328 lines
40 KiB
XML
328 lines
40 KiB
XML
<document id="4013F3375A5DFD359BA61004166CD42A" ID-DOI="10.5281/zenodo.174408" ID-GBIF-Dataset="fbbcb2e5-d5ad-4c8b-af0e-22a30f852c15" ID-ISSN="11755326" ID-Zenodo-Dep="174408" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459796571981" checkinUser="plazi" docAuthor="Schwabe, Enrico, Försterra, Günter, Häussermann, Verena, Melzer, Roland R. & Schrödl, Michael" docDate="2006" docId="155A87A4FF81FFC06051FE47FBD5AD80" docLanguage="en" docName="zt01341p027.pdf" docOrigin="Zootaxa 1341" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Tonicia calbucensis Plate 1897" docType="treatment" docVersion="9" lastPageNumber="15" masterDocId="E963FFDCFF8CFFCE6159FFA3FFDCA866" masterDocTitle="Chitons (Mollusca: Polyplacophora) from the southern Chilean Comau Fjord, with reinstatement of Tonicia calbucensis Plate, 1897" masterLastPageNumber="27" masterPageNumber="1" pageNumber="14" updateTime="1698219868725" updateUser="plazi">
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<mods:title id="A71309D9DCC2A67FA47ABC1FF554EBBC">Chitons (Mollusca: Polyplacophora) from the southern Chilean Comau Fjord, with reinstatement of Tonicia calbucensis Plate, 1897</mods:title>
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<mods:namePart id="425F8C53B2B089775CBFA7A990B9AF8F">Försterra, Günter</mods:namePart>
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<mods:namePart id="EA1B749C4F37E869D990FCFE8007646D">Häussermann, Verena</mods:namePart>
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<mods:namePart id="8CDC46048AE44EECF72C8F68D3CFD384">Melzer, Roland R.</mods:namePart>
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<mods:namePart id="3672A7F0047B9A0C5F593167DFCE53C9">Schrödl, Michael</mods:namePart>
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<mods:date id="74EFDEC7B69A96807DCA16DD7742ED89">2006</mods:date>
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<treatment id="155A87A4FF81FFC06051FE47FBD5AD80" ID-DOI="http://doi.org/10.5281/zenodo.5674880" ID-GBIF-Taxon="119351324" ID-Zenodo-Dep="5674880" LSID="urn:lsid:plazi:treatment:155A87A4FF81FFC06051FE47FBD5AD80" httpUri="http://treatment.plazi.org/id/155A87A4FF81FFC06051FE47FBD5AD80" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
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<subSubSection id="D5E96539FF81FFC36051FE47FDC7AA47" pageId="13" pageNumber="14" type="nomenclature">
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<paragraph id="9D4C36B2FF81FFC36051FE47FD5AA99B" blockId="13.[264,646,483,545]" box="[264,646,483,510]" pageId="13" pageNumber="14">
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<heading id="C60481DEFF81FFC36051FE47FD5AA99B" bold="true" box="[264,646,483,510]" fontSize="11" level="1" pageId="13" pageNumber="14" reason="1">
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<taxonomicName id="5AF34D31FF81FFC36051FE47FD5AA99B" ID-CoL="b0cde5f4-dc64-4b7c-a915-13082b27e261" authority="Plate, 1897" authorityName="Plate" authorityYear="1897" box="[264,646,483,510]" class="Polyplacophora" family="Chitonidae" genus="Tonicia" kingdom="Animalia" order="Chitonida" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="calbucensis">
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<emphasis id="AF87EAA0FF81FFC36051FE47FD5AA99B" bold="true" box="[264,646,483,510]" pageId="13" pageNumber="14">
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<emphasis id="AF87EAA0FF81FFC36051FE47FE29A998" bold="true" box="[264,501,484,510]" italics="true" pageId="13" pageNumber="14">Tonicia calbucensis</emphasis>
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<bibRefCitation id="F9624B43FF81FFC360A4FE40FD5AA99B" author="Plate" box="[509,646,483,509]" pageId="13" pageNumber="26" refString="Plate, L. H. (1897) Die Anatomie und Phylogenie der Chitonen. Fauna Chilensis 1 (1). Zoologische Jahrbucher, Abteilung fur Systematik, Okologie und Geographie der Tiere, 1, Suppl. 4, 1 - 243, pls 1 - 12." type="journal article" year="1897">Plate, 1897</bibRefCitation>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="9D4C36B2FF81FFC36051FDA4FDC7AA47" blockId="13.[264,646,483,545]" box="[264,539,519,545]" pageId="13" pageNumber="14">(Figures 10, 16, 21–30)</paragraph>
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</subSubSection>
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<subSubSection id="D5E96539FF81FFC36051FDF4FC72AB6B" pageId="13" pageNumber="14" type="reference_group">
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<paragraph id="9D4C36B2FF81FFC36051FDF4FD8AAA8B" blockId="13.[264,1323,598,782]" pageId="13" pageNumber="14">
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<taxonomicName id="5AF34D31FF81FFC36051FDF4FD49AA0B" ID-CoL="b0cde5f4-dc64-4b7c-a915-13082b27e261" authority="Plate, 1897: 205" authorityName="Plate" authorityPageNumber="205" authorityYear="1897" box="[264,661,598,622]" class="Polyplacophora" family="Chitonidae" genus="Tonicia" kingdom="Animalia" order="Chitonida" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="calbucensis">
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<emphasis id="AF87EAA0FF81FFC36051FDF4FE07AA08" box="[264,475,599,622]" italics="true" pageId="13" pageNumber="14">Tonicia calbucensis</emphasis>
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<treatmentCitation id="1C5210A3FF81FFC360BAFDF5FD49AA0B" author="Plate" box="[483,661,598,621]" page="205" pageId="13" pageNumber="14" year="1897">
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<bibRefCitation id="F9624B43FF81FFC360BAFDF5FDBCAA0B" author="Plate" box="[483,608,598,621]" pageId="13" pageNumber="26" refString="Plate, L. H. (1897) Die Anatomie und Phylogenie der Chitonen. Fauna Chilensis 1 (1). Zoologische Jahrbucher, Abteilung fur Systematik, Okologie und Geographie der Tiere, 1, Suppl. 4, 1 - 243, pls 1 - 12." type="journal article" year="1897">Plate, 1897</bibRefCitation>
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: 205
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||
</treatmentCitation>
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</taxonomicName>
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–207, pl. 12, fig. 134;
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<treatmentCitation id="1C5210A3FF81FFC362D1FDF5FB82AA0B" author="Nierstrasz" box="[904,1118,598,621]" page="98" pageId="13" pageNumber="14" year="1905">
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<bibRefCitation id="F9624B43FF81FFC362D1FDF5FBE9AA0B" author="Nierstrasz" box="[904,1077,598,621]" pageId="13" pageNumber="26" refString="Nierstrasz, H. F. (1905) Die Chitonen der Siboga-Expedition. Siboga Expeditie, 48, 1 - 112, pls 1 - 8." type="journal article" year="1905">Nierstrasz 1905</bibRefCitation>
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: 98
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</treatmentCitation>
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; 1906: 150;
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<treatmentCitation id="1C5210A3FF81FFC365BFFDF5FE46AAEB" author="Thiele" page="21" pageId="13" pageNumber="14" year="1908">
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<bibRefCitation id="F9624B43FF81FFC365BFFDF5FEADAAEB" author="Thiele" pageId="13" pageNumber="27" refString="Thiele, J. (1908) Die antarktischen und subantarktischen Chitonen. Deutsche Sudpolar Expedition 1901 - 1903, 10. Zoologie, 2, 9 - 23, pl. 1." type="journal article" year="1908">Thiele 1908</bibRefCitation>
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: 21
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</treatmentCitation>
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;
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<treatmentCitation id="1C5210A3FF81FFC360F1FDD5FD0EAAEB" author="Boudet" box="[424,722,630,653]" page="127" pageId="13" pageNumber="14" year="1945">
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<bibRefCitation id="F9624B43FF81FFC360F1FDD5FD47AAEB" author="Boudet" box="[424,667,630,653]" pageId="13" pageNumber="24" refString="Boudet Rommel, I. (1945) Los Quitones Chilenos. Revista Chilena de Historia Natural 48 (144), 122 - 140." type="journal article" year="1945">Boudet Rommel 1945</bibRefCitation>
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: 127
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</treatmentCitation>
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;
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||
<treatmentCitation id="1C5210A3FF81FFC36386FDD5FB81AAEB" author="Carcelles" box="[735,1117,630,653]" page="249" pageId="13" pageNumber="14" year="1951">
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<bibRefCitation id="F9624B43FF81FFC36386FDD5FBF9AAEB" author="Carcelles" box="[735,1061,630,653]" pageId="13" pageNumber="25" refString="Carcelles, A. R. & Williamson, S. I. (1951) Catalogo de los Moluscos marinos de la Provincia Magallanica. Revista del Instituto Nacional de Investigacion de las Ciencias Naturales Anexo al Museo Argentino de Ciencias Naturales " Bernardino Rivadavia ", Zoologicas, 2 (5), 225 - 383." type="journal article" year="1951">Carcelles & Williamson 1951</bibRefCitation>
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: 249
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||
</treatmentCitation>
|
||
;
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||
<treatmentCitation id="1C5210A3FF81FFC36533FDD5FEBEAACB" author="Castellanos" page="478" pageId="13" pageNumber="14" year="1956">
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<bibRefCitation id="F9624B43FF81FFC36533FDD5FAFAAAEB" author="Castellanos" box="[1130,1318,630,653]" pageId="13" pageNumber="25" refString="Castellanos, Z. J. A. de (1956) Catalogo de los poliplacoforos Argentinos y de aguas vecinas al E strecho de Magellanes. Revista del Museo Universidad Nacional de La Plata, Facultad de Ciencias Naturales, Zoologia, 6 (49), 465 - 486, pls 1 - 9." type="journal article" year="1956">Castellanos 1956</bibRefCitation>
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||
: 478
|
||
</treatmentCitation>
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||
, pl. 9; 1988: 22, figs. 71–73;
|
||
<treatmentCitation id="1C5210A3FF81FFC363C7FD35FC8EAACB" author="Leloup" box="[670,850,662,685]" page="78" pageId="13" pageNumber="14" year="1956">
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<bibRefCitation id="F9624B43FF81FFC363C7FD35FCF6AACB" author="Leloup" box="[670,810,662,685]" pageId="13" pageNumber="26" refString="Leloup, E. (1956) Reports of the Lund University Chile Expedition 1948 - 49. 27. Polyplacophora. Lunds Universitets Arsskriften (n. s. (2 )), 52 (15), 1 - 94." type="journal article" year="1956">Leloup 1956</bibRefCitation>
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||
: 78
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="1C5210A3FF81FFC36207FD35FB5BAACB" author="Kaas" box="[862,1159,662,685]" page="20" pageId="13" pageNumber="14" year="1980">
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<bibRefCitation id="F9624B43FF81FFC36207FD35FB83AACB" author="Kaas" box="[862,1119,662,685]" pageId="13" pageNumber="25" refString="Kaas, P. & Van Belle, R. A. (1980) Catalogue of Living Chitons (Mollusca: Polyplacophora), Dr. W. Backhuys Publisher, Rotterdam, 144 p." type="book" year="1980">Kaas & Van Belle 1980</bibRefCitation>
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||
: 20
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</treatmentCitation>
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||
; 1998: 37;
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||
<treatmentCitation id="1C5210A3FF81FFC365A6FD35FE09AAAB" author="Kilias" page="158" pageId="13" pageNumber="14" year="1995">
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<bibRefCitation id="F9624B43FF81FFC365A6FD35FE40AAAB" author="Kilias" pageId="13" pageNumber="26" refString="Kilias, R. (1995). Polyplacophora-Typen und - Typoide (Mollusca) im Zoologischen Museum in Berlin. Mitteilungen aus dem Zoologischen Museum in Berlin, 71 (1), 155 - 170." type="journal article" year="1995">Kilias 1995</bibRefCitation>
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: 158
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||
</treatmentCitation>
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(
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<typeStatus id="42488810FF81FFC360B0FD15FD91AAAB" box="[489,589,694,717]" pageId="13" pageNumber="14" type="lectotype">lectotype</typeStatus>
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designation);
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<treatmentCitation id="1C5210A3FF81FFC363ABFD15FC1CAAAB" author="Slieker" box="[754,960,694,717]" page="140" pageId="13" pageNumber="14" year="2000">
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<bibRefCitation id="F9624B43FF81FFC363ABFD15FC58AAAB" author="Slieker" box="[754,900,694,717]" pageId="13" pageNumber="27" refString="Slieker, F. J. A. (2000) Chitons of the world. An illustrated synopsis of recent Polyplacophora, L'Informatore Piceno, Ancona, i - vi, 1 - 154." type="book chapter" year="2000">Slieker 2000</bibRefCitation>
|
||
: 140
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||
</treatmentCitation>
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||
;
|
||
<treatmentCitation id="1C5210A3FF81FFC3628BFD15FAFAAAAB" author="Sirenko" box="[978,1318,694,717]" page="89" pageId="13" pageNumber="14" year="2005">
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<bibRefCitation id="F9624B43FF81FFC3628BFD15FB2BAAAB" author="Sirenko" box="[978,1271,694,717]" pageId="13" pageNumber="27" refString="Sirenko, B. I. & Gallardo, C. (2005) Chitons (Polyplacophora) of Chile. In: Poster Abstracts of the IV. International Congress of the European Malacological Societies, Oct. 10 - 14. 2005 in Naples (Italy). Notizario S. I. M., Supplem. Bollettino Malacologico, (year 23) (5 - 8), 89." type="journal article" year="2005">Sirenko & Gallardo 2005</bibRefCitation>
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: 89
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</treatmentCitation>
|
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;
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<treatmentCitation id="1C5210A3FF81FFC36061FD75FE23AA8B" author="Sirenko" box="[312,511,726,749]" page="85" pageId="13" pageNumber="14" year="2006">
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<bibRefCitation id="F9624B43FF81FFC36061FD75FE04AA8B" author="Sirenko" box="[312,472,726,749]" pageId="13" pageNumber="27" refString="Sirenko, B. I. (2006 b) Report on the present stage of our knowledge with regard to the chitons (Mollusca: Polyplacophora) of the Magellan Strait and Falkland Islands. Venus, 65 (1 - 2), 81 - 89." type="journal article" year="2006" yearSuffix="b">Sirenko 2006b</bibRefCitation>
|
||
: 85
|
||
</treatmentCitation>
|
||
, fig. 10.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="9D4C36B2FF81FFC36051FD54FC72AB6B" blockId="13.[264,1323,598,782]" box="[264,942,758,782]" pageId="13" pageNumber="14">
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<treatmentCitationGroup id="BDE3119CFF81FFC36051FD54FC72AB6B" box="[264,942,758,782]" pageId="13" pageNumber="14">
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<taxonomicName id="5AF34D31FF81FFC36051FD54FE77AB68" ID-CoL="8TPD4" box="[264,427,759,782]" class="Polyplacophora" family="Chitonidae" genus="Tonicia" kingdom="Animalia" order="Chitonida" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="lebruni">
|
||
<emphasis id="AF87EAA0FF81FFC36051FD54FE77AB68" box="[264,427,759,782]" italics="true" pageId="13" pageNumber="14">Tonicia lebruni</emphasis>
|
||
</taxonomicName>
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||
<taxonomicName id="5AF34D31FF81FFC360EBFD55FD87AB68" box="[434,603,758,782]" pageId="13" pageNumber="14" rank="variety" variety="calbucensis">
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||
var.
|
||
<emphasis id="AF87EAA0FF81FFC36086FD54FD87AB68" box="[479,603,759,782]" italics="true" pageId="13" pageNumber="14">calbucensis</emphasis>
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</taxonomicName>
|
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;
|
||
<treatmentCitation id="1C5210A3FF81FFC36333FD55FCEDAB6B" author="Forcelli" box="[618,817,758,781]" page="186" pageId="13" pageNumber="14" year="1999">
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<bibRefCitation id="F9624B43FF81FFC36333FD55FD21AB6B" author="Forcelli" box="[618,765,758,781]" pageId="13" pageNumber="25" refString="Forcelli, D. O. (1999) Moluscos Magallanicos. Guida de Moluscos de Patagonia y Sur de Chile. Vazquez Mazzni Editores, Buenos Aires, Argentinia, 200 pp." type="book" year="1999">Forcelli 1999</bibRefCitation>
|
||
: 186
|
||
</treatmentCitation>
|
||
, illustrated.
|
||
</treatmentCitationGroup>
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</paragraph>
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<subSubSection id="D5E96539FF81FFC06051FC9DFBB1A9B0" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14" type="description">
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<paragraph id="9D4C36B2FF81FFC06051FC9DFBB1A9B0" blockId="13.[264,1324,830,1897]" lastBlockId="14.[264,1324,284,1510]" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
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<emphasis id="AF87EAA0FF81FFC36051FC9DFE9FAB3E" bold="true" box="[264,323,830,856]" pageId="13" pageNumber="14">Size:</emphasis>
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Maximum length
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<quantity id="5A0B9B57FF81FFC3637BFC9CFDAAAB3F" box="[546,630,831,857]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" pageId="13" pageNumber="14" unit="mm" value="40.0">40 mm</quantity>
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(
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<bibRefCitation id="F9624B43FF81FFC363DFFC9CFCF7AB3F" author="Forcelli" box="[646,811,831,857]" pageId="13" pageNumber="25" refString="Forcelli, D. O. (1999) Moluscos Magallanicos. Guida de Moluscos de Patagonia y Sur de Chile. Vazquez Mazzni Editores, Buenos Aires, Argentinia, 200 pp." type="book" year="1999">Forcelli 1999</bibRefCitation>
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).
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<emphasis id="AF87EAA0FF81FFC36219FC9DFC36AB3E" bold="true" box="[832,1002,830,856]" pageId="13" pageNumber="14">Body outline:</emphasis>
|
||
Elongate oval, moderately elevated, subcarinated to slightly keeled (Figures 21–23).
|
||
<emphasis id="AF87EAA0FF81FFC362A9FCC5FB9DABE6" bold="true" box="[1008,1089,870,896]" pageId="13" pageNumber="14">Color:</emphasis>
|
||
Orange brown to brownish red with darker radial streaks on terminal valves and lateral areas, on central areas these streaks orientated longitudinally. Jugal area in general chestnut brown with two wider white streaks on both sides (Figures 21–26).
|
||
<emphasis id="AF87EAA0FF81FFC3622AFC7DFC16AB9E" bold="true" box="[883,970,990,1016]" pageId="13" pageNumber="14">Valves:</emphasis>
|
||
Solid and smooth except for faint growth marks arranged commarginally, more distinct at valve margins. Small granules partly extending along diagonal ridge. Head valve (Figure 24) semicircular with wide Vshaped posterior margin, weakly notched in middle. Intermediate valves rectangular, straight posterior and anterior valve margins parallel, but anterior valve margin extends anteriorly in valve ii. Apices slightly protruding. Lateral areas slightly elevated and clearly indicated by different arrangement of brownish streaks, radially in lateral areas, longitudinally in pleural areas (Figure 25). Semicircular tail valve (Figure 26) with narrow antemucronal area, mucro elevated but directed backward. Mucro situated in front of valve middle. Postmucronal slope steep and straight. Ocelli on radial streaks of lateral areas and terminal valves only (not in white zones in between).
|
||
<emphasis id="AF87EAA0FF81FFC36509FACDFAF7ADEE" bold="true" box="[1104,1323,1390,1416]" pageId="13" pageNumber="14">Articulamentum:</emphasis>
|
||
White and strongly developed. Centrally with a brownish spot. Apophyses solid, moderately large, triangular on valve ii, rectangulartrapezoidal on remaining valves (Figures 25–26). Apophyses connected by short sawed jugal sinus, laterally deeply notched. Slit formula: 8 / 1 / 8 (9 / 1 /
|
||
<quantity id="5A0B9B57FF81FFC36398F9ACFD32AE4F" box="[705,750,1551,1577]" metricMagnitude="-1" metricUnit="m" metricValue="2.286" pageId="13" pageNumber="14" unit="in" value="9.0">9 in</quantity>
|
||
original description), teeth rather short but wide. Teeth strongly pectinated on outside. Slit rays present in all valves.
|
||
<emphasis id="AF87EAA0FF81FFC36543F995FB75AE36" bold="true" box="[1050,1193,1590,1616]" pageId="13" pageNumber="14">Perinotum:</emphasis>
|
||
Very wide and fleshy. Dorsally covered with minute conical spicules (appears naked), which are smooth and sharppointed, round in cross section (
|
||
<figureCitation id="05C82A37FF81FFC36232F924FBC2AEC7" box="[875,1054,1671,1697]" captionStart="FIGURES 27 – 30" captionStartId="21.[264,382,1174,1198]" captionTargetBox="[264,1323,326,1150]" captionTargetId="figure@21.[264,1323,326,1150]" captionTargetPageId="21" captionText="FIGURES 27 – 30. Tonicia calbucensis Plate, 1897, disarticulated specimen from station 7, from a depth of 20 m (ZSM Moll 20050326). 27 – 28. Spicules of the dorsal perinotum, in situ. 29. Ventral perinotum scales, in situ. 30. Anterior portion of the radula, showing central and inner lateral teeth. Scale bars: 27 – 29 = 10 µm; 30 = 100 µm." httpUri="https://zenodo.org/record/174414/files/figure.png" pageId="13" pageNumber="14">Figures 27–28</figureCitation>
|
||
). Ventrally with radial rows of roundish to squarish scales, rounded at their free end. Scales not imbricated. Radial rows always interrupted. Scale arrangement resembles that of short pearl necklets (
|
||
<figureCitation id="05C82A37FF81FFC36049F95CFE54AF7F" box="[272,392,1791,1817]" captionStart="FIGURES 27 – 30" captionStartId="21.[264,382,1174,1198]" captionTargetBox="[264,1323,326,1150]" captionTargetId="figure@21.[264,1323,326,1150]" captionTargetPageId="21" captionText="FIGURES 27 – 30. Tonicia calbucensis Plate, 1897, disarticulated specimen from station 7, from a depth of 20 m (ZSM Moll 20050326). 27 – 28. Spicules of the dorsal perinotum, in situ. 29. Ventral perinotum scales, in situ. 30. Anterior portion of the radula, showing central and inner lateral teeth. Scale bars: 27 – 29 = 10 µm; 30 = 100 µm." httpUri="https://zenodo.org/record/174414/files/figure.png" pageId="13" pageNumber="14">Figure 29</figureCitation>
|
||
).
|
||
<emphasis id="AF87EAA0FF81FFC360C7F95DFDD6AF7E" bold="true" box="[414,522,1790,1816]" pageId="13" pageNumber="14">Ctenidia</emphasis>
|
||
(specimen
|
||
<quantity id="5A0B9B57FF81FFC363CAF95CFD21AF7F" box="[659,765,1791,1817]" metricMagnitude="-2" metricUnit="m" metricValue="2.88" pageId="13" pageNumber="14" unit="mm" value="28.8">28.8 mm</quantity>
|
||
body length): 41 ctenidia on each side of foot, arranged holobranchilly and adanally, group of largest ctenidia situated beneath valves vivii.
|
||
<emphasis id="AF87EAA0FF81FFC3606CF8EDFE4CAF0E" bold="true" box="[309,400,1870,1896]" pageId="13" pageNumber="14">Radula</emphasis>
|
||
(same specimen): Radula length
|
||
<quantity id="5A0B9B57FF81FFC3624CF8ECFCAEAF0F" box="[789,882,1871,1897]" metricMagnitude="-3" metricUnit="m" metricValue="9.4" pageId="13" pageNumber="14" unit="mm" value="9.4">9.4 mm</quantity>
|
||
, radula cartilage
|
||
<quantity id="5A0B9B57FF81FFC36561F8ECFB4DAF0F" box="[1080,1169,1871,1897]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="13" pageNumber="14" unit="mm" value="2.9">2.9 mm</quantity>
|
||
in length; 66 teeth rows, 55 already mineralized. Central tooth slightly depressed in middle, slender, with simple inwarddirected blade. First lateral tooth elongate, widened at base, on its upper end directed inward with simple blade, latter slightly notched in middle. Head of second lateral tooth discoidal, with slightly extended side margins and obtusely pointed tip of cutting edge. Major lateral tooth spoonshaped and smooth (
|
||
<figureCitation id="05C82A37FF82FFC062B0FE1FFBBCA9B0" box="[1001,1120,444,470]" captionStart="FIGURES 27 – 30" captionStartId="21.[264,382,1174,1198]" captionTargetBox="[264,1323,326,1150]" captionTargetId="figure@21.[264,1323,326,1150]" captionTargetPageId="21" captionText="FIGURES 27 – 30. Tonicia calbucensis Plate, 1897, disarticulated specimen from station 7, from a depth of 20 m (ZSM Moll 20050326). 27 – 28. Spicules of the dorsal perinotum, in situ. 29. Ventral perinotum scales, in situ. 30. Anterior portion of the radula, showing central and inner lateral teeth. Scale bars: 27 – 29 = 10 µm; 30 = 100 µm." httpUri="https://zenodo.org/record/174414/files/figure.png" pageId="14" pageNumber="15">Figure 30</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D5E96539FF82FFC06061FE40FBD8AA40" pageId="14" pageNumber="15" type="materials_examined">
|
||
<paragraph id="9D4C36B2FF82FFC06061FE40FBD8AA40" blockId="14.[264,1324,284,1510]" pageId="14" pageNumber="15">
|
||
<emphasis id="AF87EAA0FF82FFC06061FE40FE72A99B" bold="true" box="[312,430,483,509]" pageId="14" pageNumber="15">Material:</emphasis>
|
||
Thirtyfour specimens were collected from 4 stations within the fjord, covering a bathymetric range from
|
||
<quantity id="5A0B9B57FF82FFC063FFFDAFFCCAAA40" box="[678,790,524,550]" metricMagnitude="1" metricUnit="m" metricValue="1.75" metricValueMax="3.0" metricValueMin="0.5" pageId="14" pageNumber="15" unit="m" value="17.5" valueMax="30.0" valueMin="5.0">5 to 30 m</quantity>
|
||
depth (see
|
||
<tableCitation id="D0710309FF82FFC062C5FDAFFC2EAA40" box="[924,1010,524,550]" captionStart="TABLE 1" captionStartId="20.[264,352,443,467]" captionTargetBox="[272,1303,500,864]" captionTargetPageId="20" captionText="TABLE 1. Species and specimens collected by station." httpUri="http://table.plazi.org/id/C98C663AFF98FFDA6051FE18FC88A9B5" pageId="14" pageNumber="15" tableUuid="C98C663AFF98FFDA6051FE18FC88A9B5">Table 1</tableCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D5E96539FF82FFC06061FD90FB9FAA10" pageId="14" pageNumber="15" type="distribution">
|
||
<paragraph id="9D4C36B2FF82FFC06061FD90FB9FAA10" blockId="14.[264,1324,284,1510]" pageId="14" pageNumber="15">
|
||
<emphasis id="AF87EAA0FF82FFC06061FD90FE06AA2B" bold="true" box="[312,474,563,589]" pageId="14" pageNumber="15">Distribution:</emphasis>
|
||
<taxonomicName id="5AF34D31FF82FFC060B1FD97FD09AA2B" box="[488,725,564,589]" class="Polyplacophora" family="Chitonidae" genus="Tonicia" kingdom="Animalia" order="Chitonida" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="calbucensis">
|
||
<emphasis id="AF87EAA0FF82FFC060B1FD97FD09AA2B" box="[488,725,564,589]" italics="true" pageId="14" pageNumber="15">Tonicia calbucensis</emphasis>
|
||
</taxonomicName>
|
||
occurs from Puerto Montt southwards to Punta Arenas in the Straits of Magellan, from
|
||
<quantity id="5A0B9B57FF82FFC06382FDFFFC97AA10" box="[731,843,604,630]" metricMagnitude="1" metricUnit="m" metricValue="1.75" metricValueMax="3.5" metricValueMin="0.0" pageId="14" pageNumber="15" unit="m" value="17.5" valueMax="35.0" valueMin="0.0">0 to 35 m</quantity>
|
||
depth (
|
||
<bibRefCitation id="F9624B43FF82FFC062FAFDFFFBEAAA10" author="Thiele" box="[931,1078,604,630]" pageId="14" pageNumber="27" refString="Thiele, J. (1908) Die antarktischen und subantarktischen Chitonen. Deutsche Sudpolar Expedition 1901 - 1903, 10. Zoologie, 2, 9 - 23, pl. 1." type="journal article" year="1908">Thiele 1908</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D5E96539FF82FFC06061FD27FB94AB70" pageId="14" pageNumber="15" type="discussion">
|
||
<paragraph id="9D4C36B2FF82FFC06061FD27FB94AB70" blockId="14.[264,1324,284,1510]" pageId="14" pageNumber="15">
|
||
<emphasis id="AF87EAA0FF82FFC06061FD27FE6FAAF8" bold="true" box="[312,435,644,670]" pageId="14" pageNumber="15">Remarks:</emphasis>
|
||
Besides the specimens cited above, and the
|
||
<typeStatus id="42488810FF82FFC062B2FD27FB8AAAF8" box="[1003,1110,644,670]" pageId="14" pageNumber="15" type="lectotype">lectotype</typeStatus>
|
||
(Natural History Museum Berlin [ZMB] Moll 102.006, Figures 21–23), material from additional localities between Puerto Montt and Punta Arenas (ZSM) was examined. Specimens from the samples observed showed consistent valve shapes and shell coloration.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D5E96539FF82FFC06061FC87FBD5AD80" pageId="14" pageNumber="15" type="reference_group">
|
||
<paragraph id="9D4C36B2FF82FFC06061FC87FBD5AD80" blockId="14.[264,1324,284,1510]" pageId="14" pageNumber="15">
|
||
<taxonomicName id="5AF34D31FF82FFC06061FC87FDFAAB5B" ID-CoL="b0cde5f4-dc64-4b7c-a915-13082b27e261" box="[312,550,804,829]" class="Polyplacophora" family="Chitonidae" genus="Tonicia" kingdom="Animalia" order="Chitonida" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="calbucensis">
|
||
<emphasis id="AF87EAA0FF82FFC06061FC87FDFAAB5B" box="[312,550,804,829]" italics="true" pageId="14" pageNumber="15">Tonicia calbucensis</emphasis>
|
||
</taxonomicName>
|
||
was synonymized with
|
||
<taxonomicName id="5AF34D31FF82FFC06205FC87FBC9AB5B" ID-CoL="8TPD4" box="[860,1045,804,829]" class="Polyplacophora" family="Chitonidae" genus="Tonicia" kingdom="Animalia" order="Chitonida" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="lebruni">
|
||
<emphasis id="AF87EAA0FF82FFC06205FC87FBC9AB5B" box="[860,1045,804,829]" italics="true" pageId="14" pageNumber="15">Tonicia lebruni</emphasis>
|
||
</taxonomicName>
|
||
de Rochebrune, 1884 (
|
||
<figureCitation id="05C82A37FF82FFC06049FCEFFE5AAB00" box="[272,390,844,870]" captionStart="FIGURE 20" captionStartId="18.[264,368,1394,1418]" captionTargetBox="[284,1308,312,1376]" captionTargetId="figure@18.[284,1308,312,1376]" captionTargetPageId="18" captionText="FIGURE 20. Tonicia lebruni de Rochebrune, 1884, dorsal view of one syntype in MNHN, anterior at left (modified photograph kindly provided by Delphine Brabant, MNHN). 21 – 23. Tonicia calbucensis Plate, 1897, lectotype from Punta Arenas, Chile, from a depth of “ 8 Faden ” [= 14.63 m] (ZMB Moll 102.006). 21. Dorsal view of the intermediate valves, anterior at left. 22. Right lateral view, anterior at left. 23. Dorsal view of the last two valves, anterior at top. 24 – 26. Tonicia calbucensis Plate, 1897, dorsal view of a disarticulated specimen from station 7, from a depth of 20 m (ZSM Moll 20050326), anterior at top. 24. Head valve. 25. Valve ii. 26. Tail valve." httpUri="https://zenodo.org/record/174413/files/figure.png" pageId="14" pageNumber="15">Figure 20</figureCitation>
|
||
) by
|
||
<bibRefCitation id="F9624B43FF82FFC060E4FCEFFDB7AB00" author="Leloup" box="[445,619,844,870]" pageId="14" pageNumber="26" refString="Leloup, E. (1956) Reports of the Lund University Chile Expedition 1948 - 49. 27. Polyplacophora. Lunds Universitets Arsskriften (n. s. (2 )), 52 (15), 1 - 94." type="journal article" year="1956">Leloup (1956)</bibRefCitation>
|
||
. He interpreted a small specimen of
|
||
<taxonomicName id="5AF34D31FF82FFC06578FCEFFB4DAB03" ID-CoL="8TPD4" box="[1057,1169,844,869]" class="Polyplacophora" family="Chitonidae" genus="Tonicia" kingdom="Animalia" order="Chitonida" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="lebruni">
|
||
<emphasis id="AF87EAA0FF82FFC06578FCEFFB4DAB03" box="[1057,1169,844,869]" italics="true" pageId="14" pageNumber="15">T. lebruni</emphasis>
|
||
</taxonomicName>
|
||
with similar coloration, perinotum elements and radula to refer to
|
||
<taxonomicName id="5AF34D31FF82FFC06299FCD7FB6CABEB" ID-CoL="b0cde5f4-dc64-4b7c-a915-13082b27e261" box="[960,1200,884,909]" class="Polyplacophora" family="Chitonidae" genus="Tonicia" kingdom="Animalia" order="Chitonida" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="calbucensis">
|
||
<emphasis id="AF87EAA0FF82FFC06299FCD7FB6CABEB" box="[960,1200,884,909]" italics="true" pageId="14" pageNumber="15">Tonicia calbucensis</emphasis>
|
||
</taxonomicName>
|
||
. External examination of two dry, flat preserved
|
||
<typeStatus id="42488810FF82FFC06396FC3FFCEAABD0" box="[719,822,924,950]" pageId="14" pageNumber="15" type="syntype">syntypes</typeStatus>
|
||
of
|
||
<taxonomicName id="5AF34D31FF82FFC06205FC3FFC12ABD3" ID-CoL="8TPD4" box="[860,974,924,949]" class="Polyplacophora" family="Chitonidae" genus="Tonicia" kingdom="Animalia" order="Chitonida" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="lebruni">
|
||
<emphasis id="AF87EAA0FF82FFC06205FC3FFC12ABD3" box="[860,974,924,949]" italics="true" pageId="14" pageNumber="15">T. lebruni</emphasis>
|
||
</taxonomicName>
|
||
(Muséum National d'Histoire Naturelle, Paris,
|
||
<collectingCountry id="E5E47622FF82FFC0608AFC67FDFFABB8" box="[467,547,964,990]" name="France" pageId="14" pageNumber="15">France</collectingCountry>
|
||
[MNHN]) show them to reach approximately
|
||
<quantity id="5A0B9B57FF82FFC06502FC67FB6CABB8" box="[1115,1200,964,990]" metricMagnitude="-2" metricUnit="m" metricValue="4.8" pageId="14" pageNumber="15" unit="mm" value="48.0">48 mm</quantity>
|
||
in length, having all valves
|
||
<emphasis id="AF87EAA0FF82FFC06086FC4FFDF7AC63" box="[479,555,1004,1029]" italics="true" pageId="14" pageNumber="15">in situ</emphasis>
|
||
, and being chestnut with a shiny surface. Comparison between
|
||
<typeStatus id="42488810FF82FFC06051FBB7FEE6AC48" box="[264,314,1044,1070]" pageId="14" pageNumber="15">type</typeStatus>
|
||
samples of
|
||
<taxonomicName id="5AF34D31FF82FFC06082FBB7FD5BAC4B" ID-CoL="b0cde5f4-dc64-4b7c-a915-13082b27e261" box="[475,647,1044,1069]" class="Polyplacophora" family="Chitonidae" genus="Tonicia" kingdom="Animalia" order="Chitonida" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="calbucensis">
|
||
<emphasis id="AF87EAA0FF82FFC06082FBB7FD5BAC4B" box="[475,647,1044,1069]" italics="true" pageId="14" pageNumber="15">T. calbucensis</emphasis>
|
||
</taxonomicName>
|
||
(Figures 21–23) and
|
||
<taxonomicName id="5AF34D31FF82FFC062C7FBB7FBCFAC4B" ID-CoL="8TPD4" box="[926,1043,1044,1069]" class="Polyplacophora" family="Chitonidae" genus="Tonicia" kingdom="Animalia" order="Chitonida" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="lebruni">
|
||
<emphasis id="AF87EAA0FF82FFC062C7FBB7FBCFAC4B" box="[926,1043,1044,1069]" italics="true" pageId="14" pageNumber="15">T. lebruni</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="05C82A37FF82FFC06572FBB7FB7AAC48" box="[1067,1190,1044,1070]" captionStart="FIGURE 20" captionStartId="18.[264,368,1394,1418]" captionTargetBox="[284,1308,312,1376]" captionTargetId="figure@18.[284,1308,312,1376]" captionTargetPageId="18" captionText="FIGURE 20. Tonicia lebruni de Rochebrune, 1884, dorsal view of one syntype in MNHN, anterior at left (modified photograph kindly provided by Delphine Brabant, MNHN). 21 – 23. Tonicia calbucensis Plate, 1897, lectotype from Punta Arenas, Chile, from a depth of “ 8 Faden ” [= 14.63 m] (ZMB Moll 102.006). 21. Dorsal view of the intermediate valves, anterior at left. 22. Right lateral view, anterior at left. 23. Dorsal view of the last two valves, anterior at top. 24 – 26. Tonicia calbucensis Plate, 1897, dorsal view of a disarticulated specimen from station 7, from a depth of 20 m (ZSM Moll 20050326), anterior at top. 24. Head valve. 25. Valve ii. 26. Tail valve." httpUri="https://zenodo.org/record/174413/files/figure.png" pageId="14" pageNumber="15">Figure 20</figureCitation>
|
||
) reveals a number of differences: 1) the head valve has a much steeper slope in
|
||
<taxonomicName id="5AF34D31FF82FFC06504FB9FFB13AC33" ID-CoL="8TPD4" box="[1117,1231,1084,1109]" class="Polyplacophora" family="Chitonidae" genus="Tonicia" kingdom="Animalia" order="Chitonida" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="lebruni">
|
||
<emphasis id="AF87EAA0FF82FFC06504FB9FFB13AC33" box="[1117,1231,1084,1109]" italics="true" pageId="14" pageNumber="15">T. lebruni</emphasis>
|
||
</taxonomicName>
|
||
, 2) the apices in
|
||
<taxonomicName id="5AF34D31FF82FFC06025FBC7FDFFAC1B" ID-CoL="b0cde5f4-dc64-4b7c-a915-13082b27e261" box="[380,547,1124,1149]" class="Polyplacophora" family="Chitonidae" genus="Tonicia" kingdom="Animalia" order="Chitonida" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="calbucensis">
|
||
<emphasis id="AF87EAA0FF82FFC06025FBC7FDFFAC1B" box="[380,547,1124,1149]" italics="true" pageId="14" pageNumber="15">T. calbucensis</emphasis>
|
||
</taxonomicName>
|
||
are hardly protruding in the first two intermediate valves, while they protrude in all valves of
|
||
<taxonomicName id="5AF34D31FF82FFC06307FB2FFD12ACC3" ID-CoL="8TPD4" box="[606,718,1164,1189]" class="Polyplacophora" family="Chitonidae" genus="Tonicia" kingdom="Animalia" order="Chitonida" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="lebruni">
|
||
<emphasis id="AF87EAA0FF82FFC06307FB2FFD12ACC3" box="[606,718,1164,1189]" italics="true" pageId="14" pageNumber="15">T. lebruni</emphasis>
|
||
</taxonomicName>
|
||
, 3) dorsally
|
||
<taxonomicName id="5AF34D31FF82FFC06238FB2FFBDAACC3" ID-CoL="b0cde5f4-dc64-4b7c-a915-13082b27e261" box="[865,1030,1164,1189]" class="Polyplacophora" family="Chitonidae" genus="Tonicia" kingdom="Animalia" order="Chitonida" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="calbucensis">
|
||
<emphasis id="AF87EAA0FF82FFC06238FB2FFBDAACC3" box="[865,1030,1164,1189]" italics="true" pageId="14" pageNumber="15">T. calbucensis</emphasis>
|
||
</taxonomicName>
|
||
is more carinated than
|
||
<taxonomicName id="5AF34D31FF82FFC0644CFB2FFE81ACAB" ID-CoL="8TPD4" class="Polyplacophora" family="Chitonidae" genus="Tonicia" kingdom="Animalia" order="Chitonida" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="lebruni">
|
||
<emphasis id="AF87EAA0FF82FFC0644CFB2FFE81ACAB" italics="true" pageId="14" pageNumber="15">T. lebruni</emphasis>
|
||
</taxonomicName>
|
||
, 4) the lateral areas are defined more clearly in
|
||
<taxonomicName id="5AF34D31FF82FFC062FBFB17FBCFACAB" ID-CoL="8TPD4" box="[930,1043,1204,1229]" class="Polyplacophora" family="Chitonidae" genus="Tonicia" kingdom="Animalia" order="Chitonida" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="lebruni">
|
||
<emphasis id="AF87EAA0FF82FFC062FBFB17FBCFACAB" box="[930,1043,1204,1229]" italics="true" pageId="14" pageNumber="15">T. lebruni</emphasis>
|
||
</taxonomicName>
|
||
and have an anteriorly shallow slope, while the diagonal ridge in
|
||
<taxonomicName id="5AF34D31FF82FFC06255FB7FFC68AC93" ID-CoL="b0cde5f4-dc64-4b7c-a915-13082b27e261" box="[780,948,1244,1269]" class="Polyplacophora" family="Chitonidae" genus="Tonicia" kingdom="Animalia" order="Chitonida" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="calbucensis">
|
||
<emphasis id="AF87EAA0FF82FFC06255FB7FFC68AC93" box="[780,948,1244,1269]" italics="true" pageId="14" pageNumber="15">T. calbucensis</emphasis>
|
||
</taxonomicName>
|
||
is elevated immediately above the anterior slope, 5)
|
||
<taxonomicName id="5AF34D31FF82FFC0635EFAA7FDA6AD7B" ID-CoL="8TPD4" box="[519,634,1284,1309]" class="Polyplacophora" family="Chitonidae" genus="Tonicia" kingdom="Animalia" order="Chitonida" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="lebruni">
|
||
<emphasis id="AF87EAA0FF82FFC0635EFAA7FDA6AD7B" box="[519,634,1284,1309]" italics="true" pageId="14" pageNumber="15">T. lebruni</emphasis>
|
||
</taxonomicName>
|
||
has uniformly commarginal striations on the central area, while this region is longitudinally striated in
|
||
<taxonomicName id="5AF34D31FF82FFC0626AFA8FFC01AD23" ID-CoL="b0cde5f4-dc64-4b7c-a915-13082b27e261" box="[819,989,1324,1349]" class="Polyplacophora" family="Chitonidae" genus="Tonicia" kingdom="Animalia" order="Chitonida" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="calbucensis">
|
||
<emphasis id="AF87EAA0FF82FFC0626AFA8FFC01AD23" box="[819,989,1324,1349]" italics="true" pageId="14" pageNumber="15">T. calbucensis</emphasis>
|
||
</taxonomicName>
|
||
, 6) the postmucronal slope appears lesser steep in
|
||
<taxonomicName id="5AF34D31FF82FFC06373FAF7FD09AD0B" ID-CoL="b0cde5f4-dc64-4b7c-a915-13082b27e261" box="[554,725,1364,1389]" class="Polyplacophora" family="Chitonidae" genus="Tonicia" kingdom="Animalia" order="Chitonida" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="calbucensis">
|
||
<emphasis id="AF87EAA0FF82FFC06373FAF7FD09AD0B" box="[554,725,1364,1389]" italics="true" pageId="14" pageNumber="15">T. calbucensis</emphasis>
|
||
</taxonomicName>
|
||
. These characters are consistent with the newly collected specimens, and intermediate forms were not observed. This leads us to reinstate
|
||
<taxonomicName id="5AF34D31FF82FFC06051FA07FE72ADDB" ID-CoL="b0cde5f4-dc64-4b7c-a915-13082b27e261" box="[264,430,1444,1469]" class="Polyplacophora" family="Chitonidae" genus="Tonicia" kingdom="Animalia" order="Chitonida" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="calbucensis">
|
||
<emphasis id="AF87EAA0FF82FFC06051FA07FE72ADDB" box="[264,430,1444,1469]" italics="true" pageId="14" pageNumber="15">T. calbucensis</emphasis>
|
||
</taxonomicName>
|
||
as distinct from the sympatric
|
||
<taxonomicName id="5AF34D31FF82FFC06245FA07FC52ADDB" ID-CoL="8TPD4" box="[796,910,1444,1469]" class="Polyplacophora" family="Chitonidae" genus="Tonicia" kingdom="Animalia" order="Chitonida" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="lebruni">
|
||
<emphasis id="AF87EAA0FF82FFC06245FA07FC52ADDB" box="[796,910,1444,1469]" italics="true" pageId="14" pageNumber="15">T. lebruni</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<bibRefCitation id="F9624B43FF82FFC062C4FA07FB81ADD8" author="Sirenko" box="[925,1117,1444,1470]" pageId="14" pageNumber="27" refString="Sirenko, B. I. (2006 b) Report on the present stage of our knowledge with regard to the chitons (Mollusca: Polyplacophora) of the Magellan Strait and Falkland Islands. Venus, 65 (1 - 2), 81 - 89." type="journal article" year="2006" yearSuffix="b">Sirenko (2006b)</bibRefCitation>
|
||
also underscored valve and perinotum characters and came to the same conclusion.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |