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<document ID-DOI="http://dx.doi.org/10.3897/zse.96.48312" ID-GBIF-Dataset="7de3ea60-ba35-4831-ae1d-048dc362f8b0" ID-GBIF-Taxon="161939053" ID-Pensoft-Pub="1860-0743-1-25" ID-Pensoft-UUID="4EDA9B5B777551B7BBB0152DD188E5EF" ID-ZooBank="F2C8585A1268443693348B64AE20F6EE" ModsDocID="1860-0743-1-25" checkinTime="1580744411276" checkinUser="pensoft" docAuthor="Sands, Arthur F, Gloeer, Peter, Guerlek, Mustafa E, Albrecht, Christian &amp; Neubauer, Thomas A" docDate="2020" docId="0B43E9776B3A5F6A926554482B2B4A6F" docLanguage="en" docName="ZoosystEvol 96(1): 25-66" docOrigin="Zoosystematics and Evolution 96 (1)" docSource="http://dx.doi.org/10.3897/zse.96.48312" docTitle="Theodoxus gloeri Odabasi &amp; Arslan 2015" docType="treatment" docVersion="6" id="4EDA9B5B777551B7BBB0152DD188E5EF" lastPageNumber="25" masterDocId="4EDA9B5B777551B7BBB0152DD188E5EF" masterDocTitle="A revision of the extant species of Theodoxus (Gastropoda, Neritidae) in Asia, with the description of three new species" masterLastPageNumber="66" masterPageNumber="25" pageId="0" pageNumber="25" updateTime="1635972787372" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>A revision of the extant species of Theodoxus (Gastropoda, Neritidae) in Asia, with the description of three new species</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sands, Arthur F</mods:namePart>
<mods:affiliation>Justus Liebig Univrsity Giessen, Giessen, Germany</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Gloeer, Peter</mods:namePart>
<mods:affiliation>Biodiversity Research Laboratory, Hetlingen, Germany</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Guerlek, Mustafa E</mods:namePart>
<mods:affiliation>Mehmet Akif Ersoy niversity, Burdur, Turkey</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Albrecht, Christian</mods:namePart>
<mods:affiliation>Justus Liebig University Giessen, Giessen, Germany</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Neubauer, Thomas A</mods:namePart>
<mods:affiliation>Justus Liebig University Giessen, Giessen, Germany &amp; Naturalis Biodiversity Centre, Leiden, Netherlands</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>Zoosystematics and Evolution</mods:title>
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<mods:part>
<mods:date>2020</mods:date>
<mods:detail type="volume">
<mods:number>96</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>1</mods:number>
</mods:detail>
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<mods:start>25</mods:start>
<mods:end>66</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zse.96.48312</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zse.96.48312</mods:identifier>
<mods:identifier type="Pensoft-Pub">1860-0743-1-25</mods:identifier>
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<treatment ID-GBIF-Taxon="161939053" LSID="urn:lsid:plazi:treatment:0B43E9776B3A5F6A926554482B2B4A6F" httpUri="http://treatment.plazi.org/id/0B43E9776B3A5F6A926554482B2B4A6F" lastPageNumber="25" pageId="0" pageNumber="25">
<subSubSection pageId="0" pageNumber="25" type="nomenclature">
<paragraph pageId="0" pageNumber="25">
<taxonomicName LSID="0B43E977-6B3A-5F6A-9265-54482B2B4A6F" authority="Odabasi &amp; Arslan, 2015" authorityName="Odabasi &amp; Arslan" authorityYear="2015" class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus gloeri" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="gloeri">
Theodoxus gloeri
<normalizedToken originalValue="Odabaşı">Odabasi</normalizedToken>
&amp; Arslan, 2015
</taxonomicName>
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Theodoxus gloeri Odabasi &amp; Arslan, 2015. A - C. Paratype from Balikdami Wetland-Sakarya River, Eskisehir, Turkey (coll. Gloeer). Scale bar: 1 mm." figureDoi="10.3897/zse.96.48312.figure11" httpUri="https://binary.pensoft.net/fig/376460" pageId="0" pageNumber="25">Figure 11A-C</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="25" type="reference_group">
<paragraph pageId="0" pageNumber="25">
<taxonomicName authorityName="Odabasi &amp; Arslan" authorityYear="2015" class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus gloeri" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="gloeri">Theodoxus gloeri</taxonomicName>
<bibRefCitation author="Odabasi, DA" journalOrPublisher="Ecologica Montenegrina" pageId="0" pageNumber="25" pagination="327 - 333" refId="B115" refString="Odabasi, DA, Arslan, N, 2015. Description of a new subterranean nerite: Theodoxus gloeri n. sp. with some data on the freshwater gastropod fauna of Balikdami Wetland (Sakarya River, Turkey). Ecologica Montenegrina 2 (4): 327 - 333" title="Description of a new subterranean nerite: Theodoxus gloeri n. sp. with some data on the freshwater gastropod fauna of Balikdami Wetland (Sakarya River, Turkey)." volume="2" year="2015">
<normalizedToken originalValue="Odabaşı">Odabasi</normalizedToken>
and Arslan 2015
</bibRefCitation>
: 329-330, fig. 2;
<bibRefCitation author="Guerlek, ME" journalOrPublisher="Fresenius Enviromental Bulletin" pageId="0" pageNumber="25" pagination="2992 - 3013" refId="B69" refString="Guerlek, ME, Sahin, SK, Doekuemcue, N, Yildirim, MZ, 2019. Checklist of the freshwater mollusca of Turkey (Mollusca: Gastropoda, Bivalvia). Fresenius Enviromental Bulletin 28 (4): 2992 - 3013" title="Checklist of the freshwater mollusca of Turkey (Mollusca: Gastropoda, Bivalvia)." volume="28" year="2019">
<normalizedToken originalValue="Gürlek">Guerlek</normalizedToken>
et al. 2019
</bibRefCitation>
: 2993;
<bibRefCitation author="Gloeer, P" journalOrPublisher="Peter Gloeer, Hetlingen, Germany" pageId="0" pageNumber="25" refId="B62" refString="Gloeer, P, 2019. The freshwater gastropods of the West-Palaearctis. Vol. 1. Fresh- and brackish waters except spring and subterranean snails. Peter Gloeer, Hetlingen, Germany" title="The freshwater gastropods of the West-Palaearctis. Vol. 1. Fresh- and brackish waters except spring and subterranean snails." year="2019">
<normalizedToken originalValue="Glöer">Gloeer</normalizedToken>
2019
</bibRefCitation>
: 44, fig. 29.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="25" type="type locality">
<paragraph pageId="0" pageNumber="25">
<typeStatus>Type</typeStatus>
locality.
</paragraph>
<paragraph pageId="0" pageNumber="25">
<normalizedToken originalValue="Balıkdamı">Balikdami</normalizedToken>
Wetland-Sakarya River,
<normalizedToken originalValue="Eskişehir">Eskisehir</normalizedToken>
, Turkey;
<geoCoordinate degrees="39.15277" direction="north" orientation="latitude" precision="1" value="39.15277">39.15277°N</geoCoordinate>
,
<geoCoordinate degrees="31.61562" direction="east" orientation="longitude" precision="1" value="31.61562">31.61562°E</geoCoordinate>
(Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Occurrence map for Theodoxus spp. in Asia. All points conform to specimens used in the phylogenetic analyses (Suppl. material 1: Table S 1), except for T. gloeri for which no molecular data are available and only the type and paratype localities are indicated. Species are partitioned across four identical maps (A - D) to limit overlap of points as far as possible. Barring T. gloeri, closer related species are grouped together (Fig. 2)." figureDoi="10.3897/zse.96.48312.figure3" httpUri="https://binary.pensoft.net/fig/376452" pageId="0" pageNumber="25">3D</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="25" type="type material">
<paragraph pageId="0" pageNumber="25">
<typeStatus>Type</typeStatus>
material.
</paragraph>
<paragraph pageId="0" pageNumber="25">
<materialsCitation country="Turkey" typeStatus="Holotype">
<typeStatus>Holotype</typeStatus>
and seven
<typeStatus>paratypes</typeStatus>
are deposited in COMULM (coll. no. COMULM-G 0052-0053)
</materialsCitation>
.
<materialsCitation collectionCode="OGUHB" country="Turkey" typeStatus="paratypes">
Further nine
<typeStatus>paratypes</typeStatus>
are deposited in
<collectionCode collectionName="OGUHB">OGUHB</collectionCode>
(coll. no. OGUHB-01759)
</materialsCitation>
.
<materialsCitation country="Turkey" typeStatus="paratype">
An additional
<typeStatus>paratype</typeStatus>
is stored in the private collection of one of us (coll.
<normalizedToken originalValue="Glöer">Gloeer</normalizedToken>
; Fig.
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Theodoxus gloeri Odabasi &amp; Arslan, 2015. A - C. Paratype from Balikdami Wetland-Sakarya River, Eskisehir, Turkey (coll. Gloeer). Scale bar: 1 mm." figureDoi="10.3897/zse.96.48312.figure11" httpUri="https://binary.pensoft.net/fig/376460" pageId="0" pageNumber="25">11A-C</figureCitation>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="25" type="remarks">
<paragraph pageId="0" pageNumber="25">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="25">
<taxonomicName authorityName="Odabasi &amp; Arslan" authorityYear="2015" class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus gloeri" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="gloeri">
<emphasis italics="true" pageId="0" pageNumber="25">Theodoxus gloeri</emphasis>
</taxonomicName>
<normalizedToken originalValue="Odabaşı">Odabasi</normalizedToken>
&amp; Arslan, 2015 shows strong similarity to the fossil species
<taxonomicName genus="T." lsidName="T. pilidei" pageId="0" pageNumber="25" rank="species" species="pilidei">
<emphasis italics="true" pageId="0" pageNumber="25">T. pilidei</emphasis>
</taxonomicName>
(
<normalizedToken originalValue="Tournouër">Tournouer</normalizedToken>
, 1879) from Romania and
<taxonomicName genus="T." lsidName="T. lamelliferus" pageId="0" pageNumber="25" rank="species" species="lamelliferus">
<emphasis italics="true" pageId="0" pageNumber="25">T. lamelliferus</emphasis>
</taxonomicName>
(Milaschewitsch, 1912) from the Black Sea (near Alushta, Crimea) with strongly ribbed shells, but it is smaller than both (
<bibRefCitation author="Odabasi, DA" journalOrPublisher="Ecologica Montenegrina" pageId="0" pageNumber="25" pagination="327 - 333" refId="B115" refString="Odabasi, DA, Arslan, N, 2015. Description of a new subterranean nerite: Theodoxus gloeri n. sp. with some data on the freshwater gastropod fauna of Balikdami Wetland (Sakarya River, Turkey). Ecologica Montenegrina 2 (4): 327 - 333" title="Description of a new subterranean nerite: Theodoxus gloeri n. sp. with some data on the freshwater gastropod fauna of Balikdami Wetland (Sakarya River, Turkey)." volume="2" year="2015">
<normalizedToken originalValue="Odabaşı">Odabasi</normalizedToken>
and Arslan 2015
</bibRefCitation>
;
<bibRefCitation author="Gloeer, P" journalOrPublisher="Peter Gloeer, Hetlingen, Germany" pageId="0" pageNumber="25" refId="B62" refString="Gloeer, P, 2019. The freshwater gastropods of the West-Palaearctis. Vol. 1. Fresh- and brackish waters except spring and subterranean snails. Peter Gloeer, Hetlingen, Germany" title="The freshwater gastropods of the West-Palaearctis. Vol. 1. Fresh- and brackish waters except spring and subterranean snails." year="2019">
<normalizedToken originalValue="Glöer">Gloeer</normalizedToken>
2019
</bibRefCitation>
; Fig.
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Theodoxus gloeri Odabasi &amp; Arslan, 2015. A - C. Paratype from Balikdami Wetland-Sakarya River, Eskisehir, Turkey (coll. Gloeer). Scale bar: 1 mm." figureDoi="10.3897/zse.96.48312.figure11" httpUri="https://binary.pensoft.net/fig/376460" pageId="0" pageNumber="25">11</figureCitation>
). Although it is expected to be an extant subterranean species, no living material has ever been recorded and its phylogenetic position is presently unknown (
<bibRefCitation author="Odabasi, DA" journalOrPublisher="Ecologica Montenegrina" pageId="0" pageNumber="25" pagination="327 - 333" refId="B115" refString="Odabasi, DA, Arslan, N, 2015. Description of a new subterranean nerite: Theodoxus gloeri n. sp. with some data on the freshwater gastropod fauna of Balikdami Wetland (Sakarya River, Turkey). Ecologica Montenegrina 2 (4): 327 - 333" title="Description of a new subterranean nerite: Theodoxus gloeri n. sp. with some data on the freshwater gastropod fauna of Balikdami Wetland (Sakarya River, Turkey)." volume="2" year="2015">
<normalizedToken originalValue="Odabaşı">Odabasi</normalizedToken>
and Arslan 2015
</bibRefCitation>
).
</paragraph>
<caption doi="10.3897/zse.96.48312.figure11" httpUri="https://binary.pensoft.net/fig/376460" pageId="0" pageNumber="25" start="Figure 11" startId="F11">
<paragraph pageId="0" pageNumber="25">
<emphasis bold="true" pageId="0" pageNumber="25">Figure 11.</emphasis>
<taxonomicName authorityName="Odabasi &amp; Arslan" authorityYear="2015" class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus gloeri" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="gloeri">
<emphasis italics="true" pageId="0" pageNumber="25">Theodoxus gloeri</emphasis>
</taxonomicName>
<normalizedToken originalValue="Odabaşı">Odabasi</normalizedToken>
&amp; Arslan, 2015.
<emphasis bold="true" pageId="0" pageNumber="25">A</emphasis>
-
<emphasis bold="true" pageId="0" pageNumber="25">C</emphasis>
.
<typeStatus>Paratype</typeStatus>
from
<normalizedToken originalValue="Balıkdamı">Balikdami</normalizedToken>
Wetland-Sakarya River,
<normalizedToken originalValue="Eskişehir">Eskisehir</normalizedToken>
, Turkey (coll.
<normalizedToken originalValue="Glöer">Gloeer</normalizedToken>
). Scale bar:
<quantity metricMagnitude="-3" metricUnit="m" metricValue="1.0" unit="mm" value="1.0">1 mm</quantity>
.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="25" type="distribution">
<paragraph pageId="0" pageNumber="25">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="25">
Only known from two sites in very close proximity within the
<normalizedToken originalValue="Balıkdamı">Balikdami</normalizedToken>
Wetland-Sakarya River,
<normalizedToken originalValue="Eskişehir">Eskisehir</normalizedToken>
, Turkey (
<bibRefCitation author="Odabasi, DA" journalOrPublisher="Ecologica Montenegrina" pageId="0" pageNumber="25" pagination="327 - 333" refId="B115" refString="Odabasi, DA, Arslan, N, 2015. Description of a new subterranean nerite: Theodoxus gloeri n. sp. with some data on the freshwater gastropod fauna of Balikdami Wetland (Sakarya River, Turkey). Ecologica Montenegrina 2 (4): 327 - 333" title="Description of a new subterranean nerite: Theodoxus gloeri n. sp. with some data on the freshwater gastropod fauna of Balikdami Wetland (Sakarya River, Turkey)." volume="2" year="2015">
<normalizedToken originalValue="Odabaşı">Odabasi</normalizedToken>
and Arslan 2015
</bibRefCitation>
; Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Occurrence map for Theodoxus spp. in Asia. All points conform to specimens used in the phylogenetic analyses (Suppl. material 1: Table S 1), except for T. gloeri for which no molecular data are available and only the type and paratype localities are indicated. Species are partitioned across four identical maps (A - D) to limit overlap of points as far as possible. Barring T. gloeri, closer related species are grouped together (Fig. 2)." figureDoi="10.3897/zse.96.48312.figure3" httpUri="https://binary.pensoft.net/fig/376452" pageId="0" pageNumber="25">3D</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>