpha) from the Korean Peninsula and a new generic combination for Cryptostemma japonicum Miyamoto</mods:title>
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<mods:namePart id="0D77D816AF00B1D51BAB203BF39E456A">Kim, Junggon</mods:namePart>
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<mods:namePart id="916BFCB71A36F72C96F17F1DF38BEBBB">Kim, Wongun</mods:namePart>
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<mods:namePart id="B630BCAAC8ED24B48A5107F681F0AE44">Park, Jihwan</mods:namePart>
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<mods:namePart id="D7DE2C8A99E60C8B02DEA6DDC49CCF2F">Jung, Sunghoon</mods:namePart>
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<mods:title id="DFA0AF7D4F89C14E71623F2C5822226A">Zootaxa</mods:title>
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<mods:date id="6324E9E66CCFC0A3F70F8190188E256F">2019</mods:date>
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<treatment id="3449F157FFBA116117A0FB8FFBFDAD48" ID-DOI="http://doi.org/10.5281/zenodo.5944285" ID-GBIF-Taxon="154553458" ID-Zenodo-Dep="5944285" LSID="urn:lsid:plazi:treatment:3449F157FFBA116117A0FB8FFBFDAD48" httpUri="http://treatment.plazi.org/id/3449F157FFBA116117A0FB8FFBFDAD48" lastPageId="2" lastPageNumber="197" pageId="1" pageNumber="196">
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<subSubSection id="F4FA13CAFFBA116217A0FB8FFC8AABDC" box="[151,789,1140,1167]" pageId="1" pageNumber="196" type="nomenclature">
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<paragraph id="BC5F4041FFBA116217A0FB8FFC8AABDC" blockId="1.[151,789,1140,1200]" box="[151,789,1140,1167]" pageId="1" pageNumber="196">
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<heading id="E717F72DFFBA116217A0FB8FFC8AABDC" bold="true" box="[151,789,1140,1167]" fontSize="11" level="1" pageId="1" pageNumber="196" reason="1">
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<emphasis id="8E949C53FFBA116217A0FB8FFC8AABDC" bold="true" box="[151,789,1140,1167]" pageId="1" pageNumber="196">
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<taxonomicName id="7BE03BC2FFBA116217A0FB8FFD04ABDC" ID-CoL="998367a0-2360-4486-aef4-ccf7e1dc6e7b" authority="(Miyamoto, 1964)" baseAuthorityName="Miyamoto" baseAuthorityYear="1964" box="[151,667,1140,1167]" class="Insecta" family="Dipsocoridae" genus="Pachycoleus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="196" phylum="Arthropoda" rank="species" species="japonicum" status="comb. nov.">
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<emphasis id="8E949C53FFBA116217A0FB8FFE29ABDD" bold="true" box="[151,438,1140,1166]" italics="true" pageId="1" pageNumber="196">Pachycoleus japonicum</emphasis>
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(
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<bibRefCitation id="D8713DB0FFBA116216F2FB8EFD0DABDC" author="Miyamoto, M." box="[453,658,1141,1167]" pageId="1" pageNumber="196" pagination="517 - 519" refId="ref2772" refString="Miyamoto, M. (1964) A new species of Cryptostemma from Japan (Hemiptera, Dipsocoridae). Kontyu, 32 (4), 517 - 519." type="journal article" year="1964">Miyamoto, 1964</bibRefCitation>
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)
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</taxonomicName>
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<taxonomicNameLabel id="95A72128FFBA11621595FB8EFC8AABDC" box="[674,789,1141,1167]" pageId="1" pageNumber="196" rank="species">Comb. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="F4FA13CAFFBA116217A0FB6CFEF6ABE3" box="[151,361,1175,1200]" pageId="1" pageNumber="196" type="description">
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<paragraph id="BC5F4041FFBA116217A0FB6CFEF6ABE3" blockId="1.[151,789,1140,1200]" box="[151,361,1175,1200]" pageId="1" pageNumber="196">
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(
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<figureCitation id="24DB5CC4FFBA116217A8FB6CFF73ABE3" box="[159,236,1175,1200]" captionStart="FIGURES 1" captionStartId="3.[151,264,1919,1942]" captionTargetBox="[164,1409,211,1884]" captionTargetId="figure@3.[151,1435,195,1895]" captionTargetPageId="3" captionText="FIGURES 1. Dorsal habitus and diagnostic characters of Pachycoleus japonicum (Miyamoto, 1964). A: dorsal habitus, ♂; B: dorsal habitus, ♀; C: hemelytron; D: abdomen with appendages and parameres of male in dorsal view; E: left appendage; F: right appendage; G: left paramere; H: right paramere; I: abdomen of female in dorsal view; J: spermatheca. Scale bar: A–C: 0.5 mm; E–I: 0.1 mm." httpUri="https://zenodo.org/record/2628568/files/figure.png" pageId="1" pageNumber="196">Figs. 1</figureCitation>
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,
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<figureCitation id="24DB5CC4FFBA116217CDFB63FE97ABE3" box="[250,264,1176,1200]" captionStart="FIGURES 2" captionStartId="4.[151,264,1180,1203]" captionTargetBox="[153,1432,198,1154]" captionTargetId="figure@4.[150,1436,193,1159]" captionTargetPageId="4" captionText="FIGURES 2. Live individuals, habitat and type specimen of Pachycoleus japonicum (Miyamoto, 1964). A: 5th instar nymph under a rock; B: adult under rocks; C: habitat, stones along water-side; D: holotype specimen with labels." httpUri="https://zenodo.org/record/2628570/files/figure.png" pageId="1" pageNumber="196">2</figureCitation>
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A–B, D)
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</paragraph>
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</subSubSection>
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<subSubSection id="F4FA13CAFFBA116217A0FB1AFD1AABA5" box="[151,645,1248,1270]" pageId="1" pageNumber="196" type="reference_group">
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<paragraph id="BC5F4041FFBA116217A0FB1AFD1AABA5" blockId="1.[151,645,1248,1270]" box="[151,645,1248,1270]" pageId="1" pageNumber="196">
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<treatmentCitationGroup id="9CF0676FFFBA116217A0FB1AFD1AABA5" box="[151,645,1248,1270]" pageId="1" pageNumber="196">
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<taxonomicName id="7BE03BC2FFBA116217A0FB1AFD1EABA5" ID-CoL="f5968d76-ff0f-4eb6-b114-6f1851602349" authority="Miyamoto, 1964: 517" authorityName="Miyamoto" authorityPageNumber="517" authorityYear="1964" box="[151,641,1248,1270]" class="Insecta" family="Dipsocoridae" genus="Cryptostemma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="196" phylum="Arthropoda" rank="species" species="japonicum">
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<emphasis id="8E949C53FFBA116217A0FB1AFE03ABA5" box="[151,412,1249,1270]" italics="true" pageId="1" pageNumber="196">Cryptostemma japonicum</emphasis>
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<treatmentCitation id="3D416650FFBA11621695FB1BFD1EABA5" author="Miyamoto, M." box="[418,641,1248,1270]" page="517" pageId="1" pageNumber="196" year="1964">
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<bibRefCitation id="D8713DB0FFBA11621695FB1BFDD1ABA5" author="Miyamoto, M." box="[418,590,1248,1270]" pageId="1" pageNumber="196" pagination="517 - 519" refId="ref2772" refString="Miyamoto, M. (1964) A new species of Cryptostemma from Japan (Hemiptera, Dipsocoridae). Kontyu, 32 (4), 517 - 519." type="journal article" year="1964">Miyamoto, 1964</bibRefCitation>
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: 517
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</treatmentCitation>
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</taxonomicName>
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.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="F4FA13CAFFBA116217A0FAD9FBF0AAF4" pageId="1" pageNumber="196" type="diagnosis">
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<paragraph id="BC5F4041FFBA116217A0FAD9FBF0AAF4" blockId="1.[151,1437,1314,2023]" pageId="1" pageNumber="196">
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<emphasis id="8E949C53FFBA116217A0FAD9FE8CAA68" bold="true" box="[151,275,1314,1339]" pageId="1" pageNumber="196">Diagnosis:</emphasis>
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Differs from other
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<taxonomicName id="7BE03BC2FFBA116216CAFAD8FD15AA68" authorityName="Fieber" authorityYear="1860" box="[509,650,1315,1339]" class="Insecta" family="Dipsocoridae" genus="Pachycoleus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="196" phylum="Arthropoda" rank="genus">
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<emphasis id="8E949C53FFBA116216CAFAD8FD15AA68" box="[509,650,1315,1339]" italics="true" pageId="1" pageNumber="196">Pachycoleus</emphasis>
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</taxonomicName>
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species by body relatively pale brown, dull (
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<figureCitation id="24DB5CC4FFBA11621396FAD9FB7BAA68" box="[1185,1252,1314,1339]" captionStart="FIGURES 1" captionStartId="3.[151,264,1919,1942]" captionTargetBox="[164,1409,211,1884]" captionTargetId="figure@3.[151,1435,195,1895]" captionTargetPageId="3" captionText="FIGURES 1. Dorsal habitus and diagnostic characters of Pachycoleus japonicum (Miyamoto, 1964). A: dorsal habitus, ♂; B: dorsal habitus, ♀; C: hemelytron; D: abdomen with appendages and parameres of male in dorsal view; E: left appendage; F: right appendage; G: left paramere; H: right paramere; I: abdomen of female in dorsal view; J: spermatheca. Scale bar: A–C: 0.5 mm; E–I: 0.1 mm." httpUri="https://zenodo.org/record/2628568/files/figure.png" pageId="1" pageNumber="196">Fig. 1</figureCitation>
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A–B); antennae generally pale brown; hemelytron macropterous, with small cell near fracture (
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<figureCitation id="24DB5CC4FFBA11621315FABDFBE3AA0C" box="[1058,1148,1350,1375]" captionStart="FIGURES 1" captionStartId="3.[151,264,1919,1942]" captionTargetBox="[164,1409,211,1884]" captionTargetId="figure@3.[151,1435,195,1895]" captionTargetPageId="3" captionText="FIGURES 1. Dorsal habitus and diagnostic characters of Pachycoleus japonicum (Miyamoto, 1964). A: dorsal habitus, ♂; B: dorsal habitus, ♀; C: hemelytron; D: abdomen with appendages and parameres of male in dorsal view; E: left appendage; F: right appendage; G: left paramere; H: right paramere; I: abdomen of female in dorsal view; J: spermatheca. Scale bar: A–C: 0.5 mm; E–I: 0.1 mm." httpUri="https://zenodo.org/record/2628568/files/figure.png" pageId="1" pageNumber="196">Fig. 1C</figureCitation>
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); left appendage of male abdominal segment VIII with subapically curved apex, with distinct process near apex (
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<figureCitation id="24DB5CC4FFBA1162134DFA91FB4FAAD0" box="[1146,1232,1386,1411]" captionStart="FIGURES 1" captionStartId="3.[151,264,1919,1942]" captionTargetBox="[164,1409,211,1884]" captionTargetId="figure@3.[151,1435,195,1895]" captionTargetPageId="3" captionText="FIGURES 1. Dorsal habitus and diagnostic characters of Pachycoleus japonicum (Miyamoto, 1964). A: dorsal habitus, ♂; B: dorsal habitus, ♀; C: hemelytron; D: abdomen with appendages and parameres of male in dorsal view; E: left appendage; F: right appendage; G: left paramere; H: right paramere; I: abdomen of female in dorsal view; J: spermatheca. Scale bar: A–C: 0.5 mm; E–I: 0.1 mm." httpUri="https://zenodo.org/record/2628568/files/figure.png" pageId="1" pageNumber="196">Fig. 1E</figureCitation>
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); right appendage thick and roundly curved apically (
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<figureCitation id="24DB5CC4FFBA11621528FA75FDECAAF4" box="[543,627,1422,1447]" captionStart="FIGURES 1" captionStartId="3.[151,264,1919,1942]" captionTargetBox="[164,1409,211,1884]" captionTargetId="figure@3.[151,1435,195,1895]" captionTargetPageId="3" captionText="FIGURES 1. Dorsal habitus and diagnostic characters of Pachycoleus japonicum (Miyamoto, 1964). A: dorsal habitus, ♂; B: dorsal habitus, ♀; C: hemelytron; D: abdomen with appendages and parameres of male in dorsal view; E: left appendage; F: right appendage; G: left paramere; H: right paramere; I: abdomen of female in dorsal view; J: spermatheca. Scale bar: A–C: 0.5 mm; E–I: 0.1 mm." httpUri="https://zenodo.org/record/2628568/files/figure.png" pageId="1" pageNumber="196">Fig. 1F</figureCitation>
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); spermathecal duct distinctly short (
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<figureCitation id="24DB5CC4FFBA11621326FA75FBFFAAF4" box="[1041,1120,1422,1447]" captionStart="FIGURES 1" captionStartId="3.[151,264,1919,1942]" captionTargetBox="[164,1409,211,1884]" captionTargetId="figure@3.[151,1435,195,1895]" captionTargetPageId="3" captionText="FIGURES 1. Dorsal habitus and diagnostic characters of Pachycoleus japonicum (Miyamoto, 1964). A: dorsal habitus, ♂; B: dorsal habitus, ♀; C: hemelytron; D: abdomen with appendages and parameres of male in dorsal view; E: left appendage; F: right appendage; G: left paramere; H: right paramere; I: abdomen of female in dorsal view; J: spermatheca. Scale bar: A–C: 0.5 mm; E–I: 0.1 mm." httpUri="https://zenodo.org/record/2628568/files/figure.png" pageId="1" pageNumber="196">Fig. 1J</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="F4FA13CAFFBA116117F0FA49FC47AFA8" lastPageId="2" lastPageNumber="197" pageId="1" pageNumber="196" type="description">
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<paragraph id="BC5F4041FFBA116217F0FA49FC2DAA98" blockId="1.[151,1437,1314,2023]" box="[199,946,1458,1483]" pageId="1" pageNumber="196">
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<emphasis id="8E949C53FFBA116217F0FA49FEC4AA98" bold="true" box="[199,347,1458,1483]" pageId="1" pageNumber="196">Description:</emphasis>
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<emphasis id="8E949C53FFBA11621655FA48FE3BAA98" box="[354,420,1459,1483]" italics="true" pageId="1" pageNumber="196">Male:</emphasis>
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See
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<bibRefCitation id="D8713DB0FFBA116216ECFA49FD3EAA98" author="Miyamoto, M." box="[475,673,1458,1483]" pageId="1" pageNumber="196" pagination="517 - 519" refId="ref2772" refString="Miyamoto, M. (1964) A new species of Cryptostemma from Japan (Hemiptera, Dipsocoridae). Kontyu, 32 (4), 517 - 519." type="journal article" year="1964">Miyamoto (1964)</bibRefCitation>
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for original description.
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</paragraph>
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<paragraph id="BC5F4041FFBA116217F0FA2DFB85A908" blockId="1.[151,1437,1314,2023]" pageId="1" pageNumber="196">
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<emphasis id="8E949C53FFBA116217F0FA2DFEA1AABC" bold="true" box="[199,318,1494,1519]" pageId="1" pageNumber="196">Genitalia:</emphasis>
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Genital segment with asymmetrical parameres (
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<figureCitation id="24DB5CC4FFBA116214B6FA2DFC7FAABC" box="[897,992,1494,1519]" captionStart="FIGURES 1" captionStartId="3.[151,264,1919,1942]" captionTargetBox="[164,1409,211,1884]" captionTargetId="figure@3.[151,1435,195,1895]" captionTargetPageId="3" captionText="FIGURES 1. Dorsal habitus and diagnostic characters of Pachycoleus japonicum (Miyamoto, 1964). A: dorsal habitus, ♂; B: dorsal habitus, ♀; C: hemelytron; D: abdomen with appendages and parameres of male in dorsal view; E: left appendage; F: right appendage; G: left paramere; H: right paramere; I: abdomen of female in dorsal view; J: spermatheca. Scale bar: A–C: 0.5 mm; E–I: 0.1 mm." httpUri="https://zenodo.org/record/2628568/files/figure.png" pageId="1" pageNumber="196">Fig. 1D</figureCitation>
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); left paramere cross-like shape with somewhat dense setae mostly, basal swelling part with one folded-like shaped lobe (
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<figureCitation id="24DB5CC4FFBA1162134CFA01FB46A940" box="[1147,1241,1530,1555]" captionStart="FIGURES 1" captionStartId="3.[151,264,1919,1942]" captionTargetBox="[164,1409,211,1884]" captionTargetId="figure@3.[151,1435,195,1895]" captionTargetPageId="3" captionText="FIGURES 1. Dorsal habitus and diagnostic characters of Pachycoleus japonicum (Miyamoto, 1964). A: dorsal habitus, ♂; B: dorsal habitus, ♀; C: hemelytron; D: abdomen with appendages and parameres of male in dorsal view; E: left appendage; F: right appendage; G: left paramere; H: right paramere; I: abdomen of female in dorsal view; J: spermatheca. Scale bar: A–C: 0.5 mm; E–I: 0.1 mm." httpUri="https://zenodo.org/record/2628568/files/figure.png" pageId="1" pageNumber="196">Fig. 1G</figureCitation>
|
||
; arrow) and one rounded lobe, apex of paramere long and shap, slightly curved in subapical part (
|
||
<figureCitation id="24DB5CC4FFBA1162130CF9E5FB09A964" box="[1083,1174,1566,1591]" captionStart="FIGURES 1" captionStartId="3.[151,264,1919,1942]" captionTargetBox="[164,1409,211,1884]" captionTargetId="figure@3.[151,1435,195,1895]" captionTargetPageId="3" captionText="FIGURES 1. Dorsal habitus and diagnostic characters of Pachycoleus japonicum (Miyamoto, 1964). A: dorsal habitus, ♂; B: dorsal habitus, ♀; C: hemelytron; D: abdomen with appendages and parameres of male in dorsal view; E: left appendage; F: right appendage; G: left paramere; H: right paramere; I: abdomen of female in dorsal view; J: spermatheca. Scale bar: A–C: 0.5 mm; E–I: 0.1 mm." httpUri="https://zenodo.org/record/2628568/files/figure.png" pageId="1" pageNumber="196">Fig. 1G</figureCitation>
|
||
); right paramere broad with dense setae mostly, both sides with small and blunt protuberance (
|
||
<figureCitation id="24DB5CC4FFBA11621484F9B9FB92A908" box="[947,1037,1602,1627]" captionStart="FIGURES 1" captionStartId="3.[151,264,1919,1942]" captionTargetBox="[164,1409,211,1884]" captionTargetId="figure@3.[151,1435,195,1895]" captionTargetPageId="3" captionText="FIGURES 1. Dorsal habitus and diagnostic characters of Pachycoleus japonicum (Miyamoto, 1964). A: dorsal habitus, ♂; B: dorsal habitus, ♀; C: hemelytron; D: abdomen with appendages and parameres of male in dorsal view; E: left appendage; F: right appendage; G: left paramere; H: right paramere; I: abdomen of female in dorsal view; J: spermatheca. Scale bar: A–C: 0.5 mm; E–I: 0.1 mm." httpUri="https://zenodo.org/record/2628568/files/figure.png" pageId="1" pageNumber="196">Fig. 1H</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="BC5F4041FFBA116217F0F99DFC30A994" blockId="1.[151,1437,1314,2023]" pageId="1" pageNumber="196">
|
||
<emphasis id="8E949C53FFBA116217F0F99DFEBBA92D" box="[199,292,1638,1662]" italics="true" pageId="1" pageNumber="196">Female:</emphasis>
|
||
as in male except for more expanded abdomen and specialized tergite IX with appendages; anal aperture almost circular; anterior gonapophysis slender and curved slightly, tapered apically; posterior gonapophysis curved medially, apical part distinctly widened (
|
||
<figureCitation id="24DB5CC4FFBA11621464F955FC00A994" box="[851,927,1710,1735]" captionStart="FIGURES 1" captionStartId="3.[151,264,1919,1942]" captionTargetBox="[164,1409,211,1884]" captionTargetId="figure@3.[151,1435,195,1895]" captionTargetPageId="3" captionText="FIGURES 1. Dorsal habitus and diagnostic characters of Pachycoleus japonicum (Miyamoto, 1964). A: dorsal habitus, ♂; B: dorsal habitus, ♀; C: hemelytron; D: abdomen with appendages and parameres of male in dorsal view; E: left appendage; F: right appendage; G: left paramere; H: right paramere; I: abdomen of female in dorsal view; J: spermatheca. Scale bar: A–C: 0.5 mm; E–I: 0.1 mm." httpUri="https://zenodo.org/record/2628568/files/figure.png" pageId="1" pageNumber="196">
|
||
Fig.
|
||
<date id="C85E6681FFBA116214BFF954FC00A994" box="[904,927,1711,1735]" pageId="1" pageNumber="196">1I</date>
|
||
</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="BC5F4041FFBA116217F0F929FDE2A85C" blockId="1.[151,1437,1314,2023]" pageId="1" pageNumber="196">
|
||
<emphasis id="8E949C53FFBA116217F0F929FEA1A9B8" bold="true" box="[199,318,1746,1771]" pageId="1" pageNumber="196">Genitalia:</emphasis>
|
||
spermatheca at left side underneath tergite VII dorsally (
|
||
<figureCitation id="24DB5CC4FFBA116214E4F929FBBEA9B8" box="[979,1057,1746,1771]" captionStart="FIGURES 1" captionStartId="3.[151,264,1919,1942]" captionTargetBox="[164,1409,211,1884]" captionTargetId="figure@3.[151,1435,195,1895]" captionTargetPageId="3" captionText="FIGURES 1. Dorsal habitus and diagnostic characters of Pachycoleus japonicum (Miyamoto, 1964). A: dorsal habitus, ♂; B: dorsal habitus, ♀; C: hemelytron; D: abdomen with appendages and parameres of male in dorsal view; E: left appendage; F: right appendage; G: left paramere; H: right paramere; I: abdomen of female in dorsal view; J: spermatheca. Scale bar: A–C: 0.5 mm; E–I: 0.1 mm." httpUri="https://zenodo.org/record/2628568/files/figure.png" pageId="1" pageNumber="196">
|
||
Fig.
|
||
<date id="C85E6681FFBA1162133EF928FBBEA9B8" box="[1033,1057,1747,1771]" pageId="1" pageNumber="196">1I</date>
|
||
</figureCitation>
|
||
), with somewhat long gland and distinctly short spermathecal duct (
|
||
<figureCitation id="24DB5CC4FFBA11621528F90DFDF1A85C" box="[543,622,1782,1807]" captionStart="FIGURES 1" captionStartId="3.[151,264,1919,1942]" captionTargetBox="[164,1409,211,1884]" captionTargetId="figure@3.[151,1435,195,1895]" captionTargetPageId="3" captionText="FIGURES 1. Dorsal habitus and diagnostic characters of Pachycoleus japonicum (Miyamoto, 1964). A: dorsal habitus, ♂; B: dorsal habitus, ♀; C: hemelytron; D: abdomen with appendages and parameres of male in dorsal view; E: left appendage; F: right appendage; G: left paramere; H: right paramere; I: abdomen of female in dorsal view; J: spermatheca. Scale bar: A–C: 0.5 mm; E–I: 0.1 mm." httpUri="https://zenodo.org/record/2628568/files/figure.png" pageId="1" pageNumber="196">Fig. 1J</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="BC5F4041FFBA116117F0F8E1FE90AF87" blockId="1.[151,1437,1314,2023]" lastBlockId="2.[151,1437,151,539]" lastPageId="2" lastPageNumber="197" pageId="1" pageNumber="196">
|
||
<emphasis id="8E949C53FFBA116217F0F8E1FEE9A860" bold="true" box="[199,374,1818,1843]" pageId="1" pageNumber="196">Measurements</emphasis>
|
||
(In mm): Male (n = 4)/Female (n = 4) body length: 1.97–1.99/2.09–2.13; head length 0.20– 0.22/0.20–0.22; head width including compound eyes: 0.38–0.40/0.39–0.41; distance between ocelli: 0.24–0.25/ 0.24–0.26; first antennal segment: 0.08–0.09/0.09–0.10; second antennal segment: 0.15–0.16/0.15–0.16; third antennal segment: 0.40–0.41/0.42–0.44; fourth antennal segment: 0.42–0.43; 0.45–0.46; anterior pronotal width: 0.37–0.38/0.32–0.34; mesal pronotal length: 0.24–0.25/0.25–0.26; basal pronotal maximal width: 0.63–0.65/0.62– 0.65; anterior scutellar width: 0.42–0.44/0.42–0.45; mesal scutellar length: 0.23–0.24/0.23–0.24; hemelytron length: 1.61–1.64/1.63–1.67; hindleg (femur: tibia: tarsus): 0.44–0.46: 0.55–0.58: 0.19–0.20/0.44–0.47: 0.57–0.60: 0.20–0.22.
|
||
</paragraph>
|
||
<paragraph id="BC5F4041FFB9116117F0FF19FC47AFA8" blockId="2.[151,1437,151,539]" box="[199,984,223,251]" pageId="2" pageNumber="197">
|
||
<emphasis id="8E949C53FFB9116117F0FF19FEF2AFA8" bold="true" box="[199,365,226,251]" pageId="2" pageNumber="197">Distributions:</emphasis>
|
||
<collectingCountry id="C4F700D1FFB911611643FF18FE27AFA9" box="[372,440,227,250]" name="South Korea" pageId="2" pageNumber="197">Korea</collectingCountry>
|
||
<superScript id="4B95ED09FFB91161168EFF24FE5FAFBE" attach="right" box="[441,448,223,237]" fontSize="6" pageId="2" pageNumber="197">*</superScript>
|
||
,
|
||
<collectingCountry id="C4F700D1FFB9116116F9FF19FD8EAFA8" box="[462,529,226,251]" name="China" pageId="2" pageNumber="197">China</collectingCountry>
|
||
(Central East),
|
||
<collectingCountry id="C4F700D1FFB9116115F4FF18FC99AFA9" box="[707,774,227,250]" name="Japan" pageId="2" pageNumber="197">Japan</collectingCountry>
|
||
,
|
||
<collectingCountry id="C4F700D1FFB911611427FF19FCC4AFA8" box="[784,859,226,251]" name="Russia" pageId="2" pageNumber="197">Russia</collectingCountry>
|
||
(Far East).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="F4FA13CAFFB9116117F0FEFDFEEFAE34" pageId="2" pageNumber="197" type="materials_examined">
|
||
<paragraph id="BC5F4041FFB9116117F0FEFDFEEFAE34" blockId="2.[151,1437,151,539]" pageId="2" pageNumber="197">
|
||
<emphasis id="8E949C53FFB9116117F0FEFDFD39AE4C" bold="true" box="[199,678,262,287]" pageId="2" pageNumber="197">
|
||
Materials examined: [
|
||
<materialsCitation id="0C884A1CFFB9116116FAFEFDFD39AE4C" ID-GBIF-Occurrence="2234727848" box="[461,678,262,287]" collectingDate="1955-12" collectingDateMax="2017-10-13" collectingDateMin="1955-12" collectionCode="ELKU, CNU" collectorName="S. & M. Miyamoto & Eupnae-ri & Gapyeong-eup & Gapyeong-gun & J. Kim & J. Park" country="Japan" location="Chikugo" municipality="Kyushu" pageId="2" pageNumber="197" specimenCount="22" specimenCount-female="14" specimenCount-male="8" stateProvince="Fukushima" typeStatus="holotype">
|
||
<collectionCode id="DAF1D884FFB9116116FAFEFDFDBEAE4C" box="[461,545,262,287]" pageId="2" pageNumber="197">ELKU</collectionCode>
|
||
]
|
||
<typeStatus id="635BFEE3FFB911611507FEFDFD3EAE4C" box="[560,673,262,287]" pageId="2" pageNumber="197" type="holotype">Holotype</typeStatus>
|
||
:
|
||
</materialsCitation>
|
||
</emphasis>
|
||
<specimenCount id="AAE68BC8FFB911611599FEFCFD4DAE4C" box="[686,722,263,287]" pageId="2" pageNumber="197" type="male">1♂</specimenCount>
|
||
, Pref.
|
||
<collectingRegion id="7E248EA3FFB91161142AFEFDFC03AE4C" box="[797,924,262,287]" country="Japan" name="Fukushima" pageId="2" pageNumber="197">Fukushima</collectingRegion>
|
||
,
|
||
<location id="B93F169AFFB91161149FFEFDFB94AE4C" LSID="urn:lsid:plazi:treatment:3449F157FFBA116117A0FB8FFBFDAD48:B93F169AFFB91161149FFEFDFB94AE4C" box="[936,1035,262,287]" country="Japan" municipality="Kyushu" name="Chikugo" pageId="2" pageNumber="197" stateProvince="Fukushima">Chikugo</location>
|
||
,
|
||
<collectingMunicipality id="5C3BDA3BFFB91161132FFEFDFBEDAE4C" box="[1048,1138,262,287]" pageId="2" pageNumber="197">Kyushu</collectingMunicipality>
|
||
,
|
||
<collectingCountry id="C4F700D1FFB911611349FEF3FB5EAE4C" box="[1150,1217,264,287]" name="Japan" pageId="2" pageNumber="197">Japan</collectingCountry>
|
||
,
|
||
<collectingDate id="D81A9F69FFB9116113FBFEFDFAABAE4C" box="[1228,1332,262,287]" pageId="2" pageNumber="197" value="1955-12">
|
||
<date id="C85E6681FFB9116113FBFEFDFAABAE4C" box="[1228,1332,262,287]" pageId="2" pageNumber="197" value="1955-12">xii. 1955</date>
|
||
</collectingDate>
|
||
,
|
||
<collectorName id="11152597FFB911611277FEFCFE93AE10" pageId="2" pageNumber="197">S. & M. Miyamoto</collectorName>
|
||
leg.;
|
||
<emphasis id="8E949C53FFB91161167BFED1FE04AE10" bold="true" box="[332,411,298,323]" pageId="2" pageNumber="197">[CNU]</emphasis>
|
||
<specimenCount id="AAE68BC8FFB911611694FED0FE7DAE10" box="[419,482,299,323]" pageId="2" pageNumber="197" type="male">7♂♂</specimenCount>
|
||
<specimenCount id="AAE68BC8FFB9116116D5FED0FDBCAE10" box="[482,547,299,323]" pageId="2" pageNumber="197" type="female">14♀♀</specimenCount>
|
||
,
|
||
<collectorName id="11152597FFB911611507FED1FD3EAE10" box="[560,673,298,323]" pageId="2" pageNumber="197">Eupnae-ri</collectorName>
|
||
,
|
||
<collectorName id="11152597FFB91161159AFED0FCCAAE10" box="[685,853,299,323]" pageId="2" pageNumber="197">Gapyeong-eup</collectorName>
|
||
,
|
||
<collectorName id="11152597FFB911611457FED0FB96AE10" box="[864,1033,299,323]" pageId="2" pageNumber="197">Gapyeong-gun</collectorName>
|
||
, Gyeonggi-do, Korea,
|
||
<collectingDate id="D81A9F69FFB91161123CFED0FAE1AE10" box="[1291,1406,299,323]" pageId="2" pageNumber="197" value="2017-10-13">
|
||
<date id="C85E6681FFB91161123CFED0FAE1AE10" box="[1291,1406,299,323]" pageId="2" pageNumber="197" value="2017-10-13">13.x.2017</date>
|
||
</collectingDate>
|
||
,
|
||
<collectorName id="11152597FFB9116112BDFED0FF56AE34" pageId="2" pageNumber="197">J. Kim</collectorName>
|
||
&
|
||
<collectorName id="11152597FFB9116117DBFEABFEA4AE34" box="[236,315,334,359]" pageId="2" pageNumber="197">J. Park</collectorName>
|
||
, leg.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="F4FA13CAFFB9116117F0FE89FC8DAEFC" pageId="2" pageNumber="197" type="biology_ecology">
|
||
<paragraph id="BC5F4041FFB9116117F0FE89FC8DAEFC" blockId="2.[151,1437,151,539]" pageId="2" pageNumber="197">
|
||
<emphasis id="8E949C53FFB9116117F0FE89FEB5AED8" bold="true" box="[199,298,370,395]" pageId="2" pageNumber="197">Biology:</emphasis>
|
||
Live nymphs (
|
||
<figureCitation id="24DB5CC4FFB9116116E1FE89FDB0AED8" box="[470,559,370,395]" captionStart="FIGURES 2" captionStartId="4.[151,264,1180,1203]" captionTargetBox="[153,1432,198,1154]" captionTargetId="figure@4.[150,1436,193,1159]" captionTargetPageId="4" captionText="FIGURES 2. Live individuals, habitat and type specimen of Pachycoleus japonicum (Miyamoto, 1964). A: 5th instar nymph under a rock; B: adult under rocks; C: habitat, stones along water-side; D: holotype specimen with labels." httpUri="https://zenodo.org/record/2628570/files/figure.png" pageId="2" pageNumber="197">Fig. 2A</figureCitation>
|
||
) and adults (
|
||
<figureCitation id="24DB5CC4FFB9116115F6FE89FC87AED8" box="[705,792,370,395]" captionStart="FIGURES 2" captionStartId="4.[151,264,1180,1203]" captionTargetBox="[153,1432,198,1154]" captionTargetId="figure@4.[150,1436,193,1159]" captionTargetPageId="4" captionText="FIGURES 2. Live individuals, habitat and type specimen of Pachycoleus japonicum (Miyamoto, 1964). A: 5th instar nymph under a rock; B: adult under rocks; C: habitat, stones along water-side; D: holotype specimen with labels." httpUri="https://zenodo.org/record/2628570/files/figure.png" pageId="2" pageNumber="197">Fig. 2B</figureCitation>
|
||
) of this species are found under stones along rivers (
|
||
<figureCitation id="24DB5CC4FFB911611259FE89FF24AEFC" captionStart="FIGURES 2" captionStartId="4.[151,264,1180,1203]" captionTargetBox="[153,1432,198,1154]" captionTargetId="figure@4.[150,1436,193,1159]" captionTargetPageId="4" captionText="FIGURES 2. Live individuals, habitat and type specimen of Pachycoleus japonicum (Miyamoto, 1964). A: 5th instar nymph under a rock; B: adult under rocks; C: habitat, stones along water-side; D: holotype specimen with labels." httpUri="https://zenodo.org/record/2628570/files/figure.png" pageId="2" pageNumber="197">Fig. 2C</figureCitation>
|
||
), usually on the moist part of the relatively flat stone.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="F4FA13CAFFB9116117F0FE41FBFDAD48" pageId="2" pageNumber="197" type="discussion">
|
||
<paragraph id="BC5F4041FFB9116117F0FE41FBFDAD48" blockId="2.[151,1437,151,539]" pageId="2" pageNumber="197">
|
||
<emphasis id="8E949C53FFB9116117F0FE41FEA2AE80" bold="true" box="[199,317,442,467]" pageId="2" pageNumber="197">Remarks:</emphasis>
|
||
Based on male (absence of abdominal appendages anterior to segment VIII), we confirm generic position of this species as
|
||
<taxonomicName id="7BE03BC2FFB9116116DBFE24FDE6AEA4" authorityName="Fieber" authorityYear="1860" box="[492,633,479,503]" class="Insecta" family="Dipsocoridae" genus="Pachycoleus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="197" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="8E949C53FFB9116116DBFE24FDE6AEA4" box="[492,633,479,503]" italics="true" pageId="2" pageNumber="197">Pachycoleus</emphasis>
|
||
</taxonomicName>
|
||
, even though this species is strictly associated with stream like
|
||
<taxonomicName id="7BE03BC2FFB9116117A0FDF8FEA4AD49" authority="Kim & Kim & Park & Jung, 2019" authorityName="Kim & Kim & Park & Jung" authorityYear="2019" box="[151,315,515,538]" class="Insecta" family="Dipsocoridae" genus="Cryptostemma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="197" phylum="Arthropoda" rank="genus" sensu="stricto">
|
||
<emphasis id="8E949C53FFB9116117A0FDF8FEA4AD49" box="[151,315,515,538]" italics="true" pageId="2" pageNumber="197">Cryptostemma</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="95A72128FFB911611676FDF8FEE3AD49" box="[321,380,515,538]" pageId="2" pageNumber="197" sensu="stricto">s. str.</taxonomicNameLabel>
|
||
as indicated by
|
||
<bibRefCitation id="D8713DB0FFB911611506FDF9FD29AD48" author="Stys, P." box="[561,694,514,539]" pageId="2" pageNumber="197" pagination="3 - 15" refId="ref2859" refString="Stys, P. (1990) Enicocephalomorphan and dipsocoromorphan fauna of W. Palaearctic (Heteroptera): composition, distribution and biology. Scopolia, 1 (Supplement), 3 - 15." type="journal article" year="1990">Štys (1990)</bibRefCitation>
|
||
. See discussion part below for details.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |